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<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FE79FE1EB49C1CA77AB72465" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax terrigena" docType="treatment" docVersion="4" lastPageNumber="453" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="447" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prebus, Matthew M.</mods:namePart>
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States &amp; Department of Entomology &amp; Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PeerJ</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="series">
<mods:title>e 11514</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2021-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>9</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>462</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
<mods:identifier type="ISSN">2167-8359</mods:identifier>
<mods:identifier type="PMC">PMC8254503</mods:identifier>
<mods:identifier type="PubMed">34249486</mods:identifier>
<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102672" ID-GBIF-Taxon="183542636" ID-Zenodo-Dep="5102672" LSID="urn:lsid:plazi:treatment:054FDB70FE79FE1EB49C1CA77AB72465" httpUri="http://treatment.plazi.org/id/054FDB70FE79FE1EB49C1CA77AB72465" lastPageId="452" lastPageNumber="453" pageId="446" pageNumber="447">
<subSubSection box="[498,993,1722,1749]" pageId="446" pageNumber="447" type="nomenclature">
<paragraph blockId="446.[498,993,1722,1789]" box="[498,993,1722,1749]" pageId="446" pageNumber="447">
<heading box="[498,993,1722,1749]" level="2" pageId="446" pageNumber="447">
<taxonomicName authority="(Wheeler, 1903 b)" baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[498,993,1722,1749]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="446" pageNumber="447" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis bold="true" box="[498,993,1722,1749]" pageId="446" pageNumber="447">
<emphasis bold="true" box="[498,777,1722,1749]" italics="true" pageId="446" pageNumber="447">Temnothorax terrigena</emphasis>
(
<bibRefCitation author="Wheeler WM" box="[797,984,1722,1749]" journalOrPublisher="Proceedings of the Academy of Natural Sciences of: Philadelphia" pageId="446" pageNumber="447" pagination="215 - 260" refId="ref273272" refString="Wheeler WM. 1903 b. A revision of the North American ants of the genus Leptothorax Mayr. In: Proceedings of the Academy of Natural Sciences of: Philadelphia. 215 - 260." title="A revision of the North American ants of the genus Leptothorax Mayr" type="book chapter" year="1903">
<emphasis bold="true" box="[797,984,1722,1749]" italics="true" pageId="446" pageNumber="447">Wheeler, 1903b</emphasis>
</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[498,946,1762,1789]" pageId="446" pageNumber="447" type="distribution">
<paragraph blockId="446.[498,993,1722,1789]" box="[498,946,1762,1789]" pageId="446" pageNumber="447">
Distribution, worker &amp; gyne:
<figureCitation box="[847,940,1762,1788]" captionStart="Figure 170" captionStartId="446.[116,181,979,1001]" captionTargetBox="[90,1541,234,966]" captionTargetId="figure-204@446.[90,1524,234,951]" captionTargetPageId="446" captionText="Figure 170 Temnothorax terrigena. (AC) Lectotype worker (MCZENT00522078). (A) Full-face view. (B) Profile view. (C) Dorsal view. (DF) Paralectotype gyne (MCZENT00522078). (D) Full-face view. (E) Profile view. (F) Dorsal view. (G) Geographical and elevational distribution of specimens examined. Type locality is represented by a triangle, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-170" figureDoi="http://doi.org/10.5281/zenodo.5102501" httpUri="https://zenodo.org/record/5102501/files/figure.png" pageId="446" pageNumber="447">Fig. 170</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="447" lastPageNumber="448" pageId="446" pageNumber="447" type="reference_group">
<paragraph blockId="446.[498,1518,1824,1891]" pageId="446" pageNumber="447">
<treatmentCitationGroup pageId="446" pageNumber="447">
<treatmentCitation author="Wheeler WM" box="[498,998,1824,1851]" page="254" pageId="446" pageNumber="447" year="1903">
<taxonomicName authority="Wheeler, 1903 b: 254" authorityName="Wheeler" authorityPageNumber="254" authorityYear="1903" box="[498,937,1824,1850]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="446" pageNumber="447" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[498,937,1824,1850]" italics="true" pageId="446" pageNumber="447">
Leptothorax terrigena
<bibRefCitation author="Wheeler WM" box="[758,998,1824,1851]" journalOrPublisher="Proceedings of the Academy of Natural Sciences of: Philadelphia" pageId="446" pageNumber="447" pagination="215 - 260" refId="ref273272" refString="Wheeler WM. 1903 b. A revision of the North American ants of the genus Leptothorax Mayr. In: Proceedings of the Academy of Natural Sciences of: Philadelphia. 215 - 260." title="A revision of the North American ants of the genus Leptothorax Mayr" type="book chapter" year="1903">Wheeler, 1903b: 254</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[1010,1102,1825,1851]" pageId="446" pageNumber="447">Syntype</typeStatus>
workers and dealate gyne: McNeil,
<collectingRegion box="[498,567,1865,1891]" country="United States of America" name="Texas" pageId="446" pageNumber="447">Texas</collectingRegion>
,
<collectingCountry box="[581,655,1864,1890]" name="United States of America" pageId="446" pageNumber="447">U.S.A.</collectingCountry>
One
<typeStatus box="[722,811,1865,1891]" pageId="446" pageNumber="447">syntype</typeStatus>
worker here designated
<emphasis bold="true" box="[1103,1212,1864,1890]" pageId="446" pageNumber="447">
<typeStatus box="[1103,1212,1864,1890]" pageId="446" pageNumber="447">lectotype</typeStatus>
</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="447.[498,1541,234,341]" pageId="447" pageNumber="448">
<treatmentCitationGroup pageId="447" pageNumber="448">
<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" box="[498,1017,234,261]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="terrigena" subGenus="Myrafant">
<emphasis box="[498,887,234,260]" italics="true" pageId="447" pageNumber="448">Leptothorax (Myrafant) terrigena</emphasis>
(Wheeler)
</taxonomicName>
:
<treatmentCitation author="Smith DR" box="[1030,1242,234,261]" page="1395" pageId="447" pageNumber="448" year="1979">
<bibRefCitation author="Smith DR" box="[1030,1242,234,261]" editor="Krombein KV &amp; Hurd PD &amp; Smith DR &amp; Burks BD" journalOrPublisher="Washington, D. C.: Smithsonian Institution Press" pageId="447" pageNumber="448" pagination="1323 - 1467" refId="ref272728" refString="Smith DR. 1979. Superfamily Formicoidea. In: Krombein KV, Hurd PD, Smith DR, Burks BD, eds. Catalog of Hymenoptera in America north of Mexico: Apocrita (Aculeata). Vol. 2. Washington, D. C.: Smithsonian Institution Press, 1323 - 1467." title="Superfamily Formicoidea" type="book chapter" volumeTitle="Catalog of Hymenoptera in America north of Mexico: Apocrita (Aculeata). Vol. 2" year="1979">
<emphasis box="[1030,1168,234,260]" italics="true" pageId="447" pageNumber="448">
<collectingRegion box="[1030,1100,234,260]" country="Bermuda" name="Smith's" pageId="447" pageNumber="448">
<author box="[1030,1100,234,260]" fns="*" inm="true" npo="*" pageId="447" pageNumber="448">Smith</author>
</collectingRegion>
,
<year box="[1113,1168,234,260]" pageId="447" pageNumber="448">1979</year>
</emphasis>
: 1395
</bibRefCitation>
</treatmentCitation>
. First combination in
<taxonomicName authority="" box="[498,776,274,300]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="subGenus" subGenus="Myrafant">
<emphasis box="[498,776,274,300]" italics="true" pageId="447" pageNumber="448">Leptothorax (Myrafant)</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="447.[498,1541,234,341]" box="[498,1541,314,341]" pageId="447" pageNumber="448">
<treatmentCitationGroup box="[498,1541,314,341]" pageId="447" pageNumber="448">
<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[498,897,314,341]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[498,767,314,340]" italics="true" pageId="447" pageNumber="448">Temnothorax terrigena</emphasis>
(Wheeler)
</taxonomicName>
:
<treatmentCitation author="Bolton B." box="[909,1115,314,340]" page="271" pageId="447" pageNumber="448" year="2003">
<bibRefCitation author="Bolton B." box="[909,1115,314,340]" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="447" pageNumber="448" pagination="370" part="71" refId="ref270076" refString="Bolton B. 2003. Synopsis and Classification of Formicidae. Memoirs of the American Entomological Institute 71: 370." title="Synopsis and Classification of Formicidae" type="journal article" year="2003">
<emphasis box="[909,1055,314,340]" italics="true" pageId="447" pageNumber="448">
<collectingRegion box="[909,987,314,340]" country="United Kingdom" name="Bolton" pageId="447" pageNumber="448">
<author box="[909,987,314,340]" fns="*" inm="true" npo="*" pageId="447" pageNumber="448">Bolton</author>
</collectingRegion>
,
<year box="[1000,1055,314,340]" pageId="447" pageNumber="448">2003</year>
</emphasis>
: 271
</bibRefCitation>
</treatmentCitation>
. First combination in
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[1378,1535,314,340]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis box="[1378,1535,314,340]" italics="true" pageId="447" pageNumber="448">Temnothorax</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="447" pageNumber="448" type="materials_examined">
<paragraph blockId="447.[498,1542,376,1917]" pageId="447" pageNumber="448">
<emphasis bold="true" box="[530,834,376,403]" pageId="447" pageNumber="448">Type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861793" collectingDate="1902-02-22" collectionCode="MCZC" collectorName="Arcila AM &amp; M. C. Z. Type" country="United States of America" county="Travis County" location="McNeil" pageId="447" pageNumber="448" specimenCount="1" stateProvince="Texas" typeStatus="lectotype">
<emphasis box="[843,1329,376,403]" italics="true" pageId="447" pageNumber="448">
<typeStatus box="[843,951,377,402]" pageId="447" pageNumber="448">Lectotype</typeStatus>
worker of
<taxonomicName box="[1078,1329,376,402]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="terrigena">Leptothorax terrigena</taxonomicName>
</emphasis>
:
<collectingCountry box="[1344,1421,377,403]" name="United States of America" pageId="447" pageNumber="448">U.S.A.</collectingCountry>
:
<collectingRegion box="[1434,1504,377,403]" country="United States of America" name="Texas" pageId="447" pageNumber="448">Texas</collectingRegion>
:
<collectingCounty box="[498,669,416,442]" pageId="447" pageNumber="448">Travis County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:054FDB70FE79FE1EB49C1CA77AB72465:88393CBDFE78FE65B7C41BBD7A3126ED" box="[682,771,416,442]" country="United States of America" county="Travis County" name="McNeil" pageId="447" pageNumber="448" stateProvince="Texas">McNeil</location>
,
<date box="[784,992,416,442]" pageId="447" pageNumber="448" value="1902-02-22">
<collectingDate box="[784,992,416,442]" pageId="447" pageNumber="448" value="1902-02-22">22 February 1902</collectingDate>
</date>
, co-type
<collectorName box="[1101,1162,417,443]" name="Arcila AM" pageId="447" pageNumber="448">A.M.</collectorName>
N.H., bottom specimen on pin (
<collectorName box="[507,658,456,483]" pageId="447" pageNumber="448">M.C.Z. Type</collectorName>
21038, MCZENT00522078) [
<collectionCode box="[1007,1098,455,482]" pageId="447" pageNumber="448">MCZC</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph blockId="447.[498,1542,376,1917]" pageId="447" pageNumber="448">
<materialsCitation ID-GBIF-Occurrence="3321861548" collectingDate="1902-02-22" collectionCode="MCZC, LACM, USNM" collectorName="M. C. Z. Type" country="United States of America" county="Travis County" location="McNeil" pageId="447" pageNumber="448" specimenCount="14" specimenCount-worker="14" stateProvince="Texas" typeStatus="paralectotype">
<emphasis box="[530,1171,496,522]" italics="true" pageId="447" pageNumber="448">
<typeStatus box="[530,684,496,522]" pageId="447" pageNumber="448">Paralectotype</typeStatus>
workers and gyne of
<taxonomicName box="[923,1171,496,522]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="terrigena">Leptothorax terrigena</taxonomicName>
</emphasis>
: same pin as
<typeStatus box="[1323,1430,496,522]" pageId="447" pageNumber="448">lectotype</typeStatus>
, 1 dealate gyne,
<specimenCount box="[568,673,536,562]" pageId="447" pageNumber="448" type="worker">1 worker</specimenCount>
(top and middle specimens on pin) [
<collectionCode box="[1114,1204,535,562]" pageId="447" pageNumber="448">MCZC</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861889" collectingDate="1902-02-22" collectionCode="MCZC, LACM, USNM" collectorName="M. C. Z. Type" country="United States of America" county="Travis County" location="McNeil" pageId="447" pageNumber="448" specimenCount="14" specimenCount-worker="14" stateProvince="Texas" typeStatus="paralectotype">
same data as lectotype,
<specimenCount box="[498,614,576,602]" pageId="447" pageNumber="448" type="worker">4 workers</specimenCount>
(
<collectorName box="[631,782,576,602]" pageId="447" pageNumber="448">M.C.Z. Type</collectorName>
21038, MCZENT00031038) [
<collectionCode box="[1131,1219,575,602]" pageId="447" pageNumber="448">MCZC</collectionCode>
]
<specimenCount box="[1235,1350,576,602]" pageId="447" pageNumber="448" type="worker">3 workers</specimenCount>
(LACMENT181910) [
<collectionCode box="[754,842,615,642]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="447" pageNumber="448" type="Museum">LACM</collectionCode>
]
<specimenCount box="[857,973,616,642]" pageId="447" pageNumber="448" type="worker">3 workers</specimenCount>
(
<collectorName box="[990,1141,616,642]" pageId="447" pageNumber="448">M.C.Z. Type</collectorName>
21038, USNMENT00528965) [
<collectionCode box="[505,594,655,682]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="447" pageNumber="448" type="Museum">USNM</collectionCode>
]
<specimenCount box="[610,725,655,681]" pageId="447" pageNumber="448" type="worker">3 workers</specimenCount>
(CASENT0105624) [
<collectionCode box="[976,1068,655,682]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="447" pageNumber="448" type="Museum">USNM</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="447.[498,1542,376,1917]" pageId="447" pageNumber="448">
<emphasis bold="true" box="[530,888,695,721]" pageId="447" pageNumber="448">Non-type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861873" collectingDate="1947-11-07" collectionCode="USNM" country="Mexico" location="Cycad" municipality="Victoria" pageId="447" pageNumber="448" specimenCount="7" specimenCount-worker="7" stateProvince="Tamaulipas">
<collectingCountry box="[897,1012,695,721]" name="Mexico" pageId="447" pageNumber="448">MEXICO</collectingCountry>
:
<collectingRegion box="[1023,1162,695,721]" country="Mexico" name="Tamaulipas" pageId="447" pageNumber="448">Tamaulipas</collectingRegion>
:
<collectingMunicipality box="[1175,1272,695,721]" pageId="447" pageNumber="448">Victoria</collectingMunicipality>
, [intercepted in quarantine],
<date box="[647,854,735,762]" pageId="447" pageNumber="448" value="1947-11-07">
<collectingDate box="[647,854,735,762]" pageId="447" pageNumber="448" value="1947-11-07">7 November 1947</collectingDate>
</date>
, [no collector data], Lar-45405, 47-16324, on
<location LSID="urn:lsid:plazi:treatment:054FDB70FE79FE1EB49C1CA77AB72465:88393CBDFE78FE65B00F18C27C9825AE" box="[1377,1450,735,761]" country="Mexico" municipality="Victoria" name="Cycad" pageId="447" pageNumber="448" stateProvince="Tamaulipas">Cycad</location>
plant, 1 dealate gyne &amp;
<specimenCount box="[681,797,775,801]" pageId="447" pageNumber="448" type="worker">7 workers</specimenCount>
(CASENT0758302-CASENT0758305) [
<collectionCode box="[1265,1357,774,801]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="447" pageNumber="448" type="Museum">USNM</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="447.[498,1542,376,1917]" pageId="447" pageNumber="448">
<materialsCitation ID-GBIF-Occurrence="3321861395" collectingDate="1984-12-15" collectionCode="UCDC, USNM" collectorName="P. S. Ward " country="United States of America" county="Cameron County" elevation="5" location="Travis County" municipality="Laguna Atacosa National Wildlife Refuge" pageId="447" pageNumber="448" specimenCount="3" specimenCount-worker="3" stateProvince="Texas">
<collectingCountry box="[530,607,815,841]" name="United States of America" pageId="447" pageNumber="448">U.S.A.</collectingCountry>
:
<collectingRegion box="[620,690,815,841]" country="United States of America" name="Texas" pageId="447" pageNumber="448">Texas</collectingRegion>
:
<collectingCounty box="[702,910,815,841]" pageId="447" pageNumber="448">Cameron County</collectingCounty>
:
<collectingMunicipality box="[922,1411,815,841]" pageId="447" pageNumber="448">Laguna Atacosa National Wildlife Refuge</collectingMunicipality>
,
<quantity box="[1424,1471,815,842]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="447" pageNumber="448" unit="m" value="5.0">
<elevation box="[1424,1471,815,842]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="447" pageNumber="448" unit="m" value="5.0">5 m</elevation>
</quantity>
,
<date box="[498,719,854,881]" pageId="447" pageNumber="448" value="1984-12-15">
<collectingDate box="[498,719,854,881]" pageId="447" pageNumber="448" value="1984-12-15">15 December 1984</collectingDate>
</date>
,
<collectorName box="[732,851,855,881]" pageId="447" pageNumber="448">P.S. Ward</collectorName>
#7197-3, coastal mesquite scrub, ground foragers,
<specimenCount box="[1437,1542,855,881]" pageId="447" pageNumber="448" type="worker">1 worker</specimenCount>
(CASENT0104775) [
<collectionCode box="[740,829,894,921]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="447" pageNumber="448" type="Museum">UCDC</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861661" collectingDate="1984-12-15" collectionCode="UCDC, USNM" collectorName="W. M. Wheeler" country="United States of America" county="Cameron County" elevation="5" location="Travis County" municipality="Laguna Atacosa National Wildlife Refuge" pageId="447" pageNumber="448" specimenCount="3" specimenCount-worker="3" stateProvince="Texas">
<location LSID="urn:lsid:plazi:treatment:054FDB70FE79FE1EB49C1CA77AB72465:88393CBDFE78FE65B63E19637AC824CE" box="[848,1018,894,921]" country="United States of America" county="Cameron County" municipality="Laguna Atacosa National Wildlife Refuge" name="Travis County" pageId="447" pageNumber="448" stateProvince="Texas">Travis County</location>
:
<collectorName box="[1029,1109,894,920]" pageId="447" pageNumber="448">Austin</collectorName>
, [no collection date],
<collectorName box="[1363,1537,894,921]" pageId="447" pageNumber="448">W.M. Wheeler</collectorName>
,
<specimenCount box="[498,614,934,960]" pageId="447" pageNumber="448" type="worker">2 workers</specimenCount>
(CASENT0758301) [
<collectionCode box="[865,957,934,961]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="447" pageNumber="448" type="Museum">USNM</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[530,1536,974,1001]" pageId="447" pageNumber="448" type="distribution">
<paragraph blockId="447.[498,1542,376,1917]" box="[530,1536,974,1001]" pageId="447" pageNumber="448">
<emphasis bold="true" box="[530,755,974,1000]" pageId="447" pageNumber="448">Geographic range:</emphasis>
Low elevations,
<collectingRegion box="[952,1021,975,1001]" country="United States of America" name="Texas" pageId="447" pageNumber="448">Texas</collectingRegion>
,
<collectingCountry box="[1035,1109,974,1000]" name="United States of America" pageId="447" pageNumber="448">U.S.A.</collectingCountry>
to
<collectingRegion box="[1150,1288,974,1000]" country="Mexico" name="Tamaulipas" pageId="447" pageNumber="448">Tamaulipas</collectingRegion>
,
<collectingCountry box="[1301,1387,974,1000]" name="Mexico" pageId="447" pageNumber="448">Mexico</collectingCountry>
(
<figureCitation box="[1406,1520,974,1000]" captionStart="Figure 170" captionStartId="446.[116,181,979,1001]" captionTargetBox="[90,1541,234,966]" captionTargetId="figure-204@446.[90,1524,234,951]" captionTargetPageId="446" captionText="Figure 170 Temnothorax terrigena. (AC) Lectotype worker (MCZENT00522078). (A) Full-face view. (B) Profile view. (C) Dorsal view. (DF) Paralectotype gyne (MCZENT00522078). (D) Full-face view. (E) Profile view. (F) Dorsal view. (G) Geographical and elevational distribution of specimens examined. Type locality is represented by a triangle, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-170" figureDoi="http://doi.org/10.5281/zenodo.5102501" httpUri="https://zenodo.org/record/5102501/files/figure.png" pageId="447" pageNumber="448">Fig. 170G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="448" lastPageNumber="449" pageId="447" pageNumber="448" type="diagnosis">
<paragraph blockId="447.[498,1542,376,1917]" pageId="447" pageNumber="448">
<emphasis bold="true" box="[530,754,1014,1040]" pageId="447" pageNumber="448">Worker diagnosis:</emphasis>
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[763,1032,1014,1040]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[763,1032,1014,1040]" italics="true" pageId="447" pageNumber="448">Temnothorax terrigena</emphasis>
</taxonomicName>
can be separated from all other species in the
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[544,619,1054,1080]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[544,619,1054,1080]" italics="true" pageId="447" pageNumber="448">salvini</emphasis>
</taxonomicName>
clade by the following character combination: in profile view, dorsum of mesosoma weakly convex; metanotal groove not impressed; propodeum not depressed below the level of the promesonotum; propodeal spines shorter than the propodeal declivity and directed posterodorsally; hind femora weakly to moderately incrassate; petiolar node subquadrate, not leaning posteriorly, and rounded dorsally; petiolar node as long as, or only slightly longer than high; posterior face of petiolar node about the same height as the anterior face; postpetiole moderately broad, greater than 1.5 times the width of the caudal cylinder of the petiole, but less than 2.2 times; setae on head, mesosoma, waist segments and gaster erect, short, sparse and blunt (never long and tapering); integument predominantly yellow.
</paragraph>
<paragraph blockId="447.[498,1542,376,1917]" lastBlockId="448.[498,1542,234,1934]" lastPageId="448" lastPageNumber="449" pageId="447" pageNumber="448">
<emphasis bold="true" box="[530,719,1452,1478]" pageId="447" pageNumber="448">Similar species:</emphasis>
<emphasis box="[727,1411,1452,1478]" italics="true" pageId="447" pageNumber="448">
<taxonomicName box="[727,970,1452,1478]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="andrei">Temnothorax andrei</taxonomicName>
,
<taxonomicName box="[983,1125,1453,1478]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="carinatus">T. carinatus</taxonomicName>
,
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1139,1260,1453,1478]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="goniops">T. goniops</taxonomicName>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1274,1411,1452,1478]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="pilicornis" status="sp. nov.">T. pilicornis</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[1420,1513,1453,1479]" pageId="447" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis box="[498,782,1492,1518]" italics="true" pageId="447" pageNumber="448">
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[498,620,1493,1518]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="rugosus">T. rugosus</taxonomicName>
,
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[634,782,1492,1518]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="subditivus">T. subditivus</taxonomicName>
</emphasis>
(light form),
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[946,1041,1493,1518]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[946,1041,1493,1518]" italics="true" pageId="447" pageNumber="448">T. torrei</emphasis>
</taxonomicName>
.
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1057,1325,1492,1518]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[1057,1325,1492,1518]" italics="true" pageId="447" pageNumber="448">Temnothorax terrigena</emphasis>
</taxonomicName>
differs from
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,636,1532,1558]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="pilicornis" status="sp. nov.">
<emphasis box="[498,636,1532,1558]" italics="true" pageId="447" pageNumber="448">T. pilicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[645,737,1533,1559]" pageId="447" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
by the pilosity on the antennal scapes and the shape of the petiolar node: the scape setae are subdecumbent and the posterior face of the petiolar node is much shorter than the anterior face in
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[885,1023,1612,1638]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="pilicornis" status="sp. nov.">
<emphasis box="[885,1023,1612,1638]" italics="true" pageId="447" pageNumber="448">T. pilicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1032,1124,1612,1638]" pageId="447" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
The pedunculate petiole with an erect subquadrate node will separate
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[936,1072,1652,1677]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[936,1072,1652,1677]" italics="true" pageId="447" pageNumber="448">T. terrigena</emphasis>
</taxonomicName>
from
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1147,1296,1652,1678]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[1147,1296,1652,1678]" italics="true" pageId="447" pageNumber="448">T. subditivus</emphasis>
</taxonomicName>
, which has a squamiform petiolar node.
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[811,1061,1691,1717]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[811,1061,1691,1717]" italics="true" pageId="447" pageNumber="448">Temnothorax rugosus</emphasis>
</taxonomicName>
has a cuneiform to subcuneiform petiolar node, as opposed to the erect and compact petiolar node of
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1209,1345,1732,1757]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[1209,1345,1732,1757]" italics="true" pageId="447" pageNumber="448">T. terrigena</emphasis>
</taxonomicName>
.
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis italics="true" pageId="447" pageNumber="448">Temnothorax terrigena</emphasis>
</taxonomicName>
differs from
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[758,854,1772,1797]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[758,854,1772,1797]" italics="true" pageId="447" pageNumber="448">T. torrei</emphasis>
</taxonomicName>
by the presence of four erect setae on the dorsum of the petiolar node, as opposed to two.
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[898,1167,1811,1837]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[898,1167,1811,1837]" italics="true" pageId="447" pageNumber="448">Temnothorax terrigena</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[498,618,1852,1877]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="goniops">
<emphasis box="[498,618,1852,1877]" italics="true" pageId="447" pageNumber="448">T. goniops</emphasis>
</taxonomicName>
by the relatively short propodeal spines, which are shorter than the propodeal declivity (longer than the declivity in
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[943,1063,1892,1917]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="goniops">
<emphasis box="[943,1063,1892,1917]" italics="true" pageId="447" pageNumber="448">T. goniops</emphasis>
</taxonomicName>
).
<taxonomicName box="[1087,1327,1891,1917]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="447" pageNumber="448" phylum="Arthropoda" rank="species" species="andrei">
<emphasis box="[1087,1327,1891,1917]" italics="true" pageId="447" pageNumber="448">Temnothorax andrei</emphasis>
</taxonomicName>
has a relatively narrow postpetiole, and both
<taxonomicName box="[848,955,234,260]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="448" pageNumber="449" phylum="Arthropoda" rank="species" species="andrei">
<emphasis box="[848,955,234,260]" italics="true" pageId="448" pageNumber="449">T. andrei</emphasis>
</taxonomicName>
and
<taxonomicName box="[1017,1158,235,260]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="448" pageNumber="449" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis box="[1017,1158,235,260]" italics="true" pageId="448" pageNumber="449">T. carinatus</emphasis>
</taxonomicName>
differ from
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1304,1440,235,260]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="448" pageNumber="449" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[1304,1440,235,260]" italics="true" pageId="448" pageNumber="449">T. terrigena</emphasis>
</taxonomicName>
by their head sculpture, which typically has a medial strip of smooth and shining sculpture, as opposed to the uniformly areolate head of
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1004,1140,315,340]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="448" pageNumber="449" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[1004,1140,315,340]" italics="true" pageId="448" pageNumber="449">T. terrigena</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="451" lastPageNumber="452" pageId="448" pageNumber="449" type="description">
<paragraph blockId="448.[498,1542,234,1934]" pageId="448" pageNumber="449">
<emphasis bold="true" box="[530,1047,353,380]" pageId="448" pageNumber="449">
Worker measurements &amp; indices (
<emphasis bold="true" box="[950,966,354,380]" italics="true" pageId="448" pageNumber="449">n</emphasis>
= 14):
</emphasis>
SL =
<geoCoordinate box="[1119,1181,354,380]" degrees="0.349" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.349">0.349</geoCoordinate>
<geoCoordinate box="[1181,1258,354,380]" degrees="0.403" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.403">0.403</geoCoordinate>
(0.378); FRS =
<geoCoordinate box="[579,641,394,420]" degrees="0.133" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.133">0.133</geoCoordinate>
<geoCoordinate box="[642,719,393,420]" degrees="0.165" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.165">0.165</geoCoordinate>
(0.152); CW =
<geoCoordinate box="[907,969,393,420]" degrees="0.425" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.425">0.425</geoCoordinate>
<geoCoordinate box="[969,1046,394,420]" degrees="0.482" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.482">0.482</geoCoordinate>
(0.456); CWb =
<geoCoordinate box="[1249,1311,393,420]" degrees="0.395" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.395">0.395</geoCoordinate>
<geoCoordinate box="[1311,1388,393,420]" degrees="0.457" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.457">0.457</geoCoordinate>
(0.423); PoOC =
<geoCoordinate box="[604,666,434,460]" degrees="0.208" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.208">0.208</geoCoordinate>
<geoCoordinate box="[667,744,433,460]" degrees="0.233" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.233">0.233</geoCoordinate>
(0.219); CL =
<geoCoordinate box="[919,981,433,460]" degrees="0.511" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.511">0.511</geoCoordinate>
<geoCoordinate box="[981,1058,433,460]" degrees="0.561" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.561">0.561</geoCoordinate>
(0.536); EL =
<geoCoordinate box="[1231,1293,434,460]" degrees="0.102" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.102">0.102</geoCoordinate>
<geoCoordinate box="[1294,1370,433,460]" degrees="0.131" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.131">0.131</geoCoordinate>
(0.117); EW =
<geoCoordinate box="[577,639,473,499]" degrees="0.080" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.08">0.080</geoCoordinate>
<geoCoordinate box="[640,716,473,499]" degrees="0.101" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.101">0.101</geoCoordinate>
(0.089); MD =
<geoCoordinate box="[905,967,473,499]" degrees="0.104" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.104">0.104</geoCoordinate>
<geoCoordinate box="[967,1044,473,499]" degrees="0.123" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.123">0.123</geoCoordinate>
(0.112); WL =
<geoCoordinate box="[1229,1291,473,500]" degrees="0.539" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.539">0.539</geoCoordinate>
<geoCoordinate box="[1291,1368,473,499]" degrees="0.634" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.634">0.634</geoCoordinate>
(0.586); SPST =
<geoCoordinate box="[594,656,513,539]" degrees="0.128" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.128">0.128</geoCoordinate>
<geoCoordinate box="[657,734,513,539]" degrees="0.163" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.163">0.163</geoCoordinate>
(0.143); MPST =
<geoCoordinate box="[948,1010,513,539]" degrees="0.177" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.177">0.177</geoCoordinate>
<geoCoordinate box="[1011,1088,513,539]" degrees="0.222" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.222">0.222</geoCoordinate>
(0.194); PEL =
<geoCoordinate box="[1277,1339,513,539]" degrees="0.211" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.211">0.211</geoCoordinate>
<geoCoordinate box="[1339,1416,513,540]" degrees="0.265" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.265">0.265</geoCoordinate>
(0.228); NOL =
<geoCoordinate box="[591,653,553,579]" degrees="0.101" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.101">0.101</geoCoordinate>
<geoCoordinate box="[654,731,553,579]" degrees="0.134" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.134">0.134</geoCoordinate>
(0.118); NOH =
<geoCoordinate box="[937,999,553,579]" degrees="0.080" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.08">0.080</geoCoordinate>
<geoCoordinate box="[999,1076,553,579]" degrees="0.110" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.11">0.110</geoCoordinate>
(0.100); PEH =
<geoCoordinate box="[1272,1334,553,580]" degrees="0.159" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.159">0.159</geoCoordinate>
<geoCoordinate box="[1334,1411,553,579]" degrees="0.224" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.224">0.224</geoCoordinate>
(0.198); PPL =
<geoCoordinate box="[581,643,593,619]" degrees="0.140" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.14">0.140</geoCoordinate>
<geoCoordinate box="[643,720,593,619]" degrees="0.180" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.18">0.180</geoCoordinate>
(0.156); PW =
<geoCoordinate box="[906,968,593,619]" degrees="0.274" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.274">0.274</geoCoordinate>
<geoCoordinate box="[968,1045,593,619]" degrees="0.314" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.314">0.314</geoCoordinate>
(0.298); SBPA =
<geoCoordinate box="[1253,1315,593,619]" degrees="0.127" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.127">0.127</geoCoordinate>
<geoCoordinate box="[1315,1392,593,620]" degrees="0.154" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.154">0.154</geoCoordinate>
(0.141); SPTI =
<geoCoordinate box="[590,652,633,659]" degrees="0.140" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.14">0.140</geoCoordinate>
<geoCoordinate box="[653,730,633,659]" degrees="0.180" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.18">0.180</geoCoordinate>
(0.165); PEW =
<geoCoordinate box="[932,994,633,659]" degrees="0.113" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.113">0.113</geoCoordinate>
<geoCoordinate box="[994,1071,633,659]" degrees="0.146" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.146">0.146</geoCoordinate>
(0.130); PNW =
<geoCoordinate box="[1278,1340,633,659]" degrees="0.134" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.134">0.134</geoCoordinate>
<geoCoordinate box="[1340,1417,633,659]" degrees="0.171" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.171">0.171</geoCoordinate>
(0.154); PPW =
<geoCoordinate box="[590,652,673,699]" degrees="0.237" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.237">0.237</geoCoordinate>
<geoCoordinate box="[653,730,673,699]" degrees="0.284" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.284">0.284</geoCoordinate>
(0.256); HFL =
<geoCoordinate box="[917,979,673,699]" degrees="0.311" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.311">0.311</geoCoordinate>
<geoCoordinate box="[979,1056,672,699]" degrees="0.405" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.405">0.405</geoCoordinate>
(0.362); HFWmax =
<geoCoordinate box="[1306,1368,673,699]" degrees="0.102" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.102">0.102</geoCoordinate>
<geoCoordinate box="[1368,1445,673,699]" degrees="0.129" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.129">0.129</geoCoordinate>
(0.116); HFWmin =
<geoCoordinate box="[645,707,712,738]" degrees="0.038" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.038">0.038</geoCoordinate>
<geoCoordinate box="[708,785,712,739]" degrees="0.050" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.05">0.050</geoCoordinate>
(0.044); CS =
<geoCoordinate box="[958,1020,712,739]" degrees="0.651" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.651">0.651</geoCoordinate>
<geoCoordinate box="[1021,1098,712,738]" degrees="0.738" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.738">0.738</geoCoordinate>
(0.691); ES =
<geoCoordinate box="[1269,1331,712,738]" degrees="0.142" direction="north" orientation="latitude" pageId="448" pageNumber="449" precision="55" value="0.142">0.142</geoCoordinate>
<geoCoordinate box="[1331,1408,712,738]" degrees="0.178" direction="west" orientation="longitude" pageId="448" pageNumber="449" precision="55" value="-0.178">0.178</geoCoordinate>
(0.161); SI = 8393 (89); OI = 2126 (23); CI = 7682 (79); WLI = 135143 (138); SBI = 3036 (33); PSI = 2128 (24); PWI = 180215 (198); PLI = 123161 (147); NI = 98148 (118); PNWI = 106134 (119); NLI = 4361 (52); FI = 225300 (268).
</paragraph>
<paragraph blockId="448.[498,1542,234,1934]" pageId="448" pageNumber="449">
<emphasis bold="true" box="[530,777,872,898]" pageId="448" pageNumber="449">Worker description:</emphasis>
In full-face view, head subquadrate and elongate (CI 7682). Mandibles densely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin strongly convex medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about two times the maximum width of the scape (SI 8393). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment about twice as long as the preceding two in combination. Frontal carinae short, extending past the antennal toruli by about one and a half times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head very weakly convex, nearly flat, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin concave medially but rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="448.[498,1542,234,1934]" lastBlockId="449.[498,1542,234,1894]" lastPageId="449" lastPageNumber="450" pageId="448" pageNumber="449">
In profile view, compound eyes ovular and moderately large (OI 2126), with 8 ommatidia in longest row. Pronotal declivity distinct: neck and anterior face of pronotum forming a ~110
<emphasis box="[682,692,1465,1483]" italics="true" pageId="448" pageNumber="449">
<superScript attach="left" box="[682,692,1465,1483]" fontSize="8" pageId="448" pageNumber="449">°</superScript>
</emphasis>
angle; anterior face and dorsum of mesosoma forming a ~120
<emphasis box="[1433,1443,1465,1483]" italics="true" pageId="448" pageNumber="449">
<superScript attach="left" box="[1433,1443,1465,1483]" fontSize="8" pageId="448" pageNumber="449">°</superScript>
</emphasis>
angle. Mesosoma very weakly convex from where it joins the pronotal neck to the propodeal spines. Promesonotal suture extending from the posterior margin of the procoxal insertion only to the mesothoracic spiracle, which is moderately well developed. Metanotal groove visible as a disruption of the sculpture laterally from where it arises between the mid- and hind coxae to where it ends in the poorly developed metathoracic spiracle, which is nearly indistinguishable against the ground sculpture. Propodeal spiracle weakly developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines weakly developed and short (PSI 2128), about half as long as the propodeal declivity, flared at the base, triangular, and acute. Propodeal declivity flat, forming a rounded ~120
<emphasis box="[1075,1085,1863,1881]" italics="true" pageId="448" pageNumber="449">
<superScript attach="left" box="[1075,1085,1863,1881]" fontSize="8" pageId="448" pageNumber="449">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed. Metapleural gland bulla moderately large, extending from the metacoxal insertion two thirds of the way to the propodeal spiracle. Petiole short (PLI 123161), without tubercles anterodorsally. Subpetiolar process in the form of a moderately large, blunt, triangular tooth which grades evenly into the ventral margin of the petiole posteriorly; ventral margin of petiole flat posterior to it. Petiolar peduncle short: comprising about a third of the total petiole length. Petiolar node robust, erect, and rounded-subquadrate: transition between peduncle and node marked by a rounded angle of ~120
<emphasis box="[989,999,469,487]" italics="true" pageId="449" pageNumber="450">
<superScript attach="none" box="[989,999,469,487]" fontSize="8" pageId="449" pageNumber="450">°</superScript>
</emphasis>
; anterior face forming a ~100
<emphasis box="[1354,1364,469,487]" italics="true" pageId="449" pageNumber="450">
<superScript attach="left" box="[1354,1364,469,487]" fontSize="8" pageId="449" pageNumber="450">°</superScript>
</emphasis>
angle with the dorsal face, which is convex; dorsal face meeting the posterior face at a rounded ~90
<emphasis box="[1456,1466,509,527]" italics="true" pageId="449" pageNumber="450">
<superScript attach="left" box="[1456,1466,509,527]" fontSize="8" pageId="449" pageNumber="450">°</superScript>
</emphasis>
angle; posterior face forms a ~90
<emphasis box="[809,819,548,566]" italics="true" pageId="449" pageNumber="450">
<superScript attach="left" box="[809,819,548,566]" fontSize="8" pageId="449" pageNumber="450">°</superScript>
</emphasis>
angle with the caudal cylinder. Postpetiole strongly convex anteriorly, weakly convex dorsally, and weakly lobed ventrally.
</paragraph>
<paragraph blockId="449.[498,1542,234,1894]" pageId="449" pageNumber="450">In dorsal view, humeri developed and distinct: wider than the rest of the mesosoma; mesothoracic spiracles weakly protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Metanotal groove absent: mesonotum and propodeum completely fused and lateral margins converging evenly to the bases of the propodeal spines. Propodeal spines broadly approximated basally and diverging apically, their apices separated from each other by about twice their length, the negative space between them “U” shaped. Petiolar peduncle with spiracles weakly protruding past the lateral margins, but not noticeably constricted anterior to them. Petiolar node campaniform and antero-posteriorly compressed: convex anteriorly and flat posteriorly; node slightly wider than the peduncle and caudal cylinder. Postpetiole moderately broad (PWI 180215) and subtrapezoidal, articulating with the nearly the entire anterior margin of the gaster, but leaving small angles on either side exposed. Anterior margin of the postpetiole weakly convex and evenly rounds into the lateral margins; lateral margins converge slightly to the rounded posterior corners; posterior margin broadly concave. Metafemur weakly to moderately incrassate (FI 225300).</paragraph>
<paragraph blockId="449.[498,1542,234,1894]" pageId="449" pageNumber="450">Sculpture: median clypeal carina present but weak, extending posteriorly nearly to the frontal triangle, and flanked on either side by two equally strong carinae. Lateral clypeal lobes with additional, weaker carinae; ground sculpture weakly areolate. Antennal scapes weakly areolate. Cephalic dorsum and lateral surfaces of head uniformly areolate. Ventral surface of head shining through weak areolae. Pronotal neck and anterior face of the pronotum areolate. Lateral surfaces of mesosoma and shining through weak areolate sculpture. Propodeal declivity areolate. Dorsal surface of mesosoma uniformly areolate. Femora shining, with traces of weak areolate sculpture. Petiole and postpetiole areolate, with anterior and dorsal surfaces of petiolar node, and dorso-medial surface of postpetiole smooth and shining. First gastral tergite and sternite smooth and shining, without spectral iridescence.</paragraph>
<paragraph blockId="449.[498,1542,234,1894]" pageId="449" pageNumber="450">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of the head, pronotum, waist segments, and gaster with sparse, erect, blunt-tipped, nearly clavate setae, the longest of which are about the half the width of the compound eye. The head bears ~28, mesosoma ~16, petiole 6, postpetiole ~10, and first gastral tergite ~36 setae. Short, sparse pubescence present over the entire body, but difficult to detect against the lightly colored integument.</paragraph>
<paragraph blockId="450.[498,1542,234,1934]" pageId="450" pageNumber="451">Color: predominantly yellow, with masticatory margin of mandibles dark brown; antennal club testaceous yellow.</paragraph>
<paragraph blockId="450.[498,1542,234,1934]" pageId="450" pageNumber="451">
<emphasis bold="true" box="[530,1001,313,340]" pageId="450" pageNumber="451">
Gyne measurements &amp; indices (
<emphasis bold="true" box="[919,935,314,340]" italics="true" pageId="450" pageNumber="451">n</emphasis>
= 2):
</emphasis>
SL = 0.437; FRS =
<geoCoordinate box="[1232,1294,314,340]" degrees="0.190" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.19">0.190</geoCoordinate>
<geoCoordinate box="[1294,1371,314,340]" degrees="0.202" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.202">0.202</geoCoordinate>
(0.196); CW =
<geoCoordinate box="[580,642,354,381]" degrees="0.596" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.596">0.596</geoCoordinate>
<geoCoordinate box="[642,719,354,381]" degrees="0.605" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.605">0.605</geoCoordinate>
(0.601); CWb =
<geoCoordinate box="[922,984,354,381]" degrees="0.541" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.541">0.541</geoCoordinate>
<geoCoordinate box="[984,1061,354,381]" degrees="0.554" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.554">0.554</geoCoordinate>
(0.548); PoOC =
<geoCoordinate box="[1274,1336,354,380]" degrees="0.214" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.214">0.214</geoCoordinate>
<geoCoordinate box="[1336,1413,354,380]" degrees="0.221" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.221">0.221</geoCoordinate>
(0.218); CL =
<geoCoordinate box="[567,629,393,420]" degrees="0.599" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.599">0.599</geoCoordinate>
<geoCoordinate box="[630,706,394,420]" degrees="0.613" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.613">0.613</geoCoordinate>
(0.606); EL =
<geoCoordinate box="[879,941,394,420]" degrees="0.189" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.189">0.189</geoCoordinate>
<geoCoordinate box="[942,1019,394,420]" degrees="0.209" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.209">0.209</geoCoordinate>
(0.199); EW = 0.154; MD =
<geoCoordinate box="[1363,1425,394,420]" degrees="0.108" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.108">0.108</geoCoordinate>
<geoCoordinate box="[1425,1502,394,420]" degrees="0.112" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.112">0.112</geoCoordinate>
(0.110); WL =
<geoCoordinate box="[674,736,433,459]" degrees="0.969" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.969">0.969</geoCoordinate>
<geoCoordinate box="[736,813,433,460]" degrees="0.989" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.989">0.989</geoCoordinate>
(0.979); SPST =
<geoCoordinate box="[1016,1078,434,460]" degrees="0.191" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.191">0.191</geoCoordinate>
<geoCoordinate box="[1078,1155,433,460]" degrees="0.207" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.207">0.207</geoCoordinate>
(0.199); MPST =
<geoCoordinate box="[1370,1432,434,460]" degrees="0.249" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.249">0.249</geoCoordinate>
<geoCoordinate box="[1432,1509,433,459]" degrees="0.266" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.266">0.266</geoCoordinate>
(0.258); PEL =
<geoCoordinate box="[678,740,473,499]" degrees="0.310" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.31">0.310</geoCoordinate>
<geoCoordinate box="[741,818,473,499]" degrees="0.319" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.319">0.319</geoCoordinate>
(0.315); NOL =
<geoCoordinate box="[1017,1079,473,500]" degrees="0.152" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.152">0.152</geoCoordinate>
<geoCoordinate box="[1080,1157,473,500]" degrees="0.165" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.165">0.165</geoCoordinate>
(0.159); NOH =
<geoCoordinate box="[1363,1425,473,500]" degrees="0.115" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.115">0.115</geoCoordinate>
<geoCoordinate box="[1425,1502,473,499]" degrees="0.133" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.133">0.133</geoCoordinate>
(0.124); PEH =
<geoCoordinate box="[677,739,513,539]" degrees="0.247" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.247">0.247</geoCoordinate>
<geoCoordinate box="[739,816,513,539]" degrees="0.268" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.268">0.268</geoCoordinate>
(0.258); PPL =
<geoCoordinate box="[994,1056,513,539]" degrees="0.163" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.163">0.163</geoCoordinate>
<geoCoordinate box="[1056,1133,513,539]" degrees="0.211" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.211">0.211</geoCoordinate>
(0.187); PW =
<geoCoordinate box="[1307,1369,513,540]" degrees="0.575" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.575">0.575</geoCoordinate>
<geoCoordinate box="[1370,1447,513,539]" degrees="0.601" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.601">0.601</geoCoordinate>
(0.588); SBPA =
<geoCoordinate box="[600,662,553,579]" degrees="0.288" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.288">0.288</geoCoordinate>
<geoCoordinate box="[662,739,553,579]" degrees="0.306" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.306">0.306</geoCoordinate>
(0.297); SPTI =
<geoCoordinate box="[938,1000,553,580]" degrees="0.257" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.257">0.257</geoCoordinate>
<geoCoordinate box="[1000,1077,553,579]" degrees="0.268" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.268">0.268</geoCoordinate>
(0.263); PEW =
<geoCoordinate box="[1279,1341,553,579]" degrees="0.161" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.161">0.161</geoCoordinate>
<geoCoordinate box="[1341,1418,553,579]" degrees="0.202" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.202">0.202</geoCoordinate>
(0.182); PNW =
<geoCoordinate box="[599,661,593,619]" degrees="0.216" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.216">0.216</geoCoordinate>
<geoCoordinate box="[661,738,593,619]" degrees="0.217" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.217">0.217</geoCoordinate>
(0.217); PPW =
<geoCoordinate box="[940,1002,593,619]" degrees="0.339" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.339">0.339</geoCoordinate>
<geoCoordinate box="[1002,1079,593,619]" degrees="0.346" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.346">0.346</geoCoordinate>
(0.343); HFL =
<geoCoordinate box="[1273,1335,593,619]" degrees="0.466" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.466">0.466</geoCoordinate>
<geoCoordinate box="[1335,1412,593,619]" degrees="0.482" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.482">0.482</geoCoordinate>
(0.474); HFWmax =
<geoCoordinate box="[648,710,633,659]" degrees="0.099" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.099">0.099</geoCoordinate>
<geoCoordinate box="[711,788,633,659]" degrees="0.124" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.124">0.124</geoCoordinate>
(0.112); HFWmin =
<geoCoordinate box="[1041,1103,633,659]" degrees="0.043" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.043">0.043</geoCoordinate>
<geoCoordinate box="[1104,1181,633,659]" degrees="0.044" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.044">0.044</geoCoordinate>
(0.044); CS =
<geoCoordinate box="[1354,1416,633,659]" degrees="0.841" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.841">0.841</geoCoordinate>
<geoCoordinate box="[1417,1494,633,659]" degrees="0.861" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.861">0.861</geoCoordinate>
(0.851); ES =
<geoCoordinate box="[658,720,672,698]" degrees="0.266" direction="north" orientation="latitude" pageId="450" pageNumber="451" precision="55" value="0.266">0.266</geoCoordinate>
<geoCoordinate box="[720,797,672,699]" degrees="0.286" direction="west" orientation="longitude" pageId="450" pageNumber="451" precision="55" value="-0.286">0.286</geoCoordinate>
(0.276); SI = 7981 (80); OI = 3233 (32); CI = 90; WLI = 179; SBI = 5355 (54); PSI = 1921 (20); PWI = 168215 (191); PLI = 147196 (171); NI = 124132 (128); PNWI = 107135 (121); NLI = 4952 (50); FI = 230282 (256).
</paragraph>
<paragraph blockId="450.[498,1542,234,1934]" pageId="450" pageNumber="451">
<emphasis bold="true" box="[530,747,792,818]" pageId="450" pageNumber="451">Gyne description:</emphasis>
In full-face view, head subquadrate, slightly longer than broad (CI 90). Mandibles densely, finely striate but shining and armed with five teeth: the apical-most well developed, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin strongly convex medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about twice the maximum scape width (SI 7981). Antennae 12-segmented; antennal club composed of three segments, with the apical-most segment about twice as long as the preceding two in combination. Frontal carinae short, extending past the antennal toruli by about the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head evenly convex, converging from below the compound eyes to the mandibular insertions. Posterior head margin flat, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="450.[498,1542,234,1934]" lastBlockId="451.[498,1542,234,1895]" lastPageId="451" lastPageNumber="452" pageId="450" pageNumber="451">
In profile view, compound eyes ovular and large (OI 3233), with 16 ommatidia in longest row. Mesoscutum rounded evenly anteriorly, covering the dorsal surface of the pronotum, and flat dorsally. Mesoscutellum on the same plane as the mesoscutum. Posterior margin of metanotum extending past the posterior margin of the mesoscutum. Propodeal spiracle moderately well developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines stout and moderately well developed, but short (PSI ~ 20), about a third as long as the propodeal declivity, flared at the base, triangular, and acute. Propodeal declivity straight and flat, forming a ~120
<emphasis box="[677,687,1584,1602]" italics="true" pageId="450" pageNumber="451">
<superScript attach="left" box="[677,687,1584,1602]" fontSize="8" pageId="450" pageNumber="451">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and very weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole moderately long (PLI 147196), without tubercles anterodorsally. Subpetiolar process in the form of a moderately large, triangular, blunt tooth, which grades evenly into the ventral margin of the petiole posteriorly. Petiolar peduncle moderately long: comprising about half of the total petiole length. Petiolar node erect and squamiform: transition between peduncle and node a rounded ~130
<emphasis box="[664,674,1863,1881]" italics="true" pageId="450" pageNumber="451">
<superScript attach="left" box="[664,674,1863,1881]" fontSize="8" pageId="450" pageNumber="451">°</superScript>
</emphasis>
angle; anterior face forming a ~90
<emphasis box="[1079,1089,1863,1881]" italics="true" pageId="450" pageNumber="451">
<superScript attach="left" box="[1079,1089,1863,1881]" fontSize="8" pageId="450" pageNumber="451">°</superScript>
</emphasis>
angle with the brief dorsal face; dorsal face meeting the posterior face at a ~90
<emphasis box="[957,967,1903,1921]" italics="true" pageId="450" pageNumber="451">
<superScript attach="left" box="[957,967,1903,1921]" fontSize="8" pageId="450" pageNumber="451">°</superScript>
</emphasis>
angle; posterior face forms a ~90
<emphasis box="[1358,1368,1903,1921]" italics="true" pageId="450" pageNumber="451">
<superScript attach="left" box="[1358,1368,1903,1921]" fontSize="8" pageId="450" pageNumber="451">°</superScript>
</emphasis>
angle with the caudal cylinder. Postpetiole flat anteriorly, bulging slightly anterodorsally, flattened posterodorsally; ventral surface weakly lobed.
</paragraph>
<paragraph blockId="451.[498,1542,234,1895]" pageId="451" pageNumber="452">In dorsal view, mesoscutum covering pronotum anteriorly, but humeri visible laterally as angulate sclerites. Propodeal spines parallel apically, their apices separated from each other by about three times their length. Petiolar peduncle with spiracles weakly protruding past the lateral margins. Petiolar node transversely campaniform: rounded anteriorly and flattened posteriorly; subquadrate when viewed posterodorsally, the dorsum of the node weakly convex. Petiolar node slightly broader than the peduncle and caudal cylinder. Postpetiole moderately broad (PWI 168215), anteroposteriorly compressed, and subtrapezoidal, articulating with most of the anterior margin of the gaster, leaving small, angulate margins on each side exposed. Anterior margin of postpetiole flat, with corners marked by rounded angles as it transitions to the lateral margins, which converge slightly to the angulate posterior corners; posterior margin flat. Metafemur weakly to moderately incrassate (FI 230282).</paragraph>
<paragraph blockId="451.[498,1542,234,1895]" pageId="451" pageNumber="452">Sculpture: median clypeal carina present, extending from the anterior margin nearly to frontal triangle, and flanked by two weaker, indistinct carinae. Lateral clypeal lobes with additional weaker carinae; ground sculpture weakly areolate. Antennal scapes weakly areolate. Cephalic dorsum areolate, with weak costulae overlying the ground sculpture. Lateral surfaces of head areolate, with fine costulae between the compound eye and mandibular insertion. Ventral surface of head smooth and shining, with weak costulae. Pronotal neck and anterior face of pronotum areolate. Pronotum with areolate ground sculpture arranged into longitudinal rows by superficial costulae. Katepisternum shining medially, otherwise weakly areolate. Anepisternum areolate. Metapleurae and lateral face of propodeum areolate, with costulae that become stronger over the metapleural gland bulla. Propodeal declivity areolate. Mesoscutum predominantly with costulae over areolate ground sculpture; a patch of smooth and shining sculpture anteromedially. Mesoscutellum with costulae over areolate ground sculpture. Femora shining, with traces of weak areolate sculpture. Petiole and postpetiole uniformly areolate, but sculpture becomes weaker on the anterior face of the petiolar node and anterior face of postpetiole. First gastral tergite and sternite smooth and shining, without spectral iridescence.</paragraph>
<paragraph blockId="451.[498,1542,234,1895]" pageId="451" pageNumber="452">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of the head, pronotum, waist segments, and gaster with sparse, erect, blunt-tipped, nearly clavate setae, the longest of which are about a third of the width of the compound eye. Short, sparse pubescence present over the entire body, but difficult to detect against the lightly colored integument.</paragraph>
<paragraph blockId="451.[498,1542,234,1895]" pageId="451" pageNumber="452">Color: predominantly yellow; with antennae, dorsomedial surface of head, posteromedial surface of mesoscutum, metanotum, propodeal declivity, caudal cylinder of the petiole, caudal cylinder of the postpetiole, and distal quarters of the gastral sclerites testaceous yellow. Wing bases and masticatory margin of mandibles dark brown.</paragraph>
<paragraph blockId="451.[498,1542,234,1895]" box="[530,730,1788,1814]" pageId="451" pageNumber="452">
<emphasis bold="true" box="[530,597,1788,1814]" pageId="451" pageNumber="452">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="451" pageNumber="452" type="etymology">
<paragraph blockId="451.[498,1542,234,1895]" pageId="451" pageNumber="452">
<emphasis bold="true" box="[530,668,1828,1854]" pageId="451" pageNumber="452">Etymology:</emphasis>
Behavioural, from the Latin terra (= earth) + genus (= born), presumably a reference to the terrestrial nesting and foraging habits of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="452" pageNumber="453" type="discussion">
<paragraph blockId="452.[498,1542,234,819]" pageId="452" pageNumber="453">
<emphasis bold="true" box="[530,670,234,260]" pageId="452" pageNumber="453">Comments:</emphasis>
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[677,943,234,260]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="452" pageNumber="453" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[677,943,234,260]" italics="true" pageId="452" pageNumber="453">Temnothorax terrigena</emphasis>
</taxonomicName>
, which inhabits a range from the Chisos Mountains in southwestern Texas through central Texas, and south to
<collectingRegion box="[1201,1399,274,301]" country="Mexico" name="Tamaulipas" pageId="452" pageNumber="453">Tamaulipas state</collectingRegion>
in
<collectingCountry box="[1440,1528,274,300]" name="Mexico" pageId="452" pageNumber="453">Mexico</collectingCountry>
, apparently inhabits a broad range of habitats, including pinyon forests, desert canyons, high Douglas fir/pine forests, and semiarid plains (
<bibRefCitation author="van Pelt A." box="[1099,1264,354,380]" journalOrPublisher="The Southwestern Naturalist" pageId="452" pageNumber="453" pagination="137 - 142" part="28" refId="ref273009" refString="van Pelt A. 1983. Ants of the Chisos Mountains, Texas (Hymenoptera: Formicidae). The Southwestern Naturalist 28 (2): 137 - 142 DOI 10.2307 / 3671381." title="Ants of the Chisos Mountains, Texas (Hymenoptera: Formicidae)" type="journal article" year="1983">
<emphasis box="[1099,1264,354,380]" italics="true" pageId="452" pageNumber="453">van Pelt, 1983</emphasis>
</bibRefCitation>
,
<bibRefCitation author="Wheeler WM" box="[1279,1458,354,380]" journalOrPublisher="Psyche: A Journal of Entomology" pageId="452" pageNumber="453" pagination="93 - 111" part="10" refId="ref273239" refString="Wheeler WM. 1903 a. A decad of Texan Formicidae. Psyche: A Journal of Entomology 10 (323): 93 - 111 DOI 10.1155 / 1903 / 67840." title="A decad of Texan Formicidae" type="journal article" year="1903">
<emphasis box="[1279,1458,354,380]" italics="true" pageId="452" pageNumber="453">Wheeler, 1903a</emphasis>
</bibRefCitation>
). It is common in the
<typeStatus box="[685,735,394,420]" pageId="452" pageNumber="453">type</typeStatus>
locality of Austin, Texas, where it has been collected in litter samples at the Brackenridge Field Laboratory (
<bibRefCitation author="Feener DH" box="[946,1092,434,460]" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="452" pageNumber="453" pagination="269 - 277" part="54" refId="ref270833" refString="Feener DH. 1981. Notes on the biology of Pheidole lamia (Hymenoptera: Formicidae) at its type locality (Austin, Texas). Journal of the Kansas Entomological Society 54: 269 - 277." title="Notes on the biology of Pheidole lamia (Hymenoptera: Formicidae) at its type locality (Austin, Texas)" type="journal article" year="1981">
<emphasis box="[946,1092,434,460]" italics="true" pageId="452" pageNumber="453">Feener, 1981</emphasis>
</bibRefCitation>
). In the original description (
<bibRefCitation author="Wheeler WM" journalOrPublisher="Proceedings of the Academy of Natural Sciences of: Philadelphia" pageId="452" pageNumber="453" pagination="215 - 260" refId="ref273272" refString="Wheeler WM. 1903 b. A revision of the North American ants of the genus Leptothorax Mayr. In: Proceedings of the Academy of Natural Sciences of: Philadelphia. 215 - 260." title="A revision of the North American ants of the genus Leptothorax Mayr" type="book chapter" year="1903">
<emphasis box="[1440,1541,433,459]" italics="true" pageId="452" pageNumber="453">Wheeler,</emphasis>
<emphasis box="[498,567,473,499]" italics="true" pageId="452" pageNumber="453">1903b</emphasis>
</bibRefCitation>
),
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[592,727,474,499]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="452" pageNumber="453" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[592,727,474,499]" italics="true" pageId="452" pageNumber="453">T. terrigena</emphasis>
</taxonomicName>
was noted as nesting under stones in black clay soil.
<bibRefCitation author="Wheeler WM" box="[1351,1541,473,499]" journalOrPublisher="Proceedings of the Academy of Natural Sciences of: Philadelphia" pageId="452" pageNumber="453" pagination="215 - 260" refId="ref273272" refString="Wheeler WM. 1903 b. A revision of the North American ants of the genus Leptothorax Mayr. In: Proceedings of the Academy of Natural Sciences of: Philadelphia. 215 - 260." title="A revision of the North American ants of the genus Leptothorax Mayr" type="book chapter" year="1903">
<emphasis box="[1351,1541,473,499]" italics="true" pageId="452" pageNumber="453">Wheeler (1903b)</emphasis>
</bibRefCitation>
observed workers foraging diurnally and that this species may possibly be pleisiobiotic with other myrmicines: Wheeler collected a nest from a midden pile of the fungus farming ant
<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" box="[544,969,593,620]" class="Insecta" family="Formicidae" genus="Mycetomoellerius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="452" pageNumber="453" phylum="Arthropoda" rank="species" species="turrifex">
<emphasis box="[544,841,593,619]" italics="true" pageId="452" pageNumber="453">Mycetomoellerius turrifex</emphasis>
(Wheeler)
</taxonomicName>
, and another from a
<taxonomicName authority="Pergande" authorityName="Pergande" authorityYear="1896" box="[1221,1542,593,619]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="452" pageNumber="453" phylum="Arthropoda" rank="species" species="tepicana">
<emphasis box="[1221,1424,593,619]" italics="true" pageId="452" pageNumber="453">Pheidole tepicana</emphasis>
Pergande
</taxonomicName>
colony. The references to
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[804,939,633,658]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="452" pageNumber="453" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[804,939,633,658]" italics="true" pageId="452" pageNumber="453">T. terrigena</emphasis>
</taxonomicName>
from
<collectingRegion box="[1015,1169,633,659]" country="Mexico" name="Hidalgo" pageId="452" pageNumber="453">Hidalgo state</collectingRegion>
by
<emphasis box="[1215,1486,633,659]" italics="true" pageId="452" pageNumber="453">
Hernández &amp;
<emphasis box="[498,676,673,699]" italics="true" pageId="452" pageNumber="453">Castaño-Meneses (2010)</emphasis>
</emphasis>
,
<bibRefCitation author="Hernandez FV" box="[691,901,673,699]" journalOrPublisher="Revista Peruana de Entomologia" pageId="452" pageNumber="453" pagination="1 - 8" part="48" refId="ref271413" refString="Hernandez FV. 2013. Variacion temporal de las comunidades de hormigas en matorral xerofilo con dominancia de Cephalocereus senilis y Stenocereus dumortieri en la Reserva de la Biosfera Barranca de Metztitlan, Hidalgo, Mexico. Revista Peruana de Entomologia 48: 1 - 8." title="Variacion temporal de las comunidades de hormigas en matorral xerofilo con dominancia de Cephalocereus senilis y Stenocereus dumortieri en la Reserva de la Biosfera Barranca de Metztitlan, Hidalgo, Mexico" type="journal article" year="2013">
<emphasis box="[691,901,673,699]" italics="true" pageId="452" pageNumber="453">Hernández (2013)</emphasis>
</bibRefCitation>
,
<bibRefCitation author="Vasquez-Bolanos M." box="[916,1196,673,699]" journalOrPublisher="Metodos en Ecologia y Sistematica" pageId="452" pageNumber="453" pagination="1 - 53" part="10" refId="ref273069" refString="Vasquez-Bolanos M. 2015. Taxonomia de Formicidae (Hymenoptera) para Mexico. Metodos en Ecologia y Sistematica 10 (1): 1 - 53." title="Taxonomia de Formicidae (Hymenoptera) para Mexico" type="journal article" year="2015">
<emphasis box="[916,1196,673,699]" italics="true" pageId="452" pageNumber="453">Vásquez-Bolaños (2015)</emphasis>
</bibRefCitation>
are incorrect: the exemplar specimen from the study by Hernández &amp; Castaño-Meneses deposited in the UNAM collection proved to be an undescribed species of the
<taxonomicName box="[1131,1266,753,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="452" pageNumber="453" phylum="Arthropoda" rank="species" species="tricarinatus">
<emphasis box="[1131,1266,753,778]" italics="true" pageId="452" pageNumber="453">tricarinatus</emphasis>
</taxonomicName>
species group (sensu
<bibRefCitation author="Mackay WP" box="[498,660,792,818]" journalOrPublisher="Sociobiology" pageId="452" pageNumber="453" pagination="265 - 444" part="36" refId="ref271737" refString="Mackay WP. 2000. A review of the new world ants of the subgenus Myrafant (genus Leptothorax) (Hymenoptera: Formicidae). Sociobiology 36: 265 - 444." title="A review of the new world ants of the subgenus Myrafant (genus Leptothorax) (Hymenoptera: Formicidae)" type="journal article" year="2000">
<emphasis box="[498,660,792,818]" italics="true" pageId="452" pageNumber="453">Mackay, 2000</emphasis>
</bibRefCitation>
) upon examination.
</paragraph>
</subSubSection>
</treatment>
</document>