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<document id="7F3C2B3115521428110AC96AF07C259F" ID-DOI="10.11646/zootaxa.3893.4.8" ID-GBIF-Dataset="0a404643-b1ad-422a-b274-96c8c472b7bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="227638" ID-ZooBank="F7114368-60C9-4DAC-A718-3148AE31A39E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460996691588" checkinUser="plazi" docAuthor="Wang, Yang &amp; Zhang, Yalin" docDate="2014" docId="052AD3409162E74AFAFEB79C2D9B74CD" docLanguage="en" docName="zt03893p594.pdf" docOrigin="Zootaxa 3893 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Angustuma flavimacula Wang &amp; Zhang, 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="593" masterDocId="F913AB389164E74CFA69B70B2E367726" masterDocTitle="Three new species in the leafhopper genus Angustuma (Hemiptera: Cicadellidae: Evacanthinae) from Thailand" masterLastPageNumber="594" masterPageNumber="587" pageNumber="593" updateTime="1698562951739" updateUser="plazi">
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<mods:title id="E704BCE4000A98305FAE262BA3D4EB3D">Three new species in the leafhopper genus Angustuma (Hemiptera: Cicadellidae: Evacanthinae) from Thailand</mods:title>
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<mods:namePart id="34396A973C6F206569A3914D6819961A">Wang, Yang</mods:namePart>
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<mods:namePart id="174DE5E882D1EC0E2C4E087326C8F260">Zhang, Yalin</mods:namePart>
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<mods:title id="2C188A8C46740A3DB30E020E493B8F3C">Zootaxa</mods:title>
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<mods:date id="5AC58FEB0C3DC0DBA61E8636B30542A5">2014</mods:date>
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<treatment id="052AD3409162E74AFAFEB79C2D9B74CD" ID-DOI="http://doi.org/10.5281/zenodo.6137487" ID-GBIF-Taxon="119597315" ID-Zenodo-Dep="6137487" LSID="urn:lsid:plazi:treatment:052AD3409162E74AFAFEB79C2D9B74CD" httpUri="http://treatment.plazi.org/id/052AD3409162E74AFAFEB79C2D9B74CD" lastPageNumber="593" pageId="6" pageNumber="593">
<subSubSection id="C59931DD9162E74AFAFEB79C2F1077F5" pageId="6" pageNumber="593" type="nomenclature">
<paragraph id="8D3C62569162E74AFAFEB79C2C287794" blockId="6.[151,542,151,211]" box="[151,542,151,178]" pageId="6" pageNumber="593">
<heading id="D674D53A9162E74AFAFEB79C2C287794" bold="true" box="[151,542,151,178]" fontSize="11" level="1" pageId="6" pageNumber="593" reason="1">
<emphasis id="BFF7BE449162E74AFAFEB79C2C287794" bold="true" box="[151,542,151,178]" pageId="6" pageNumber="593">
<taxonomicName id="4A8319D59162E74AFAFEB79C2F8F7797" ID-CoL="8JPF9" box="[151,441,151,177]" class="Insecta" family="Cicadellidae" genus="Angustuma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="flavimacula" status="sp. nov.">
<emphasis id="BFF7BE449162E74AFAFEB79C2F8F7797" bold="true" box="[151,441,151,177]" italics="true" pageId="6" pageNumber="593">Angustuma flavimacula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4C4033F9162E74AFBA8B7932C287794" box="[449,542,152,178]" pageId="6" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8D3C62569162E74AFAFEB7B12F1077F5" blockId="6.[151,542,151,211]" box="[151,294,186,211]" pageId="6" pageNumber="593">
(
<figureCitation id="15B87ED39162E74AFAF6B7B12F2877F5" box="[159,286,186,211]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Figs 2131</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C59931DD9162E74AFAFEB6092B1A75ED" pageId="6" pageNumber="593" type="description">
<paragraph id="8D3C62569162E74AFAFEB6092FF176D5" blockId="6.[151,1437,258,1003]" pageId="6" pageNumber="593">
<emphasis id="BFF7BE449162E74AFAFEB6092F1F763D" bold="true" box="[151,297,258,283]" pageId="6" pageNumber="593">Description.</emphasis>
Body length (including forewing): ♂,
<quantity id="4A7BCFB39162E74AF89BB6082D79763C" box="[754,847,259,283]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="6" pageNumber="593" unit="mm" value="7.2">7.2 mm</quantity>
; ♀,
<quantity id="4A7BCFB39162E74AF9EFB6082DD5763C" box="[902,995,259,283]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="6" pageNumber="593" unit="mm" value="7.9">7.9 mm</quantity>
. Dorsum (
<figureCitation id="15B87ED39162E74AFE36B6092A82763D" box="[1119,1204,258,283]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Fig. 21</figureCitation>
) blackish ochreous, marked with off-white and yellow spots. Crown (
<figureCitation id="15B87ED39162E74AF8D4B62D2D0A7619" box="[701,828,294,319]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Figs 21, 23</figureCitation>
) with four irregular off-white spots on lateral, anterior and posterior margin. Face (
<figureCitation id="15B87ED39162E74AFBBAB6412C137645" box="[467,549,330,355]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Fig. 24</figureCitation>
) with median longitudinal carina blackish ochreous, gradually narrowing from apex to base, with pair of narrow blackish ochreous maculae just mesad of eyes. Lorum (
<figureCitation id="15B87ED39162E74AFE13B6652A8B76A1" box="[1146,1213,366,391]" captionStart="FIGURES 1 10" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[186,1415,203,1890]" captionTargetId="figure@3.[166,1420,190,1902]" captionTargetPageId="3" captionText="FIGURES 1 10. Angustuma basinigra sp. nov.: 1. male habitus, dorsal view; 2. male habitus, lateral view; 3. male head and thorax, dorsal view; 4. male face; 5. male pygofer, lateral view; 6. male pygofer side, lateral view; 7. aedeagal, lateral view; 8. aedeagal, ventral view; 9. connective and style, ventral view; 10. subgenital plate, ventral view." httpUri="https://zenodo.org/record/227639/files/figure.png" pageId="6" pageNumber="593">Fig. 4</figureCitation>
) blackish ochreous. Gena (
<figureCitation id="15B87ED39162E74AFA8DB6992F01768D" box="[228,311,402,427]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Fig. 24</figureCitation>
) with blackish ochreous stripes on the middle portion. Pronotum (
<figureCitation id="15B87ED39162E74AFE5DB6992A8E768D" box="[1076,1208,402,427]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Figs 21, 23</figureCitation>
) blackish ochreous, marked with off-white on middle lateral margin. Forewing (
<figureCitation id="15B87ED39162E74AF924B6BD2DE576E9" box="[845,979,438,463]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Figs 21, 22</figureCitation>
) with large yellow spots on clavus and lateral margin respectively.
</paragraph>
<paragraph id="8D3C62569162E74AFAAEB6F52CD975A5" blockId="6.[151,1437,258,1003]" pageId="6" pageNumber="593">
Aedeagal (
<figureCitation id="15B87ED39162E74AFB29B6F52FA47531" box="[320,402,510,535]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Fig. 28</figureCitation>
) ventral apophysis in lateral view finlike, apex pointed. Style (
<figureCitation id="15B87ED39162E74AFE31B6F52A9C7531" box="[1112,1194,510,535]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Fig. 29</figureCitation>
) apophysis in ventral view elongate and acuminate. Connective (
<figureCitation id="15B87ED39162E74AF814B5292CF9751D" box="[637,719,546,571]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Fig. 27</figureCitation>
) stem broadened near apex with pair of short retrorse preapical spines. Subgenital plate (
<figureCitation id="15B87ED39162E74AFBD0B54D2C3B7579" box="[441,525,582,607]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Fig. 30</figureCitation>
) knifelike, broad in middle portion, macrosetae in single row through most of length, row of elongate setae near interior submargin.
</paragraph>
<paragraph id="8D3C62569162E74AFAAEB5852B1A75ED" blockId="6.[151,1437,258,1003]" pageId="6" pageNumber="593">
Female and male (
<figureCitation id="15B87ED39162E74AFBF5B5852FC67581" box="[412,496,654,679]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Fig. 25</figureCitation>
) are slightly dimorphic in coloration and shape of marks. Female seventh sternite (
<figureCitation id="15B87ED39162E74AFAF6B5B92EC775ED" box="[159,241,690,715]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Fig. 31</figureCitation>
) longer than the sixth sternite, posterior margin well produced laterally and almost flat medially.
</paragraph>
</subSubSection>
<subSubSection id="C59931DD9162E74AFAAEB5DD2F367411" pageId="6" pageNumber="593" type="materials_examined">
<paragraph id="8D3C62569162E74AFAAEB5DD2F367411" blockId="6.[151,1437,258,1003]" pageId="6" pageNumber="593">
<emphasis id="BFF7BE449162E74AFAAEB5DD2F4875C9" bold="true" box="[199,382,726,751]" pageId="6" pageNumber="593">
<typeStatus id="5238DCF49162E74AFAAEB5DD2F3475C9" box="[199,258,726,751]" pageId="6" pageNumber="593">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5238DCF49162E74AFBE7B5DD2FCD75C9" box="[398,507,726,751]" pageId="6" pageNumber="593" type="holotype">Holotype</typeStatus>
: ♂ (
<collectionCode id="EB92FA939162E74AF855B5DC2CBA75C9" box="[572,652,727,751]" pageId="6" pageNumber="593">QSBG</collectionCode>
),
<collectingCountry id="F59422C69162E74AF8C1B5DC2D0B75C9" box="[680,829,727,751]" name="Thailand" pageId="6" pageNumber="593">THAILAND</collectingCountry>
: Chiang Mai Doi Phahompok NP Doi phaluang,
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,
<quantity id="4A7BCFB39162E74AFBF4B5F02FC27434" box="[413,500,763,787]" metricMagnitude="3" metricUnit="m" metricValue="1.449" pageId="6" pageNumber="593" unit="m" value="1449.0">1449 m</quantity>
, Malaise trap,
<date id="F93D44969162E74AF8F3B5F02D667435" box="[666,848,763,787]" pageId="6" pageNumber="593" value="2007-07-20" valueMax="2007-07-27" valueMin="2007-07-20">2027.VII.2007</date>
, Wongchai P.
<typeStatus id="5238DCF49162E74AF987B5F02A637435" box="[1006,1109,763,787]" pageId="6" pageNumber="593" type="paratype">Paratype</typeStatus>
: 1♀ (
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), same data as
<typeStatus id="5238DCF49162E74AFAFEB4152ECB7411" box="[151,253,798,823]" pageId="6" pageNumber="593" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C59931DD9162E74AFAAEB4492A457485" pageId="6" pageNumber="593" type="discussion">
<paragraph id="8D3C62569162E74AFAAEB4492A457485" blockId="6.[151,1437,258,1003]" pageId="6" pageNumber="593">
<emphasis id="BFF7BE449162E74AFAAEB4492F0D747D" bold="true" box="[199,315,834,859]" pageId="6" pageNumber="593">Remarks.</emphasis>
This species is easy distinguished from other species of
<emphasis id="BFF7BE449162E74AF9D7B4482A0A747C" box="[958,1084,835,858]" italics="true" pageId="6" pageNumber="593">Angustuma</emphasis>
<bibRefCitation id="E9121FA79162E74AFE2DB4492B34747D" author="Xing" box="[1092,1282,834,859]" pageId="6" pageNumber="593" refString="Xing, J. C. &amp; Li, Z. Z. (2013) New replacement name for Angustella Li, 1986 (Hemiptera: Cicadellidae: Evacanthinae: Evacanthini), with description of a new species. Zootaxa, 3702 (4), 386 - 390." type="journal article" year="2013">Xing &amp; Li, 2013</bibRefCitation>
by the dorsal coloration and marks. Its male genitalia are very similar to those of
<taxonomicName id="4A8319D59162E74AF9DDB46C2B0B7459" authority="Li, 1986" authorityName="Li" authorityYear="1986" box="[948,1341,870,895]" class="Insecta" family="Cicadellidae" genus="Angustuma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="nigricarina">
<emphasis id="BFF7BE449162E74AF9DDB46C2A8A7458" box="[948,1212,871,894]" italics="true" pageId="6" pageNumber="593">Angustuma nigricarina</emphasis>
(
<bibRefCitation id="E9121FA79162E74AFEA7B46D2B037459" author="Li" box="[1230,1333,870,895]" pageId="6" pageNumber="593" refString="Li, Z. Z. (1986) On a new genus and a new species of Evacanthidae from Guizhou Province, China (Homoptera: Cicadelloidea). Acta Zootaxonomica Sinica, 11 (4), 309 - 311." type="journal article" year="1986">Li, 1986</bibRefCitation>
)
</taxonomicName>
, but the shapes of the aedeagal shaft and ventral apophysis (
<figureCitation id="15B87ED39162E74AF8B3B4812D1D7485" box="[730,811,906,931]" captionStart="FIGURES 21 31" captionStartId="5.[151,264,1913,1936]" captionTargetBox="[178,1396,198,1855]" captionTargetId="figure@5.[162,1406,193,1876]" captionTargetPageId="5" captionText="FIGURES 21 31. Angustuma flavimacula sp. nov.: 21. male habitus, dorsal view; 22. male habitus, lateral view; 23. male head and thorax, dorsal view; 24. male face; 25. female habitus, dorsal view; 26. male pygofer side, lateral view; 27. aedeagal, lateral view; 28. aedeagal, ventral view; 29. connective and style, ventral view; 30. subgenital plate, ventral view; 31. apex of female abdomen, ventral view." httpUri="https://zenodo.org/record/227641/files/figure.png" pageId="6" pageNumber="593">Fig. 28</figureCitation>
) in lateral view are different.
</paragraph>
</subSubSection>
<subSubSection id="C59931DD9162E74AFAAEB4A52D9B74CD" pageId="6" pageNumber="593" type="etymology">
<paragraph id="8D3C62569162E74AFAAEB4A52D9B74CD" blockId="6.[151,1437,258,1003]" pageId="6" pageNumber="593">
<emphasis id="BFF7BE449162E74AFAAEB4A52F7874E1" bold="true" box="[199,334,942,967]" pageId="6" pageNumber="593">Etymology.</emphasis>
The specific epithet, a combination of “flavus”, meaning yellow, and “macula”, meaning spot, refers to the forewing having a large yellow spot on the lateral margin.
</paragraph>
</subSubSection>
</treatment>
</document>