486 lines
54 KiB
XML
486 lines
54 KiB
XML
<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FF6FFF77B49C1CE57DFC2190" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax skwarrae" docType="treatment" docVersion="4" lastPageNumber="174" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="169" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prebus, Matthew M.</mods:namePart>
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<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States & Department of Entomology & Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
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<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PeerJ</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="series">
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<mods:title>e 11514</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2021-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>9</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>462</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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||
<mods:location>
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<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
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<mods:identifier type="ISSN">2167-8359</mods:identifier>
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<mods:identifier type="PMC">PMC8254503</mods:identifier>
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<mods:identifier type="PubMed">34249486</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102523" ID-GBIF-Taxon="183542610" ID-Zenodo-Dep="5102523" LSID="urn:lsid:plazi:treatment:054FDB70FF6FFF77B49C1CE57DFC2190" httpUri="http://treatment.plazi.org/id/054FDB70FF6FFF77B49C1CE57DFC2190" lastPageId="173" lastPageNumber="174" pageId="168" pageNumber="169">
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<subSubSection box="[498,979,1784,1811]" pageId="168" pageNumber="169" type="nomenclature">
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||
<paragraph blockId="168.[498,1076,1784,1851]" box="[498,979,1784,1811]" pageId="168" pageNumber="169">
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<heading box="[498,979,1784,1811]" level="2" pageId="168" pageNumber="169">
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<taxonomicName authority="(Wheeler, 1931)" baseAuthorityName="Wheeler" baseAuthorityYear="1931" box="[498,979,1784,1811]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="skwarrae">
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<emphasis bold="true" box="[498,979,1784,1811]" pageId="168" pageNumber="169">
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<emphasis bold="true" box="[498,777,1784,1811]" italics="true" pageId="168" pageNumber="169">Temnothorax skwarrae</emphasis>
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(
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<bibRefCitation author="Wheeler WM" box="[797,969,1784,1811]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="168" pageNumber="169" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">
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<emphasis bold="true" box="[797,969,1784,1811]" italics="true" pageId="168" pageNumber="169">Wheeler, 1931</emphasis>
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</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[498,1076,1825,1851]" pageId="168" pageNumber="169" type="distribution">
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<paragraph blockId="168.[498,1076,1784,1851]" box="[498,1076,1825,1851]" pageId="168" pageNumber="169">
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Distribution:
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<figureCitation box="[657,771,1825,1851]" captionStart="Figure 107" captionStartId="148.[524,589,1194,1216]" captionTargetBox="[498,1541,235,1181]" captionTargetId="figure-181@148.[498,1534,234,1166]" captionTargetPageId="148" captionText="Figure 107 Geographical and elevational distribution of the fuscatus group. (A) Temnothorax fuscatus (B) T. nebliselva sp. nov. (C) T. ocarinae (D) T. skwarrae. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-107" figureDoi="http://doi.org/10.5281/zenodo.5102311" httpUri="https://zenodo.org/record/5102311/files/figure.png" pageId="168" pageNumber="169">Fig. 107D</figureCitation>
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; worker & gyne:
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<figureCitation box="[977,1070,1825,1851]" captionStart="Figure 113" captionStartId="168.[524,589,1387,1409]" captionTargetBox="[498,1541,234,1374]" captionTargetId="figure-100@168.[498,1534,234,1359]" captionTargetPageId="168" captionText="Figure 113 Temnothorax skwarrae. (A–C) Lectotype worker (MCZENT00016358). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D–F) Paralectotype gyne (MCZENT00577117). (D) Full-face view. (E) Profile view. (F) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-113" figureDoi="http://doi.org/10.5281/zenodo.5102327" httpUri="https://zenodo.org/record/5102327/files/figure.png" pageId="168" pageNumber="169">Fig. 113</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="169" pageNumber="170" type="reference_group">
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<paragraph blockId="169.[498,1542,234,1934]" pageId="169" pageNumber="170">
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<treatmentCitationGroup pageId="169" pageNumber="170">
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<treatmentCitation author="Wheeler WM" box="[530,1018,234,261]" page="10" pageId="169" pageNumber="170" year="1931">
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<taxonomicName authority="Wheeler, 1931: 10" authorityName="Wheeler" authorityPageNumber="10" authorityYear="1931" box="[530,973,234,260]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="skwarrae">
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<emphasis box="[530,973,234,260]" italics="true" pageId="169" pageNumber="170">
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Macromischa skwarrae
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<bibRefCitation author="Wheeler WM" box="[808,1018,234,261]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="169" pageNumber="170" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">
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<author box="[808,905,234,260]" fns="*" inm="true" npo="*" pageId="169" pageNumber="170">Wheeler</author>
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,
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<year box="[918,973,234,260]" pageId="169" pageNumber="170">1931</year>
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||
: 10
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</bibRefCitation>
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||
</emphasis>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus box="[1031,1123,234,260]" pageId="169" pageNumber="170">Syntype</typeStatus>
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workers and gyne. Cuernavaca,
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<collectingCountry box="[498,587,274,300]" name="Mexico" pageId="169" pageNumber="170">Mexico</collectingCountry>
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. One
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<typeStatus box="[658,747,275,301]" pageId="169" pageNumber="170">syntype</typeStatus>
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worker here designated
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<emphasis bold="true" box="[1039,1148,274,300]" pageId="169" pageNumber="170">
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<typeStatus box="[1039,1148,274,300]" pageId="169" pageNumber="170">lectotype</typeStatus>
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</emphasis>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="169.[498,1542,234,1934]" pageId="169" pageNumber="170">
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<treatmentCitationGroup pageId="169" pageNumber="170">
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<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" baseAuthorityYear="1931" box="[530,913,314,341]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="skwarrae">
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<emphasis box="[530,782,314,340]" italics="true" pageId="169" pageNumber="170">Leptothorax skwarrae</emphasis>
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(Wheeler)
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</taxonomicName>
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:
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<treatmentCitation author="Baroni Urbani C." box="[925,1226,314,341]" page="503" pageId="169" pageNumber="170" year="1978">
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<bibRefCitation author="Baroni Urbani C." box="[925,1226,314,341]" journalOrPublisher="Entomologica Basiliensia" pageId="169" pageNumber="170" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
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<emphasis box="[925,1166,314,340]" italics="true" pageId="169" pageNumber="170">
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<author box="[925,1097,314,340]" case="TC" fns="*" inm="true" npo="*" pageId="169" pageNumber="170">Baroni Urbani</author>
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,
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<year box="[1111,1166,314,340]" pageId="169" pageNumber="170">1978</year>
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</emphasis>
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: 503
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</bibRefCitation>
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</treatmentCitation>
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. First combination in
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<taxonomicName authority="Mayr" box="[498,638,354,380]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis box="[498,638,354,380]" italics="true" pageId="169" pageNumber="170">Leptothorax</emphasis>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="169.[498,1542,234,1934]" pageId="169" pageNumber="170">
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<treatmentCitationGroup pageId="169" pageNumber="170">
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<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" box="[530,930,394,421]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="skwarrae">
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<emphasis box="[530,800,394,420]" italics="true" pageId="169" pageNumber="170">Temnothorax skwarrae</emphasis>
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(Wheeler)
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</taxonomicName>
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:
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<treatmentCitation author="Bolton B." box="[942,1148,394,420]" page="272" pageId="169" pageNumber="170" year="2003">
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<bibRefCitation author="Bolton B." box="[942,1148,394,420]" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="169" pageNumber="170" pagination="370" part="71" refId="ref270076" refString="Bolton B. 2003. Synopsis and Classification of Formicidae. Memoirs of the American Entomological Institute 71: 370." title="Synopsis and Classification of Formicidae" type="journal article" year="2003">
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<emphasis box="[942,1088,394,420]" italics="true" pageId="169" pageNumber="170">
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||
<collectingRegion box="[942,1020,394,420]" country="United Kingdom" name="Bolton" pageId="169" pageNumber="170">
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||
<author box="[942,1020,394,420]" fns="*" inm="true" npo="*" pageId="169" pageNumber="170">Bolton</author>
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</collectingRegion>
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,
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<year box="[1033,1088,394,420]" pageId="169" pageNumber="170">2003</year>
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</emphasis>
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: 272
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</bibRefCitation>
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</treatmentCitation>
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. First combination in
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<taxonomicName authorityName="Mayr" authorityYear="1861" box="[498,655,433,459]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis box="[498,655,433,459]" italics="true" pageId="169" pageNumber="170">Temnothorax</emphasis>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="169" pageNumber="170" type="materials_examined">
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<paragraph blockId="169.[498,1542,234,1934]" pageId="169" pageNumber="170">
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<emphasis bold="true" box="[530,834,473,499]" pageId="169" pageNumber="170">Type material examined:</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3321861312" collectingDate="1929-06-26" collectionCode="MCZC" collectorName="E. Skwarra " country="Mexico" location="Cuernavaca" pageId="169" pageNumber="170" specimenCount="1" specimenCount-worker="1" stateProvince="Morelos" typeStatus="lectotype">
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<emphasis box="[843,1039,473,499]" italics="true" pageId="169" pageNumber="170">
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<typeStatus box="[843,951,474,499]" pageId="169" pageNumber="170">Lectotype</typeStatus>
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worker
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</emphasis>
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:
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<collectingCountry box="[1054,1169,473,499]" name="Mexico" pageId="169" pageNumber="170">MEXICO</collectingCountry>
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:
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<collectingRegion box="[1181,1280,473,499]" country="Mexico" name="Morelos" pageId="169" pageNumber="170">Morelos</collectingRegion>
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:
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<location LSID="urn:lsid:plazi:treatment:054FDB70FF6FFF77B49C1CE57DFC2190:88393CBDFF6EFF73B0651BC47CA426A4" box="[1291,1430,473,499]" country="Mexico" name="Cuernavaca" pageId="169" pageNumber="170" stateProvince="Morelos">Cuernavaca</location>
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,
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<date pageId="169" pageNumber="170" value="1929-06-26">
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<collectingDate pageId="169" pageNumber="170" value="1929-06-26">26 June 1929</collectingDate>
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</date>
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,
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<collectorName box="[569,698,513,540]" pageId="169" pageNumber="170">E. Skwarra</collectorName>
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#Z857a/
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<collectorName box="[792,826,513,540]" pageId="169" pageNumber="170">Sk.</collectorName>
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<specimenCount box="[834,939,513,539]" pageId="169" pageNumber="170" type="worker">1 worker</specimenCount>
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(
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<collectorName box="[957,1015,513,539]" name="Menozzi C" pageId="169" pageNumber="170">M.C.</collectorName>
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Z. co-type 10 16358, MCZENT00016358) [
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<collectionCode box="[505,596,552,579]" pageId="169" pageNumber="170">MCZC</collectionCode>
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||
].
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</materialsCitation>
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</paragraph>
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<paragraph blockId="169.[498,1542,234,1934]" pageId="169" pageNumber="170">
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<emphasis box="[530,897,593,619]" italics="true" pageId="169" pageNumber="170">
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<typeStatus box="[530,684,593,619]" pageId="169" pageNumber="170">Paralectotype</typeStatus>
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workers and gyne
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</emphasis>
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:
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<materialsCitation ID-GBIF-Occurrence="3321861591" collectionCode="MCZC" collectorName="E. Skwarra" pageId="169" pageNumber="170" specimenCount="1" typeStatus="paralectotype">
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same data as previous, except:
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<collectorName box="[1274,1403,593,619]" pageId="169" pageNumber="170">E. Skwarra</collectorName>
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#Z879/
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<collectorName box="[1486,1520,593,619]" pageId="169" pageNumber="170">Sk.</collectorName>
|
||
, ex
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||
<taxonomicName box="[533,774,633,659]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="circinnata">
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||
<emphasis box="[533,774,633,659]" italics="true" pageId="169" pageNumber="170">Tillandsia circinnata</emphasis>
|
||
</taxonomicName>
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||
, 1 dealate gyne (
|
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<collectorName box="[974,1033,633,659]" name="Menozzi C" pageId="169" pageNumber="170">M.C.</collectorName>
|
||
Z. co-type 916358, MCZENT00577117) [
|
||
<collectionCode box="[505,595,672,699]" pageId="169" pageNumber="170">MCZC</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3321861684" collectionCode="SMND" collectorName="E. Skwarra & FoCol" location="FoCol" pageId="169" pageNumber="170" specimenCount="1" specimenCount-worker="1" typeStatus="paralectotype">
|
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same data as previous, except:
|
||
<collectorName box="[978,1106,673,699]" pageId="169" pageNumber="170">E. Skwarra</collectorName>
|
||
#Z877, ex
|
||
<taxonomicName box="[1227,1468,673,699]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="circinnata">
|
||
<emphasis box="[1227,1468,673,699]" italics="true" pageId="169" pageNumber="170">Tillandsia circinnata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount box="[498,603,712,738]" pageId="169" pageNumber="170" type="worker">1 worker</specimenCount>
|
||
(images of
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||
<collectorName box="[743,815,712,738]" pageId="169" pageNumber="170">FoCol</collectorName>
|
||
0411 examined on antweb.org) [
|
||
<collectionCode box="[1194,1283,712,739]" pageId="169" pageNumber="170">SMND</collectionCode>
|
||
] 1 teneral worker (images of
|
||
<location LSID="urn:lsid:plazi:treatment:054FDB70FF6FFF77B49C1CE57DFC2190:88393CBDFF6EFF73B71B18ED7B8E245D" box="[629,700,752,778]" name="FoCol" pageId="169" pageNumber="170">FoCol</location>
|
||
0412 examined on antweb.org) [
|
||
<collectionCode box="[1079,1170,752,779]" pageId="169" pageNumber="170">SMND</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
;
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||
<materialsCitation ID-GBIF-Occurrence="3321861320" collectionCode="USNM" collectorName="E. Skwarra" location="Smithsonian National Museum of Natural History Department" pageId="169" pageNumber="170" specimenCount="1" typeStatus="paralectotype">
|
||
same data as previous, except:
|
||
<collectorName box="[498,628,792,819]" pageId="169" pageNumber="170">E. Skwarra</collectorName>
|
||
#Z807a/
|
||
<collectorName box="[723,757,792,818]" pageId="169" pageNumber="170">Sk.</collectorName>
|
||
, ex
|
||
<taxonomicName box="[804,1045,792,818]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="circinnata">
|
||
<emphasis box="[804,1045,792,818]" italics="true" pageId="169" pageNumber="170">Tillandsia circinnata</emphasis>
|
||
</taxonomicName>
|
||
(images of
|
||
<collectorName box="[1185,1243,792,818]" name="Menozzi C" pageId="169" pageNumber="170">M.C.</collectorName>
|
||
Z. cotype 4-5 16358, S.N.M.U. type no. co 59607, USNMENT00532047 examined on the
|
||
<location LSID="urn:lsid:plazi:treatment:054FDB70FF6FFF77B49C1CE57DFC2190:88393CBDFF6EFF73B069195D7AE324D5" name="Smithsonian National Museum of Natural History Department" pageId="169" pageNumber="170">Smithsonian National Museum of Natural History Department</location>
|
||
of Entomology Collections website) [
|
||
<collectionCode box="[1420,1512,871,898]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="169" pageNumber="170" type="Museum">USNM</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="169" pageNumber="170" type="distribution">
|
||
<paragraph blockId="169.[498,1542,234,1934]" pageId="169" pageNumber="170">
|
||
<emphasis bold="true" box="[530,833,911,937]" pageId="169" pageNumber="170">Geographic distribution:</emphasis>
|
||
Mid elevations of south-central
|
||
<collectingCountry box="[1219,1305,912,938]" name="Mexico" pageId="169" pageNumber="170">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion box="[1321,1421,912,939]" country="Mexico" name="Morelos" pageId="169" pageNumber="170">Morelos</collectingRegion>
|
||
) (
|
||
<figureCitation box="[508,623,951,978]" captionStart="Figure 107" captionStartId="148.[524,589,1194,1216]" captionTargetBox="[498,1541,235,1181]" captionTargetId="figure-181@148.[498,1534,234,1166]" captionTargetPageId="148" captionText="Figure 107 Geographical and elevational distribution of the fuscatus group. (A) Temnothorax fuscatus (B) T. nebliselva sp. nov. (C) T. ocarinae (D) T. skwarrae. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-107" figureDoi="http://doi.org/10.5281/zenodo.5102311" httpUri="https://zenodo.org/record/5102311/files/figure.png" pageId="169" pageNumber="170">Fig. 107D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="170" lastPageNumber="171" pageId="169" pageNumber="170" type="diagnosis">
|
||
<paragraph blockId="169.[498,1542,234,1934]" pageId="169" pageNumber="170">
|
||
<emphasis bold="true" box="[530,751,991,1018]" pageId="169" pageNumber="170">Worker diagnosis:</emphasis>
|
||
<taxonomicName box="[757,1024,991,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="skwarrae">
|
||
<emphasis box="[757,1024,991,1017]" italics="true" pageId="169" pageNumber="170">Temnothorax skwarrae</emphasis>
|
||
</taxonomicName>
|
||
can be separated from all other species in the
|
||
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[498,573,1031,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis box="[498,573,1031,1057]" italics="true" pageId="169" pageNumber="170">salvini</emphasis>
|
||
</taxonomicName>
|
||
clade by the following character combination: antennal scapes short: failing to reach the posterior margin of the head by about the maximum width of the antennal scape (SI 92); body elongate (WLI> 150); metanotal groove not strongly impressed; propodeal spines directed posterodorsally, never directly upward; propodeal spines short, much shorter than the length of the propodeal declivity (PSI 18); hind femora strongly incrassate (FI> 350); petiolar node low, elongate and rounded dorsally (NI 244); postpetiole narrow: greater than one and a half times as wide as the petiole in dorsal view, but less than 2 times as wide (PWI 198); setae on head, mesosoma, waist segments and gaster erect, moderately long, moderately dense and blunt (never long and tapering); integument bicolored: dark brown with a yellow gaster.
|
||
</paragraph>
|
||
<paragraph blockId="169.[498,1542,234,1934]" lastBlockId="170.[498,1542,234,1934]" lastPageId="170" lastPageNumber="171" pageId="169" pageNumber="170">
|
||
<emphasis bold="true" box="[530,719,1429,1455]" pageId="169" pageNumber="170">Similar species:</emphasis>
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[727,1046,1430,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
|
||
<emphasis box="[727,1046,1430,1456]" italics="true" pageId="169" pageNumber="170">Temnothorax acutispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1055,1148,1430,1456]" pageId="169" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<emphasis italics="true" pageId="169" pageNumber="170">
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1161,1313,1430,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="subditivus">T. subditivus</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1326,1512,1430,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="tenuisculptus">T. tenuisculptus</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,640,1470,1496]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">T. tuxtlanus</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[649,742,1470,1496]" pageId="169" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, species of the
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[921,1017,1470,1495]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="annexus">
|
||
<emphasis box="[921,1017,1470,1495]" italics="true" pageId="169" pageNumber="170">annexus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[1032,1116,1470,1495]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[1032,1116,1470,1495]" italics="true" pageId="169" pageNumber="170">augusti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[1131,1223,1470,1496]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="fuscatus">
|
||
<emphasis box="[1131,1223,1470,1496]" italics="true" pageId="169" pageNumber="170">fuscatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1292,1367,1470,1496]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis box="[1292,1367,1470,1496]" italics="true" pageId="169" pageNumber="170">salvini</emphasis>
|
||
</taxonomicName>
|
||
groups.
|
||
<taxonomicName box="[498,768,1509,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="skwarrae">
|
||
<emphasis box="[498,768,1509,1535]" italics="true" pageId="169" pageNumber="170">Temnothorax skwarrae</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1092,1234,1509,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
|
||
<emphasis box="[1092,1234,1509,1535]" italics="true" pageId="169" pageNumber="170">T. tuxtlanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1243,1334,1510,1536]" pageId="169" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the incrassate femora (FI <
|
||
<quantity box="[654,726,1549,1575]" metricMagnitude="0" metricUnit="m" metricValue="7.112" pageId="169" pageNumber="170" unit="in" value="280.0">280 in</quantity>
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[734,874,1549,1575]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
|
||
<emphasis box="[734,874,1549,1575]" italics="true" pageId="169" pageNumber="170">T. tuxtlanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[881,974,1550,1576]" pageId="169" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). The low, elongate petiolar node (NI> 180) will separate
|
||
<taxonomicName box="[602,739,1589,1615]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="skwarrae">
|
||
<emphasis box="[602,739,1589,1615]" italics="true" pageId="169" pageNumber="170">T. skwarrae</emphasis>
|
||
</taxonomicName>
|
||
from all species listed above, aside from the
|
||
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1272,1347,1589,1615]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis box="[1272,1347,1589,1615]" italics="true" pageId="169" pageNumber="170">salvini</emphasis>
|
||
</taxonomicName>
|
||
group and most other members of the
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[829,921,1629,1655]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="fuscatus">
|
||
<emphasis box="[829,921,1629,1655]" italics="true" pageId="169" pageNumber="170">fuscatus</emphasis>
|
||
</taxonomicName>
|
||
group. The moderately long, blunt-tipped setae will separate
|
||
<taxonomicName box="[600,735,1669,1695]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="skwarrae">
|
||
<emphasis box="[600,735,1669,1695]" italics="true" pageId="169" pageNumber="170">T. skwarrae</emphasis>
|
||
</taxonomicName>
|
||
from the species of the
|
||
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1010,1085,1669,1695]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis box="[1010,1085,1669,1695]" italics="true" pageId="169" pageNumber="170">salvini</emphasis>
|
||
</taxonomicName>
|
||
group, which have long, tapering setae. The short propodeal spines, which are much shorter than the propodeal declivity, as well as the bicolored integument, will distinguish
|
||
<taxonomicName box="[1085,1222,1748,1774]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="skwarrae">
|
||
<emphasis box="[1085,1222,1748,1774]" italics="true" pageId="169" pageNumber="170">T. skwarrae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[1297,1422,1749,1775]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="fuscatus">
|
||
<emphasis box="[1297,1422,1749,1775]" italics="true" pageId="169" pageNumber="170">T. fuscatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,642,1788,1814]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="nebliselva" status="sp. nov.">
|
||
<emphasis box="[498,642,1788,1814]" italics="true" pageId="169" pageNumber="170">T. nebliselva</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[651,744,1789,1815]" pageId="169" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, which have longer propodeal spines and are uniformly colored.
|
||
<taxonomicName box="[498,768,1828,1854]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="skwarrae">
|
||
<emphasis box="[498,768,1828,1854]" italics="true" pageId="169" pageNumber="170">Temnothorax skwarrae</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName authorityName="" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1047,1179,1829,1854]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="ocarinae">
|
||
<emphasis box="[1047,1179,1829,1854]" italics="true" pageId="169" pageNumber="170">T. ocarinae</emphasis>
|
||
</taxonomicName>
|
||
by the relatively large, oval compound eye, which is about as long as the distance between the compound eye and the mandibular insertion (shorter than the distance between the compound eye and mandibular insertion in
|
||
<taxonomicName authorityName="" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[783,913,235,260]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="species" species="ocarinae">
|
||
<emphasis box="[783,913,235,260]" italics="true" pageId="170" pageNumber="171">T. ocarinae</emphasis>
|
||
</taxonomicName>
|
||
). Furthermore,
|
||
<taxonomicName box="[1099,1234,234,260]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="species" species="skwarrae">
|
||
<emphasis box="[1099,1234,234,260]" italics="true" pageId="170" pageNumber="171">T. skwarrae</emphasis>
|
||
</taxonomicName>
|
||
has a bright yellow gaster, as opposed to medium brown in
|
||
<taxonomicName authorityName="" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[892,1024,275,300]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="species" species="ocarinae">
|
||
<emphasis box="[892,1024,275,300]" italics="true" pageId="170" pageNumber="171">T. ocarinae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="173" lastPageNumber="174" pageId="170" pageNumber="171" type="description">
|
||
<paragraph blockId="170.[498,1542,234,1934]" pageId="170" pageNumber="171">
|
||
<emphasis bold="true" box="[530,1032,313,340]" pageId="170" pageNumber="171">
|
||
Worker measurements & indices (
|
||
<emphasis bold="true" box="[950,966,314,340]" italics="true" pageId="170" pageNumber="171">n</emphasis>
|
||
= 1):
|
||
</emphasis>
|
||
SL = 0.737; FRS = 0.247; CW = 0.857; CWb = 0.804; PoOC = 0.387; CL = 0.965; EL = 0.219; EW = 0.174; MD = 0.218; WL = 1.270; SPST = 0.231; MPST = 0.368; PEL = 0.388; NOL = 0.342; NOH = 0.140; PEH = 0.248; PPL = 0.264; PPH = 0.291; PW = 0.558; SBPA = 0.164; SPTI = 0.189; PEW = 0.177; PNW = 0.145; PPW = 0.350; HFL = 0.880; HFWmax = 0.255; HFWmin = 0.069; CS = 1.287; ES = 0.306; SI = 92; OI = 24; CI = 83; WLI = 158; SBI = 20; PSI = 18; PWI = 198; PLI = 147; NI = 244; PNWI = 82; NLI = 88; FI = 370.
|
||
</paragraph>
|
||
<paragraph blockId="170.[498,1542,234,1934]" pageId="170" pageNumber="171">
|
||
<emphasis bold="true" box="[530,777,593,619]" pageId="170" pageNumber="171">Worker description:</emphasis>
|
||
In full-face view, head subquadrate, longer than broad (CI 83). Mandibles densely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal weakly emarginate medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about the width of the antennal scape (SI 92). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment slightly longer than the preceding two in combination. Frontal carinae moderately long, extending past the antennal toruli by about two times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head weakly convex, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin very weakly concave but rounding evenly into the lateral margins.
|
||
</paragraph>
|
||
<paragraph blockId="170.[498,1542,234,1934]" lastBlockId="171.[498,1542,230,1934]" lastPageId="171" lastPageNumber="172" pageId="170" pageNumber="171">
|
||
In profile view, compound eyes ovular and moderately large (OI 24), with 14 ommatidia in longest row. Pronotal declivity distinct, neck and anterior face of pronotum forming a rounded ~90
|
||
<emphasis box="[672,682,1186,1204]" italics="true" pageId="170" pageNumber="171">
|
||
<superScript attach="left" box="[672,682,1186,1204]" fontSize="8" pageId="170" pageNumber="171">°</superScript>
|
||
</emphasis>
|
||
angle; transition between the anterior and dorsal faces delimited by a transverse carina. Dorsal face of pronotum evenly rounding into the rest of the mesosoma, which is flat to the propodeal spines. Promesonotal suture extending from the posterior margin of the procoxal insertion only to the mesothoracic spiracle, which is moderately well developed. Metanotal groove visible as a disruption of the sculpture laterally from where it arises between the mid- and hind coxae to where it ends in the poorly developed metathoracic spiracle, which is nearly indistinguishable against the ground sculpture. Propodeal spiracle moderately well developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines weakly developed and short (PSI 18), about a third as long as the propodeal declivity, tapering evenly from the base, weakly downcurved, and acute. Propodeal declivity weakly concave, forming a rounded ~120
|
||
<emphasis box="[1092,1102,1624,1642]" italics="true" pageId="170" pageNumber="171">
|
||
<superScript attach="left" box="[1092,1102,1624,1642]" fontSize="8" pageId="170" pageNumber="171">°</superScript>
|
||
</emphasis>
|
||
angle with the base of the propodeal spines. Propodeal lobes dorsally angulate and small. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole moderately long (PLI 147), with tubercles anterodorsally. Subpetiolar process in the form of a very small, blunt tooth; ventral margin of petiole flat posterior to it. Petiolar peduncle short: comprising about a quarter of the total length of the petiole. Petiolar node low and dorsally rounded transition between peduncle and node an even grade, resulting in a weakly concave anterior node face; anterior face meeting the dorsal face at a rounded ~120
|
||
<emphasis box="[557,567,230,248]" italics="true" pageId="171" pageNumber="172">
|
||
<superScript attach="left" box="[557,567,230,248]" fontSize="8" pageId="171" pageNumber="172">°</superScript>
|
||
</emphasis>
|
||
angle; dorsal face rounding evenly into the posterior face. Postpetiole long; anterior and dorsal faces forming an even convexity; lobed ventrally.
|
||
</paragraph>
|
||
<paragraph blockId="171.[498,1542,230,1934]" pageId="171" pageNumber="172">In dorsal view, humeri developed and distinct: evenly rounded and wider than the rest of the mesosoma; mesothoracic spiracles weakly protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Metanotal groove absent: mesonotum and propodeum completely fused and lateral margins converging evenly to the bases of the propodeal spines. Propodeal spines narrowly approximated basally and weakly diverging apically, their apices separated from each other by about their length, the negative space between them broadly “U” shaped. Petiolar peduncle with spiracles not protruding past the lateral margins. Petiolar node ovular, slightly longitudinally elongate, and narrowed anteriorly; node the same width as the peduncle and the caudal cylinder. Postpetiole narrow (PWI 198) and campaniform. Anterior margin of the postpetiole convex and evenly rounds into the lateral margins, which are parallel to each other; posterior corners angulate; posterior margin weakly emarginate medially. Metafemur strongly incrassate (FI 370).</paragraph>
|
||
<paragraph blockId="171.[498,1542,230,1934]" pageId="171" pageNumber="172">Sculpture: median clypeal carina present, extending posteriorly nearly to the frontal triangle, and flanked on either side by two equally strong carinae. Lateral clypeal lobes with additional, weaker carinae; ground sculpture areolate. Antennal scapes areolate. Cephalic dorsum densely areolate, with overlying reticulate sculpture; cross-reticulations weaker between the frontal carinae. Lateral surfaces of head densely areolate, with dense reticulate sculpture overlying the ground sculpture. Ventral surface of head longitudinally coarsely areolate-costulate. Pronotal neck areolate. Lateral surface of the mesosoma densely, finely areolate, with coarse longitudinal striae overlying the ground sculpture. Propodeal declivity weakly transversely areolate-costulate. Dorsal surface of mesosoma densely areolate, with coarse rugose sculpture overlying the ground sculpture on the pronotum and propodeum; mesonotal area without overlying sculpture. Femora areolate. Petiole densely areolate, with a coarse longitudinal carina extending from the anterodorsal flange to the caudal cylinder; reticulate sculpture overlying the areolate ground sculpture dorsal to the carina. Postpetiole densely areolate, with reticulate sculpture overlying the ground sculpture. First gastral sternite uniformly weakly areolate, with weak spectral iridescence. First gastral sternite smooth and shining, without spectral iridescence.</paragraph>
|
||
<paragraph blockId="171.[498,1542,230,1934]" pageId="171" pageNumber="172">Setae: antennal scapes and funiculi with short, decumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with sparse, erect, blunt-tipped setae, the longest of which are about the length of the compound eye. The head bears ~34, mesosoma ~30, petiole 8, postpetiole ~16, and first gastral tergite ~30 setae. Short, sparse pubescence present over the entire body, but difficult to detect against the densely sculptured integument.</paragraph>
|
||
<paragraph blockId="171.[498,1542,230,1934]" pageId="171" pageNumber="172">Color: bicolored. Predominantly dark brown, with a testaceous yellow gaster. Mandibles and legs testaceous brown.</paragraph>
|
||
<paragraph blockId="171.[498,1542,230,1934]" lastBlockId="172.[498,1542,234,1934]" lastPageId="172" lastPageNumber="173" pageId="171" pageNumber="172">
|
||
<emphasis bold="true" box="[530,1001,1788,1815]" pageId="171" pageNumber="172">
|
||
Gyne measurements & indices (
|
||
<emphasis bold="true" box="[919,935,1789,1815]" italics="true" pageId="171" pageNumber="172">n</emphasis>
|
||
= 1):
|
||
</emphasis>
|
||
SL = 0.751; FRS = 0.335; CW = 1.056; CWb = 1.005; PoOC = 0.456; CL = 1.121; EL = 0.286; EW = 0.217; MD = 0.220; WL = 1.881; SPST = 0.430; MPST = 0.497; PEL = 0.525; NOL = 0.287; NOH = 0.239; PEH = 0.387; PPL = 0.379; PPH = 0.437; PW = 1.018; SBPA = 0.417; SPTI = 0.392; PEW = 0.263; PNW = 0.186; PPW = 0.576; HFL = 0.979; HFWmax = 0.267; HFWmin = 0.078; CS = 1.566; ES = 0.395; SI = 75; OI = 25; CI = 90; WLI = 187; SBI = 41; PSI = 23; PWI = 219; PLI = 139; NI = 120; PNWI = 71; NLI = 55; FI = 342.
|
||
</paragraph>
|
||
<paragraph blockId="172.[498,1542,234,1934]" pageId="172" pageNumber="173">
|
||
<emphasis bold="true" box="[530,747,353,379]" pageId="172" pageNumber="173">Gyne description:</emphasis>
|
||
In full-face view, head subquadrate, longer than broad (CI 90). Mandibles densely striate but shining and armed with five teeth: the apical-most well developed, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin weakly emarginated medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about three times the maximum width of the scape (SI 75). Antennae 12-segmented; antennal club composed of three segments, with the apical-most segment slightly longer than the preceding two in combination. Frontal carinae moderately long, extending past the antennal toruli by about three times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head convex posterior to the compound eyes, flat below the compound eyes to the mandibular insertions. Posterior head margin flat, rounding evenly into the lateral margins.
|
||
</paragraph>
|
||
<paragraph blockId="172.[498,1542,234,1934]" pageId="172" pageNumber="173">
|
||
In profile view, compound eyes ovular and moderately large (OI 25), with 17 ommatidia in longest row. Mesoscutum rounded evenly anteriorly, not fully covering the dorsal surface of the pronotum, and flat dorsally. Mesoscutellum on the same level as the mesoscutum but sloping posteriorly. Metanotum on the same plane as the mesoscutellum. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines stout and well developed, but short (PSI 23), about a third as long as the propodeal declivity, tapering evenly from the base, directed posterodorsally, straight, and acute. Propodeal declivity straight and flat, forming a rounded ~110
|
||
<emphasis box="[991,1001,1186,1204]" italics="true" pageId="172" pageNumber="173">
|
||
<superScript attach="left" box="[991,1001,1186,1204]" fontSize="8" pageId="172" pageNumber="173">°</superScript>
|
||
</emphasis>
|
||
angle with the base of the propodeal spines. Propodeal lobes dorsally angulate and small. Metapleural gland bulla moderately large, extending from the metacoxal insertion two thirds of the way to the propodeal spiracle. Petiole moderately long (PLI 139), with tubercles anterodorsally. Subpetiolar process in the form of a small, acute tooth, which grades evenly into the ventral margin of the petiole posteriorly. Petiolar peduncle short: comprising less than a quarter of the total length of the petiole. Petiolar node erect and cuneiform: transition between peduncle and node evenly rounded, resulting in a concave anterior node face; anterior face forming a very sharp ~90
|
||
<emphasis box="[615,625,1505,1523]" italics="true" pageId="172" pageNumber="173">
|
||
<superScript attach="left" box="[615,625,1505,1523]" fontSize="8" pageId="172" pageNumber="173">°</superScript>
|
||
</emphasis>
|
||
angle with the dorsal face, which rounds evenly into the posterior face. Caudal cylinder long, about one and a half times the maximum width of the antennal scape. Postpetiole evenly rounded anterodorsally, bulging slightly before it transitions into the flattened dorsal face; ventral surface lobed.
|
||
</paragraph>
|
||
<paragraph blockId="172.[498,1542,234,1934]" lastBlockId="173.[498,1542,234,1735]" lastPageId="173" lastPageNumber="174" pageId="172" pageNumber="173">In dorsal view, mesoscutum not fully covering the pronotum anteriorly; humeri visible laterally as angulate sclerites. Propodeal spines flat dorsally, weakly diverging apically, their apices separated from each other by about two times their length. Petiolar peduncle with spiracles not protruding past the lateral margins. Petiolar node, when viewed posterodorsally, subquadrate: apex narrower apically than the base, which is the same width as the caudal cylinder and peduncle. Postpetiole moderately broad (PWI 219) and campaniform. Anterior margin of postpetiole convex, rounding evenly into the lateral margins, which are parallel to each other; posterior corners angulate; posterior margin flat. Metafemur strongly incrassate (FI 342).</paragraph>
|
||
<paragraph blockId="173.[498,1542,234,1735]" pageId="173" pageNumber="174">Sculpture: median clypeal carina present, extending from the anterior margin nearly to frontal triangle, and flanked by two equally strong carinae; lateral margins of median clypeal lobe with two carinae that are as strong as the median carina. Lateral clypeal lobes with additional weaker carinae; ground sculpture finely areolate. Antennal scapes areolate-costulate. Cephalic dorsum densely areolate, with overlying rugose sculpture, becoming costate between the frontal carinae. Lateral surfaces of head densely areolate, with dense predominantly costate sculpture overlying the ground sculpture. Ventral surface of head areolate-costulate. Pronotal neck areolate. Pronotum areolate anteriorly, lateral face areolate, with coarse costae overlying the ground sculpture. Anepisternum and katepisternum shining on their anterior quarters, transitioning into areolate-costulate sculpture posteriorly. Metapleuron and propodeum costate. Propodeal declivity weakly areolate. Mesoscutum and mesoscutellum costate, with areolate-rugulose sculpture between them. Metanotum areolate. Dorsum of propodeum strigulate over areolate ground sculpture. Femora weakly areolate. Petiole weakly areolate ventrally, as well as on the dorsal surface of the peduncle and node; coarse longitudinal carina extending from the anterodorsal flange to the caudal cylinder; rugose sculpture overlying the areolate ground sculpture dorsal to the carina. Postpetiole areolate, with concentric costulae overlying the ground sculpture. First gastral tergite densely, finely areolate, without spectral iridescence. Surface of the first gastral sternite smooth and shining, without spectral iridescence.</paragraph>
|
||
<paragraph blockId="173.[498,1542,234,1735]" pageId="173" pageNumber="174">Setae: antennal scapes and funiculi with short, decumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with sparse, erect, blunt-tipped setae, the longest of which are about half the width of the compound eye. Short, sparse pubescence present over the entire body, but difficult to detect against the densely sculptured integument.</paragraph>
|
||
<paragraph blockId="173.[498,1542,234,1735]" pageId="173" pageNumber="174">Color: bicolored. Predominantly dark brown, with a testaceous yellow gaster. Mandibles and legs testaceous brown.</paragraph>
|
||
<paragraph blockId="173.[498,1542,234,1735]" box="[530,730,1350,1376]" pageId="173" pageNumber="174">
|
||
<emphasis bold="true" box="[530,597,1350,1376]" pageId="173" pageNumber="174">Male:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[530,1517,1390,1416]" pageId="173" pageNumber="174" type="etymology">
|
||
<paragraph blockId="173.[498,1542,234,1735]" box="[530,1517,1390,1416]" pageId="173" pageNumber="174">
|
||
<emphasis bold="true" box="[530,668,1390,1416]" pageId="173" pageNumber="174">Etymology:</emphasis>
|
||
Matronym, named for Elizabeth Skwarra, who collected the
|
||
<typeStatus box="[1388,1438,1390,1416]" pageId="173" pageNumber="174">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="173" pageNumber="174" type="discussion">
|
||
<paragraph blockId="173.[498,1542,234,1735]" pageId="173" pageNumber="174">
|
||
<emphasis bold="true" box="[530,670,1429,1455]" pageId="173" pageNumber="174">Comments:</emphasis>
|
||
The
|
||
<typeStatus box="[732,782,1430,1456]" pageId="173" pageNumber="174">type</typeStatus>
|
||
series of
|
||
<taxonomicName box="[893,1162,1430,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="skwarrae">
|
||
<emphasis box="[893,1162,1430,1456]" italics="true" pageId="173" pageNumber="174">Temnothorax skwarrae</emphasis>
|
||
</taxonomicName>
|
||
was collected from the epiphyte
|
||
<taxonomicName box="[498,740,1470,1496]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="173" pageNumber="174" phylum="Tracheophyta" rank="species" species="circinnata">
|
||
<emphasis box="[498,740,1470,1496]" italics="true" pageId="173" pageNumber="174">Tillandsia circinnata</emphasis>
|
||
</taxonomicName>
|
||
near Cuernavaca in
|
||
<collectingRegion box="[986,1146,1470,1496]" country="Mexico" name="Morelos" pageId="173" pageNumber="174">Morelos state</collectingRegion>
|
||
,
|
||
<collectingCountry box="[1159,1248,1470,1496]" name="Mexico" pageId="173" pageNumber="174">Mexico</collectingCountry>
|
||
. Very little is known about the biology of this species, but it resembles
|
||
<taxonomicName authorityName="" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1087,1219,1510,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="ocarinae">
|
||
<emphasis box="[1087,1219,1510,1535]" italics="true" pageId="173" pageNumber="174">T. ocarinae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion box="[1293,1432,1509,1536]" country="Mexico" name="Jalisco" pageId="173" pageNumber="174">Jalisco state</collectingRegion>
|
||
, and the cloud forest species
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[735,860,1549,1575]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="fuscatus">
|
||
<emphasis box="[735,860,1549,1575]" italics="true" pageId="173" pageNumber="174">T. fuscatus</emphasis>
|
||
</taxonomicName>
|
||
, which has a range in Central America from south of the Isthmus Tehuantepec to
|
||
<collectingCountry box="[837,966,1589,1615]" name="Costa Rica" pageId="173" pageNumber="174">Costa Rica</collectingCountry>
|
||
. This species, like
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[1184,1309,1589,1615]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="fuscatus">
|
||
<emphasis box="[1184,1309,1589,1615]" italics="true" pageId="173" pageNumber="174">T. fuscatus</emphasis>
|
||
</taxonomicName>
|
||
, is apparently arboreally nesting.
|
||
<taxonomicName box="[721,989,1629,1655]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="skwarrae">
|
||
<emphasis box="[721,989,1629,1655]" italics="true" pageId="173" pageNumber="174">Temnothorax skwarrae</emphasis>
|
||
</taxonomicName>
|
||
has also been reported from tropical dry forest habitat in Estacion de Biologia Chamela in Jalisco state (
|
||
<emphasis box="[1147,1358,1669,1695]" italics="true" pageId="173" pageNumber="174">Dáttilo et al., 2020</emphasis>
|
||
), but I have not personally investigated the specimens, so I exclude them here.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |