664 lines
74 KiB
XML
664 lines
74 KiB
XML
<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FE7FFE64B49C1FF17DC0225D" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax tenuisculptus" docType="treatment" docVersion="4" lastPageNumber="446" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="441" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Prebus, Matthew M.</mods:namePart>
|
||
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States & Department of Entomology & Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
|
||
<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
|
||
<mods:title>PeerJ</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2021</mods:date>
|
||
<mods:detail type="series">
|
||
<mods:title>e 11514</mods:title>
|
||
</mods:detail>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2021-06-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>9</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>462</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
|
||
<mods:identifier type="ISSN">2167-8359</mods:identifier>
|
||
<mods:identifier type="PMC">PMC8254503</mods:identifier>
|
||
<mods:identifier type="PubMed">34249486</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102664" ID-GBIF-Taxon="183542638" ID-Zenodo-Dep="5102664" LSID="urn:lsid:plazi:treatment:054FDB70FE7FFE64B49C1FF17DC0225D" httpUri="http://treatment.plazi.org/id/054FDB70FE7FFE64B49C1FF17DC0225D" lastPageId="446" lastPageNumber="446" pageId="440" pageNumber="441">
|
||
<subSubSection box="[498,1105,1516,1543]" pageId="440" pageNumber="441" type="nomenclature">
|
||
<paragraph blockId="440.[498,1105,1516,1583]" box="[498,1105,1516,1543]" pageId="440" pageNumber="441">
|
||
<heading box="[498,1105,1516,1543]" level="2" pageId="440" pageNumber="441">
|
||
<taxonomicName authority="(Baroni Urbani, 1978)" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[498,1105,1516,1543]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="440" pageNumber="441" phylum="Arthropoda" rank="species" species="tenuisculptus">
|
||
<emphasis bold="true" box="[498,1105,1516,1543]" pageId="440" pageNumber="441">
|
||
<emphasis bold="true" box="[498,826,1516,1543]" italics="true" pageId="440" pageNumber="441">Temnothorax tenuisculptus</emphasis>
|
||
(
|
||
<bibRefCitation author="Baroni Urbani C." box="[845,1095,1516,1543]" journalOrPublisher="Entomologica Basiliensia" pageId="440" pageNumber="441" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
|
||
<emphasis bold="true" box="[845,1095,1516,1543]" italics="true" pageId="440" pageNumber="441">Baroni Urbani, 1978</emphasis>
|
||
</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[498,946,1557,1583]" pageId="440" pageNumber="441" type="distribution">
|
||
<paragraph blockId="440.[498,1105,1516,1583]" box="[498,946,1557,1583]" pageId="440" pageNumber="441">
|
||
Distribution, worker & gyne:
|
||
<figureCitation box="[847,940,1557,1583]" captionStart="Figure 169" captionStartId="440.[524,589,947,969]" captionTargetBox="[498,1541,234,935]" captionTargetId="figure-172@440.[498,1534,234,919]" captionTargetPageId="440" captionText="Figure 169 Temnothorax tenuisculptus. (A–C) Holotype worker (MCZENT00032435). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D–G) Gyne (CASENT0756078). (D) Full-face view. (E) Profile view. (F) Dorsal view. (G) Wings. (H) Geographical and elevational distribution of specimens examined. Type locality is represented by a triangle, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-169" figureDoi="http://doi.org/10.5281/zenodo.5102499" httpUri="https://zenodo.org/record/5102499/files/figure.png" pageId="440" pageNumber="441">Fig. 169</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="440" pageNumber="441" type="reference_group">
|
||
<paragraph blockId="440.[498,1505,1619,1765]" pageId="440" pageNumber="441">
|
||
<treatmentCitationGroup pageId="440" pageNumber="441">
|
||
<treatmentCitation author="Baroni Urbani C." box="[498,1106,1619,1646]" page="517" pageId="440" pageNumber="441" year="1978">
|
||
<taxonomicName authority="Baroni Urbani, 1978: 517" authorityName="Baroni Urbani" authorityPageNumber="517" authorityYear="1978" box="[498,1047,1619,1645]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="440" pageNumber="441" phylum="Arthropoda" rank="species" species="tenuisculptus">
|
||
<emphasis box="[498,1047,1619,1645]" italics="true" pageId="440" pageNumber="441">
|
||
Leptothorax tenuisculptus
|
||
<bibRefCitation author="Baroni Urbani C." box="[806,1106,1619,1646]" journalOrPublisher="Entomologica Basiliensia" pageId="440" pageNumber="441" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">Baroni Urbani, 1978: 517</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
<typeStatus box="[1119,1227,1619,1645]" pageId="440" pageNumber="441">Holotype</typeStatus>
|
||
worker.
|
||
<location LSID="urn:lsid:plazi:treatment:054FDB70FE7FFE64B49C1FF17DC0225D:88393CBDFE7FFE62B05B1C4E7CEF2139" box="[1333,1501,1619,1646]" country="Mexico" name="Pueblo Nuevo" pageId="440" pageNumber="441" stateProvince="Veracruz de Ignacio de la Llave">Pueblo Nuevo</location>
|
||
,
|
||
<collectingRegion box="[498,606,1660,1686]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="440" pageNumber="441">Veracruz</collectingRegion>
|
||
,
|
||
<collectingCountry box="[620,709,1659,1685]" name="Mexico" pageId="440" pageNumber="441">Mexico</collectingCountry>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="440.[498,1505,1619,1765]" pageId="440" pageNumber="441">
|
||
<treatmentCitationGroup pageId="440" pageNumber="441">
|
||
<taxonomicName authority="(Baroni Urbani)" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[498,1016,1699,1726]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="440" pageNumber="441" phylum="Arthropoda" rank="species" species="tenuisculptus">
|
||
<emphasis box="[498,815,1699,1725]" italics="true" pageId="440" pageNumber="441">Temnothorax tenuisculptus</emphasis>
|
||
(Baroni Urbani)
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation author="Bolton B." box="[1028,1234,1699,1726]" page="272" pageId="440" pageNumber="441" year="2003">
|
||
<bibRefCitation author="Bolton B." box="[1028,1234,1699,1726]" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="440" pageNumber="441" pagination="370" part="71" refId="ref270076" refString="Bolton B. 2003. Synopsis and Classification of Formicidae. Memoirs of the American Entomological Institute 71: 370." title="Synopsis and Classification of Formicidae" type="journal article" year="2003">
|
||
<emphasis box="[1028,1174,1699,1725]" italics="true" pageId="440" pageNumber="441">
|
||
<collectingRegion box="[1028,1106,1699,1725]" country="United Kingdom" name="Bolton" pageId="440" pageNumber="441">
|
||
<author box="[1028,1106,1699,1725]" fns="*" inm="true" npo="*" pageId="440" pageNumber="441">Bolton</author>
|
||
</collectingRegion>
|
||
,
|
||
<year box="[1119,1174,1699,1725]" pageId="440" pageNumber="441">2003</year>
|
||
</emphasis>
|
||
: 272
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. First combination in
|
||
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[498,655,1739,1765]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="440" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[498,655,1739,1765]" italics="true" pageId="440" pageNumber="441">Temnothorax</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="441" lastPageNumber="442" pageId="440" pageNumber="441" type="materials_examined">
|
||
<paragraph blockId="440.[498,1530,1801,1908]" pageId="440" pageNumber="441">
|
||
<emphasis bold="true" box="[530,834,1801,1827]" pageId="440" pageNumber="441">Type material examined:</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3321861342" collectingDate="1953-08-09" collectionCode="MCZC" collectorName="E. O. Wilson" country="Mexico" location="Pueblo Nuevo" pageId="440" pageNumber="441" specimenCode="MCZENT00032435" specimenCount="1" specimenCount-worker="1" stateProvince="Veracruz" typeStatus="holotype">
|
||
<emphasis box="[843,1041,1801,1827]" italics="true" pageId="440" pageNumber="441">
|
||
<typeStatus box="[843,946,1801,1827]" pageId="440" pageNumber="441">Holotype</typeStatus>
|
||
worker:
|
||
</emphasis>
|
||
<collectingCountry box="[1049,1164,1801,1827]" name="Mexico" pageId="440" pageNumber="441">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion box="[1176,1285,1802,1828]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="440" pageNumber="441">Veracruz</collectingRegion>
|
||
:
|
||
<location LSID="urn:lsid:plazi:treatment:054FDB70FE7FFE64B49C1FF17DC0225D:88393CBDFE7FFE62B07C1D147C882073" box="[1298,1466,1801,1828]" country="Mexico" name="Pueblo Nuevo" pageId="440" pageNumber="441" stateProvince="Veracruz">Pueblo Nuevo</location>
|
||
,
|
||
<locationDeviation pageId="440" pageNumber="441">near Tetzonapa</locationDeviation>
|
||
,
|
||
<collectorName box="[636,779,1841,1867]" pageId="440" pageNumber="441">E.O. Wilson</collectorName>
|
||
<date box="[788,961,1841,1868]" pageId="440" pageNumber="441" value="1953-08-09">
|
||
<collectingDate box="[788,961,1841,1868]" pageId="440" pageNumber="441" value="1953-08-09">9 August 1953</collectingDate>
|
||
</date>
|
||
, rainforest,
|
||
<specimenCount box="[1102,1206,1841,1867]" pageId="440" pageNumber="441" type="worker">1 worker</specimenCount>
|
||
(M.C.Z. Type 32435,
|
||
<specimenCode box="[498,731,1881,1908]" pageId="440" pageNumber="441">MCZENT00032435</specimenCode>
|
||
, top specimen on pin) [
|
||
<collectionCode box="[1013,1104,1880,1907]" pageId="440" pageNumber="441">MCZC</collectionCode>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="441.[498,1542,234,1895]" pageId="441" pageNumber="442">
|
||
<materialsCitation ID-GBIF-Occurrence="3321861393" collectingDate="1953-08-09" collectionCode="MCZC, NHMB" collectorName="M. C. Z. Type" country="Mexico" location="Pueblo Nuevo" pageId="441" pageNumber="442" specimenCount="3" specimenCount-worker="3" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
|
||
<emphasis box="[530,737,234,260]" italics="true" pageId="441" pageNumber="442">
|
||
<typeStatus box="[530,632,235,260]" pageId="441" pageNumber="442">Paratype</typeStatus>
|
||
workers:
|
||
</emphasis>
|
||
same pin as
|
||
<typeStatus box="[890,994,234,260]" pageId="441" pageNumber="442">holotype</typeStatus>
|
||
,
|
||
<specimenCount box="[1006,1110,234,260]" pageId="441" pageNumber="442" type="worker">1 worker</specimenCount>
|
||
(bottom specimen on pin) [
|
||
<collectionCode box="[1441,1531,234,261]" pageId="441" pageNumber="442">MCZC</collectionCode>
|
||
]; same data as holotype,
|
||
<specimenCount box="[774,878,274,300]" pageId="441" pageNumber="442" type="worker">1 worker</specimenCount>
|
||
(
|
||
<collectorName box="[896,1047,274,301]" pageId="441" pageNumber="442">M.C.Z. Type</collectorName>
|
||
32435, MCZENT00578578) [
|
||
<collectionCode box="[1396,1484,273,300]" pageId="441" pageNumber="442">MCZC</collectionCode>
|
||
]
|
||
<specimenCount pageId="441" pageNumber="442" type="worker">1 worker</specimenCount>
|
||
(images of CASENT0913005 examined on antweb.org) [
|
||
<collectionCode box="[1251,1347,313,340]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="441" pageNumber="442">NHMB</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="441.[498,1542,234,1895]" pageId="441" pageNumber="442">
|
||
<emphasis bold="true" box="[530,886,353,380]" pageId="441" pageNumber="442">Non-type material examined:</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3321861735" collectingDate="1976-04-13" collectionCode="USNM" collectorName="J. Kline & R. Eads" country="Mexico" location="San Luis Potosi" pageId="441" pageNumber="442" specimenCode="ANTC43726, CASENT0756076, CASENT0756084" specimenCount="10" specimenCount-worker="3" stateProvince="San Luis Potosi">
|
||
<collectingCountry box="[894,1009,354,380]" name="Mexico" pageId="441" pageNumber="442">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion box="[1020,1203,354,380]" country="Mexico" name="San Luis Potosi" pageId="441" pageNumber="442">San Luis Potosí</collectingRegion>
|
||
: [quarantine],
|
||
<date box="[1375,1537,354,380]" pageId="441" pageNumber="442" value="1976-04-13">
|
||
<collectingDate box="[1375,1537,354,380]" pageId="441" pageNumber="442" value="1976-04-13">13 April 1976</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[498,712,394,420]" pageId="441" pageNumber="442">J. Kline & R. Eads</collectorName>
|
||
#
|
||
<specimenCode box="[735,885,394,420]" pageId="441" pageNumber="442">ANTC43726</specimenCode>
|
||
,
|
||
<taxonomicName box="[898,1034,394,420]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="441" pageNumber="442" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis box="[898,993,394,420]" italics="true" pageId="441" pageNumber="442">Quercus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName box="[1103,1287,394,420]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="441" pageNumber="442" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis box="[1103,1246,394,420]" italics="true" pageId="441" pageNumber="442">Epidendrum</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
orchid plant,
|
||
<specimenCount pageId="441" pageNumber="442">7 alate gynes</specimenCount>
|
||
&
|
||
<specimenCount box="[602,718,433,460]" pageId="441" pageNumber="442" type="worker">3 workers</specimenCount>
|
||
(
|
||
<specimenCode box="[734,944,433,460]" pageId="441" pageNumber="442">CASENT0756076</specimenCode>
|
||
-
|
||
<specimenCode box="[952,1162,433,460]" pageId="441" pageNumber="442">CASENT0756084</specimenCode>
|
||
) [
|
||
<collectionCode box="[1186,1278,433,460]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="441" pageNumber="442" type="Museum">USNM</collectionCode>
|
||
].
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3321861477" collectingDate="2016-06-04" collectionCode="UCDC" collectorName="M. M. Prebus" elevation="970" latitude="18.51232" location="San Andres Tuxtla" longLatPrecision="1" longitude="-95.16729" municipality="San Andres Tuxtla" pageId="441" pageNumber="442" specimenCode="CASENT0757196, CASENT0757198, CASENT0758683" specimenCount="4" specimenCount-worker="4" stateProvince="Veracruz">
|
||
<collectingRegion box="[1296,1405,434,460]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="441" pageNumber="442">Veracruz</collectingRegion>
|
||
:
|
||
<locationDeviation bearing="45.0" box="[1418,1525,433,460]" distance="9000.0" pageId="441" pageNumber="442">
|
||
<quantity box="[1418,1478,433,460]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="441" pageNumber="442" unit="km" value="9.0">9 km</quantity>
|
||
NE
|
||
</locationDeviation>
|
||
<collectingMunicipality box="[498,721,473,499]" pageId="441" pageNumber="442">San Andrés Tuxtla</collectingMunicipality>
|
||
,
|
||
<geoCoordinate box="[734,871,469,500]" degrees="18.51232" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="1" value="18.51232">
|
||
18.51232
|
||
<emphasis box="[838,848,469,487]" italics="true" pageId="441" pageNumber="442">
|
||
<superScript attach="right" box="[838,848,469,487]" fontSize="8" pageId="441" pageNumber="442">°</superScript>
|
||
</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
<geoCoordinate box="[880,1024,469,500]" degrees="95.16729" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="1" value="-95.16729">
|
||
95.16729
|
||
<emphasis box="[984,994,469,487]" italics="true" pageId="441" pageNumber="442">
|
||
<superScript attach="right" box="[984,994,469,487]" fontSize="8" pageId="441" pageNumber="442">°</superScript>
|
||
</emphasis>
|
||
W
|
||
</geoCoordinate>
|
||
±
|
||
<quantity box="[1058,1106,473,500]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="441" pageNumber="442" unit="m" value="4.0">4 m</quantity>
|
||
,
|
||
<quantity box="[1120,1194,473,500]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="441" pageNumber="442" unit="m" value="970.0">
|
||
<elevation box="[1120,1194,473,500]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="441" pageNumber="442" unit="m" value="970.0">970 m</elevation>
|
||
</quantity>
|
||
±
|
||
<quantity box="[1228,1275,473,500]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="441" pageNumber="442" unit="m" value="5.0">5 m</quantity>
|
||
<date box="[1284,1426,473,500]" pageId="441" pageNumber="442" value="2016-06-04">
|
||
<collectingDate box="[1284,1426,473,500]" pageId="441" pageNumber="442" value="2016-06-04">4 June 2016</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName pageId="441" pageNumber="442">M.M. Prebus</collectorName>
|
||
#
|
||
<accessionNumber box="[593,732,513,539]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MMP02671" pageId="441" pageNumber="442">MMP02671</accessionNumber>
|
||
, mixed tropical temperate mesic forest, between leaves and stalk of
|
||
<taxonomicName box="[498,614,553,579]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="441" pageNumber="442" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[498,614,553,579]" italics="true" pageId="441" pageNumber="442">Tillandsia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount box="[629,745,553,579]" pageId="441" pageNumber="442" type="worker">4 workers</specimenCount>
|
||
(
|
||
<specimenCode box="[761,972,553,580]" pageId="441" pageNumber="442">CASENT0757196</specimenCode>
|
||
-
|
||
<specimenCode box="[979,1190,553,580]" pageId="441" pageNumber="442">CASENT0757198</specimenCode>
|
||
,
|
||
<specimenCode box="[1202,1412,553,580]" pageId="441" pageNumber="442">CASENT0758683</specimenCode>
|
||
) [
|
||
<collectionCode box="[1435,1525,552,579]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="441" pageNumber="442" type="Museum">UCDC</collectionCode>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[530,1462,593,619]" pageId="441" pageNumber="442" type="distribution">
|
||
<paragraph blockId="441.[498,1542,234,1895]" box="[530,1462,593,619]" pageId="441" pageNumber="442">
|
||
<emphasis bold="true" box="[530,755,593,619]" pageId="441" pageNumber="442">Geographic range:</emphasis>
|
||
Low elevations of southern
|
||
<collectingCountry box="[1090,1176,593,619]" name="Mexico" pageId="441" pageNumber="442">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion box="[1193,1303,593,619]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="441" pageNumber="442">Veracruz</collectingRegion>
|
||
) (
|
||
<figureCitation box="[1330,1446,593,619]" captionStart="Figure 169" captionStartId="440.[524,589,947,969]" captionTargetBox="[498,1541,234,935]" captionTargetId="figure-172@440.[498,1534,234,919]" captionTargetPageId="440" captionText="Figure 169 Temnothorax tenuisculptus. (A–C) Holotype worker (MCZENT00032435). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D–G) Gyne (CASENT0756078). (D) Full-face view. (E) Profile view. (F) Dorsal view. (G) Wings. (H) Geographical and elevational distribution of specimens examined. Type locality is represented by a triangle, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-169" figureDoi="http://doi.org/10.5281/zenodo.5102499" httpUri="https://zenodo.org/record/5102499/files/figure.png" pageId="441" pageNumber="442">Fig. 169H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="441" pageNumber="442" type="diagnosis">
|
||
<paragraph blockId="441.[498,1542,234,1895]" pageId="441" pageNumber="442">
|
||
<emphasis bold="true" box="[530,754,632,659]" pageId="441" pageNumber="442">Worker diagnosis:</emphasis>
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[762,1078,633,659]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="tenuisculptus">
|
||
<emphasis box="[762,1078,633,659]" italics="true" pageId="441" pageNumber="442">Temnothorax tenuisculptus</emphasis>
|
||
</taxonomicName>
|
||
can be separated from all other species in the
|
||
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[576,651,673,699]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis box="[576,651,673,699]" italics="true" pageId="441" pageNumber="442">salvini</emphasis>
|
||
</taxonomicName>
|
||
clade by the following character combination: in profile view, dorsum of mesosoma strongly sinuate; propodeum depressed below the level of the promesonotum; hind femora strongly incrassate; petiolar node subquadrate, not leaning posteriorly; postpetiole moderately broad, greater than 1.5 times the width of the caudal cylinder of the petiole, but less than 2.2 times; setae on head, mesosoma, waist segments and gaster erect, moderately long, sparse and blunt (never long and tapering);integument predominantly testaceous brown, with apex of mandibles, pedicel of antenna, pronotal neck, tibiae, tarsi, and sting testaceous yellow.
|
||
</paragraph>
|
||
<paragraph blockId="441.[498,1542,234,1895]" pageId="441" pageNumber="442">
|
||
<emphasis bold="true" box="[530,717,991,1017]" pageId="441" pageNumber="442">Similar species:</emphasis>
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[724,1041,991,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
|
||
<emphasis box="[724,1041,991,1017]" italics="true" pageId="441" pageNumber="442">Temnothorax acutispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1048,1140,992,1018]" pageId="441" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1151,1291,991,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
|
||
<emphasis box="[1151,1291,991,1017]" italics="true" pageId="441" pageNumber="442">T. tuxtlanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1298,1389,992,1018]" pageId="441" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1401,1502,992,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="xincai" status="sp. nov.">
|
||
<emphasis box="[1401,1502,992,1017]" italics="true" pageId="441" pageNumber="442">T. xincai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="441" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, members of the
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[757,841,1032,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[757,841,1032,1057]" italics="true" pageId="441" pageNumber="442">augusti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[902,1018,1031,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="subditivus">
|
||
<emphasis box="[902,1018,1031,1057]" italics="true" pageId="441" pageNumber="442">subditivus</emphasis>
|
||
</taxonomicName>
|
||
groups.
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1122,1439,1031,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="tenuisculptus">
|
||
<emphasis box="[1122,1439,1031,1057]" italics="true" pageId="441" pageNumber="442">Temnothorax tenuisculptus</emphasis>
|
||
</taxonomicName>
|
||
can be separated from all of the above species, except for the
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[1120,1204,1072,1097]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[1120,1204,1072,1097]" italics="true" pageId="441" pageNumber="442">augusti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1261,1377,1071,1097]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="subditivus">
|
||
<emphasis box="[1261,1377,1071,1097]" italics="true" pageId="441" pageNumber="442">subditivus</emphasis>
|
||
</taxonomicName>
|
||
groups by the strongly incrassate hind femora. The petiolar node, which is erect in
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1313,1497,1111,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="tenuisculptus">
|
||
<emphasis box="[1313,1497,1111,1137]" italics="true" pageId="441" pageNumber="442">T. tenuisculptus</emphasis>
|
||
</taxonomicName>
|
||
, can be used to distinguish it from similar appearing members of the
|
||
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[1297,1381,1152,1177]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[1297,1381,1152,1177]" italics="true" pageId="441" pageNumber="442">augusti</emphasis>
|
||
</taxonomicName>
|
||
group, which have posteriorly leaning petiolar nodes. Due to the smooth integument of the head,
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[498,682,1230,1256]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="tenuisculptus">
|
||
<emphasis box="[498,682,1230,1256]" italics="true" pageId="441" pageNumber="442">T. tenuisculptus</emphasis>
|
||
</taxonomicName>
|
||
may be confused with
|
||
<taxonomicName box="[958,1068,1230,1256]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="politus">
|
||
<emphasis box="[958,1068,1230,1256]" italics="true" pageId="441" pageNumber="442">T. politus</emphasis>
|
||
</taxonomicName>
|
||
, but the short, blunt tipped setae and sinuate mesosoma of the former will separate it from
|
||
<taxonomicName box="[1130,1239,1270,1296]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="politus">
|
||
<emphasis box="[1130,1239,1270,1296]" italics="true" pageId="441" pageNumber="442">T. politus</emphasis>
|
||
</taxonomicName>
|
||
, which has long, tapering setae and an arched mesosoma. Similarly,
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[982,1163,1310,1336]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="tenuisculptus">
|
||
<emphasis box="[982,1163,1310,1336]" italics="true" pageId="441" pageNumber="442">T. tenuisculptus</emphasis>
|
||
</taxonomicName>
|
||
may be mistaken for some of the more lightly sculptured specimens of
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[942,1091,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="subditivus">
|
||
<emphasis box="[942,1091,1350,1376]" italics="true" pageId="441" pageNumber="442">T. subditivus</emphasis>
|
||
</taxonomicName>
|
||
but differs from it by the sinuate mesosoma (uniformly convex in
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[888,1037,1390,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="subditivus">
|
||
<emphasis box="[888,1037,1390,1416]" italics="true" pageId="441" pageNumber="442">T. subditivus</emphasis>
|
||
</taxonomicName>
|
||
) and the subquadrate petiolar node (squamiform in
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[692,840,1430,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="subditivus">
|
||
<emphasis box="[692,840,1430,1456]" italics="true" pageId="441" pageNumber="442">T. subditivus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="445" lastPageNumber="446" pageId="441" pageNumber="442" type="description">
|
||
<paragraph blockId="441.[498,1542,234,1895]" lastBlockId="442.[498,1542,234,1934]" lastPageId="442" lastPageNumber="443" pageId="441" pageNumber="442">
|
||
<emphasis bold="true" box="[530,1018,1469,1496]" pageId="441" pageNumber="442">
|
||
Worker measurements & indices (
|
||
<emphasis bold="true" box="[941,957,1470,1496]" italics="true" pageId="441" pageNumber="442">n</emphasis>
|
||
= 9):
|
||
</emphasis>
|
||
SL =
|
||
<geoCoordinate box="[1084,1146,1469,1496]" degrees="0.501" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.501">0.501</geoCoordinate>
|
||
<geoCoordinate box="[1146,1223,1469,1496]" degrees="0.592" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.592">–0.592</geoCoordinate>
|
||
(0.554); FRS =
|
||
<geoCoordinate box="[1402,1464,1470,1496]" degrees="0.171" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.171">0.171</geoCoordinate>
|
||
<geoCoordinate box="[1464,1541,1469,1496]" degrees="0.205" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.205">–0.205</geoCoordinate>
|
||
(0.191); CW =
|
||
<geoCoordinate box="[678,740,1509,1536]" degrees="0.575" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.575">0.575</geoCoordinate>
|
||
<geoCoordinate box="[740,817,1509,1535]" degrees="0.698" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.698">–0.698</geoCoordinate>
|
||
(0.654); CWb =
|
||
<geoCoordinate box="[1020,1082,1509,1536]" degrees="0.516" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.516">0.516</geoCoordinate>
|
||
<geoCoordinate box="[1082,1159,1509,1535]" degrees="0.643" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.643">–0.643</geoCoordinate>
|
||
(0.604); PoOC =
|
||
<geoCoordinate box="[1371,1433,1509,1536]" degrees="0.255" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.255">0.255</geoCoordinate>
|
||
<geoCoordinate box="[1434,1511,1509,1535]" degrees="0.312" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.312">–0.312</geoCoordinate>
|
||
(0.283); CL =
|
||
<geoCoordinate box="[665,727,1549,1575]" degrees="0.647" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.647">0.647</geoCoordinate>
|
||
<geoCoordinate box="[727,804,1549,1575]" degrees="0.767" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.767">–0.767</geoCoordinate>
|
||
(0.720); EL =
|
||
<geoCoordinate box="[977,1039,1549,1575]" degrees="0.134" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.134">0.134</geoCoordinate>
|
||
<geoCoordinate box="[1039,1116,1549,1575]" degrees="0.169" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.169">–0.169</geoCoordinate>
|
||
(0.154); EW =
|
||
<geoCoordinate box="[1302,1364,1549,1575]" degrees="0.096" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.096">0.096</geoCoordinate>
|
||
<geoCoordinate box="[1364,1441,1549,1575]" degrees="0.127" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.127">–0.127</geoCoordinate>
|
||
(0.114); MD =
|
||
<geoCoordinate box="[580,642,1589,1616]" degrees="0.158" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.158">0.158</geoCoordinate>
|
||
<geoCoordinate box="[642,719,1589,1615]" degrees="0.189" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.189">–0.189</geoCoordinate>
|
||
(0.170); WL =
|
||
<geoCoordinate box="[904,966,1589,1615]" degrees="0.748" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.748">0.748</geoCoordinate>
|
||
<geoCoordinate box="[966,1043,1589,1615]" degrees="0.919" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.919">–0.919</geoCoordinate>
|
||
(0.857); SPST =
|
||
<geoCoordinate box="[1245,1307,1589,1615]" degrees="0.216" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.216">0.216</geoCoordinate>
|
||
<geoCoordinate box="[1308,1385,1589,1615]" degrees="0.317" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.317">–0.317</geoCoordinate>
|
||
(0.267); MPST =
|
||
<geoCoordinate box="[607,669,1629,1655]" degrees="0.244" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.244">0.244</geoCoordinate>
|
||
<geoCoordinate box="[669,746,1629,1655]" degrees="0.320" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.32">–0.320</geoCoordinate>
|
||
(0.279); PEL =
|
||
<geoCoordinate box="[935,997,1629,1655]" degrees="0.300" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.3">0.300</geoCoordinate>
|
||
<geoCoordinate box="[997,1074,1629,1655]" degrees="0.388" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.388">–0.388</geoCoordinate>
|
||
(0.353); NOL =
|
||
<geoCoordinate box="[1274,1336,1629,1655]" degrees="0.147" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.147">0.147</geoCoordinate>
|
||
<geoCoordinate box="[1336,1413,1629,1655]" degrees="0.199" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.199">–0.199</geoCoordinate>
|
||
(0.176); NOH =
|
||
<geoCoordinate box="[598,660,1669,1695]" degrees="0.119" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.119">0.119</geoCoordinate>
|
||
<geoCoordinate box="[660,737,1669,1696]" degrees="0.159" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.159">–0.159</geoCoordinate>
|
||
(0.139); PEH =
|
||
<geoCoordinate box="[933,995,1669,1695]" degrees="0.196" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.196">0.196</geoCoordinate>
|
||
<geoCoordinate box="[995,1072,1669,1696]" degrees="0.255" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.255">–0.255</geoCoordinate>
|
||
(0.222); PPL =
|
||
<geoCoordinate box="[1262,1324,1669,1696]" degrees="0.156" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.156">0.156</geoCoordinate>
|
||
<geoCoordinate box="[1324,1401,1669,1696]" degrees="0.215" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.215">–0.215</geoCoordinate>
|
||
(0.178); PW =
|
||
<geoCoordinate box="[577,639,1709,1735]" degrees="0.372" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.372">0.372</geoCoordinate>
|
||
<geoCoordinate box="[640,716,1708,1735]" degrees="0.456" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.456">–0.456</geoCoordinate>
|
||
(0.420); SBPA =
|
||
<geoCoordinate box="[925,987,1709,1735]" degrees="0.133" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.133">0.133</geoCoordinate>
|
||
<geoCoordinate box="[987,1064,1709,1735]" degrees="0.194" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.194">–0.194</geoCoordinate>
|
||
(0.166); SPTI =
|
||
<geoCoordinate box="[1263,1325,1708,1735]" degrees="0.205" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.205">0.205</geoCoordinate>
|
||
<geoCoordinate box="[1325,1402,1709,1735]" degrees="0.274" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.274">–0.274</geoCoordinate>
|
||
(0.250); PEW =
|
||
<geoCoordinate box="[594,656,1749,1775]" degrees="0.131" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.131">0.131</geoCoordinate>
|
||
<geoCoordinate box="[656,733,1748,1775]" degrees="0.164" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.164">–0.164</geoCoordinate>
|
||
(0.153); PNW =
|
||
<geoCoordinate box="[940,1002,1748,1774]" degrees="0.136" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.136">0.136</geoCoordinate>
|
||
<geoCoordinate box="[1002,1079,1749,1775]" degrees="0.202" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.202">–0.202</geoCoordinate>
|
||
(0.170); PPW =
|
||
<geoCoordinate box="[1281,1343,1749,1775]" degrees="0.234" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.234">0.234</geoCoordinate>
|
||
<geoCoordinate box="[1344,1421,1749,1775]" degrees="0.288" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.288">–0.288</geoCoordinate>
|
||
(0.266); HFL =
|
||
<geoCoordinate box="[586,648,1788,1815]" degrees="0.535" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.535">0.535</geoCoordinate>
|
||
<geoCoordinate box="[648,725,1788,1814]" degrees="0.668" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.668">–0.668</geoCoordinate>
|
||
(0.621); HFWmax =
|
||
<geoCoordinate box="[981,1043,1788,1815]" degrees="0.151" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.151">0.151</geoCoordinate>
|
||
<geoCoordinate box="[1044,1121,1788,1814]" degrees="0.199" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.199">–0.199</geoCoordinate>
|
||
(0.180); HFWmin =
|
||
<geoCoordinate box="[1374,1436,1788,1814]" degrees="0.042" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.042">0.042</geoCoordinate>
|
||
<geoCoordinate box="[1437,1514,1788,1815]" degrees="0.056" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.056">–0.056</geoCoordinate>
|
||
(0.050); CS =
|
||
<geoCoordinate box="[660,722,1828,1854]" degrees="0.840" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.84">0.840</geoCoordinate>
|
||
<geoCoordinate box="[722,799,1828,1854]" degrees="1.023" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-1.023">–1.023</geoCoordinate>
|
||
(0.964); ES =
|
||
<geoCoordinate box="[967,1029,1828,1854]" degrees="0.182" direction="north" orientation="latitude" pageId="441" pageNumber="442" precision="55" value="0.182">0.182</geoCoordinate>
|
||
<geoCoordinate box="[1029,1106,1828,1854]" degrees="0.230" direction="west" orientation="longitude" pageId="441" pageNumber="442" precision="55" value="-0.23">–0.230</geoCoordinate>
|
||
(0.211); SI = 89–97 (92); OI = 21–23 (22); CI = 80–86 (84); WLI = 138–145 (142); SBI = 23–30 (27); PSI = 27–34 (31); PWI = 167–180 (174); PLI = 174–233 (199); NI = 111–147 (128); PNWI = 94–126 (111); NLI = 40–57 (50); FI = 348–392 (361).
|
||
</paragraph>
|
||
<paragraph blockId="442.[498,1542,234,1934]" pageId="442" pageNumber="443">
|
||
<emphasis bold="true" box="[530,775,314,340]" pageId="442" pageNumber="443">Worker description:</emphasis>
|
||
In full-face view, head subquadrate, longer than broad (CI 80–86). Mandibles densely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin evenly convex medially. Antennal scapes moderately long: when fully retracted, failing to reach the posterior margin of the head capsule by about the width of the scape (SI 89–97). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment about one and a half times as long as the preceding two in combination. Frontal carinae moderately long, extending past the antennal toruli by about twice the maximum width of the antennal scape. Compound eyes weakly protruding past the lateral margins of the head capsule. Lateral margin of head weakly convex, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin flat but rounding evenly into the lateral margins.
|
||
</paragraph>
|
||
<paragraph blockId="442.[498,1542,234,1934]" pageId="442" pageNumber="443">
|
||
In profile view, compound eyes ovular and moderately large (OI 21–23), with 10 ommatidia in longest row. Pronotal declivity indistinct, neck and anterior face of pronotum forming a ~120
|
||
<emphasis box="[809,819,907,925]" italics="true" pageId="442" pageNumber="443">
|
||
<superScript attach="left" box="[809,819,907,925]" fontSize="8" pageId="442" pageNumber="443">°</superScript>
|
||
</emphasis>
|
||
angle. Mesosoma evenly convex from where it joins the pronotal neck to the propodeum. Promesonotal suture extending from the posterior margin of the procoxal insertion to the mesothoracic spiracle, which is moderately well developed, then continuing dorsally as a disruption in the ground sculpture. Metanotal groove visible as a weak sulcus and disruption of the sculpture laterally from where it arises between the mid- and hind coxae to where it ends in the poorly developed metathoracic spiracle, which is nearly indistinguishable against the ground sculpture. Propodeal spiracle moderately well developed, directed posterolaterally, and separated from the propodeal declivity by about three and a half spiracle diameters. Propodeal spines well developed and very long (PSI 27–34), slightly longer than the propodeal declivity, flared at the base, straight, and acute. Propodeal declivity weakly concave, forming a rounded ~120
|
||
<emphasis box="[557,567,1345,1363]" italics="true" pageId="442" pageNumber="443">
|
||
<superScript attach="left" box="[557,567,1345,1363]" fontSize="8" pageId="442" pageNumber="443">°</superScript>
|
||
</emphasis>
|
||
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed. Metapleural gland bulla large, extending from the metacoxal insertion three quarters of the way to the propodeal spiracle. Petiole moderately long (PLI 174–233), without tubercles anterodorsally. Subpetiolar process in the form of a small, acute, spiniform tooth; ventral margin of petiole flat posterior to it. Petiolar peduncle moderately long: comprising a little more than a third of the total petiole length. Petiolar node robust, erect, and subquadrate: transition between peduncle and node marked by a rounded angle of ~110
|
||
<emphasis box="[658,668,1624,1642]" italics="true" pageId="442" pageNumber="443">
|
||
<superScript attach="none" box="[658,668,1624,1642]" fontSize="8" pageId="442" pageNumber="443">°</superScript>
|
||
</emphasis>
|
||
; anterior face forming a ~100
|
||
<emphasis box="[1022,1032,1624,1642]" italics="true" pageId="442" pageNumber="443">
|
||
<superScript attach="left" box="[1022,1032,1624,1642]" fontSize="8" pageId="442" pageNumber="443">°</superScript>
|
||
</emphasis>
|
||
angle with the dorsal face, which is weakly convex; dorsal face meeting the posterior face at a rounded ~90
|
||
<emphasis box="[1220,1230,1664,1682]" italics="true" pageId="442" pageNumber="443">
|
||
<superScript attach="left" box="[1220,1230,1664,1682]" fontSize="8" pageId="442" pageNumber="443">°</superScript>
|
||
</emphasis>
|
||
angle; posterior face forms a ~90
|
||
<emphasis box="[565,575,1704,1722]" italics="true" pageId="442" pageNumber="443">
|
||
<superScript attach="left" box="[565,575,1704,1722]" fontSize="8" pageId="442" pageNumber="443">°</superScript>
|
||
</emphasis>
|
||
angle with the caudal cylinder. Postpetiole evenly rounded anterodorsally; weakly lobed ventrally.
|
||
</paragraph>
|
||
<paragraph blockId="442.[498,1542,234,1934]" lastBlockId="443.[498,1542,234,1935]" lastPageId="443" lastPageNumber="444" pageId="442" pageNumber="443">
|
||
In dorsal view, humeri developed and distinct: evenly rounded and slightly wider than the rest of the mesosoma; mesothoracic spiracles weakly protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Promesonotal suture visible as a disruption in the ground sculpture. Metanotal groove absent: mesonotum and propodeum completely fused and lateral margins converging evenly to the bases of the propodeal spines. Propodeal spines narrowly approximated basally and diverging apically, their apices separated from each other by slightly less than their length, the negative space between them “V” shaped. Petiolar peduncle with spiracles slightly protruding past the lateral margins, and slightly constricted anterior to them. Petiolar node ovular, slightly narrower anteriorly and flattened posteriorly; node the same width as the peduncle, and evenly grading into the caudal cylinder, which is about the same width as the node. Postpetiole narrow (PWI 167–180) and subquadrate. Anterior margin of the postpetiole weakly convex, meeting the lateral margins at a rounded ~90
|
||
<emphasis box="[886,896,588,606]" italics="true" pageId="443" pageNumber="444">
|
||
<superScript attach="left" box="[886,896,588,606]" fontSize="8" pageId="443" pageNumber="444">°</superScript>
|
||
</emphasis>
|
||
angle; lateral margins parallel to each other; lateral margin meeting the posterior margin at a ~90
|
||
<emphasis box="[1042,1052,628,646]" italics="true" pageId="443" pageNumber="444">
|
||
<superScript attach="left" box="[1042,1052,628,646]" fontSize="8" pageId="443" pageNumber="444">°</superScript>
|
||
</emphasis>
|
||
angle; posterior margin flat. Metafemur strongly incrassate (FI 348–392).
|
||
</paragraph>
|
||
<paragraph blockId="443.[498,1542,234,1935]" pageId="443" pageNumber="444">Sculpture: median clypeal carina present, extending posteriorly to the level of the antennal insertions, and flanked on either side by one slightly weaker carina. Lateral clypeal lobes with additional, weaker carinae; ground sculpture smooth and shining. Antennal scapes finely areolate-costulate. Cephalic dorsum weakly sculptured: finely costulate between the frontal carinae; fine concentric costulae surrounding the antennal insertions; weak areolate sculpture on the dorsolateral margins behind the compound eyes and on the posterior quarter of the head, otherwise shining. Lateral surfaces of head weakly areolate, with fine concentric costulae around the compound eyes, and fine rugulose sculpture between the compound eye and mandibular insertion. Ventral surface of head smooth and shining medially, with weak costulae and areolae laterally. Pronotal neck areolate. Lateral face of the pronotum areolate; the remainder of the lateral surface of the mesosoma finely rugulose. Propodeal declivity weakly areolate. Dorsal surface of mesosoma finely areolate, with rugulae overlying the ground sculpture; rugulose sculpture much weaker on the propodeum. Femora shining through weak areolate sculpture. Petiole predominantly weakly areolate, but smooth and shining on the ventral surface of the peduncle. Postpetiole weakly areolate, with sculpture becoming stronger on the posterior quarter and the lateral faces. First gastral tergite and sternite smooth and shining, with weak spectral iridescence.</paragraph>
|
||
<paragraph blockId="443.[498,1542,234,1935]" pageId="443" pageNumber="444">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, stout, blunt-tipped setae, the longest of which are about the width of the compound eye. The head bears ~30, mesosoma ~22, petiole 6, postpetiole ~8, and first gastral tergite ~18 setae. Short, sparse, adpressed pubescence present over the entire body, but difficult to detect against the densely sculptured integument.</paragraph>
|
||
<paragraph blockId="443.[498,1542,234,1935]" pageId="443" pageNumber="444">Color: predominantly testaceous brown, with apex of mandibles, pedicel of antenna, pronotal neck, tibiae, tarsi, and sting testaceous yellow.</paragraph>
|
||
<paragraph blockId="443.[498,1542,234,1935]" lastBlockId="444.[498,1542,234,1934]" lastPageId="444" lastPageNumber="445" pageId="443" pageNumber="444">
|
||
<emphasis bold="true" box="[530,1001,1748,1775]" pageId="443" pageNumber="444">
|
||
Gyne measurements & indices (
|
||
<emphasis bold="true" box="[919,935,1749,1775]" italics="true" pageId="443" pageNumber="444">n</emphasis>
|
||
= 2):
|
||
</emphasis>
|
||
SL =
|
||
<geoCoordinate box="[1073,1135,1748,1775]" degrees="0.625" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.625">0.625</geoCoordinate>
|
||
<geoCoordinate box="[1136,1213,1748,1775]" degrees="0.634" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.634">–0.634</geoCoordinate>
|
||
(0.630); FRS =
|
||
<geoCoordinate box="[1400,1462,1748,1774]" degrees="0.236" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.236">0.236</geoCoordinate>
|
||
<geoCoordinate box="[1463,1539,1749,1775]" degrees="0.249" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.249">–0.249</geoCoordinate>
|
||
(0.243); CW =
|
||
<geoCoordinate box="[678,740,1788,1815]" degrees="0.815" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.815">0.815</geoCoordinate>
|
||
<geoCoordinate box="[740,817,1788,1814]" degrees="0.834" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.834">–0.834</geoCoordinate>
|
||
(0.825); CWb =
|
||
<geoCoordinate box="[1020,1082,1788,1815]" degrees="0.756" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.756">0.756</geoCoordinate>
|
||
<geoCoordinate box="[1082,1159,1788,1815]" degrees="0.757" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.757">–0.757</geoCoordinate>
|
||
(0.757); PoOC =
|
||
<geoCoordinate box="[1372,1434,1788,1814]" degrees="0.328" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.328">0.328</geoCoordinate>
|
||
<geoCoordinate box="[1434,1511,1788,1814]" degrees="0.333" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.333">–0.333</geoCoordinate>
|
||
(0.331); CL =
|
||
<geoCoordinate box="[665,727,1828,1854]" degrees="0.826" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.826">0.826</geoCoordinate>
|
||
<geoCoordinate box="[727,804,1828,1854]" degrees="0.829" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.829">–0.829</geoCoordinate>
|
||
(0.828); EL =
|
||
<geoCoordinate box="[977,1039,1828,1854]" degrees="0.230" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.23">0.230</geoCoordinate>
|
||
<geoCoordinate box="[1039,1116,1828,1854]" degrees="0.247" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.247">–0.247</geoCoordinate>
|
||
(0.239); EW =
|
||
<geoCoordinate box="[1302,1364,1828,1854]" degrees="0.187" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.187">0.187</geoCoordinate>
|
||
<geoCoordinate box="[1364,1441,1828,1854]" degrees="0.204" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.204">–0.204</geoCoordinate>
|
||
(0.196); MD =
|
||
<geoCoordinate box="[580,642,1868,1895]" degrees="0.155" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.155">0.155</geoCoordinate>
|
||
<geoCoordinate box="[642,719,1868,1894]" degrees="0.171" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.171">–0.171</geoCoordinate>
|
||
(0.163); WL =
|
||
<geoCoordinate box="[904,966,1868,1894]" degrees="1.321" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="1.321">1.321</geoCoordinate>
|
||
<geoCoordinate box="[966,1043,1868,1894]" degrees="1.322" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-1.322">–1.322</geoCoordinate>
|
||
(1.322); SPST =
|
||
<geoCoordinate box="[1245,1307,1868,1894]" degrees="0.340" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.34">0.340</geoCoordinate>
|
||
<geoCoordinate box="[1308,1385,1868,1894]" degrees="0.344" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.344">–0.344</geoCoordinate>
|
||
(0.342); MPST =
|
||
<geoCoordinate box="[607,669,1908,1934]" degrees="0.344" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.344">0.344</geoCoordinate>
|
||
<geoCoordinate box="[669,746,1908,1935]" degrees="0.351" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.351">–0.351</geoCoordinate>
|
||
(0.348); PEL =
|
||
<geoCoordinate box="[935,997,1908,1934]" degrees="0.478" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.478">0.478</geoCoordinate>
|
||
<geoCoordinate box="[997,1074,1908,1934]" degrees="0.483" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.483">–0.483</geoCoordinate>
|
||
(0.481); NOL =
|
||
<geoCoordinate box="[1274,1336,1908,1934]" degrees="0.216" direction="north" orientation="latitude" pageId="443" pageNumber="444" precision="55" value="0.216">0.216</geoCoordinate>
|
||
<geoCoordinate box="[1336,1413,1908,1934]" degrees="0.219" direction="west" orientation="longitude" pageId="443" pageNumber="444" precision="55" value="-0.219">–0.219</geoCoordinate>
|
||
(0.218); NOH =
|
||
<geoCoordinate box="[598,660,234,260]" degrees="0.173" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.173">0.173</geoCoordinate>
|
||
<geoCoordinate box="[660,737,234,260]" degrees="0.193" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.193">–0.193</geoCoordinate>
|
||
(0.183); PEH =
|
||
<geoCoordinate box="[933,995,234,260]" degrees="0.317" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.317">0.317</geoCoordinate>
|
||
<geoCoordinate box="[995,1072,234,261]" degrees="0.325" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.325">–0.325</geoCoordinate>
|
||
(0.321); PPL =
|
||
<geoCoordinate box="[1262,1324,234,260]" degrees="0.229" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.229">0.229</geoCoordinate>
|
||
<geoCoordinate box="[1324,1401,234,261]" degrees="0.235" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.235">–0.235</geoCoordinate>
|
||
(0.232); PW =
|
||
<geoCoordinate box="[577,639,274,300]" degrees="0.832" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.832">0.832</geoCoordinate>
|
||
<geoCoordinate box="[640,716,274,300]" degrees="0.833" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.833">–0.833</geoCoordinate>
|
||
(0.833); SBPA =
|
||
<geoCoordinate box="[925,987,274,300]" degrees="0.361" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.361">0.361</geoCoordinate>
|
||
<geoCoordinate box="[987,1064,274,300]" degrees="0.370" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.37">–0.370</geoCoordinate>
|
||
(0.366); SPTI =
|
||
<geoCoordinate box="[1263,1325,274,300]" degrees="0.366" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.366">0.366</geoCoordinate>
|
||
<geoCoordinate box="[1325,1402,274,300]" degrees="0.376" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.376">–0.376</geoCoordinate>
|
||
(0.371); PEW =
|
||
<geoCoordinate box="[589,650,314,340]" degrees="0.214" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.214">0.214</geoCoordinate>
|
||
<geoCoordinate box="[650,725,314,340]" degrees="0.224" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.224">–0.224</geoCoordinate>
|
||
(0.219); PNW = 0.216; PPW =
|
||
<geoCoordinate box="[1086,1147,314,340]" degrees="0.392" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.392">0.392</geoCoordinate>
|
||
<geoCoordinate box="[1147,1222,314,340]" degrees="0.408" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.408">–0.408</geoCoordinate>
|
||
(0.400); HFL =
|
||
<geoCoordinate box="[1405,1466,314,340]" degrees="0.774" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.774">0.774</geoCoordinate>
|
||
<geoCoordinate box="[1466,1542,314,340]" degrees="0.798" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.798">–0.798</geoCoordinate>
|
||
(0.786); HFWmax =
|
||
<geoCoordinate box="[734,795,354,380]" degrees="0.171" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.171">0.171</geoCoordinate>
|
||
<geoCoordinate box="[795,870,354,380]" degrees="0.193" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.193">–0.193</geoCoordinate>
|
||
(0.182); HFWmin =
|
||
<geoCoordinate box="[1109,1170,354,381]" degrees="0.058" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.058">0.058</geoCoordinate>
|
||
<geoCoordinate box="[1170,1245,354,380]" degrees="0.076" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.076">–0.076</geoCoordinate>
|
||
(0.067); CS =
|
||
<geoCoordinate box="[1405,1466,354,380]" degrees="1.170" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="1.17">1.170</geoCoordinate>
|
||
<geoCoordinate box="[1466,1542,354,380]" degrees="1.171" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-1.171">–1.171</geoCoordinate>
|
||
(1.170); ES =
|
||
<geoCoordinate box="[657,718,394,420]" degrees="0.324" direction="north" orientation="latitude" pageId="444" pageNumber="445" precision="55" value="0.324">0.324</geoCoordinate>
|
||
<geoCoordinate box="[718,793,394,420]" degrees="0.349" direction="west" orientation="longitude" pageId="444" pageNumber="445" precision="55" value="-0.349">–0.349</geoCoordinate>
|
||
(0.336); SI = 83–84 (83); OI = 28–30 (29); CI = 91–92 (91); WLI = 175; SBI = 48–49 (48); PSI = 26; PWI = 182–183 (183); PLI = 203–211 (207); NI = 113–125 (119); PNWI = 96–101 (99); NLI = 45–45 (45); FI = 254–295 (274).
|
||
</paragraph>
|
||
<paragraph blockId="444.[498,1542,234,1934]" pageId="444" pageNumber="445">
|
||
<emphasis bold="true" box="[530,747,513,539]" pageId="444" pageNumber="445">Gyne description:</emphasis>
|
||
In full-face view, head subquadrate, slightly longer than broad (CI 91–92). Mandibles densely striate but shining and armed with five teeth: the apical-most well developed, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin weakly emarginated medially. Antennal scapes moderately long: when fully retracted, just reaching the posterior margin of the head capsule (SI 83–84). Antennae 12-segmented; antennal club composed of three segments, with the apical-most segment about one and a half times as long as the preceding two in combination. Frontal carinae moderately long, extending past the antennal toruli by about two and a half times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margins of head convex behind the compound eyes, but parallel to each other between the compound eyes and the mandibular insertions. Posterior head margin flat, rounding evenly into the lateral margins.
|
||
</paragraph>
|
||
<paragraph blockId="444.[498,1542,234,1934]" pageId="444" pageNumber="445">
|
||
In profile view, compound eyes ovular and large (OI 28–30), with 16 ommatidia in longest row. Mesoscutum rounded evenly anteriorly, covering the dorsal surface of the pronotum, and weakly convex dorsally, nearly flat. Mesoscutellum very slightly depressed below the level of the mesoscutum and gently sloping posteriorly. Posterior margin of metanotum extending slightly past the posterior margin of the mesoscutum. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines stout and well developed, but short (PSI 26), about as two thirds as long as the propodeal declivity, tapering evenly from the base, directed posterodorsally, straight, and acute. Propodeal declivity straight and flat, forming a rounded ~120
|
||
<emphasis box="[832,842,1385,1403]" italics="true" pageId="444" pageNumber="445">
|
||
<superScript attach="left" box="[832,842,1385,1403]" fontSize="8" pageId="444" pageNumber="445">°</superScript>
|
||
</emphasis>
|
||
angle with the base of the propodeal spines. Propodeal lobes rounded and very weakly developed. Metapleural gland bulla large, extending from the metacoxal insertion three quarters of the way to the propodeal spiracle. Petiole long (PLI 203–211), without tubercles anterodorsally. Subpetiolar process in the form of a small, triangular tooth, which grades evenly into the ventral margin of the petiole posteriorly. Petiolar peduncle moderately long: comprising a little more than a third of the total petiole length. Petiolar node robust and erect: transition between peduncle and node an evenly rounded ~130
|
||
<emphasis box="[785,795,1664,1682]" italics="true" pageId="444" pageNumber="445">
|
||
<superScript attach="left" box="[785,795,1664,1682]" fontSize="8" pageId="444" pageNumber="445">°</superScript>
|
||
</emphasis>
|
||
angle, resulting in a very slightly concave anterior node face; anterior face forming a rounded ~100
|
||
<emphasis box="[937,947,1704,1722]" italics="true" pageId="444" pageNumber="445">
|
||
<superScript attach="left" box="[937,947,1704,1722]" fontSize="8" pageId="444" pageNumber="445">°</superScript>
|
||
</emphasis>
|
||
angle with the dorsal face, which is flat; dorsal face meeting the posterior face at a ~100
|
||
<emphasis box="[925,935,1744,1762]" italics="true" pageId="444" pageNumber="445">
|
||
<superScript attach="left" box="[925,935,1744,1762]" fontSize="8" pageId="444" pageNumber="445">°</superScript>
|
||
</emphasis>
|
||
angle; posterior face forms a ~90
|
||
<emphasis box="[1333,1343,1744,1762]" italics="true" pageId="444" pageNumber="445">
|
||
<superScript attach="left" box="[1333,1343,1744,1762]" fontSize="8" pageId="444" pageNumber="445">°</superScript>
|
||
</emphasis>
|
||
angle with the caudal cylinder. Postpetiole evenly rounded anterodorsally, bulging slightly before it transitions into the flattened posterodorsal face; ventral surface weakly lobed.
|
||
</paragraph>
|
||
<paragraph blockId="444.[498,1542,234,1934]" lastBlockId="445.[498,1542,234,1895]" lastPageId="445" lastPageNumber="446" pageId="444" pageNumber="445">In dorsal view, mesoscutum covering pronotum anteriorly, but humeri visible laterally as rounded sclerites. Propodeal spines weakly diverging apically, their apices separated from each other by about one and a half times their length. Petiolar peduncle with spiracles not protruding past the lateral margins, but slightly narrowed anterior to them. Petiolar node subquadrate, slightly broadened transversely. Petiolar node slightly narrower than the peduncle, and evenly grading into the caudal cylinder, which is about the same width as the node. Postpetiole narrow (PWI 182–183), anteroposteriorly compressed, and subquadrate. Anterior margin of postpetiole flat, with corners marked by rounded angles as it transitions to the lateral margins; lateral margins parallel to each other; posterior corners angulate; posterior margin broadly flat. Metafemur moderately incrassate (FI 254–295).</paragraph>
|
||
<paragraph blockId="445.[498,1542,234,1895]" pageId="445" pageNumber="446">Sculpture: median clypeal carina present, extending from the anterior margin nearly to frontal triangle, and flanked by weaker, indistinct carinae. Lateral clypeal lobes with additional weaker carinae; ground sculpture smooth and shining. Antennal scapes finely areolate-costulate. Cephalic dorsum areolate, with rugose sculpture that becomes costate between the frontal carinae; concentric costulae surround the antennal insertions. Lateral surfaces of head areolate, with rugose sculpture overlying the ground sculpture. Ventral surface of head smooth and shining medially, with weak areolate-rugulose sculpture laterally and posteriorly. Pronotal neck areolate. Anterior face of pronotum shining and weakly, indistinctly sculptured. Lateral face of pronotum costate. Anepisternum and katepisternum shining on their anterior halves, transitioning into weak areolate-costulate sculpture posteriorly. Metapleuron areolate, with costulae. Propodeum weakly areolate laterally. Propodeal declivity weakly areolate. Mesoscutum with costulae over areolate ground sculpture; a patch of finely areolate sculpture present anteromedially. Mesoscutellum longitudinally areolate-costulate. Metanotum areolate. Propodeum areolate, with weak rugae. Femora shining, with traces of weak areolate sculpture. Petiole uniformly finely areolate, with parallel rugae that are dorsoventrally oriented around the petiolar node; dorsum of node rugose. Postpetiole shining through weak areolate sculpture anteriorly, areolate laterally and dorsally; fine concentric costulae dorsally. First gastral tergite and sternite smooth and shining, with weak spectral iridescence.</paragraph>
|
||
<paragraph blockId="445.[498,1542,234,1895]" pageId="445" pageNumber="446">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, stout, blunt-tipped setae, the longest of which are about half the width of the compound eye. Short, sparse, adpressed pubescence present over the entire body, but difficult to detect against the densely sculptured integument.</paragraph>
|
||
<paragraph blockId="445.[498,1542,234,1895]" pageId="445" pageNumber="446">Color; predominantly testaceous brown, with apex of mandibles, pedicel of antenna, pronotal neck, tibiae, tarsi, and sting testaceous yellow.</paragraph>
|
||
<paragraph blockId="445.[498,1542,234,1895]" box="[530,730,1669,1695]" pageId="445" pageNumber="446">
|
||
<emphasis bold="true" box="[530,597,1669,1695]" pageId="445" pageNumber="446">Male:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="445" pageNumber="446" type="etymology">
|
||
<paragraph blockId="445.[498,1542,234,1895]" pageId="445" pageNumber="446">
|
||
<emphasis bold="true" box="[530,668,1709,1735]" pageId="445" pageNumber="446">Etymology:</emphasis>
|
||
Morphological, from the Latin ‘tenuis’ (= weak) + ‘sculptus’ (= sculptured), probably in reference to the smooth and shining integument of the head.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="446" lastPageNumber="447" pageId="445" pageNumber="446" type="discussion">
|
||
<paragraph blockId="445.[498,1542,234,1895]" lastBlockId="446.[498,1542,1144,1290]" lastPageId="446" lastPageNumber="447" pageId="445" pageNumber="446">
|
||
<emphasis bold="true" box="[530,670,1788,1814]" pageId="445" pageNumber="446">Comments:</emphasis>
|
||
This species is only known from a few collections, including lowland rainforest in
|
||
<collectingRegion box="[651,759,1829,1855]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="445" pageNumber="446">Veracruz</collectingRegion>
|
||
,
|
||
<collectingCountry box="[773,861,1828,1854]" name="Mexico" pageId="445" pageNumber="446">Mexico</collectingCountry>
|
||
; a nest from between the leaves of
|
||
<taxonomicName box="[1276,1392,1828,1854]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="445" pageNumber="446" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[1276,1392,1828,1854]" italics="true" pageId="445" pageNumber="446">Tillandsia</emphasis>
|
||
</taxonomicName>
|
||
in mid-elevation mesic forest in
|
||
<collectingRegion box="[847,955,1869,1895]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="445" pageNumber="446">Veracruz</collectingRegion>
|
||
, and a nest in an oak branch bearing
|
||
<taxonomicName box="[1398,1541,1868,1894]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="445" pageNumber="446" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[1398,1541,1868,1894]" italics="true" pageId="445" pageNumber="446">Epidendrum</emphasis>
|
||
</taxonomicName>
|
||
orchids, which was intercepted in a shipment from
|
||
<collectingCountry box="[1101,1187,1144,1170]" name="Mexico" pageId="446" pageNumber="447">Mexico</collectingCountry>
|
||
at the
|
||
<collectingCountry box="[1269,1317,1144,1170]" name="United States of America" pageId="446" pageNumber="447">U.S.</collectingCountry>
|
||
border by customs officials. This is an arboreally nesting species, similar to the species of the
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1358,1454,1185,1210]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="446" pageNumber="447" phylum="Arthropoda" rank="species" species="annexus">
|
||
<emphasis box="[1358,1454,1185,1210]" italics="true" pageId="446" pageNumber="447">annexus</emphasis>
|
||
</taxonomicName>
|
||
species group, with which it is closely related. Very little morphological variation can be seen between the three collections of this species that I have observed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |