671 lines
72 KiB
XML
671 lines
72 KiB
XML
<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FF69FF69B49C1CE07CB322EA" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax achii Prebus 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="180" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="175" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prebus, Matthew M.</mods:namePart>
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<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States & Department of Entomology & Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
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<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PeerJ</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="series">
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<mods:title>e 11514</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2021-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>9</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>462</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
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<mods:identifier type="ISSN">2167-8359</mods:identifier>
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<mods:identifier type="PMC">PMC8254503</mods:identifier>
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<mods:identifier type="PubMed">34249486</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102527" ID-GBIF-Taxon="183542612" ID-Zenodo-Dep="5102527" LSID="urn:lsid:zoobank.org:act:0340AA27-740C-47D9-9925-CEE1136FD8BF" httpUri="http://treatment.plazi.org/id/054FDB70FF69FF69B49C1CE07CB322EA" lastPageId="179" lastPageNumber="180" pageId="174" pageNumber="175">
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||
<subSubSection box="[498,832,1789,1816]" pageId="174" pageNumber="175" type="nomenclature">
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||
<paragraph blockId="174.[498,982,1789,1856]" box="[498,832,1789,1816]" pageId="174" pageNumber="175">
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||
<heading box="[498,832,1789,1816]" level="2" pageId="174" pageNumber="175">
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<emphasis bold="true" box="[498,832,1789,1816]" pageId="174" pageNumber="175">
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<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,728,1789,1816]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
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<emphasis bold="true" box="[498,728,1789,1816]" italics="true" pageId="174" pageNumber="175">Temnothorax achii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[737,832,1789,1815]" pageId="174" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[498,982,1829,1856]" pageId="174" pageNumber="175" type="distribution">
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<paragraph blockId="174.[498,982,1789,1856]" box="[498,982,1829,1856]" pageId="174" pageNumber="175">
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Distribution:
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<figureCitation box="[657,770,1829,1855]" captionStart="Figure 114" captionStartId="174.[524,589,1195,1217]" captionTargetBox="[498,1541,234,1182]" captionTargetId="figure-143@174.[498,1534,234,1167]" captionTargetPageId="174" captionText="Figure 114 Geographical and elevational distribution of the goniops group. (A) Temnothorax achii sp. nov. (B) T. goniops (C) T. ixili (D) T. xincai sp. nov. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Map for T. huehuetenangoi not included due to the uncertainty of the collection locality. Full-size DOI: 10.7717/peerj.11514/fig-114" figureDoi="http://doi.org/10.5281/zenodo.5102329" httpUri="https://zenodo.org/record/5102329/files/figure.png" pageId="174" pageNumber="175">Fig. 114A</figureCitation>
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; worker:
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<figureCitation box="[882,976,1829,1856]" captionStart="Figure 115" captionStartId="175.[524,589,1138,1160]" captionTargetBox="[498,1541,234,1125]" captionTargetId="figure-197@175.[498,1295,234,1109]" captionTargetPageId="175" captionText="Figure 115 Temnothorax achii sp. nov. holotype worker (JTLC000009876). (A) Profile view. (B) Full-face view. (C) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-115" figureDoi="http://doi.org/10.5281/zenodo.5102333" httpUri="https://zenodo.org/record/5102333/files/figure.png" pageId="174" pageNumber="175">Fig. 115</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection box="[498,1117,1877,1903]" pageId="174" pageNumber="175" type="reference_group">
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<paragraph blockId="174.[498,1117,1877,1903]" box="[498,1117,1877,1903]" pageId="174" pageNumber="175">
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<treatmentCitationGroup box="[498,1117,1877,1903]" pageId="174" pageNumber="175">
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<taxonomicName authorityName="Mayr" authorityYear="1861" box="[498,655,1877,1903]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="genus">
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<emphasis box="[498,655,1877,1903]" italics="true" pageId="174" pageNumber="175">Temnothorax</emphasis>
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</taxonomicName>
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mmp04
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<treatmentCitation author="Prebus M." box="[763,1114,1877,1903]" page="8" pageId="174" pageNumber="175" year="2017">
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<bibRefCitation author="Prebus M." box="[763,942,1877,1903]" journalOrPublisher="BMC Evolutionary Biology" pageId="174" pageNumber="175" pagination="250" part="17" refId="ref272195" refString="Prebus M. 2017. Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae). BMC Evolutionary Biology 17 (1): 250 DOI 10.1186 / s 12862 - 017 - 1095 - 8." title="Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae)" type="journal article" year="2017">
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<emphasis box="[763,911,1877,1903]" italics="true" pageId="174" pageNumber="175">Prebus, 2017</emphasis>
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: 8
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</bibRefCitation>
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. In phylogeny
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102333" ID-Zenodo-Dep="5102333" httpUri="https://zenodo.org/record/5102333/files/figure.png" pageId="175" pageNumber="176" startId="175.[524,589,1138,1160]" targetBox="[498,1541,234,1125]" targetPageId="175">
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<paragraph blockId="175.[524,1516,1137,1189]" pageId="175" pageNumber="176">
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<emphasis bold="true" box="[524,1328,1137,1160]" pageId="175" pageNumber="176">
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Figure 115
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<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[651,846,1137,1159]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
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<emphasis bold="true" box="[651,846,1137,1159]" italics="true" pageId="175" pageNumber="176">Temnothorax achii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[861,945,1138,1159]" pageId="175" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
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holotype worker (JTLC000009876).
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</emphasis>
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(A) Profile view. (B) Full-face view. (C) Dorsal view. Scale bars
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||
<quantity box="[969,1044,1167,1188]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="175" pageNumber="176" unit="mm" value="0.2">0.2 mm</quantity>
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. Full-size DOI: 10.7717/peerj.11514/fig-115
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</paragraph>
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</caption>
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<subSubSection pageId="175" pageNumber="176" type="materials_examined">
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<paragraph blockId="175.[498,1542,1244,1908]" pageId="175" pageNumber="176">
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<emphasis bold="true" box="[530,834,1244,1270]" pageId="175" pageNumber="176">Type material examined:</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3321861760" collectingDate="2007-07-07" collectionCode="CASC" collectorName="J. Longino" country="Guatemala" elevation="460" latitude="14.9051" location="Morazan" longLatPrecision="5" longitude="-90.15362" pageId="175" pageNumber="176" specimenCode="6002-s, JTLC000009876" specimenCount="1" stateProvince="El Progreso" typeStatus="holotype">
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<emphasis box="[843,1041,1244,1270]" italics="true" pageId="175" pageNumber="176">
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||
<typeStatus box="[843,946,1244,1270]" pageId="175" pageNumber="176">Holotype</typeStatus>
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<specimenCount box="[954,1034,1244,1270]" pageId="175" pageNumber="176" type="worker">worker</specimenCount>
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:
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</emphasis>
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<collectingCountry box="[1049,1230,1244,1270]" name="Guatemala" pageId="175" pageNumber="176">GUATEMALA</collectingCountry>
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:
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<collectingRegion box="[1243,1381,1244,1271]" country="Guatemala" name="El Progreso" pageId="175" pageNumber="176">El Progreso</collectingRegion>
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||
:
|
||
<locationDeviation box="[1393,1525,1244,1270]" pageId="175" pageNumber="176">
|
||
<quantity box="[1393,1474,1244,1270]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="175" pageNumber="176" unit="km" value="3.7">3.7 km</quantity>
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SW
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||
</locationDeviation>
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||
<location LSID="urn:lsid:plazi:treatment:054FDB70FF69FF69B49C1CE07CB322EA:88393CBDFF68FF75B49C1F197B6F2249" box="[498,605,1284,1310]" country="Guatemala" latitude="14.9051" longLatPrecision="5" longitude="-90.15362" name="Morazan" pageId="175" pageNumber="176" stateProvince="El Progreso">Morazán</location>
|
||
,
|
||
<geoCoordinate box="[618,742,1279,1311]" degrees="14.9051" direction="north" orientation="latitude" pageId="175" pageNumber="176" precision="5" value="14.9051">
|
||
14.9051
|
||
<emphasis box="[708,718,1279,1297]" italics="true" pageId="175" pageNumber="176">
|
||
<superScript attach="right" box="[708,718,1279,1297]" fontSize="8" pageId="175" pageNumber="176">°</superScript>
|
||
</emphasis>
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||
N
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||
</geoCoordinate>
|
||
-
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||
<geoCoordinate box="[758,906,1279,1311]" degrees="90.15362" direction="west" orientation="longitude" pageId="175" pageNumber="176" precision="1" value="-90.15362">
|
||
90.15362
|
||
<emphasis box="[865,875,1279,1297]" italics="true" pageId="175" pageNumber="176">
|
||
<superScript attach="right" box="[865,875,1279,1297]" fontSize="8" pageId="175" pageNumber="176">°</superScript>
|
||
</emphasis>
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||
W
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||
</geoCoordinate>
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||
,
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||
<quantity box="[920,996,1284,1311]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="175" pageNumber="176" unit="m" value="460.0">
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||
<elevation box="[920,996,1284,1311]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="175" pageNumber="176" unit="m" value="460.0">460 m</elevation>
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||
</quantity>
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||
,
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<date box="[1009,1144,1284,1311]" pageId="175" pageNumber="176" value="2007-07-07">
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||
<collectingDate box="[1009,1144,1284,1311]" pageId="175" pageNumber="176" value="2007-07-07">7 July 2007</collectingDate>
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||
</date>
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,
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<collectorName box="[1157,1279,1284,1311]" pageId="175" pageNumber="176">J. Longino</collectorName>
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<specimenCode box="[1288,1381,1284,1310]" pageId="175" pageNumber="176">#6002-s</specimenCode>
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, tropical dry forest, ex sifted leaf litter (
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<specimenCode box="[806,999,1324,1351]" collectionCode="JTLC" pageId="175" pageNumber="176">JTLC000009876</specimenCode>
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) [
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<collectionCode box="[1022,1103,1323,1350]" pageId="175" pageNumber="176">CASC</collectionCode>
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||
].
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</materialsCitation>
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||
</paragraph>
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<paragraph blockId="175.[498,1542,1244,1908]" pageId="175" pageNumber="176">
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<materialsCitation ID-GBIF-Occurrence="3321861543" collectingDate="2007-07-07" collectionCode="MCZC, USNM, LACM, UVGC, UCDC" collectorName="M. G. Branstetter" country="Guatemala" elevation="460" latitude="14.9051" location="Morazan" longLatPrecision="5" longitude="-90.15362" pageId="175" pageNumber="176" specimenCode="JTLC000009871, JTLC000009906" specimenCount="5" specimenCount-worker="5" stateProvince="El Progreso" typeStatus="paratype">
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<emphasis box="[530,735,1364,1390]" italics="true" pageId="175" pageNumber="176">
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<typeStatus box="[530,632,1364,1389]" pageId="175" pageNumber="176">Paratype</typeStatus>
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workers:
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</emphasis>
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same data as holotype,
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<specimenCount box="[1008,1110,1364,1390]" pageId="175" pageNumber="176" type="worker">1 worker</specimenCount>
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(
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<specimenCode box="[1124,1318,1364,1390]" collectionCode="JTLC" pageId="175" pageNumber="176">JTLC000009871</specimenCode>
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) [
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<collectionCode box="[1338,1426,1363,1390]" pageId="175" pageNumber="176">MCZC</collectionCode>
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||
]
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<specimenCount box="[1439,1542,1364,1390]" pageId="175" pageNumber="176" type="worker">1 worker</specimenCount>
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(
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<specimenCode box="[506,699,1403,1430]" collectionCode="JTLC" pageId="175" pageNumber="176">JTLC000009906</specimenCode>
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||
) [
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<collectionCode box="[722,811,1403,1430]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="175" pageNumber="176" type="Museum">USNM</collectionCode>
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||
]
|
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<specimenCount box="[826,931,1403,1430]" pageId="175" pageNumber="176" type="worker">1 worker</specimenCount>
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||
(CASENT0869159) [
|
||
<collectionCode box="[1182,1270,1403,1430]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="175" pageNumber="176" type="Museum">LACM</collectionCode>
|
||
]
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||
<specimenCount box="[1285,1390,1403,1430]" pageId="175" pageNumber="176" type="worker">1 worker</specimenCount>
|
||
(CASENT0869160) [
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<collectionCode box="[741,831,1443,1470]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="175" pageNumber="176">UVGC</collectionCode>
|
||
];
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3321861678" collectingDate="2007-07-07" collectionCode="MCZC, USNM, LACM, UVGC, UCDC" collectorName="M. G. Branstetter" country="Guatemala" elevation="460" latitude="14.9051" location="Morazan" longLatPrecision="5" longitude="-90.15362" pageId="175" pageNumber="176" specimenCode="JTLC000009871, JTLC000009906" specimenCount="5" specimenCount-worker="5" stateProvince="El Progreso" typeStatus="paratype">
|
||
same data as
|
||
<typeStatus box="[1009,1113,1443,1469]" pageId="175" pageNumber="176">holotype</typeStatus>
|
||
, except:
|
||
<collectorName box="[1215,1411,1443,1470]" pageId="175" pageNumber="176">M.G. Branstetter</collectorName>
|
||
#626-12, tropical dry scrub, ex sifted leaf litter,
|
||
<specimenCount box="[949,1054,1483,1509]" pageId="175" pageNumber="176" type="worker">1 worker</specimenCount>
|
||
(CASENT0600922) [
|
||
<collectionCode box="[1305,1395,1483,1510]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="175" pageNumber="176" type="Museum">UCDC</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="175.[498,1542,1244,1908]" pageId="175" pageNumber="176">
|
||
<emphasis bold="true" box="[530,888,1523,1549]" pageId="175" pageNumber="176">Non-type material examined:</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3321861877" collectingDate="2014-04-27" collectionCode="UCDC" collectorName="M. Barrios" country="Guatemala" elevation="0" latitude="14.77968" location="Chiquimula" longLatPrecision="1" longitude="-89.67686" pageId="175" pageNumber="176" specimenCount="1" specimenCount-worker="1" stateProvince="Chiquimula">
|
||
<collectingCountry box="[897,1078,1523,1549]" name="Guatemala" pageId="175" pageNumber="176">GUATEMALA</collectingCountry>
|
||
:
|
||
<collectingRegion box="[1090,1232,1523,1549]" country="Guatemala" name="Chiquimula" pageId="175" pageNumber="176">Chiquimula</collectingRegion>
|
||
:
|
||
<quantity box="[1244,1319,1523,1550]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="175" pageNumber="176" unit="km" value="15.0">15 km</quantity>
|
||
W
|
||
<collectingRegion box="[1364,1505,1523,1549]" country="Guatemala" name="Chiquimula" pageId="175" pageNumber="176">Chiquimula</collectingRegion>
|
||
,
|
||
<geoCoordinate box="[498,636,1558,1589]" degrees="14.77968" direction="north" orientation="latitude" pageId="175" pageNumber="176" precision="1" value="14.77968">
|
||
14.77968
|
||
<emphasis box="[603,613,1558,1576]" italics="true" pageId="175" pageNumber="176">
|
||
<superScript attach="right" box="[603,613,1558,1576]" fontSize="8" pageId="175" pageNumber="176">°</superScript>
|
||
</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
<geoCoordinate box="[644,788,1558,1589]" degrees="89.67686" direction="west" orientation="longitude" pageId="175" pageNumber="176" precision="1" value="-89.67686">
|
||
89.67686
|
||
<emphasis box="[749,759,1558,1576]" italics="true" pageId="175" pageNumber="176">
|
||
<superScript attach="right" box="[749,759,1558,1576]" fontSize="8" pageId="175" pageNumber="176">°</superScript>
|
||
</emphasis>
|
||
W
|
||
</geoCoordinate>
|
||
±
|
||
<elevation box="[823,995,1563,1590]" isEnumeration="true" pageId="175" pageNumber="176">
|
||
<quantity box="[823,885,1563,1590]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="175" pageNumber="176" unit="m" value="50.0">50 m</quantity>
|
||
,
|
||
<quantity box="[898,995,1563,1590]" metricMagnitude="3" metricUnit="m" metricValue="1.735" pageId="175" pageNumber="176" unit="m" value="1735.0">1,735 m</quantity>
|
||
</elevation>
|
||
,
|
||
<date box="[1008,1173,1563,1589]" pageId="175" pageNumber="176" value="2014-04-27">
|
||
<collectingDate box="[1008,1173,1563,1589]" pageId="175" pageNumber="176" value="2014-04-27">27 April 2014</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1186,1311,1563,1589]" pageId="175" pageNumber="176">M. Barrios</collectorName>
|
||
#MB1314, second growth oak cloud forest, ex sifted leaf litter,
|
||
<specimenCount box="[1020,1125,1603,1629]" pageId="175" pageNumber="176" type="worker">1 worker</specimenCount>
|
||
(CASENT0758686) [
|
||
<collectionCode box="[1376,1465,1602,1629]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="175" pageNumber="176" type="Museum">UCDC</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3321861337" collectingDate="2003-11-19" collectionCode="UCDC" collectorName="P. S. Ward" country="Guatemala" elevation="1480" latitude="15.033333" location="La Cumbre" longLatPrecision="1" longitude="-90.21667" pageId="175" pageNumber="176" specimenCount="3" specimenCount-worker="3" stateProvince="Baja Verapaz">
|
||
<collectingRegion country="Guatemala" name="Baja Verapaz" pageId="175" pageNumber="176">Baja Verapaz</collectingRegion>
|
||
:
|
||
<quantity box="[608,666,1642,1669]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="175" pageNumber="176" unit="km" value="1.0">1 km</quantity>
|
||
SSW
|
||
<location LSID="urn:lsid:plazi:treatment:054FDB70FF69FF69B49C1CE07CB322EA:88393CBDFF68FF75B78E1C767A5721D3" box="[736,869,1642,1669]" country="Guatemala" latitude="15.033333" longLatPrecision="1" longitude="-90.21667" name="La Cumbre" pageId="175" pageNumber="176" stateProvince="Baja Verapaz">La Cumbre</location>
|
||
,
|
||
<geoCoordinate box="[879,1031,1638,1669]" degrees="15.033333" direction="north" orientation="latitude" pageId="175" pageNumber="176" precision="1" value="15.033333">
|
||
15.033333
|
||
<emphasis box="[998,1008,1638,1656]" italics="true" pageId="175" pageNumber="176">
|
||
<superScript attach="right" box="[998,1008,1638,1656]" fontSize="8" pageId="175" pageNumber="176">°</superScript>
|
||
</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
<geoCoordinate box="[1038,1197,1638,1669]" degrees="90.216667" direction="west" orientation="longitude" pageId="175" pageNumber="176" precision="1" value="-90.21667">
|
||
90.216667
|
||
<emphasis box="[1156,1166,1638,1656]" italics="true" pageId="175" pageNumber="176">
|
||
<superScript attach="right" box="[1156,1166,1638,1656]" fontSize="8" pageId="175" pageNumber="176">°</superScript>
|
||
</emphasis>
|
||
W
|
||
</geoCoordinate>
|
||
,
|
||
<quantity box="[1209,1303,1643,1669]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="175" pageNumber="176" unit="m" value="1480.0">
|
||
<elevation box="[1209,1303,1643,1669]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="175" pageNumber="176" unit="m" value="1480.0">1,480 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date box="[1315,1537,1642,1669]" pageId="175" pageNumber="176" value="2003-11-19">
|
||
<collectingDate box="[1315,1537,1642,1669]" pageId="175" pageNumber="176" value="2003-11-19">19 November 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[498,619,1682,1709]" pageId="175" pageNumber="176">P.S. Ward</collectorName>
|
||
#15097-14, mixed tropical/temperate mesic forest, sifted litter (leaf mold, rotten wood),
|
||
<specimenCount box="[667,783,1722,1748]" pageId="175" pageNumber="176" type="worker">3 workers</specimenCount>
|
||
(CASENT0915976) [
|
||
<collectionCode box="[1034,1124,1722,1749]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="175" pageNumber="176" type="Museum">UCDC</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="175" pageNumber="176" type="distribution">
|
||
<paragraph blockId="175.[498,1542,1244,1908]" pageId="175" pageNumber="176">
|
||
<emphasis bold="true" box="[530,755,1762,1788]" pageId="175" pageNumber="176">Geographic range:</emphasis>
|
||
Low-to-mid elevations in central
|
||
<collectingCountry box="[1157,1283,1762,1788]" name="Guatemala" pageId="175" pageNumber="176">Guatemala</collectingCountry>
|
||
(
|
||
<collectingRegion box="[1300,1459,1762,1789]" country="Guatemala" name="Baja Verapaz" pageId="175" pageNumber="176">Baja Verapaz</collectingRegion>
|
||
,
|
||
<collectingRegion box="[498,639,1802,1828]" country="Guatemala" name="Chiquimula" pageId="175" pageNumber="176">Chiquimula</collectingRegion>
|
||
,
|
||
<collectingRegion box="[652,790,1802,1829]" country="Guatemala" name="El Progreso" pageId="175" pageNumber="176">El Progreso</collectingRegion>
|
||
) (
|
||
<figureCitation box="[817,930,1802,1828]" captionStart="Figure 114" captionStartId="174.[524,589,1195,1217]" captionTargetBox="[498,1541,234,1182]" captionTargetId="figure-143@174.[498,1534,234,1167]" captionTargetPageId="174" captionText="Figure 114 Geographical and elevational distribution of the goniops group. (A) Temnothorax achii sp. nov. (B) T. goniops (C) T. ixili (D) T. xincai sp. nov. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Map for T. huehuetenangoi not included due to the uncertainty of the collection locality. Full-size DOI: 10.7717/peerj.11514/fig-114" figureDoi="http://doi.org/10.5281/zenodo.5102329" httpUri="https://zenodo.org/record/5102329/files/figure.png" pageId="175" pageNumber="176">Fig. 114A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="176" lastPageNumber="177" pageId="175" pageNumber="176" type="diagnosis">
|
||
<paragraph blockId="175.[498,1542,1244,1908]" lastBlockId="176.[498,1542,234,1935]" lastPageId="176" lastPageNumber="177" pageId="175" pageNumber="176">
|
||
<emphasis bold="true" box="[530,754,1842,1868]" pageId="175" pageNumber="176">Worker diagnosis:</emphasis>
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[761,982,1842,1868]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[761,982,1842,1868]" italics="true" pageId="175" pageNumber="176">Temnothorax achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[990,1081,1842,1868]" pageId="175" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be separated from all other species in the
|
||
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[575,650,1882,1908]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis box="[575,650,1882,1908]" italics="true" pageId="175" pageNumber="176">salvini</emphasis>
|
||
</taxonomicName>
|
||
clade by the following character combination: antennal scapes short, failing to reach the posterior margin of the head capsule by about the maximum width of the scape (SI 85–90); compound eyes small (OI 22–26); body moderately compact (WLI 137–143); metanotal groove absent; propodeum not strongly depressed; propodeal spines shorter than the propodeal declivity and dentate (PSI 25–28); propodeal spines broadly approximated in dorsal view (SBI 28–34); hind femora weakly to moderately incrassate (FI 180–303); petiolar node erect and dorsally rounded, not overhanging the caudal cylinder in profile view; petiolar node compact, not elongate (NI 143–170); petiolar node very slightly broader than caudal cylinder in dorsal view (PNWI <115); head sculpture areolate, with areolae arranged into columns by costulae; dorsum of postpetiole sculptured; antennal scapes with short, decumbent pilosity; setae on head, mesosoma, waist segments and gaster erect, short, sparse and blunt (never long and tapering); integument predominantly dark brown, with trochanters, meso- and metacoxae, tibiae, tarsi, and sting very light yellow.
|
||
</paragraph>
|
||
<paragraph blockId="176.[498,1542,234,1935]" pageId="176" pageNumber="177">
|
||
<emphasis bold="true" box="[530,717,752,778]" pageId="176" pageNumber="177">Similar species:</emphasis>
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[724,1041,752,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
|
||
<emphasis box="[724,1041,752,778]" italics="true" pageId="176" pageNumber="177">Temnothorax acutispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1049,1141,753,779]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1152,1291,752,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
|
||
<emphasis box="[1152,1291,752,778]" italics="true" pageId="176" pageNumber="177">T. altinodus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1298,1390,753,779]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1401,1542,752,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
|
||
<emphasis box="[1401,1542,752,778]" italics="true" pageId="176" pageNumber="177">T. tuxtlanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[498,590,793,819]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[602,749,792,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="subditivus">
|
||
<emphasis box="[602,749,792,818]" italics="true" pageId="176" pageNumber="177">T. subditivus</emphasis>
|
||
</taxonomicName>
|
||
, species of the
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[925,1009,793,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[925,1009,793,818]" italics="true" pageId="176" pageNumber="177">augusti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1022,1109,793,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="goniops">
|
||
<emphasis box="[1022,1109,793,818]" italics="true" pageId="176" pageNumber="177">goniops</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1173,1286,792,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis box="[1173,1286,792,818]" italics="true" pageId="176" pageNumber="177">pulchellus</emphasis>
|
||
</taxonomicName>
|
||
groups, species of the
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[498,555,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis box="[498,555,832,858]" italics="true" pageId="176" pageNumber="177">sallei</emphasis>
|
||
</taxonomicName>
|
||
clade.
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[639,861,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[639,861,832,858]" italics="true" pageId="176" pageNumber="177">Temnothorax achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[870,961,832,858]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from similar appearing members of the
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[690,747,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis box="[690,747,872,898]" italics="true" pageId="176" pageNumber="177">sallei</emphasis>
|
||
</taxonomicName>
|
||
clade (e.g.,
|
||
<emphasis box="[886,1359,872,898]" italics="true" pageId="176" pageNumber="177">
|
||
<taxonomicName box="[886,996,873,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="manni">T. manni</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1009,1165,873,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="mexicanus">T. mexicanus</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="" baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1178,1359,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="punctithorax">T. punctithorax</taxonomicName>
|
||
</emphasis>
|
||
) by the petiole, which lacks a transverse carina anterodorsally. Unlike the long, dorsally directed propodeal spines of
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[611,797,952,977]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
|
||
<emphasis box="[611,797,952,977]" italics="true" pageId="176" pageNumber="177">T. acutispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[806,899,952,978]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[912,1001,951,977]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[912,1001,951,977]" italics="true" pageId="176" pageNumber="177">T. achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1010,1101,952,978]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has short propodeal teeth that are directed posterodorsally.
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[790,1009,991,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[790,1009,991,1017]" italics="true" pageId="176" pageNumber="177">Temnothorax achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1015,1104,992,1018]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a moderately broad postpetiole in dorsal view, whereas
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[743,883,1031,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
|
||
<emphasis box="[743,883,1031,1057]" italics="true" pageId="176" pageNumber="177">T. tuxtlanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[890,980,1032,1058]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a relatively narrow postpetiole (PWI <150).
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,771,1071,1097]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
|
||
<emphasis box="[498,771,1071,1097]" italics="true" pageId="176" pageNumber="177">Temnothorax altinodus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[780,871,1072,1098]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has comparatively strongly incrassate hind femora (FI 327).
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[566,787,1111,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[566,787,1111,1137]" italics="true" pageId="176" pageNumber="177">Temnothorax achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[796,887,1111,1137]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be separated from
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1166,1315,1111,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="subditivus">
|
||
<emphasis box="[1166,1315,1111,1137]" italics="true" pageId="176" pageNumber="177">T. subditivus</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[1422,1506,1112,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[1422,1506,1112,1137]" italics="true" pageId="176" pageNumber="177">augusti</emphasis>
|
||
</taxonomicName>
|
||
group by the structure of the petiolar node, which is erect, dorsally rounded, and only slightly broader than the caudal cylinder in dorsal view, as opposed to squamiform in
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[498,647,1230,1256]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="subditivus">
|
||
<emphasis box="[498,647,1230,1256]" italics="true" pageId="176" pageNumber="177">T. subditivus</emphasis>
|
||
</taxonomicName>
|
||
, or leaning posteriorly over the caudal cylinder in the
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[1294,1378,1231,1256]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[1294,1378,1231,1256]" italics="true" pageId="176" pageNumber="177">augusti</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,720,1270,1296]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[498,720,1270,1296]" italics="true" pageId="176" pageNumber="177">Temnothorax achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[729,820,1271,1297]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from the
|
||
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1189,1302,1270,1296]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis box="[1189,1302,1270,1296]" italics="true" pageId="176" pageNumber="177">pulchellus</emphasis>
|
||
</taxonomicName>
|
||
group by the combination of a relatively longer mesosoma (WLI> 135 vs. <
|
||
<quantity box="[1248,1322,1310,1337]" metricMagnitude="0" metricUnit="m" metricValue="3.429" pageId="176" pageNumber="177" unit="in" value="135.0">135 in</quantity>
|
||
most
|
||
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1398,1511,1310,1336]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis box="[1398,1511,1310,1336]" italics="true" pageId="176" pageNumber="177">pulchellus</emphasis>
|
||
</taxonomicName>
|
||
group members). Some members of the
|
||
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[969,1082,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis box="[969,1082,1350,1376]" italics="true" pageId="176" pageNumber="177">pulchellus</emphasis>
|
||
</taxonomicName>
|
||
group may approach these dimensions (e.g.,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[562,702,1390,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="agavicola" status="sp. nov.">
|
||
<emphasis box="[562,702,1390,1416]" italics="true" pageId="176" pageNumber="177">T. agavicola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[711,804,1390,1416]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<emphasis box="[817,1292,1390,1416]" italics="true" pageId="176" pageNumber="177">
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[817,968,1390,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="albispinus">T. albispinus</taxonomicName>
|
||
,
|
||
<taxonomicName box="[981,1129,1390,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="flavidulus">T. flavidulus</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1142,1292,1390,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="hippolytus" status="sp. nov.">T. hippolytus</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[1301,1394,1390,1416]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1408,1525,1390,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="laticrus" status="sp. nov.">
|
||
<emphasis box="[1408,1525,1390,1416]" italics="true" pageId="176" pageNumber="177">T. laticrus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[498,592,1430,1456]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<emphasis box="[605,844,1430,1456]" italics="true" pageId="176" pageNumber="177">
|
||
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[605,736,1430,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="terricola">T. terricola</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[748,844,1430,1455]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="torrei">T. torrei</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[912,1026,1430,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="wilsoni" status="sp. nov.">
|
||
<emphasis box="[912,1026,1430,1456]" italics="true" pageId="176" pageNumber="177">T. wilsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1035,1129,1430,1456]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), but they can be separated from
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,586,1470,1496]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[498,586,1470,1496]" italics="true" pageId="176" pageNumber="177">T. achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[593,683,1470,1496]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by their very broad postpetioles (PWI> 220).
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1225,1445,1470,1496]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[1225,1445,1470,1496]" italics="true" pageId="176" pageNumber="177">Temnothorax achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1452,1542,1470,1496]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from other
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[884,971,1510,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="goniops">
|
||
<emphasis box="[884,971,1510,1535]" italics="true" pageId="176" pageNumber="177">goniops</emphasis>
|
||
</taxonomicName>
|
||
group members by the short, dentate propodeal spines (longer than the propodeal declivity and spiniform in
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1346,1466,1550,1575]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="goniops">
|
||
<emphasis box="[1346,1466,1550,1575]" italics="true" pageId="176" pageNumber="177">T. goniops</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[498,714,1589,1615]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="huehuetenangoi">
|
||
<emphasis box="[498,714,1589,1615]" italics="true" pageId="176" pageNumber="177">T. huehuetenangoi</emphasis>
|
||
</taxonomicName>
|
||
), the areolate-rugulose head and postpetiole (smooth and shining in
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,599,1630,1655]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="xincai" status="sp. nov.">
|
||
<emphasis box="[498,599,1630,1655]" italics="true" pageId="176" pageNumber="177">T. xincai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[605,697,1630,1656]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), and the relatively narrow, rugulose petiolar node (narrower, on average, in
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[530,619,1669,1695]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[530,619,1669,1695]" italics="true" pageId="176" pageNumber="177">T. achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[628,720,1669,1695]" pageId="176" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and uniformly areolate).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="178" lastPageNumber="179" pageId="176" pageNumber="177" type="description">
|
||
<paragraph blockId="176.[498,1542,234,1935]" lastBlockId="177.[498,1542,234,1934]" lastPageId="177" lastPageNumber="178" pageId="176" pageNumber="177">
|
||
<emphasis bold="true" box="[530,1018,1708,1735]" pageId="176" pageNumber="177">
|
||
Worker measurements & indices (
|
||
<emphasis bold="true" box="[941,957,1709,1735]" italics="true" pageId="176" pageNumber="177">n</emphasis>
|
||
= 6):
|
||
</emphasis>
|
||
SL =
|
||
<geoCoordinate box="[1084,1146,1708,1735]" degrees="0.375" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.375">0.375</geoCoordinate>
|
||
<geoCoordinate box="[1146,1223,1709,1735]" degrees="0.444" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.444">–0.444</geoCoordinate>
|
||
(0.418); FRS =
|
||
<geoCoordinate box="[1402,1464,1708,1735]" degrees="0.165" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.165">0.165</geoCoordinate>
|
||
<geoCoordinate box="[1464,1541,1709,1735]" degrees="0.181" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.181">–0.181</geoCoordinate>
|
||
(0.174); CW =
|
||
<geoCoordinate box="[678,740,1748,1775]" degrees="0.485" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.485">0.485</geoCoordinate>
|
||
<geoCoordinate box="[740,817,1748,1775]" degrees="0.561" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.561">–0.561</geoCoordinate>
|
||
(0.524); CWb =
|
||
<geoCoordinate box="[1020,1082,1749,1775]" degrees="0.442" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.442">0.442</geoCoordinate>
|
||
<geoCoordinate box="[1082,1159,1748,1775]" degrees="0.519" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.519">–0.519</geoCoordinate>
|
||
(0.479); PoOC =
|
||
<geoCoordinate box="[1372,1434,1749,1775]" degrees="0.220" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.22">0.220</geoCoordinate>
|
||
<geoCoordinate box="[1434,1511,1748,1775]" degrees="0.253" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.253">–0.253</geoCoordinate>
|
||
(0.235); CL =
|
||
<geoCoordinate box="[665,727,1788,1815]" degrees="0.541" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.541">0.541</geoCoordinate>
|
||
<geoCoordinate box="[727,804,1788,1814]" degrees="0.633" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.633">–0.633</geoCoordinate>
|
||
(0.582); EL =
|
||
<geoCoordinate box="[977,1039,1788,1814]" degrees="0.114" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.114">0.114</geoCoordinate>
|
||
<geoCoordinate box="[1039,1116,1788,1814]" degrees="0.146" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.146">–0.146</geoCoordinate>
|
||
(0.136); EW =
|
||
<geoCoordinate box="[1302,1364,1788,1814]" degrees="0.090" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.09">0.090</geoCoordinate>
|
||
<geoCoordinate box="[1364,1441,1788,1814]" degrees="0.112" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.112">–0.112</geoCoordinate>
|
||
(0.104); MD =
|
||
<geoCoordinate box="[580,642,1828,1854]" degrees="0.113" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.113">0.113</geoCoordinate>
|
||
<geoCoordinate box="[642,719,1828,1854]" degrees="0.142" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.142">–0.142</geoCoordinate>
|
||
(0.121); WL =
|
||
<geoCoordinate box="[904,966,1828,1854]" degrees="0.611" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.611">0.611</geoCoordinate>
|
||
<geoCoordinate box="[966,1043,1828,1854]" degrees="0.744" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.744">–0.744</geoCoordinate>
|
||
(0.668); SPST =
|
||
<geoCoordinate box="[1245,1307,1828,1855]" degrees="0.157" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.157">0.157</geoCoordinate>
|
||
<geoCoordinate box="[1308,1385,1828,1854]" degrees="0.201" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.201">–0.201</geoCoordinate>
|
||
(0.177); MPST =
|
||
<geoCoordinate box="[607,669,1868,1894]" degrees="0.196" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.196">0.196</geoCoordinate>
|
||
<geoCoordinate box="[669,746,1868,1894]" degrees="0.244" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.244">–0.244</geoCoordinate>
|
||
(0.220); PEL =
|
||
<geoCoordinate box="[935,997,1868,1895]" degrees="0.258" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.258">0.258</geoCoordinate>
|
||
<geoCoordinate box="[997,1074,1868,1894]" degrees="0.308" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.308">–0.308</geoCoordinate>
|
||
(0.286); NOL =
|
||
<geoCoordinate box="[1274,1336,1868,1894]" degrees="0.142" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.142">0.142</geoCoordinate>
|
||
<geoCoordinate box="[1336,1413,1868,1894]" degrees="0.184" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.184">–0.184</geoCoordinate>
|
||
(0.161); NOH =
|
||
<geoCoordinate box="[598,660,1908,1934]" degrees="0.098" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.098">0.098</geoCoordinate>
|
||
<geoCoordinate box="[660,737,1908,1934]" degrees="0.109" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.109">–0.109</geoCoordinate>
|
||
(0.102); PEH =
|
||
<geoCoordinate box="[933,995,1908,1934]" degrees="0.168" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.168">0.168</geoCoordinate>
|
||
<geoCoordinate box="[995,1072,1908,1934]" degrees="0.198" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.198">–0.198</geoCoordinate>
|
||
(0.185); PPL =
|
||
<geoCoordinate box="[1262,1324,1908,1935]" degrees="0.156" direction="north" orientation="latitude" pageId="176" pageNumber="177" precision="55" value="0.156">0.156</geoCoordinate>
|
||
<geoCoordinate box="[1324,1401,1908,1935]" degrees="0.205" direction="west" orientation="longitude" pageId="176" pageNumber="177" precision="55" value="-0.205">–0.205</geoCoordinate>
|
||
(0.176); PPH =
|
||
<geoCoordinate box="[588,650,234,260]" degrees="0.176" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.176">0.176</geoCoordinate>
|
||
<geoCoordinate box="[650,727,234,260]" degrees="0.212" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.212">–0.212</geoCoordinate>
|
||
(0.195); PW =
|
||
<geoCoordinate box="[912,974,234,260]" degrees="0.308" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.308">0.308</geoCoordinate>
|
||
<geoCoordinate box="[975,1052,234,260]" degrees="0.389" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.389">–0.389</geoCoordinate>
|
||
(0.340); SBPA =
|
||
<geoCoordinate box="[1260,1322,234,260]" degrees="0.128" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.128">0.128</geoCoordinate>
|
||
<geoCoordinate box="[1322,1399,234,261]" degrees="0.175" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.175">–0.175</geoCoordinate>
|
||
(0.145); SPTI =
|
||
<geoCoordinate box="[590,652,274,300]" degrees="0.169" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.169">0.169</geoCoordinate>
|
||
<geoCoordinate box="[653,730,274,301]" degrees="0.235" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.235">–0.235</geoCoordinate>
|
||
(0.194); PEW =
|
||
<geoCoordinate box="[932,994,274,300]" degrees="0.136" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.136">0.136</geoCoordinate>
|
||
<geoCoordinate box="[994,1071,274,301]" degrees="0.152" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.152">–0.152</geoCoordinate>
|
||
(0.146); PNW =
|
||
<geoCoordinate box="[1278,1340,274,300]" degrees="0.148" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.148">0.148</geoCoordinate>
|
||
<geoCoordinate box="[1340,1417,274,300]" degrees="0.184" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.184">–0.184</geoCoordinate>
|
||
(0.164); PPW =
|
||
<geoCoordinate box="[590,652,314,341]" degrees="0.265" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.265">0.265</geoCoordinate>
|
||
<geoCoordinate box="[653,730,314,340]" degrees="0.309" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.309">–0.309</geoCoordinate>
|
||
(0.293); HFL =
|
||
<geoCoordinate box="[917,979,314,340]" degrees="0.402" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.402">0.402</geoCoordinate>
|
||
<geoCoordinate box="[979,1056,314,340]" degrees="0.474" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.474">–0.474</geoCoordinate>
|
||
(0.443); HFWmax =
|
||
<geoCoordinate box="[1306,1368,314,340]" degrees="0.081" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.081">0.081</geoCoordinate>
|
||
<geoCoordinate box="[1368,1445,314,340]" degrees="0.129" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.129">–0.129</geoCoordinate>
|
||
(0.112); HFWmin =
|
||
<geoCoordinate box="[645,707,354,380]" degrees="0.040" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.04">0.040</geoCoordinate>
|
||
<geoCoordinate box="[708,785,354,380]" degrees="0.046" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.046">–0.046</geoCoordinate>
|
||
(0.044); CS =
|
||
<geoCoordinate box="[958,1020,354,380]" degrees="0.713" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.713">0.713</geoCoordinate>
|
||
<geoCoordinate box="[1021,1098,354,380]" degrees="0.836" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.836">–0.836</geoCoordinate>
|
||
(0.77); ES =
|
||
<geoCoordinate box="[1254,1316,354,381]" degrees="0.159" direction="north" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="0.159">0.159</geoCoordinate>
|
||
<geoCoordinate box="[1317,1394,354,380]" degrees="0.199" direction="west" orientation="longitude" pageId="177" pageNumber="178" precision="55" value="-0.199">–0.199</geoCoordinate>
|
||
(0.188); SI = 85–90 (87); OI = 22–26 (24); CI = 82–85 (82); WLI = 137–143 (139); SBI = 28–34 (30); PSI = 25–28 (26); PWI = 191–208 (200); PLI = 150–170 (163); NI = 143–170 (157); PNWI = 109–121 (112); NLI = 50–60 (56); FI = 180–303 (256).
|
||
</paragraph>
|
||
<paragraph blockId="177.[498,1542,234,1934]" pageId="177" pageNumber="178">
|
||
<emphasis bold="true" box="[530,775,513,539]" pageId="177" pageNumber="178">Worker description:</emphasis>
|
||
In full-face view, head subquadrate, longer than broad (CI 82–85). Mandibles densely, finely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin evenly convex medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about the maximum width of the scape (SI 85–90). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment longer than the preceding two in combination. Frontal carinae moderately long, extending past the antennal toruli by about three times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head very weakly convex, nearly flat, forming a continuous low arc from the mandibular insertions to the posterior margin of the head. Posterior head margin weakly concave medially but rounding evenly into the lateral margins.
|
||
</paragraph>
|
||
<paragraph blockId="177.[498,1542,234,1934]" lastBlockId="178.[498,1542,230,1894]" lastPageId="178" lastPageNumber="179" pageId="177" pageNumber="178">
|
||
In profile view, compound eyes ovular, longitudinally elongate, and moderately large (OI 22–26), with 10 ommatidia in longest row. Pronotal declivity indistinct, but neck and anterior face of pronotum forming a ~120
|
||
<emphasis box="[997,1007,1106,1124]" italics="true" pageId="177" pageNumber="178">
|
||
<superScript attach="left" box="[997,1007,1106,1124]" fontSize="8" pageId="177" pageNumber="178">°</superScript>
|
||
</emphasis>
|
||
angle. Mesosoma convex from where it joins the pronotal neck to the propodeal spines; propodeum slightly flatter, giving the dorsum of the mesosoma a weakly sinuate profile. Promesonotal suture extending from the posterior margin of the procoxal insertion to the mesothoracic spiracle, which is moderately well developed; continuing dorsally as a weak sulcus. Metanotal groove visible as a disruption of the sculpture laterally from where it arises between the mid- and hind coxae to where it ends in the poorly developed metathoracic spiracle, which is nearly indistinguishable against the ground sculpture. Propodeal spiracle moderately well developed, directed posterolaterally, and separated from the propodeal declivity by about two and a half spiracle diameters. Propodeal spines poorly developed and short (PSI 25–28), about two thirds the length of the propodeal declivity, flared at the base, straight, and acute. Propodeal declivity with weak carinae extending between the base of the propodeal spines and the propodeal lobes; declivity straight, forming a rounded ~120
|
||
<emphasis box="[1530,1540,1584,1602]" italics="true" pageId="177" pageNumber="178">
|
||
<superScript attach="none" box="[1530,1540,1584,1602]" fontSize="8" pageId="177" pageNumber="178">°</superScript>
|
||
</emphasis>
|
||
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole moderately long (PLI 150–170), without tubercles anterodorsally. Subpetiolar process in the form of a small, triangular, acute tooth; ventral margin of petiole flat posteriorly to it. Petiolar peduncle short: comprising about a third of the petiole length. Petiolar node robust and rounded: node transitioning evenly into the petiolar peduncle, resulting in a concave anterior node face; anterior face rounding evenly into the dorsal face; dorsal face rounding evenly into the short posterior face, which forms a ~110
|
||
<emphasis box="[655,665,230,248]" italics="true" pageId="178" pageNumber="179">
|
||
<superScript attach="left" box="[655,665,230,248]" fontSize="8" pageId="178" pageNumber="179">°</superScript>
|
||
</emphasis>
|
||
angle with the caudal cylinder. Postpetiole evenly rounded anteriorly, convex dorsally, and weakly lobed ventrally.
|
||
</paragraph>
|
||
<paragraph blockId="178.[498,1542,230,1894]" pageId="178" pageNumber="179">In dorsal view, humeri poorly developed: evenly rounded and only slightly wider than the rest of the mesosoma; mesothoracic spiracles not protruding past the lateral margins of the mesosoma. Promesonotal suture visible as a weak sulcus. Metanotal groove absent: mesonotum and propodeum completely fused and lateral margins converging evenly to the bases of the propodeal spines. Propodeal spines broadly approximated basally and diverging apically, their apices separated from each other by about their length, the negative space between them “U” shaped. Petiolar peduncle with spiracles very weakly protruding past the lateral margins, not noticeably constricted anterior to them. Petiolar node campaniform: evenly convex anteriorly, flattened posteriorly; node slightly wider than the peduncle, but narrower than the caudal cylinder. Postpetiole moderately broad (PWI 191–208) and campaniform, articulating with most of the anterior margin of the gaster, leaving small, angulate margins on each side exposed. Anterior margin of the postpetiole flat and evenly rounds into the lateral margins, which diverge slightly to the angulate posterior corners; posterior margin flat. Metafemur weakly to moderately incrassate (FI 180–303).</paragraph>
|
||
<paragraph blockId="178.[498,1542,230,1894]" pageId="178" pageNumber="179">Sculpture: median clypeal carina present, extending posteriorly nearly to the frontal triangle, and flanked on either side by a slightly weaker carinae. Lateral clypeal lobes with additional, weaker carinae; ground sculpture very weakly areolate. Antennal scapes weakly areolate. Cephalic dorsum finely areolate, but areolae arranged into longitudinal columns by costulae which flank the frontal carinae medially and extend to the posterior margin of the head; concentric costulae surrounding the antennal insertions. Lateral surfaces of head uniformly weakly areolate-costulate, with dense rugose sculpture between the compound eye and the mandibular insertion. Ventral surface of head smooth and shining anteriorly, weakly areolate posteriorly. Mesosoma uniformly areolate-rugulose, but sculpture surrounding the propodeal spiracle costulate; propodeal declivity shining through very weak areolate sculpture. Femora shining through traces of shallow areolate sculpture, which becomes stronger distally. First gastral tergite and sternite smooth and shining, with weak spectral iridescence.</paragraph>
|
||
<paragraph blockId="178.[498,1542,230,1894]" pageId="178" pageNumber="179">Setae: antennal scapes and funiculi with short, decumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, blunt-tipped setae, the longest of which are about the width of the compound eye. The head bears ~28, mesosoma ~20, petiole 6, postpetiole ~12, and first gastral tergite ~46 setae. Longer, flexuous setae on the gular region, and the ventrites of the gaster. Short, sparse pubescence present over the entire body, but difficult to detect against the densely sculptured integument.</paragraph>
|
||
<paragraph blockId="178.[498,1542,230,1894]" pageId="178" pageNumber="179">Color: predominantly dark brown, with mandibles and antennae testaceous yellow. Femora medium brown. Trochanters, Meso- and metacoxae, tibiae, tarsi, and sting very light yellow, nearly white.</paragraph>
|
||
<paragraph blockId="178.[498,1542,230,1894]" box="[530,734,1828,1854]" pageId="178" pageNumber="179">
|
||
<emphasis bold="true" box="[530,601,1828,1854]" pageId="178" pageNumber="179">Gyne:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph blockId="178.[498,1542,230,1894]" box="[530,730,1868,1894]" pageId="178" pageNumber="179">
|
||
<emphasis bold="true" box="[530,597,1868,1894]" pageId="178" pageNumber="179">Male:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102335" ID-Zenodo-Dep="5102335" httpUri="https://zenodo.org/record/5102335/files/figure.png" pageId="179" pageNumber="180" startId="179.[524,589,1058,1080]" targetBox="[498,1541,234,1045]" targetPageId="179">
|
||
<paragraph blockId="179.[524,1515,1058,1109]" pageId="179" pageNumber="180">
|
||
<emphasis bold="true" box="[524,1256,1058,1080]" pageId="179" pageNumber="180">
|
||
Figure 116
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[644,859,1058,1080]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="goniops">
|
||
<emphasis bold="true" box="[644,859,1058,1080]" italics="true" pageId="179" pageNumber="180">Temnothorax goniops</emphasis>
|
||
</taxonomicName>
|
||
holotype worker (MCZENT00032436).
|
||
</emphasis>
|
||
(A) Profile view. (B) Fullface view. (C) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-116
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="179" pageNumber="180" type="etymology">
|
||
<paragraph blockId="179.[498,1542,1164,1470]" pageId="179" pageNumber="180">
|
||
<emphasis bold="true" box="[530,668,1165,1191]" pageId="179" pageNumber="180">Etymology:</emphasis>
|
||
Geographical, from ‘achi’, one of the many Mayan dialects spoken by the indigenous people of
|
||
<collectingCountry box="[752,881,1204,1230]" name="Guatemala" pageId="179" pageNumber="180">Guatemala</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="179" pageNumber="180" type="discussion">
|
||
<paragraph blockId="179.[498,1542,1164,1470]" pageId="179" pageNumber="180">
|
||
<emphasis bold="true" box="[530,670,1244,1270]" pageId="179" pageNumber="180">Comments:</emphasis>
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[679,901,1244,1270]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[679,901,1244,1270]" italics="true" pageId="179" pageNumber="180">Temnothorax achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[910,1001,1245,1271]" pageId="179" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has been collected from Winkler leaf litter extractions in several localities in Central
|
||
<collectingCountry box="[992,1120,1284,1310]" name="Guatemala" pageId="179" pageNumber="180">Guatemala</collectingCountry>
|
||
, spanning low-to-mid elevation tropical and mesic forests. This species is closely related to
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1198,1275,1324,1350]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="ixili">
|
||
<emphasis box="[1198,1275,1324,1350]" italics="true" pageId="179" pageNumber="180">T. ixili</emphasis>
|
||
</taxonomicName>
|
||
, which is also known from low to mid elevation leaf litter extractions and nests in
|
||
<taxonomicName box="[1199,1315,1364,1390]" class="Liliopsida" family="Orchidaceae" genus="Oncidium" kingdom="Fungi" order="Asparagales" pageId="179" pageNumber="180" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[1199,1315,1364,1390]" italics="true" pageId="179" pageNumber="180">Oncidium</emphasis>
|
||
</taxonomicName>
|
||
orchids. They have not been collected at the same locality, but collections of
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1161,1236,1403,1429]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="ixili">
|
||
<emphasis box="[1161,1236,1403,1429]" italics="true" pageId="179" pageNumber="180">T. ixili</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1295,1382,1403,1429]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
|
||
<emphasis box="[1295,1382,1403,1429]" italics="true" pageId="179" pageNumber="180">T. achii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1390,1480,1404,1430]" pageId="179" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were taken about
|
||
<quantity box="[646,720,1443,1470]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="179" pageNumber="180" unit="km" value="15.0">15 km</quantity>
|
||
apart from each other. I consider them sympatric species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |