377 lines
42 KiB
XML
377 lines
42 KiB
XML
<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FF9FFF81B49C19ED7B0C2030" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax altinodus Prebus 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="92" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="89" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prebus, Matthew M.</mods:namePart>
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<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States & Department of Entomology & Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
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<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PeerJ</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="series">
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<mods:title>e 11514</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2021-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>9</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>462</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
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<mods:identifier type="ISSN">2167-8359</mods:identifier>
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<mods:identifier type="PMC">PMC8254503</mods:identifier>
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<mods:identifier type="PubMed">34249486</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102455" ID-GBIF-Taxon="183542578" ID-Zenodo-Dep="5102455" LSID="urn:lsid:plazi:treatment:054FDB70FF9FFF81B49C19ED7B0C2030" httpUri="http://treatment.plazi.org/id/054FDB70FF9FFF81B49C19ED7B0C2030" lastPageId="91" lastPageNumber="92" pageId="88" pageNumber="89">
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<subSubSection box="[498,885,1008,1035]" pageId="88" pageNumber="89" type="nomenclature">
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<paragraph blockId="88.[498,885,1008,1075]" box="[498,885,1008,1035]" pageId="88" pageNumber="89">
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<heading box="[498,885,1008,1035]" level="2" pageId="88" pageNumber="89">
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<emphasis bold="true" box="[498,885,1008,1035]" pageId="88" pageNumber="89">
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<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,781,1008,1035]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
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<emphasis bold="true" box="[498,781,1008,1035]" italics="true" pageId="88" pageNumber="89">Temnothorax altinodus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[790,885,1009,1035]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[498,862,1048,1075]" pageId="88" pageNumber="89" type="distribution">
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<paragraph blockId="88.[498,885,1008,1075]" box="[498,862,1048,1075]" pageId="88" pageNumber="89">
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Distribution & worker:
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<figureCitation box="[777,856,1048,1075]" captionStart="Figure 94" captionStartId="89.[524,589,1098,1120]" captionTargetBox="[498,1541,234,1085]" captionTargetId="figure-238@89.[498,1534,234,1070]" captionTargetPageId="89" captionText="Figure 94 Temnothorax altinodus sp. nov. holotype worker (CASENT0617708). (A) Profile view. (B) Full-face view. (C) Dorsal view. (D) Geographical and elevational distribution of specimens examined. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-94" figureDoi="http://doi.org/10.5281/zenodo.5102273" httpUri="https://zenodo.org/record/5102273/files/figure.png" pageId="88" pageNumber="89">Fig. 94</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="88" pageNumber="89" type="materials_examined">
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<paragraph blockId="88.[498,1542,1111,1934]" pageId="88" pageNumber="89">
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<emphasis bold="true" box="[530,834,1111,1137]" pageId="88" pageNumber="89">Type material examined:</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3321861839" collectingDate="2010-06-01" collectionCode="CASC" country="Honduras" elevation="1290" latitude="15.48896" location="Parque Nacional Cusuco" longLatPrecision="1" longitude="-88.23439" pageId="88" pageNumber="89" specimenCode="LLAMAGo-C-06-4-03, CASENT0617708" specimenCount="1" stateProvince="Cortes" typeStatus="holotype">
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<emphasis box="[843,1041,1111,1137]" italics="true" pageId="88" pageNumber="89">
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<typeStatus box="[843,946,1111,1137]" pageId="88" pageNumber="89">Holotype</typeStatus>
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<specimenCount box="[954,1034,1111,1137]" pageId="88" pageNumber="89" type="worker">worker</specimenCount>
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:
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</emphasis>
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<collectingCountry box="[1049,1212,1111,1137]" name="Honduras" pageId="88" pageNumber="89">HONDURAS</collectingCountry>
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:
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<collectingRegion box="[1224,1303,1111,1137]" country="Honduras" name="Cortes" pageId="88" pageNumber="89">Cortés</collectingRegion>
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:
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<location LSID="urn:lsid:plazi:treatment:054FDB70FF9FFF81B49C19ED7B0C2030:88393CBDFF9FFF82B04D1E457B7923CE" country="Honduras" latitude="15.48896" longLatPrecision="1" longitude="-88.23439" name="Parque Nacional Cusuco" pageId="88" pageNumber="89" stateProvince="Cortes">Parque Nacional Cusuco</location>
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,
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<geoCoordinate box="[600,738,1146,1178]" degrees="15.48896" direction="north" orientation="latitude" pageId="88" pageNumber="89" precision="1" value="15.48896">
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15.48896
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<emphasis box="[704,714,1146,1164]" italics="true" pageId="88" pageNumber="89">
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<superScript attach="right" box="[704,714,1146,1164]" fontSize="8" pageId="88" pageNumber="89">°</superScript>
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</emphasis>
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N
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</geoCoordinate>
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<geoCoordinate box="[745,889,1146,1177]" degrees="88.23439" direction="west" orientation="longitude" pageId="88" pageNumber="89" precision="1" value="-88.23439">
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88.23439
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<emphasis box="[850,860,1146,1164]" italics="true" pageId="88" pageNumber="89">
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<superScript attach="right" box="[850,860,1146,1164]" fontSize="8" pageId="88" pageNumber="89">°</superScript>
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</emphasis>
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W
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</geoCoordinate>
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±
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<quantity box="[922,983,1151,1177]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="88" pageNumber="89" unit="m" value="70.0">70 m</quantity>
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,
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<quantity box="[997,1092,1151,1177]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="88" pageNumber="89" unit="m" value="1290.0">
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<elevation box="[997,1092,1151,1177]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="88" pageNumber="89" unit="m" value="1290.0">1,290 m</elevation>
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</quantity>
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,
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<date box="[1105,1246,1151,1177]" pageId="88" pageNumber="89" value="2010-06-01">
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<collectingDate box="[1105,1246,1151,1177]" pageId="88" pageNumber="89" value="2010-06-01">1 June 2010</collectingDate>
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</date>
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,
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<specimenCode box="[1259,1537,1151,1177]" pageId="88" pageNumber="89">LLAMA#Go-C-06-4-03</specimenCode>
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, mesophyll forest, beating vegetation (
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<specimenCode box="[937,1149,1191,1218]" pageId="88" pageNumber="89">CASENT0617708</specimenCode>
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) [
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<collectionCode box="[1172,1253,1190,1217]" pageId="88" pageNumber="89">CASC</collectionCode>
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].
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="88" pageNumber="89" type="distribution">
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<paragraph blockId="88.[498,1542,1111,1934]" pageId="88" pageNumber="89">
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<emphasis bold="true" box="[530,752,1230,1257]" pageId="88" pageNumber="89">Geographic range:</emphasis>
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<collectingCountry box="[758,877,1230,1256]" name="Honduras" pageId="88" pageNumber="89">Honduras</collectingCountry>
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, mid elevations of the Central American Nucleus (
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<collectingRegion box="[1454,1534,1231,1257]" country="Honduras" name="Cortes" pageId="88" pageNumber="89">Cortés</collectingRegion>
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) (
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<figureCitation box="[508,608,1270,1296]" captionStart="Figure 94" captionStartId="89.[524,589,1098,1120]" captionTargetBox="[498,1541,234,1085]" captionTargetId="figure-238@89.[498,1534,234,1070]" captionTargetPageId="89" captionText="Figure 94 Temnothorax altinodus sp. nov. holotype worker (CASENT0617708). (A) Profile view. (B) Full-face view. (C) Dorsal view. (D) Geographical and elevational distribution of specimens examined. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-94" figureDoi="http://doi.org/10.5281/zenodo.5102273" httpUri="https://zenodo.org/record/5102273/files/figure.png" pageId="88" pageNumber="89">Fig. 94D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89" type="diagnosis">
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<paragraph blockId="88.[498,1542,1111,1934]" pageId="88" pageNumber="89">
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<emphasis bold="true" box="[530,754,1310,1336]" pageId="88" pageNumber="89">Worker diagnosis:</emphasis>
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<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[763,1036,1310,1336]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
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<emphasis box="[763,1036,1310,1336]" italics="true" pageId="88" pageNumber="89">Temnothorax altinodus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1045,1136,1311,1337]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
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can be separated from all other species in the
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<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[662,737,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="salvini">
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<emphasis box="[662,737,1350,1376]" italics="true" pageId="88" pageNumber="89">salvini</emphasis>
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</taxonomicName>
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clade by the following character combination: antennal scapes short, failing to reach the posterior margin of the head capsule by about the maximum width of the scape (SI 83); mesosoma relatively elongate (WLI 148); dorsum of propodeum without erect setae; propodeal spines directed dorsally; hind femora strongly incrassate (FI 327); petiole with a weak transverse carina anterodorsally; petiolar node compact (NI 143); in dorsal view, apex of petiolar node narrower than caudal cylinder; petiolar node dorsally rounded; postpetiole moderately broad (PWI 202); integument predominantly black; setae on head, mesosoma, waist segments and gaster erect, moderately long, sparse and blunt (never long and tapering).
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</paragraph>
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<paragraph blockId="88.[498,1542,1111,1934]" lastBlockId="89.[498,1542,1292,1877]" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
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<emphasis bold="true" box="[530,716,1708,1734]" pageId="88" pageNumber="89">Similar species:</emphasis>
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<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[722,941,1709,1735]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
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<emphasis box="[722,941,1709,1735]" italics="true" pageId="88" pageNumber="89">Temnothorax achii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[947,1037,1709,1735]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1048,1213,1710,1735]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
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<emphasis box="[1048,1213,1710,1735]" italics="true" pageId="88" pageNumber="89">T. acuminatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1219,1310,1709,1735]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1320,1503,1710,1735]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
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<emphasis box="[1320,1503,1710,1735]" italics="true" pageId="88" pageNumber="89">T. acutispinosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<emphasis box="[562,951,1748,1774]" italics="true" pageId="88" pageNumber="89">
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<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[562,638,1748,1774]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="ixili">T. ixili</taxonomicName>
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,
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<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[650,800,1748,1774]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="subditivus">T. subditivus</taxonomicName>
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,
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<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[811,951,1748,1774]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">T. tuxtlanus</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[958,1049,1749,1775]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1060,1161,1749,1774]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="xincai" status="sp. nov.">
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<emphasis box="[1060,1161,1749,1774]" italics="true" pageId="88" pageNumber="89">T. xincai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1168,1259,1749,1775]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
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, members of the
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<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[1458,1542,1749,1774]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="augusti">
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<emphasis box="[1458,1542,1749,1774]" italics="true" pageId="88" pageNumber="89">augusti</emphasis>
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</taxonomicName>
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and
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<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[551,664,1788,1814]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="pulchellus">
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<emphasis box="[551,664,1788,1814]" italics="true" pageId="88" pageNumber="89">pulchellus</emphasis>
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</taxonomicName>
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groups, and species of the
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<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[987,1044,1788,1814]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sallei">
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<emphasis box="[987,1044,1788,1814]" italics="true" pageId="88" pageNumber="89">sallei</emphasis>
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</taxonomicName>
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clade.
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<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1128,1400,1788,1814]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
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<emphasis box="[1128,1400,1788,1814]" italics="true" pageId="88" pageNumber="89">Temnothorax altinodus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1409,1501,1789,1815]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
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can be distinguished from similar appearing members of the
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<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[1221,1278,1828,1854]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sallei">
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<emphasis box="[1221,1278,1828,1854]" italics="true" pageId="88" pageNumber="89">sallei</emphasis>
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</taxonomicName>
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clade (e.g.
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<emphasis italics="true" pageId="88" pageNumber="89">
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<taxonomicName box="[1412,1523,1829,1854]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="manni">T. manni</taxonomicName>
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,
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<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[498,654,1869,1894]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="mexicanus">T. mexicanus</taxonomicName>
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,
|
||
<taxonomicName authorityName="" baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[666,847,1868,1894]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="punctithorax">T. punctithorax</taxonomicName>
|
||
</emphasis>
|
||
) by the sculpture of the head: in similar looking
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[1417,1474,1868,1894]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis box="[1417,1474,1868,1894]" italics="true" pageId="88" pageNumber="89">sallei</emphasis>
|
||
</taxonomicName>
|
||
clade species, the head is mostly smooth and shining, with concentric costulae surrounding the antennal insertions; additionally, the medial lobe of the clypeus is typically smooth, with three distinct carinae, whereas it is densely costulate-rugulose in
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1264,1404,1332,1358]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
|
||
<emphasis box="[1264,1404,1332,1358]" italics="true" pageId="89" pageNumber="90">T. altinodus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1412,1504,1333,1359]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
In contrast to all of the remaining species above, the petiole of
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1236,1375,1372,1398]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
|
||
<emphasis box="[1236,1375,1372,1398]" italics="true" pageId="89" pageNumber="90">T. altinodus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1383,1474,1373,1399]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a weak transverse carina anterodorsally, a feature not found in other members of the
|
||
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1467,1542,1412,1438]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis box="[1467,1542,1412,1438]" italics="true" pageId="89" pageNumber="90">salvini</emphasis>
|
||
</taxonomicName>
|
||
clade, except for
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[699,866,1453,1478]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
|
||
<emphasis box="[699,866,1453,1478]" italics="true" pageId="89" pageNumber="90">T. acuminatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[875,969,1452,1478]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[982,1123,1452,1478]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
|
||
<emphasis box="[982,1123,1452,1478]" italics="true" pageId="89" pageNumber="90">T. tuxtlanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1132,1225,1452,1478]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1238,1358,1453,1478]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="rugosus">
|
||
<emphasis box="[1238,1358,1453,1478]" italics="true" pageId="89" pageNumber="90">T. rugosus</emphasis>
|
||
</taxonomicName>
|
||
, and members of the
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[576,672,1492,1517]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="annexus">
|
||
<emphasis box="[576,672,1492,1517]" italics="true" pageId="89" pageNumber="90">annexus</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[764,1037,1492,1518]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
|
||
<emphasis box="[764,1037,1492,1518]" italics="true" pageId="89" pageNumber="90">Temnothorax altinodus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1046,1137,1492,1518]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is easily separated from
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,684,1532,1557]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
|
||
<emphasis box="[498,684,1532,1557]" italics="true" pageId="89" pageNumber="90">T. acutispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[693,786,1532,1558]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[799,967,1532,1557]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
|
||
<emphasis box="[799,967,1532,1557]" italics="true" pageId="89" pageNumber="90">T. acuminatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[976,1068,1532,1558]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1129,1271,1532,1558]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
|
||
<emphasis box="[1129,1271,1532,1558]" italics="true" pageId="89" pageNumber="90">T. tuxtlanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1279,1371,1532,1558]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the hind femora, which are incrassate in
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[874,1014,1571,1597]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
|
||
<emphasis box="[874,1014,1571,1597]" italics="true" pageId="89" pageNumber="90">T. altinodus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1023,1116,1572,1598]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; it may be distinguished from
|
||
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[498,618,1612,1637]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="rugosus">
|
||
<emphasis box="[498,618,1612,1637]" italics="true" pageId="89" pageNumber="90">T. rugosus</emphasis>
|
||
</taxonomicName>
|
||
and the members of the
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[916,1012,1612,1637]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="annexus">
|
||
<emphasis box="[916,1012,1612,1637]" italics="true" pageId="89" pageNumber="90">annexus</emphasis>
|
||
</taxonomicName>
|
||
group by the anterior margin of the clypeus, which entire in
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[686,826,1651,1677]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
|
||
<emphasis box="[686,826,1651,1677]" italics="true" pageId="89" pageNumber="90">T. altinodus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[834,927,1652,1678]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, as opposed to medially emarginate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102273" ID-Zenodo-Dep="5102273" httpUri="https://zenodo.org/record/5102273/files/figure.png" pageId="89" pageNumber="90" startId="89.[524,589,1098,1120]" targetBox="[498,1541,234,1085]" targetPageId="89">
|
||
<paragraph blockId="89.[524,1516,1098,1237]" pageId="89" pageNumber="90">
|
||
<emphasis bold="true" box="[524,1341,1098,1120]" pageId="89" pageNumber="90">
|
||
Figure 94
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[634,869,1098,1120]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
|
||
<emphasis bold="true" box="[634,869,1098,1120]" italics="true" pageId="89" pageNumber="90">Temnothorax altinodus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[878,958,1099,1120]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
holotype worker (CASENT0617708).
|
||
</emphasis>
|
||
(A) Profile view. (B) Full-face view. (C) Dorsal view. (D) Geographical and elevational distribution of specimens examined. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-94
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="91" lastPageNumber="92" pageId="89" pageNumber="90" type="description">
|
||
<paragraph blockId="89.[498,1542,1292,1877]" lastBlockId="90.[498,1542,234,1934]" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
|
||
<emphasis bold="true" box="[530,1032,1691,1717]" pageId="89" pageNumber="90">
|
||
Worker measurements & indices (
|
||
<emphasis bold="true" box="[950,966,1691,1717]" italics="true" pageId="89" pageNumber="90">n</emphasis>
|
||
= 1):
|
||
</emphasis>
|
||
SL = 0.607; FRS = 0.223; CW = 0.773; CWb = 0.727; PoOC = 0.343; CL = 0.823; EL = 0.188; EW = 0.131; MD = 0.164; WL = 1.079; SPST = 0.252; MPST = 0.353; PEL = 0.38; NOL = 0.22; NOH = 0.154; PEH = 0.272; PPL = 0.190; PPH = 0.276; PW = 0.499; SBPA = 0.197; SPTI = 0.266; PEW = 0.157; PNW = 0.146; PPW = 0.317; HFL = 0.715; HFWmax = 0.209; HFWmin = 0.064; CS = 1.1385; ES = 0.254; SI = 83; OI = 22; CI = 88; WLI = 148; SBI = 27; PSI = 23; PWI = 202; PLI = 200; NI = 143; PNWI = 93; NLI = 58; FI = 327.
|
||
</paragraph>
|
||
<paragraph blockId="90.[498,1542,234,1934]" pageId="90" pageNumber="91">
|
||
<emphasis bold="true" box="[530,777,314,340]" pageId="90" pageNumber="91">Worker description:</emphasis>
|
||
In full-face view, head subquadrate, longer than broad (CI 88). Mandibles densely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin flat medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about the maximum width of the scape (SI 83). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment about one and a half times as long as the preceding two in combination. Frontal carinae moderately long, extending past the antennal toruli by about two times the maximum width of the antennal scape. Compound eyes weakly protruding past the lateral margins of the head capsule. Lateral margin of head weakly convex, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin flat but rounding evenly into the lateral margins.
|
||
</paragraph>
|
||
<paragraph blockId="90.[498,1542,234,1934]" pageId="90" pageNumber="91">
|
||
In profile view, compound eyes ovular and small (OI 22), with 11 ommatidia in longest row. Pronotal declivity indistinct, neck and anterior face of pronotum forming a ~120
|
||
<emphasis box="[1514,1524,867,885]" italics="true" pageId="90" pageNumber="91">
|
||
<superScript attach="left" box="[1514,1524,867,885]" fontSize="8" pageId="90" pageNumber="91">°</superScript>
|
||
</emphasis>
|
||
angle. Mesosoma evenly rounded anteriorly and flat dorsally to the bases of the propodeal spines. Promesonotal suture extending from the posterior margin of the procoxal insertion to the mesothoracic spiracle, which is moderately well developed. Metanotal groove visible as a disruption of the sculpture laterally from where it arises between the mid- and hind coxae to where it ends in the poorly developed metathoracic spiracle, which is nearly indistinguishable against the ground sculpture. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about three spiracle diameters. Propodeal spines moderately well developed (PSI 23), about as long as the propodeal declivity, tapering evenly from the base, straight, and acute. Propodeal declivity weakly concave, forming a rounded ~120
|
||
<emphasis box="[986,996,1266,1284]" italics="true" pageId="90" pageNumber="91">
|
||
<superScript attach="left" box="[986,996,1266,1284]" fontSize="8" pageId="90" pageNumber="91">°</superScript>
|
||
</emphasis>
|
||
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed. Metapleural gland bulla moderately large, extending from the metacoxal insertion two thirds of the way to the propodeal spiracle. Petiole moderately long (PLI 200), with moderately well-developed tubercles anterodorsally. Subpetiolar process in the form of a small, posteriorly directed, blunt tooth; ventral margin of petiole flat concave posterior to it. Petiolar peduncle short: comprising about a third of the total length of the petiole. Petiolar node erect: transition between peduncle and node marked by a rounded angle of ~140
|
||
<emphasis box="[1155,1165,1545,1563]" italics="true" pageId="90" pageNumber="91">
|
||
<superScript attach="left" box="[1155,1165,1545,1563]" fontSize="8" pageId="90" pageNumber="91">°</superScript>
|
||
</emphasis>
|
||
, resulting in a weakly concave anterior node face; anterior face forming a ~90
|
||
<emphasis box="[1037,1047,1584,1602]" italics="true" pageId="90" pageNumber="91">
|
||
<superScript attach="left" box="[1037,1047,1584,1602]" fontSize="8" pageId="90" pageNumber="91">°</superScript>
|
||
</emphasis>
|
||
angle with the dorsal face, which is weakly convex; dorsal face meeting the posterior face at a ~100
|
||
<emphasis box="[1156,1166,1624,1642]" italics="true" pageId="90" pageNumber="91">
|
||
<superScript attach="left" box="[1156,1166,1624,1642]" fontSize="8" pageId="90" pageNumber="91">°</superScript>
|
||
</emphasis>
|
||
angle; posterior face forming a ~90
|
||
<emphasis box="[543,553,1664,1682]" italics="true" pageId="90" pageNumber="91">
|
||
<superScript attach="left" box="[543,553,1664,1682]" fontSize="8" pageId="90" pageNumber="91">°</superScript>
|
||
</emphasis>
|
||
angle with the caudal cylinder. Postpetiole evenly rounded anterodorsally, bulging before flattening posterodorsally; lobed ventrally.
|
||
</paragraph>
|
||
<paragraph blockId="90.[498,1542,234,1934]" lastBlockId="91.[498,1542,234,1895]" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">In dorsal view, humeri developed and distinct: evenly rounded and wider than the rest of the mesosoma; mesothoracic spiracles weakly protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Promesonotal suture visible as a slight disruption in the ground sculpture. Metanotal groove absent: mesonotum and propodeum completely fused and lateral margins converging evenly to the bases of the propodeal spines. Propodeal spines broadly approximated basally and diverging apically, their apices separated from each other by about their length, the negative space between them “U” shaped. Petiolar peduncle with spiracles protruding past the lateral margins; peduncle broadened where they arise. Petiolar node subquadrate; node slightly broader than the peduncle, and evenly grading into the caudal cylinder, which is slightly wider than the node. Postpetiole moderately broad (PWI 202) and campaniform, articulating with the nearly the entire anterior margin of the gaster but leaving small angulate corners exposed. Anterior margin of the postpetiole weakly convex and evenly rounds into the lateral margins, which diverge slightly to the angulate posterior corners; posterior margin flat. Metafemur incrassate (FI 327).</paragraph>
|
||
<paragraph blockId="91.[498,1542,234,1895]" pageId="91" pageNumber="92">Sculpture: median clypeal carina present, extending posteriorly to the frontal triangle, and flanked on either side by two equally strong carinae. Lateral clypeal lobes with additional, weaker carinae; ground sculpture weakly areolate. Antennal scapes areolatecostulate. Cephalic dorsum densely areolate-rugulose, with fine rugose sculpture overlying it. Lateral surfaces of sculptured similarly to the dorsum, but with areolate-rugulose sculpture becoming stronger between compound eye and mandibular insertion. Ventral surface of head longitudinally areolate-costulate. Pronotal neck weakly areolate. Lateral surfaces of mesosoma densely areolate, with fine rugose-costulate sculpture overlying it. Propodeal declivity weakly areolate. Dorsal surface of mesosoma densely areolate, with fine rugose overlying it. Femora shining through weak areolate sculpture. Petiole uniformly areolate; a weak carina present laterally, extending longitudinally from the anterodorsal flange to the caudal cylinder, passing through the petiolar spiracle. Postpetiole uniformly areolate. First gastral tergite weakly areolate on the basal quarter, otherwise smooth and shining, with weak spectral iridescence. First gastral sternite smooth and shining.</paragraph>
|
||
<paragraph blockId="91.[498,1542,234,1895]" pageId="91" pageNumber="92">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of the head, pronotum, waist segments, and gaster with sparse, short, erect, blunt-tipped setae, the longest of which are slightly less than the width of the compound eye. The head bears ~16, mesosoma ~8 restricted to the promesonotum, petiole 2, postpetiole ~4, and first gastral tergite ~20 setae. Short, sparse pubescence present over the entire body, but difficult to detect against the densely sculptured integument.</paragraph>
|
||
<paragraph blockId="91.[498,1542,234,1895]" pageId="91" pageNumber="92">Color: predominantly black, with dark brown antennae, mandibles, and legs (excluding tarsi). Tarsi and sting testaceous yellow.</paragraph>
|
||
<paragraph blockId="91.[498,1542,234,1895]" box="[530,734,1509,1535]" pageId="91" pageNumber="92">
|
||
<emphasis bold="true" box="[530,601,1509,1535]" pageId="91" pageNumber="92">Gyne:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph blockId="91.[498,1542,234,1895]" box="[530,730,1549,1575]" pageId="91" pageNumber="92">
|
||
<emphasis bold="true" box="[530,597,1549,1575]" pageId="91" pageNumber="92">Male:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="91" pageNumber="92" type="etymology">
|
||
<paragraph blockId="91.[498,1542,234,1895]" pageId="91" pageNumber="92">
|
||
<emphasis bold="true" box="[530,668,1589,1615]" pageId="91" pageNumber="92">Etymology:</emphasis>
|
||
Morphological, from the Latin ‘altus’ (= high, tall) + ‘nodus’ (= node), a reference to the very tall petiolar node.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="91" pageNumber="92" type="discussion">
|
||
<paragraph blockId="91.[498,1542,234,1895]" pageId="91" pageNumber="92">
|
||
<emphasis bold="true" box="[530,670,1669,1695]" pageId="91" pageNumber="92">Comments:</emphasis>
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[679,951,1669,1695]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
|
||
<emphasis box="[679,951,1669,1695]" italics="true" pageId="91" pageNumber="92">Temnothorax altinodus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[959,1050,1669,1695]" pageId="91" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known only from the
|
||
<typeStatus box="[1343,1445,1669,1695]" pageId="91" pageNumber="92">holotype</typeStatus>
|
||
worker, collected via beating vegetation in a mid-elevation mesophyll forest in northwestern
|
||
<collectingCountry box="[498,618,1748,1774]" name="Honduras" pageId="91" pageNumber="92">Honduras</collectingCountry>
|
||
. This species is most closely related to the exclusively arboreally nesting
|
||
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1449,1541,1749,1775]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fuscatus">
|
||
<emphasis box="[1449,1541,1749,1775]" italics="true" pageId="91" pageNumber="92">fuscatus</emphasis>
|
||
</taxonomicName>
|
||
species group, which ranges from Central Mexico to
|
||
<collectingCountry box="[1114,1240,1788,1814]" name="Costa Rica" pageId="91" pageNumber="92">Costa Rica</collectingCountry>
|
||
, and the
|
||
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1895" box="[1347,1459,1788,1814]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="pergandei">
|
||
<emphasis box="[1347,1459,1788,1814]" italics="true" pageId="91" pageNumber="92">pergandei</emphasis>
|
||
</taxonomicName>
|
||
group, ground nesting species ranging from the northern
|
||
<collectingCountry box="[1092,1248,1828,1854]" name="United States of America" pageId="91" pageNumber="92">United States</collectingCountry>
|
||
to
|
||
<collectingCountry box="[1287,1406,1828,1854]" name="Nicaragua" pageId="91" pageNumber="92">Nicaragua</collectingCountry>
|
||
(Prebus, in prep.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |