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<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FF44FF5DB49C1FC97C5323BE" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax leucacanthoides Prebus 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="136" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="132" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prebus, Matthew M.</mods:namePart>
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States &amp; Department of Entomology &amp; Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PeerJ</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="series">
<mods:title>e 11514</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2021-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>9</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>462</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
<mods:identifier type="ISSN">2167-8359</mods:identifier>
<mods:identifier type="PMC">PMC8254503</mods:identifier>
<mods:identifier type="PubMed">34249486</mods:identifier>
<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102507" ID-GBIF-Taxon="183542622" ID-Zenodo-Dep="5102507" LSID="urn:lsid:zoobank.org:act:25BBD7A1-CB88-45EA-8808-452CB6466A4F" httpUri="http://treatment.plazi.org/id/054FDB70FF44FF5DB49C1FC97C5323BE" lastPageId="135" lastPageNumber="136" pageId="131" pageNumber="132">
<subSubSection box="[498,963,1492,1519]" pageId="131" pageNumber="132" type="nomenclature">
<paragraph blockId="131.[498,981,1492,1559]" box="[498,963,1492,1519]" pageId="131" pageNumber="132">
<heading box="[498,963,1492,1519]" level="2" pageId="131" pageNumber="132">
<emphasis bold="true" box="[498,963,1492,1519]" pageId="131" pageNumber="132">
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,859,1492,1519]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis bold="true" box="[498,859,1492,1519]" italics="true" pageId="131" pageNumber="132">Temnothorax leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[868,963,1492,1518]" pageId="131" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[498,981,1532,1559]" pageId="131" pageNumber="132" type="distribution">
<paragraph blockId="131.[498,981,1492,1559]" box="[498,981,1532,1559]" pageId="131" pageNumber="132">
Distribution:
<figureCitation box="[657,770,1532,1558]" captionStart="Figure 101" captionStartId="121.[524,589,1194,1216]" captionTargetBox="[498,1541,234,1181]" captionTargetId="figure-149@121.[498,1534,234,1166]" captionTargetPageId="121" captionText="Figure 101 Geographical and elevational distribution of the augusti group. (A) Temnothorax augusti (B) T. aureus sp. nov. (C) T. leucacanthoides sp. nov. (D) T. leucacanthus. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-101" figureDoi="http://doi.org/10.5281/zenodo.5102293" httpUri="https://zenodo.org/record/5102293/files/figure.png" pageId="131" pageNumber="132">Fig. 101C</figureCitation>
; worker:
<figureCitation box="[882,975,1532,1558]" captionStart="Figure 104" captionStartId="132.[524,589,1134,1156]" captionTargetBox="[498,1541,234,1121]" captionTargetId="figure-226@132.[498,1295,234,1106]" captionTargetPageId="132" captionText="Figure 104 Temnothorax leucacanthoides sp. nov. holotype worker (CASENT0756102). (A) Full-face view. (B) Profile view. (C) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-104" figureDoi="http://doi.org/10.5281/zenodo.5102301" httpUri="https://zenodo.org/record/5102301/files/figure.png" pageId="131" pageNumber="132">Fig. 104</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[498,1160,1580,1607]" pageId="131" pageNumber="132" type="reference_group">
<paragraph blockId="131.[498,1160,1580,1607]" box="[498,1160,1580,1607]" pageId="131" pageNumber="132">
<treatmentCitationGroup box="[498,1160,1580,1607]" pageId="131" pageNumber="132">
<taxonomicName authority="Prebus, 2017: 8" authorityName="Prebus" authorityPageNumber="8" authorityYear="2017" box="[498,831,1580,1607]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[498,655,1580,1606]" italics="true" pageId="131" pageNumber="132">Temnothorax</emphasis>
sp. nr.
<emphasis box="[747,987,1580,1606]" italics="true" pageId="131" pageNumber="132">
augusti
<treatmentCitation author="Prebus M." box="[839,1156,1580,1607]" page="8" pageId="131" pageNumber="132" year="2017">
<bibRefCitation author="Prebus M." box="[839,1018,1580,1607]" journalOrPublisher="BMC Evolutionary Biology" pageId="131" pageNumber="132" pagination="250" part="17" refId="ref272195" refString="Prebus M. 2017. Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae). BMC Evolutionary Biology 17 (1): 250 DOI 10.1186 / s 12862 - 017 - 1095 - 8." title="Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae)" type="journal article" year="2017">
<author box="[839,919,1580,1606]" fns="*" inm="true" npo="*" pageId="131" pageNumber="132">Prebus</author>
,
<year box="[932,987,1580,1606]" pageId="131" pageNumber="132">2017</year>
: 8
</bibRefCitation>
. Phylogeny
</treatmentCitation>
</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="131" pageNumber="132" type="materials_examined">
<paragraph blockId="131.[498,1542,1642,1908]" pageId="131" pageNumber="132">
<emphasis bold="true" box="[530,834,1642,1669]" pageId="131" pageNumber="132">Type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861439" collectingDate="2006-03-22" collectionCode="CASC" collectorName="P. S. Ward" country="Mexico" elevation="20" latitude="15.666667" location="Puerto Angel" longLatPrecision="1" longitude="-96.55" pageId="131" pageNumber="132" specimenCode="15573, CASENT0756102" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
<emphasis box="[843,1041,1642,1668]" italics="true" pageId="131" pageNumber="132">
<typeStatus box="[843,946,1642,1668]" pageId="131" pageNumber="132">Holotype</typeStatus>
<specimenCount box="[954,1034,1642,1668]" pageId="131" pageNumber="132" type="worker">worker</specimenCount>
:
</emphasis>
<collectingCountry box="[1049,1164,1643,1669]" name="Mexico" pageId="131" pageNumber="132">MEXICO</collectingCountry>
:
<collectingRegion box="[1176,1265,1643,1669]" country="Mexico" name="Oaxaca" pageId="131" pageNumber="132">Oaxaca</collectingRegion>
:
<collectingRegion box="[1277,1422,1643,1669]" pageId="131" pageNumber="132">Cerro Largo</collectingRegion>
,
<locationDeviation box="[1435,1533,1642,1669]" pageId="131" pageNumber="132">
<quantity box="[1435,1496,1642,1668]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="131" pageNumber="132" unit="km" value="6.0">6 km</quantity>
W
</locationDeviation>
<location LSID="urn:lsid:plazi:treatment:054FDB70FF44FF5DB49C1FC97C5323BE:88393CBDFF44FF59B49C1C8E7BBC21FB" box="[498,654,1682,1709]" country="Mexico" latitude="15.666667" longLatPrecision="1" longitude="-96.55" name="Puerto Angel" pageId="131" pageNumber="132" stateProvince="Oaxaca">Puerto Angel</location>
,
<quantity box="[664,724,1682,1709]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="131" pageNumber="132" unit="m" value="20.0">
<elevation box="[664,724,1682,1709]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="131" pageNumber="132" unit="m" value="20.0">20 m</elevation>
</quantity>
,
<geoCoordinate box="[735,887,1678,1709]" degrees="15.666667" direction="north" orientation="latitude" pageId="131" pageNumber="132" precision="1" value="15.666667">
15.666667
<emphasis box="[854,864,1678,1696]" italics="true" pageId="131" pageNumber="132">
<superScript attach="right" box="[854,864,1678,1696]" fontSize="8" pageId="131" pageNumber="132">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[893,1053,1678,1709]" degrees="96.550000" direction="west" orientation="longitude" pageId="131" pageNumber="132" precision="1" value="-96.55">
96.550000
<emphasis box="[1011,1021,1678,1696]" italics="true" pageId="131" pageNumber="132">
<superScript attach="right" box="[1011,1021,1678,1696]" fontSize="8" pageId="131" pageNumber="132">°</superScript>
</emphasis>
W
</geoCoordinate>
,
<date box="[1064,1240,1682,1708]" pageId="131" pageNumber="132" value="2006-03-22">
<collectingDate box="[1064,1240,1682,1708]" pageId="131" pageNumber="132" value="2006-03-22">22 March 2006</collectingDate>
</date>
,
<collectorName box="[1251,1367,1682,1709]" pageId="131" pageNumber="132">P.S. Ward</collectorName>
<specimenCode box="[1367,1453,1682,1709]" pageId="131" pageNumber="132">#15573</specimenCode>
, littoral vegetation, ex dead stalk of
<taxonomicName box="[822,946,1722,1748]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="131" pageNumber="132" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
(
<specimenCode box="[961,1173,1722,1749]" pageId="131" pageNumber="132">CASENT0756102</specimenCode>
, top specimen on pin) [
<collectionCode box="[1450,1531,1721,1748]" pageId="131" pageNumber="132">CASC</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph blockId="131.[498,1542,1642,1908]" pageId="131" pageNumber="132">
<materialsCitation ID-GBIF-Occurrence="3321861388" collectingDate="2006-03-22" collectionCode="CASC, UCDC, MCZC" collectorName="P. S. Ward" country="Mexico" elevation="20" latitude="15.666667" location="Cerro Largo" longLatPrecision="1" longitude="-96.55" pageId="131" pageNumber="132" specimenCount="6" specimenCount-worker="6" stateProvince="Oaxaca" typeStatus="paratype">
<emphasis box="[530,737,1762,1788]" italics="true" pageId="131" pageNumber="132">
<typeStatus box="[530,632,1763,1788]" pageId="131" pageNumber="132">Paratype</typeStatus>
workers:
</emphasis>
same pin as
<typeStatus box="[894,998,1762,1788]" pageId="131" pageNumber="132">holotype</typeStatus>
,
<specimenCount box="[1011,1115,1762,1788]" pageId="131" pageNumber="132" type="worker">1 worker</specimenCount>
(CASENT0756102, bottom specimen on pin) [
<collectionCode box="[713,794,1801,1828]" pageId="131" pageNumber="132">CASC</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861445" collectingDate="2006-03-22" collectionCode="CASC, UCDC, MCZC" collectorName="P. S. Ward" country="Mexico" elevation="20" latitude="15.666667" location="Cerro Largo" longLatPrecision="1" longitude="-96.55" pageId="131" pageNumber="132" specimenCount="6" specimenCount-worker="6" stateProvince="Oaxaca" typeStatus="paratype">
same data as holotype,
<specimenCount box="[1078,1180,1802,1828]" pageId="131" pageNumber="132" type="worker">1 worker</specimenCount>
(CASENT0732606) [
<collectionCode box="[1427,1515,1801,1828]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="131" pageNumber="132" type="Museum">UCDC</collectionCode>
]
<specimenCount pageId="131" pageNumber="132" type="worker">1 worker</specimenCount>
(CASENT0758708) [
<collectionCode box="[832,920,1841,1868]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="131" pageNumber="132" type="Museum">UCDC</collectionCode>
]
<specimenCount box="[934,1050,1842,1868]" pageId="131" pageNumber="132" type="worker">3 workers</specimenCount>
(CASENT0916001) [
<collectionCode box="[1301,1392,1841,1868]" pageId="131" pageNumber="132">MCZC</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[530,1438,1881,1908]" pageId="131" pageNumber="132" type="distribution">
<paragraph blockId="131.[498,1542,1642,1908]" box="[530,1438,1881,1908]" pageId="131" pageNumber="132">
<emphasis bold="true" box="[530,755,1881,1908]" pageId="131" pageNumber="132">Geographic range:</emphasis>
Low elevations of southern
<collectingCountry box="[1090,1176,1882,1908]" name="Mexico" pageId="131" pageNumber="132">Mexico</collectingCountry>
(
<collectingRegion box="[1193,1283,1882,1908]" country="Mexico" name="Oaxaca" pageId="131" pageNumber="132">Oaxaca</collectingRegion>
) (
<figureCitation box="[1309,1422,1882,1908]" captionStart="Figure 101" captionStartId="121.[524,589,1194,1216]" captionTargetBox="[498,1541,234,1181]" captionTargetId="figure-149@121.[498,1534,234,1166]" captionTargetPageId="121" captionText="Figure 101 Geographical and elevational distribution of the augusti group. (A) Temnothorax augusti (B) T. aureus sp. nov. (C) T. leucacanthoides sp. nov. (D) T. leucacanthus. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-101" figureDoi="http://doi.org/10.5281/zenodo.5102293" httpUri="https://zenodo.org/record/5102293/files/figure.png" pageId="131" pageNumber="132">Fig. 101C</figureCitation>
).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102301" ID-Zenodo-Dep="5102301" httpUri="https://zenodo.org/record/5102301/files/figure.png" pageId="132" pageNumber="133" startId="132.[524,589,1134,1156]" targetBox="[498,1541,234,1121]" targetPageId="132">
<paragraph blockId="132.[524,1515,1134,1214]" pageId="132" pageNumber="133">
<emphasis bold="true" box="[524,1388,1134,1156]" pageId="132" pageNumber="133">
Figure 104
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[642,935,1134,1156]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis bold="true" box="[642,935,1134,1156]" italics="true" pageId="132" pageNumber="133">Temnothorax leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[941,1017,1134,1155]" pageId="132" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
holotype worker (CASENT0756102).
</emphasis>
(A) Full-face view. (B) Profile view. (C) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-104
</paragraph>
</caption>
<subSubSection lastPageId="133" lastPageNumber="134" pageId="132" pageNumber="133" type="diagnosis">
<paragraph blockId="132.[498,1542,1269,1894]" pageId="132" pageNumber="133">
<emphasis bold="true" box="[530,754,1269,1296]" pageId="132" pageNumber="133">Worker diagnosis:</emphasis>
The following character combination separates
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis italics="true" pageId="132" pageNumber="133">Temnothorax leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[688,780,1310,1336]" pageId="132" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
from all other species in the
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1128,1203,1309,1335]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[1128,1203,1309,1335]" italics="true" pageId="132" pageNumber="133">salvini</emphasis>
</taxonomicName>
clade: head relatively broad (CI 8489); moderately long antennal scapes, which just reach the posterior margin of the head (SI 8696); mesosoma about one and a half times as long as the width of the head (WLI 136143); propodeal spines about as long as the length of the propodeal declivity (PSI 3942); petiolar node subquadrate, overhanging the caudal cylinder of the petiole; postpetiole moderately to very broad, more than two times the width of the petiole in dorsal view (PWI 202224); integument medium brown; erect setae on the dorsal surface of the head.
</paragraph>
<paragraph blockId="132.[498,1542,1269,1894]" lastBlockId="133.[498,1542,234,1894]" lastPageId="133" lastPageNumber="134" pageId="132" pageNumber="133">
<emphasis bold="true" box="[530,719,1628,1654]" pageId="132" pageNumber="133">Similar species:</emphasis>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[727,1046,1628,1654]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis box="[727,1046,1628,1654]" italics="true" pageId="132" pageNumber="133">Temnothorax acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1055,1148,1629,1655]" pageId="132" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis italics="true" pageId="132" pageNumber="133">
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1161,1313,1628,1654]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="subditivus">T. subditivus</taxonomicName>
,
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1326,1512,1628,1654]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="tenuisculptus">T. tenuisculptus</taxonomicName>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,640,1668,1694]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">T. tuxtlanus</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[649,742,1669,1695]" pageId="132" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
, species of the
<emphasis box="[921,1117,1669,1694]" italics="true" pageId="132" pageNumber="133">
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[921,1019,1669,1694]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="annexus">annexus</taxonomicName>
,
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[1033,1117,1669,1694]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="augusti">augusti</taxonomicName>
</emphasis>
,
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1132,1224,1668,1694]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis box="[1132,1224,1668,1694]" italics="true" pageId="132" pageNumber="133">fuscatus</emphasis>
</taxonomicName>
, and
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1292,1367,1668,1694]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[1292,1367,1668,1694]" italics="true" pageId="132" pageNumber="133">salvini</emphasis>
</taxonomicName>
groups.
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,844,1708,1734]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis box="[498,844,1708,1734]" italics="true" pageId="132" pageNumber="133">Temnothorax leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[852,942,1708,1734]" pageId="132" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from the first three species by the shape of the dorsal margin of the mesosoma in profile, which is weakly, evenly convex (with the metanotal groove slightly impressed in some specimens): in
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1326,1475,1787,1813]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[1326,1475,1787,1813]" italics="true" pageId="132" pageNumber="133">T. subditivus</emphasis>
</taxonomicName>
, the mesosoma is strongly arched, while
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[925,1109,1827,1853]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="tenuisculptus">
<emphasis box="[925,1109,1827,1853]" italics="true" pageId="132" pageNumber="133">T. tenuisculptus</emphasis>
</taxonomicName>
and
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1170,1311,1827,1853]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
<emphasis box="[1170,1311,1827,1853]" italics="true" pageId="132" pageNumber="133">T. tuxtlanus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1320,1411,1828,1854]" pageId="132" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
have a sinuate profile. Petiolar node shape is also a useful character for separating
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,712,234,260]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis box="[498,712,234,260]" italics="true" pageId="133" pageNumber="134">T. leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[721,813,235,261]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
from the above taxa:
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1072,1221,234,260]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[1072,1221,234,260]" italics="true" pageId="133" pageNumber="134">T. subditivus</emphasis>
</taxonomicName>
and the potentially co-occurring members of the
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[850,925,274,300]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[850,925,274,300]" italics="true" pageId="133" pageNumber="134">salvini</emphasis>
</taxonomicName>
group have a squamiform petiolar node, which is much broader than the peduncle in dorsal view (only slightly wider in
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1328,1541,314,340]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis box="[1328,1541,314,340]" italics="true" pageId="133" pageNumber="134">T. leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[498,593,354,380]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
), and the nodes of the
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[867,963,355,380]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[867,963,355,380]" italics="true" pageId="133" pageNumber="134">annexus</emphasis>
</taxonomicName>
group are erect and subquadrate, as opposed to leaning posteriorly in
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[756,969,394,420]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis box="[756,969,394,420]" italics="true" pageId="133" pageNumber="134">T. leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[977,1070,394,420]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1082,1264,394,420]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="tenuisculptus">
<emphasis box="[1082,1264,394,420]" italics="true" pageId="133" pageNumber="134">T. tenuisculptus</emphasis>
</taxonomicName>
, like
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1328,1541,394,420]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis box="[1328,1541,394,420]" italics="true" pageId="133" pageNumber="134">T. leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[498,592,434,460]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
, has a subquadrate petiolar node, but it does not lean posteriorly over the caudal cylinder;
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[691,832,473,499]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
<emphasis box="[691,832,473,499]" italics="true" pageId="133" pageNumber="134">T. tuxtlanus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[840,931,474,500]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[991,1176,474,499]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis box="[991,1176,474,499]" italics="true" pageId="133" pageNumber="134">T. acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1185,1276,474,500]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
have rounded petiolar nodes in profile view.
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[748,1092,513,539]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis box="[748,1092,513,539]" italics="true" pageId="133" pageNumber="134">Temnothorax leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1098,1187,514,540]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
can also be distinguished from
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,640,553,579]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
<emphasis box="[498,640,553,579]" italics="true" pageId="133" pageNumber="134">T. tuxtlanus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[649,740,554,580]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[801,987,554,579]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis box="[801,987,554,579]" italics="true" pageId="133" pageNumber="134">T. acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[996,1088,554,580]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
by the presence of erect setae on the dorsum of the propodeum. The potentially co-occurring members of the
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1364,1456,593,619]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis box="[1364,1456,593,619]" italics="true" pageId="133" pageNumber="134">fuscatus</emphasis>
</taxonomicName>
group have a low, elongate petiolar node in profile view. Finally,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1181,1394,633,659]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis box="[1181,1394,633,659]" italics="true" pageId="133" pageNumber="134">T. leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1402,1493,633,659]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from fellow members of the
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[991,1075,673,698]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="augusti">
<emphasis box="[991,1075,673,698]" italics="true" pageId="133" pageNumber="134">augusti</emphasis>
</taxonomicName>
group by the combination of long propodeal spines, which are about as long as the length of the propodeal declivity (shorter than the declivity in
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[744,861,753,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="augusti">
<emphasis box="[744,861,753,778]" italics="true" pageId="133" pageNumber="134">T. augusti</emphasis>
</taxonomicName>
and
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[922,1068,753,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="casanovai" status="sp. nov.">
<emphasis box="[922,1068,753,778]" italics="true" pageId="133" pageNumber="134">T. casanovai</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1077,1171,753,779]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
), the relatively broad head (narrower in all other
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[763,847,793,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="augusti">
<emphasis box="[763,847,793,818]" italics="true" pageId="133" pageNumber="134">augusti</emphasis>
</taxonomicName>
group species: CI&gt; 84 vs. &lt;82); relatively short antennal scapes (SI 8696 vs.&gt;
<quantity box="[766,840,832,859]" metricMagnitude="0" metricUnit="m" metricValue="2.667" pageId="133" pageNumber="134" unit="in" value="105.0">105 in</quantity>
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[849,966,833,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="augusti">
<emphasis box="[849,966,833,858]" italics="true" pageId="133" pageNumber="134">T. augusti</emphasis>
</taxonomicName>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[981,1091,833,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="aureus" status="sp. nov.">
<emphasis box="[981,1091,833,858]" italics="true" pageId="133" pageNumber="134">T. aureus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1100,1193,832,858]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1206,1352,833,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="casanovai" status="sp. nov.">
<emphasis box="[1206,1352,833,858]" italics="true" pageId="133" pageNumber="134">T. casanovai</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1361,1456,832,858]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="135" lastPageNumber="136" pageId="133" pageNumber="134" type="description">
<paragraph blockId="133.[498,1542,234,1894]" pageId="133" pageNumber="134">
<emphasis bold="true" box="[530,1018,871,898]" pageId="133" pageNumber="134">
Worker measurements &amp; indices (
<emphasis bold="true" box="[941,957,872,898]" italics="true" pageId="133" pageNumber="134">n</emphasis>
= 7):
</emphasis>
SL =
<geoCoordinate box="[1084,1146,872,899]" degrees="0.521" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.521">0.521</geoCoordinate>
<geoCoordinate box="[1146,1223,872,899]" degrees="0.573" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.573">0.573</geoCoordinate>
(0.547); FRS =
<geoCoordinate box="[1402,1464,872,898]" degrees="0.182" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.182">0.182</geoCoordinate>
<geoCoordinate box="[1464,1541,872,898]" degrees="0.197" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.197">0.197</geoCoordinate>
(0.189); CW =
<geoCoordinate box="[678,740,911,937]" degrees="0.618" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.618">0.618</geoCoordinate>
<geoCoordinate box="[740,817,911,938]" degrees="0.658" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.658">0.658</geoCoordinate>
(0.64); CWb =
<geoCoordinate box="[1006,1068,911,938]" degrees="0.578" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.578">0.578</geoCoordinate>
<geoCoordinate box="[1068,1145,911,938]" degrees="0.611" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.611">0.611</geoCoordinate>
(0.596); PoOC =
<geoCoordinate box="[1358,1420,911,938]" degrees="0.251" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.251">0.251</geoCoordinate>
<geoCoordinate box="[1420,1497,911,938]" degrees="0.263" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.263">0.263</geoCoordinate>
(0.256); CL =
<geoCoordinate box="[665,727,951,977]" degrees="0.680" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.68">0.680</geoCoordinate>
<geoCoordinate box="[727,804,952,978]" degrees="0.704" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.704">0.704</geoCoordinate>
(0.693); EL =
<geoCoordinate box="[977,1039,951,978]" degrees="0.158" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.158">0.158</geoCoordinate>
<geoCoordinate box="[1039,1116,951,978]" degrees="0.166" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.166">0.166</geoCoordinate>
(0.162); EW =
<geoCoordinate box="[1302,1364,952,978]" degrees="0.112" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.112">0.112</geoCoordinate>
<geoCoordinate box="[1364,1441,951,978]" degrees="0.126" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.126">0.126</geoCoordinate>
(0.120); MD =
<geoCoordinate box="[580,642,992,1018]" degrees="0.171" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.171">0.171</geoCoordinate>
<geoCoordinate box="[642,719,991,1018]" degrees="0.188" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.188">0.188</geoCoordinate>
(0.179); WL =
<geoCoordinate box="[904,966,991,1018]" degrees="0.815" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.815">0.815</geoCoordinate>
<geoCoordinate box="[966,1043,991,1017]" degrees="0.863" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.863">0.863</geoCoordinate>
(0.836); SPST =
<geoCoordinate box="[1245,1307,992,1018]" degrees="0.319" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.319">0.319</geoCoordinate>
<geoCoordinate box="[1308,1385,991,1017]" degrees="0.360" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.36">0.360</geoCoordinate>
(0.335); MPST =
<geoCoordinate box="[607,669,1031,1057]" degrees="0.249" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.249">0.249</geoCoordinate>
<geoCoordinate box="[669,746,1031,1057]" degrees="0.276" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.276">0.276</geoCoordinate>
(0.261); PEL =
<geoCoordinate box="[935,997,1031,1057]" degrees="0.349" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.349">0.349</geoCoordinate>
<geoCoordinate box="[997,1074,1031,1057]" degrees="0.376" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.376">0.376</geoCoordinate>
(0.363); NOL =
<geoCoordinate box="[1274,1336,1031,1057]" degrees="0.181" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.181">0.181</geoCoordinate>
<geoCoordinate box="[1336,1413,1031,1057]" degrees="0.206" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.206">0.206</geoCoordinate>
(0.192); NOH =
<geoCoordinate box="[598,660,1071,1097]" degrees="0.122" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.122">0.122</geoCoordinate>
<geoCoordinate box="[660,737,1071,1098]" degrees="0.150" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.15">0.150</geoCoordinate>
(0.137); PEH =
<geoCoordinate box="[933,995,1071,1097]" degrees="0.240" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.24">0.240</geoCoordinate>
<geoCoordinate box="[995,1072,1071,1097]" degrees="0.260" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.26">0.260</geoCoordinate>
(0.251); PPL =
<geoCoordinate box="[1262,1324,1071,1098]" degrees="0.154" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.154">0.154</geoCoordinate>
<geoCoordinate box="[1324,1401,1071,1097]" degrees="0.187" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.187">0.187</geoCoordinate>
(0.170); PPH =
<geoCoordinate box="[588,650,1111,1137]" degrees="0.198" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.198">0.198</geoCoordinate>
<geoCoordinate box="[650,727,1111,1138]" degrees="0.225" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.225">0.225</geoCoordinate>
(0.211); PW =
<geoCoordinate box="[912,974,1111,1137]" degrees="0.420" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.42">0.420</geoCoordinate>
<geoCoordinate box="[975,1052,1111,1138]" degrees="0.454" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.454">0.454</geoCoordinate>
(0.437); SBPA =
<geoCoordinate box="[1260,1322,1111,1138]" degrees="0.159" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.159">0.159</geoCoordinate>
<geoCoordinate box="[1322,1399,1111,1137]" degrees="0.184" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.184">0.184</geoCoordinate>
(0.171); SPTI =
<geoCoordinate box="[590,652,1151,1178]" degrees="0.259" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.259">0.259</geoCoordinate>
<geoCoordinate box="[653,730,1151,1177]" degrees="0.280" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.28">0.280</geoCoordinate>
(0.267); PEW =
<geoCoordinate box="[932,994,1151,1178]" degrees="0.150" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.15">0.150</geoCoordinate>
<geoCoordinate box="[994,1071,1151,1178]" degrees="0.165" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.165">0.165</geoCoordinate>
(0.156); PNW =
<geoCoordinate box="[1278,1340,1151,1177]" degrees="0.174" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.174">0.174</geoCoordinate>
<geoCoordinate box="[1340,1417,1151,1177]" degrees="0.192" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.192">0.192</geoCoordinate>
(0.184); PPW =
<geoCoordinate box="[590,652,1191,1217]" degrees="0.317" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.317">0.317</geoCoordinate>
<geoCoordinate box="[653,730,1191,1217]" degrees="0.342" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.342">0.342</geoCoordinate>
(0.330); HFL =
<geoCoordinate box="[917,979,1190,1217]" degrees="0.555" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.555">0.555</geoCoordinate>
<geoCoordinate box="[979,1056,1191,1217]" degrees="0.603" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.603">0.603</geoCoordinate>
(0.583); HFWmax =
<geoCoordinate box="[1306,1368,1190,1217]" degrees="0.145" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.145">0.145</geoCoordinate>
<geoCoordinate box="[1368,1445,1191,1217]" degrees="0.173" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.173">0.173</geoCoordinate>
(0.162); HFWmin =
<geoCoordinate box="[645,707,1231,1257]" degrees="0.044" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.044">0.044</geoCoordinate>
<geoCoordinate box="[708,785,1230,1257]" degrees="0.054" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.054">0.054</geoCoordinate>
(0.048); CS =
<geoCoordinate box="[958,1020,1231,1257]" degrees="0.920" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.92">0.920</geoCoordinate>
<geoCoordinate box="[1021,1098,1230,1257]" degrees="0.957" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.957">0.957</geoCoordinate>
(0.942); ES =
<geoCoordinate box="[1269,1331,1231,1257]" degrees="0.214" direction="north" orientation="latitude" pageId="133" pageNumber="134" precision="55" value="0.214">0.214</geoCoordinate>
<geoCoordinate box="[1331,1408,1230,1257]" degrees="0.229" direction="west" orientation="longitude" pageId="133" pageNumber="134" precision="55" value="-0.229">0.229</geoCoordinate>
(0.221); SI = 8696 (92); OI = 2324 (24); CI = 8489 (86); WLI = 136143 (140); SBI = 2830 (29); PSI = 3942 (40); PWI = 202224 (211); PLI = 198244 (215); NI = 127155 (141); PNWI = 112123 (118); NLI = 5055 (53); FI = 302393 (341).
</paragraph>
<paragraph blockId="133.[498,1542,234,1894]" pageId="133" pageNumber="134">
<emphasis bold="true" box="[530,775,1390,1416]" pageId="133" pageNumber="134">Worker description:</emphasis>
In full-face view, head subquadrate, longer than broad (CI 8489). Mandibles densely, finely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin flat medially. Antennal scapes moderately long: when fully retracted, just reaching the posterior margin of the head capsule (SI 8696). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment slightly longer than the preceding two in combination. Frontal carinae long, extending past the antennal toruli by about three times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head weakly convex, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin medially weakly emarginate, but predominantly flat, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="134.[498,1542,234,1934]" pageId="134" pageNumber="135">
In profile view, compound eyes ovular and moderately large (OI 2324), with 10 ommatidia in longest row. Pronotal declivity distinct, neck and anterior face of pronotum forming a ~120
<emphasis box="[680,690,309,327]" italics="true" pageId="134" pageNumber="135">
<superScript attach="left" box="[680,690,309,327]" fontSize="8" pageId="134" pageNumber="135">°</superScript>
</emphasis>
angle; anterior face and dorsal face meeting at a ~120
<emphasis box="[1322,1332,309,327]" italics="true" pageId="134" pageNumber="135">
<superScript attach="left" box="[1322,1332,309,327]" fontSize="8" pageId="134" pageNumber="135">°</superScript>
</emphasis>
angle. Mesosoma evenly convex from where it joins the pronotal declivity to the propodeal spines, but the metanotal groove slightly impressed in some specimens. Promesonotal suture extending from the posterior margin of the procoxal insertion only to the mesothoracic spiracle, which is moderately well developed. Propodeal spiracle moderately well developed, directed posterolaterally, and separated from the propodeal declivity by about three and a half spiracle diameters. Propodeal spines well developed, and moderately long (PSI 3942), about as long as the propodeal declivity, tapering evenly from the base, upturned at the tips, and acute. Propodeal declivity flat, forming a rounded ~110
<emphasis box="[1454,1464,628,646]" italics="true" pageId="134" pageNumber="135">
<superScript attach="left" box="[1454,1464,628,646]" fontSize="8" pageId="134" pageNumber="135">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole long (PLI 198244), without tubercles anterodorsally. Subpetiolar process in the form of an acute tooth, ventral margin of petiole very weakly bulging medially. Petiolar peduncle moderately long: half the length of the petiole. Petiolar node robust and erect, nearly squamiform: transition between peduncle and node marked by a rounded angle of ~120
<emphasis box="[825,835,907,925]" italics="true" pageId="134" pageNumber="135">
<superScript attach="none" box="[825,835,907,925]" fontSize="8" pageId="134" pageNumber="135">°</superScript>
</emphasis>
, resulting in a concave anterior node face; anterior face rounding evenly into the dorsal face, which is evenly convex; dorsal face rounding evenly into the posterior face, which forms a ~80
<emphasis box="[996,1006,987,1005]" italics="true" pageId="134" pageNumber="135">
<superScript attach="left" box="[996,1006,987,1005]" fontSize="8" pageId="134" pageNumber="135">°</superScript>
</emphasis>
angle with the caudal cylinder, overhanging it. Postpetiole flat anteriorly, rounding evenly into the flattened dorsal face; weakly lobed ventrally.
</paragraph>
<paragraph blockId="134.[498,1542,234,1934]" pageId="134" pageNumber="135">In dorsal view, dorsal margin of pronotum delimited from the pronotal declivity by a carina. Humeri developed: evenly rounded and wider than the rest of the mesosoma; mesothoracic spiracles very weakly protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Metanotal groove visible in some specimens as a slight disruption in the sculpture. Propodeal spines narrowly approximated basally, but diverging rapidly and becoming parallel apically, their apices separated from each other by slightly less than their length, the negative space between them “V” shaped. Petiolar peduncle with spiracles protruding past the lateral margins; peduncle constricted anterior to them. Petiolar node campaniform: very weakly convex posteriorly, nearly flat; node broader than the peduncle, and slightly wider than the caudal cylinder. Postpetiole subquadrate and moderately to strongly broad (PWI 202224), articulating with most of the anterior margin of the gaster, leaving small, angulate margins on each side exposed. Anterior margin of the postpetiole very weakly convex, nearly flat, and evenly rounding into the lateral margins, which weakly diverge to the angulate posterior corners; posterior margin broadly concave. Metafemur strongly incrassate (FI 302393).</paragraph>
<paragraph blockId="134.[498,1542,234,1934]" lastBlockId="135.[498,1542,234,1257]" lastPageId="135" lastPageNumber="136" pageId="134" pageNumber="135">Sculpture: median clypeal carina present, extending posteriorly to the level of the antennal toruli, and flanked on either side by two equally strong carinae. Lateral clypeal lobes with additional, weaker carinae; ground sculpture weakly areolate. Antennal scapes areolate. Cephalic dorsum areolate, but with rugae overlying the ground sculpture, becoming costate between the frontal carinae; concentric costulae surrounding the antennal insertions. Lateral surfaces of head sculptured similarly to the dorsum of the head, but with rugae becoming weaker posterior to the compound eye. Ventral surface of head predominantly smooth and shining. Mesosoma with areolate sculpture on the pronotal neck. Lateral surface of the mesosoma strongly areolate, with overlying rugose-costate sculpture on the pronotum, on the border of the meso- and metapleurae, and on the propodeal declivity. Dorsal surface of mesosoma areolate-rugulose, with rugae overlying the ground sculpture. Femora shining through weakly areolate sculpture. Petiole and postpetiole predominantly areolate, but sculpture is weaker on the petiolar peduncle, dorsal face of the petiolar node, and the anteromedial part of the postpetiole. First gastral tergite smooth and shining, with weak spectral iridescence. First gastral sternite smooth and shining, without spectral iridescence.</paragraph>
<paragraph blockId="135.[498,1542,234,1257]" pageId="135" pageNumber="136">Setae: antennal scapes and funiculi with short, decumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, blunt-tipped setae, the longest of which are about the width of the compound eye. The head bears ~32, mesosoma ~38, petiole 10, postpetiole ~20, and first gastral tergite ~72 setae. Short, sparse pubescence present over the entire body, but difficult to detect against the ground sculpture.</paragraph>
<paragraph blockId="135.[498,1542,234,1257]" pageId="135" pageNumber="136">Color: predominantly medium brown, with mandibles, antennae, pronotal neck, tibiae, tarsi, propodeal spines, and sting testaceous yellow.</paragraph>
<paragraph blockId="135.[498,1542,234,1257]" box="[530,734,991,1017]" pageId="135" pageNumber="136">
<emphasis bold="true" box="[530,601,991,1017]" pageId="135" pageNumber="136">Gyne:</emphasis>
Unknown.
</paragraph>
<paragraph blockId="135.[498,1542,234,1257]" box="[530,730,1031,1057]" pageId="135" pageNumber="136">
<emphasis bold="true" box="[530,597,1031,1057]" pageId="135" pageNumber="136">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection box="[530,1522,1071,1098]" pageId="135" pageNumber="136" type="etymology">
<paragraph blockId="135.[498,1542,234,1257]" box="[530,1522,1071,1098]" pageId="135" pageNumber="136">
<emphasis bold="true" box="[530,668,1071,1097]" pageId="135" pageNumber="136">Etymology:</emphasis>
Morphological, for the close resemblance to
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1201,1516,1071,1097]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="leucacanthus">
<emphasis box="[1201,1516,1071,1097]" italics="true" pageId="135" pageNumber="136">Temnothorax leucacanthus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="135" pageNumber="136" type="discussion">
<paragraph blockId="135.[498,1542,234,1257]" pageId="135" pageNumber="136">
<emphasis bold="true" box="[530,670,1111,1137]" pageId="135" pageNumber="136">Comments:</emphasis>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[679,1026,1111,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis box="[679,1026,1111,1137]" italics="true" pageId="135" pageNumber="136">Temnothorax leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1035,1126,1111,1137]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
is known from a single collection made by Phil Ward in
<collectingRegion box="[765,913,1151,1177]" country="Mexico" name="Oaxaca" pageId="135" pageNumber="136">Oaxaca state</collectingRegion>
,
<collectingCountry box="[926,1015,1151,1177]" name="Mexico" pageId="135" pageNumber="136">Mexico</collectingCountry>
. The
<typeStatus box="[1079,1129,1151,1177]" pageId="135" pageNumber="136">type</typeStatus>
series was nesting in the dead stalk of a plant of the
<taxonomicName box="[691,815,1191,1217]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="135" pageNumber="136" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
in littoral vegetation.
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1073,1418,1191,1217]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="leucacanthoides" status="sp. nov.">
<emphasis box="[1073,1418,1191,1217]" italics="true" pageId="135" pageNumber="136">Temnothorax leucacanthoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1426,1516,1191,1217]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
is morphologically close to
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[795,977,1230,1256]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="leucacanthus">
<emphasis box="[795,977,1230,1256]" italics="true" pageId="135" pageNumber="136">T. leucacanthus</emphasis>
</taxonomicName>
from neighboring
<collectingRegion box="[1202,1373,1231,1257]" country="Mexico" name="Guerrero" pageId="135" pageNumber="136">Guerrero state</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>