545 lines
78 KiB
XML
545 lines
78 KiB
XML
<document id="E86EAC02D7131BCB5E0279B829903F45" ID-DOI="10.5281/zenodo.204752" ID-GBIF-Dataset="8587cb2c-0974-472f-a27e-f0f062d95643" ID-ISSN="1175-5326" ID-Zenodo-Dep="204752" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460294193991" checkinUser="plazi" docAuthor="Fontoura, Paulo, Pilato, Giovanni & Lisi, Oscar" docDate="2011" docId="051C87C3B611FFB5FF482A21FDDCFC7F" docLanguage="en" docName="zt02838p040.pdf" docOrigin="Zootaxa 2838" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Echiniscus clavispinosus Fontoura, Pilato & Lisi, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="F925FFBBB610FFB3FFDF2F2BFFC9FFC8" masterDocTitle="Tardigrada from Santo Antão Island (Archipelago of Cape Verde, West Africa) with the description of a new species" masterLastPageNumber="40" masterPageNumber="30" pageNumber="31" updateTime="1698251923419" updateUser="plazi">
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<mods:mods id="C79085DAD0D5CAB6203B8E431216CFF3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8B82BBEA76E49A6BAA1A7F056FD353B5">
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<mods:title id="D06D5D819487DECC6D1040D4050074BC">Tardigrada from Santo Antão Island (Archipelago of Cape Verde, West Africa) with the description of a new species</mods:title>
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<mods:name id="D7465DEE832E939FEB50B8D3092F1B92" type="personal">
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<mods:namePart id="9C4A3A93DC8670ACC01E7A94CC7BEA6B">Fontoura, Paulo</mods:namePart>
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<mods:roleTerm id="2D9CBC48FC1AD3BEAF7DB10654E4B4D3">Author</mods:roleTerm>
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<mods:namePart id="AD61547E03A2EE3BF149F84DE42B0BD2">Pilato, Giovanni</mods:namePart>
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<mods:name id="732FCFFE0585FF42A94C30BE16B9C73A" type="personal">
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<mods:roleTerm id="48A1E7FB7B558BC93849F2C155F78E80">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C1C3AB47FA878C973A443902AB66D497">Lisi, Oscar</mods:namePart>
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<mods:typeOfResource id="7D6652AC70A872098ACC7C10D0A3F009">text</mods:typeOfResource>
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<mods:title id="895C4A0A08598D7E5EDAA066AD95554E">Zootaxa</mods:title>
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<mods:part id="1C834C0CF9C4FE237C7A0A380EBF7625">
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<mods:date id="532A81115E77F788393EDBC1FD8588BE">2011</mods:date>
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<mods:detail id="2EC512E2DB4DC165DAF12DB7ABC02FF9" type="volume">
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<mods:number id="5E110612515D51B08CABD509A98049FB">2838</mods:number>
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<mods:classification id="10D92EE5FA9E444E9FE61F9005961993">journal article</mods:classification>
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<mods:identifier id="A24384440ED4A064B4264B4E9A380005" type="DOI">10.5281/zenodo.204752</mods:identifier>
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<mods:identifier id="C4A51F63E062D70959D722435B92B182" type="GBIF-Dataset">8587cb2c-0974-472f-a27e-f0f062d95643</mods:identifier>
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<mods:identifier id="A58D9B5718D9D5714A15A586B8CDF4EA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4BD4AF586A9B1F404D014AE1313BBE7F" type="Zenodo-Dep">204752</mods:identifier>
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<treatment id="051C87C3B611FFB5FF482A21FDDCFC7F" ID-DOI="http://doi.org/10.5281/zenodo.5677065" ID-GBIF-Taxon="119399071" ID-Zenodo-Dep="5677065" LSID="urn:lsid:zoobank.org:act:3D83C255-B4CF-4A3A-8D7A-965673839016" httpUri="http://treatment.plazi.org/id/051C87C3B611FFB5FF482A21FDDCFC7F" lastPageId="6" lastPageNumber="36" pageId="1" pageNumber="31">
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<subSubSection id="C5AF655EB611FFB2FF482A21FD4DFA7A" pageId="1" pageNumber="31" type="nomenclature">
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<paragraph id="8D0A36D5B611FFB2FF482A21FDD1FAEA" blockId="1.[151,536,1290,1350]" box="[151,536,1290,1315]" pageId="1" pageNumber="31">
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<heading id="D64281B9B611FFB2FF482A21FDD1FAEA" bold="true" box="[151,536,1290,1315]" fontSize="10" level="2" pageId="1" pageNumber="31" reason="2">
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<emphasis id="BFC1EAC7B611FFB2FF482A21FDD1FAEA" bold="true" box="[151,536,1290,1315]" pageId="1" pageNumber="31">
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<taxonomicName id="4AB54D56B611FFB2FF482A21FE7EFAEB" box="[151,439,1290,1315]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="1" pageNumber="31" phylum="Tardigrada" rank="species" species="clavispinosus" status="sp. nov.">
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<emphasis id="BFC1EAC7B611FFB2FF482A21FE7EFAEB" bold="true" box="[151,439,1290,1315]" italics="true" pageId="1" pageNumber="31">Echiniscus clavispinosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A4F257BCB611FFB2FE612A20FDD1FAEA" box="[446,536,1291,1315]" pageId="1" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8D0A36D5B611FFB2FF482A05FEBAFA8E" blockId="1.[151,536,1290,1350]" box="[151,371,1326,1350]" pageId="1" pageNumber="31">
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(
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<figureCitation id="158E2A50B611FFB2FF402A05FF24FA8E" box="[159,237,1326,1350]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="1" pageNumber="31">Figs. 1</figureCitation>
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,
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<figureCitation id="158E2A50B611FFB2FF252A05FEC2FA8E" box="[250,267,1326,1350]" captionStart="FIGURE 2" captionStartId="3.[151,250,1463,1486]" captionTargetBox="[153,1429,195,1462]" captionTargetId="figure@3.[151,1436,163,1468]" captionTargetPageId="3" captionText="FIGURE 2. Echiniscus clavispinosus sp. nov. A, anterior portion of the body; arrow a: indicates the pointed clava; arrow b: indicates a small spine on the first pair of legs. B, anterior portion of the body; arrow indicates the pointed clava. C, Posterior portion of the body; arrow a: indicates the papilla on the hind legs; arrow b: indicates the spur of one of the internal claws. D, anterior portion of a two-clawed larva; it is evident that the plate ornamentation lacks dark tubercles. Arrow indicates the pointed clava. Scale bars = 10 µm. A, C, D: PCM images; B: DIC image." httpUri="https://zenodo.org/record/204754/files/figure.png" pageId="1" pageNumber="31">2</figureCitation>
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;
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<tableCitation id="C037036EB611FFB2FEC82A05FEA1FA8E" box="[279,360,1326,1350]" captionStart="TABLE 1" captionStartId="4.[151,235,506,527]" captionTargetBox="[151,1436,627,1187]" captionTargetPageId="4" captionText="TABLE 1. Measurements of the holotype, the smallest and the largest specimens of Echiniscus clavispinosus sp. nov. (% bo = percentage of body length; % sc = percentage of scapular plate length)." httpUri="http://table.plazi.org/id/D9CA665DB614FFB7FF482ED1FCA1FDE7" pageId="1" pageNumber="31" tableUuid="D9CA665DB614FFB7FF482ED1FCA1FDE7">Table 1</tableCitation>
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)
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</paragraph>
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<paragraph id="8D0A36D5B611FFB2FF482A5CFD4DFA7A" blockId="1.[151,1437,1398,2034]" pageId="1" pageNumber="31">
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<emphasis id="BFC1EAC7B611FFB2FF482A5CFE8BFA47" bold="true" box="[151,322,1398,1423]" pageId="1" pageNumber="31">Locus typicus.</emphasis>
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Xôxô, Ribeira da Torre, Municipality of Ribeira Grande, Santo Antão Island, Archipelago of
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<collectingCountry id="F5A27645B611FFB2FABD2A5DFF12FA7A" name="Cape Verde" pageId="1" pageNumber="31">Cape Verde</collectingCountry>
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, in a lichen sample growing on rocks.
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</paragraph>
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</subSubSection>
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<subSubSection id="C5AF655EB611FFB2FF1A2A95FE7EF8AA" pageId="1" pageNumber="31" type="materials_examined">
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<paragraph id="8D0A36D5B611FFB2FF1A2A95FB99FA1E" blockId="1.[151,1437,1398,2034]" box="[197,1104,1470,1495]" pageId="1" pageNumber="31">
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<emphasis id="BFC1EAC7B611FFB2FF1A2A95FE67FA1F" bold="true" box="[197,430,1470,1495]" pageId="1" pageNumber="31">Material examined.</emphasis>
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Thirteen specimens, of which four were two-clawed larvae.
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</paragraph>
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<paragraph id="8D0A36D5B611FFB2FF1A2AC8FC14F9AE" blockId="1.[151,1437,1398,2034]" pageId="1" pageNumber="31">
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<emphasis id="BFC1EAC7B611FFB2FF1A2AC8FE4FFA33" bold="true" box="[197,390,1506,1531]" pageId="1" pageNumber="31">
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<typeStatus id="520E8877B611FFB2FF1A2AC8FF37FA33" box="[197,254,1507,1531]" pageId="1" pageNumber="31">Type</typeStatus>
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repository.
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</emphasis>
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The
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<typeStatus id="520E8877B611FFB2FE1C2AC9FDECFA32" box="[451,549,1506,1530]" pageId="1" pageNumber="31" type="holotype">holotype</typeStatus>
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(slide N. 5432) and one two-clawed larva (slide N. 5433) are deposited in the collection of Binda & Pilato (Museum of the Department of Animal Biology “Marcello La Greca”, University of Catania,
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<collectingCountry id="F5A27645B611FFB2FF232901FEFDF98A" box="[252,308,1578,1602]" name="Italy" pageId="1" pageNumber="31">Italy</collectingCountry>
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). The other
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<typeStatus id="520E8877B611FFB2FE1E2900FDE5F98B" box="[449,556,1579,1603]" pageId="1" pageNumber="31" type="paratype">paratypes</typeStatus>
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(slides CIV88 - CIV96) are deposited in the collection of P. Fontoura at the Department of Biology, Faculty of Sciences, University of Porto,
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<collectingCountry id="F5A27645B611FFB2FCA92965FC10F9AE" box="[886,985,1614,1638]" name="Portugal" pageId="1" pageNumber="31">Portugal</collectingCountry>
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.
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</paragraph>
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<paragraph id="8D0A36D5B611FFB2FF1A2959FE7EF8AA" blockId="1.[151,1437,1398,2034]" pageId="1" pageNumber="31">
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<emphasis id="BFC1EAC7B611FFB2FF1A2959FE56F943" bold="true" box="[197,415,1650,1675]" pageId="1" pageNumber="31">Specific diagnosis.</emphasis>
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Colour green; eye spots absent; dorsal plates well marked. Median plate 3 absent; cuticular ornamentation consisting of slightly raised dark tubercle and fine dots. At a higher focal position very dense, fine dots and small light spots appear. Distance between the dark tubercles shorter than the diameter of the tubercles. Anterior portion of median plates 1 and 2 and area between paired plate III and terminal plate unsculptured. Buccal cirri and cirrus
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<emphasis id="BFC1EAC7B611FFB2FE9A2828FE9FF8D3" box="[325,342,1795,1819]" italics="true" pageId="1" pageNumber="31">A</emphasis>
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short, elongated clavae with pointed apices; no other lateral or dorsal appendages present. Spine present on the first pair of legs; papilla and dentate collar on the hind legs; claws well developed; internal claws with well developed spur.
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</paragraph>
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</subSubSection>
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<subSubSection id="C5AF655EB611FFB7FF1A2845FADBFDA8" lastPageId="4" lastPageNumber="34" pageId="1" pageNumber="31" type="description">
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<paragraph id="8D0A36D5B611FFB1FF1A2845FEF0FD97" blockId="1.[151,1437,1398,2034]" lastBlockId="2.[151,1437,151,607]" lastPageId="2" lastPageNumber="32" pageId="1" pageNumber="31">
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<emphasis id="BFC1EAC7B611FFB2FF1A2845FDD9F84F" bold="true" box="[197,528,1902,1927]" pageId="1" pageNumber="31">
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Description of the
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<typeStatus id="520E8877B611FFB2FE7D2844FDC2F84F" box="[418,523,1903,1927]" pageId="1" pageNumber="31" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Body length 220 µm; plate colour green in specimens observed with transmitted light, after mounting on a slide, but reddish brown when observed with reflected light; eye spots not observed in either living nor slide mounted specimens. A narrow finely dotted transverse band was present anterior to the cephalic plate (
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<figureCitation id="158E2A50B611FFB2FE9F28F1FEB6F83A" box="[320,383,2010,2034]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="1" pageNumber="31">Fig. 1</figureCitation>
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A, D); this plate with an anterior notch; a true neck plate was absent but there was a thickened and faint ornamented transverse area between the cephalic plate and the scapular plate (
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<figureCitation id="158E2A50B612FFB1FBA02FBCFB76FF67" box="[1151,1215,151,175]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="2" pageNumber="32">Fig. 1</figureCitation>
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A, D); the scapular plate, 43.6 µm long in the medium line, was subdivided into a wide central portion and two small lateral, almost triangular, portions (
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<figureCitation id="158E2A50B612FFB1FEBB2FF4FE6DFF3F" box="[356,420,223,247]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="2" pageNumber="32">Fig. 1</figureCitation>
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A, D). Median plate 2 with unsculptured anterior portion (
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<figureCitation id="158E2A50B612FFB1FBE72FF4FBB1FF3F" box="[1080,1144,223,247]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="2" pageNumber="32">Fig. 1</figureCitation>
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B); median plate 3 absent and the area between paired plate III and terminal plate unsculptured (
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<figureCitation id="158E2A50B612FFB1FC1D2E2FFBCAFED4" box="[962,1027,260,284]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="2" pageNumber="32">Fig. 1</figureCitation>
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B, C); terminal plate unfaceted and with two indentations (
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<figureCitation id="158E2A50B612FFB1FE432E0CFE14FEF7" box="[412,477,295,319]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="2" pageNumber="32">Fig. 1</figureCitation>
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B). Plate sculpture of the adults comprised of slightly raised dark tubercles and very fine, dense dots (
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<figureCitation id="158E2A50B612FFB1FE862E67FE54FEAC" box="[345,413,332,356]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="2" pageNumber="32">Fig. 1</figureCitation>
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A–D); at a higher focal position small light spots appear (
|
||
<figureCitation id="158E2A50B612FFB1FBE62E67FBB0FEAC" box="[1081,1145,332,356]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="2" pageNumber="32">Fig. 1</figureCitation>
|
||
D). Distance between the dark tubercles shorter than the diameter of the tubercles; largest tubercles (diameter
|
||
<emphasis id="BFC1EAC7B612FFB1FB872E5AFBA2FE40" box="[1112,1131,369,392]" italics="true" pageId="2" pageNumber="32">c.</emphasis>
|
||
2.3 µm) are present in the central portion of the scapular plate. Cephalic plate sculptured with only fine dots (
|
||
<figureCitation id="158E2A50B612FFB1FBE72EBFFBBEFE64" box="[1080,1143,404,428]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="2" pageNumber="32">Fig. 1</figureCitation>
|
||
D). In the anterior portion of the paired plates dark tubercles (
|
||
<figureCitation id="158E2A50B612FFB1FDFE2E9CFDA8FE07" box="[545,609,439,463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1883,1906]" captionTargetBox="[181,1420,654,1882]" captionTargetId="figure@2.[169,1429,625,1890]" captionTargetPageId="2" captionText="FIGURE 1. Echiniscus clavispinosus sp. nov. A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image." httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="2" pageNumber="32">Fig. 1</figureCitation>
|
||
B, C) were present (and a transverse area with almost invisible dots). Leg plates sculptured with only fine dots. Ventral body surface had an extremely fine granulation more visible in the cephalic region. Small spine (2.4 µm long) was present on the first pair of legs (
|
||
<figureCitation id="158E2A50B612FFB1FBCF2ED4FB9AFDDF" box="[1040,1107,511,535]" captionStart="FIGURE 2" captionStartId="3.[151,250,1463,1486]" captionTargetBox="[153,1429,195,1462]" captionTargetId="figure@3.[151,1436,163,1468]" captionTargetPageId="3" captionText="FIGURE 2. Echiniscus clavispinosus sp. nov. A, anterior portion of the body; arrow a: indicates the pointed clava; arrow b: indicates a small spine on the first pair of legs. B, anterior portion of the body; arrow indicates the pointed clava. C, Posterior portion of the body; arrow a: indicates the papilla on the hind legs; arrow b: indicates the spur of one of the internal claws. D, anterior portion of a two-clawed larva; it is evident that the plate ornamentation lacks dark tubercles. Arrow indicates the pointed clava. Scale bars = 10 µm. A, C, D: PCM images; B: DIC image." httpUri="https://zenodo.org/record/204754/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
|
||
A); small papilla 3.3 µm long and dentate collar with about 10 unequal, sharp teeth (the longest
|
||
<emphasis id="BFC1EAC7B612FFB1FCA72D0EFC42FDF4" box="[888,907,549,572]" italics="true" pageId="2" pageNumber="32">c.</emphasis>
|
||
2.2 µm long) were present in the fourth pair of legs (
|
||
<figureCitation id="158E2A50B612FFB1FF0C2D6CFEDEFD97" box="[211,279,583,607]" captionStart="FIGURE 2" captionStartId="3.[151,250,1463,1486]" captionTargetBox="[153,1429,195,1462]" captionTargetId="figure@3.[151,1436,163,1468]" captionTargetPageId="3" captionText="FIGURE 2. Echiniscus clavispinosus sp. nov. A, anterior portion of the body; arrow a: indicates the pointed clava; arrow b: indicates a small spine on the first pair of legs. B, anterior portion of the body; arrow indicates the pointed clava. C, Posterior portion of the body; arrow a: indicates the papilla on the hind legs; arrow b: indicates the spur of one of the internal claws. D, anterior portion of a two-clawed larva; it is evident that the plate ornamentation lacks dark tubercles. Arrow indicates the pointed clava. Scale bars = 10 µm. A, C, D: PCM images; B: DIC image." httpUri="https://zenodo.org/record/204754/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
<caption id="D9CA665DB612FFB1FF482870FB9EF825" httpUri="https://zenodo.org/record/204753/files/figure.png" pageId="2" pageNumber="32" targetBox="[181,1420,654,1882]" targetPageId="2">
|
||
<paragraph id="8D0A36D5B612FFB1FF482870FB9EF825" blockId="2.[151,1436,1883,2029]" pageId="2" pageNumber="32">
|
||
<emphasis id="BFC1EAC7B612FFB1FF482870FEDBF8B9" bold="true" box="[151,274,1883,1906]" pageId="2" pageNumber="32">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4AB54D56B612FFB1FEC72877FDDDF8BA" box="[280,532,1884,1906]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="2" pageNumber="32" phylum="Tardigrada" rank="species" species="clavispinosus" status="sp. nov.">
|
||
<emphasis id="BFC1EAC7B612FFB1FEC72877FDDDF8BA" box="[280,532,1884,1906]" italics="true" pageId="2" pageNumber="32">Echiniscus clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BFC1EAC7B612FFB1FDC42877FDA2F8B9" bold="true" box="[539,619,1884,1905]" pageId="2" pageNumber="32">
|
||
<taxonomicNameLabel id="A4F257BCB612FFB1FDC42877FDA2F8B9" box="[539,619,1884,1905]" pageId="2" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, anterior portion of the body. The arrow indicates the sculptured band anterior to the cephalic plate. B, posterior portion of the body. The arrow indicates that median plate 3 is absent and the corresponding area lacks sculpture. C, details of the plate sculpture; arrows a: indicate dark tubercles; arrow b: indicates small dots. D, anterior portion of the body; it is evident that cephalic plate lacks dark tubercles. Arrow a: indicates a dark tubercle; arrow b: small dots; arrows c: indicate small light spots. Scale bars = 10 µm. A, B, D: PCM images; C: DIC image.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D9CA665DB613FFB0FF482A9CFCB6F980" httpUri="https://zenodo.org/record/204754/files/figure.png" pageId="3" pageNumber="33" targetBox="[153,1429,195,1462]" targetPageId="3">
|
||
<paragraph id="8D0A36D5B613FFB0FF482A9CFCB6F980" blockId="3.[151,1436,1463,1608]" pageId="3" pageNumber="33">
|
||
<emphasis id="BFC1EAC7B613FFB0FF482A9CFEDDFA05" bold="true" box="[151,276,1463,1486]" pageId="3" pageNumber="33">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4AB54D56B613FFB0FEC32A9CFDD0FA05" box="[284,537,1463,1485]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="33" phylum="Tardigrada" rank="species" species="clavispinosus" status="sp. nov.">
|
||
<emphasis id="BFC1EAC7B613FFB0FEC32A9CFDD0FA05" box="[284,537,1463,1485]" italics="true" pageId="3" pageNumber="33">Echiniscus clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A4F257BCB613FFB0FDFE2A93FDBAFA05" box="[545,627,1464,1485]" pageId="3" pageNumber="33" rank="species">
|
||
<emphasis id="BFC1EAC7B613FFB0FDFE2A93FDBAFA05" bold="true" box="[545,627,1464,1485]" pageId="3" pageNumber="33">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A, anterior portion of the body; arrow a: indicates the pointed clava; arrow b: indicates a small spine on the first pair of legs. B, anterior portion of the body; arrow indicates the pointed clava. C, Posterior portion of the body; arrow a: indicates the papilla on the hind legs; arrow b: indicates the spur of one of the internal claws. D, anterior portion of a two-clawed larva; it is evident that the plate ornamentation lacks dark tubercles. Arrow indicates the pointed clava. Scale bars = 10 µm. A, C, D: PCM images; B: DIC image.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8D0A36D5B613FFB0FF182944FB13F907" blockId="3.[151,1436,1647,2031]" pageId="3" pageNumber="33">
|
||
Internal buccal cirri 9.6 µm long; external cephalic cirri slightly longer (11.9 µm); cephalic papillae 6.8 µm long (
|
||
<figureCitation id="158E2A50B613FFB0FF0529B8FED6F963" box="[218,287,1683,1707]" captionStart="FIGURE 2" captionStartId="3.[151,250,1463,1486]" captionTargetBox="[153,1429,195,1462]" captionTargetId="figure@3.[151,1436,163,1468]" captionTargetPageId="3" captionText="FIGURE 2. Echiniscus clavispinosus sp. nov. A, anterior portion of the body; arrow a: indicates the pointed clava; arrow b: indicates a small spine on the first pair of legs. B, anterior portion of the body; arrow indicates the pointed clava. C, Posterior portion of the body; arrow a: indicates the papilla on the hind legs; arrow b: indicates the spur of one of the internal claws. D, anterior portion of a two-clawed larva; it is evident that the plate ornamentation lacks dark tubercles. Arrow indicates the pointed clava. Scale bars = 10 µm. A, C, D: PCM images; B: DIC image." httpUri="https://zenodo.org/record/204754/files/figure.png" pageId="3" pageNumber="33">Fig. 2</figureCitation>
|
||
A); clavae, 7.2 µm long, had a distinctive sharp-tipped, conical shape rather than the rounded apices (
|
||
<figureCitation id="158E2A50B613FFB0FF40299CFF2AF907" box="[159,227,1719,1743]" captionStart="FIGURE 2" captionStartId="3.[151,250,1463,1486]" captionTargetBox="[153,1429,195,1462]" captionTargetId="figure@3.[151,1436,163,1468]" captionTargetPageId="3" captionText="FIGURE 2. Echiniscus clavispinosus sp. nov. A, anterior portion of the body; arrow a: indicates the pointed clava; arrow b: indicates a small spine on the first pair of legs. B, anterior portion of the body; arrow indicates the pointed clava. C, Posterior portion of the body; arrow a: indicates the papilla on the hind legs; arrow b: indicates the spur of one of the internal claws. D, anterior portion of a two-clawed larva; it is evident that the plate ornamentation lacks dark tubercles. Arrow indicates the pointed clava. Scale bars = 10 µm. A, C, D: PCM images; B: DIC image." httpUri="https://zenodo.org/record/204754/files/figure.png" pageId="3" pageNumber="33">Fig. 2</figureCitation>
|
||
B). Cirri
|
||
<emphasis id="BFC1EAC7B613FFB0FE98299CFE91F907" box="[327,344,1719,1743]" italics="true" pageId="3" pageNumber="33">A</emphasis>
|
||
, a short filament 26.0 µm long (
|
||
<figureCitation id="158E2A50B613FFB0FD1C299CFCCCF907" box="[707,773,1719,1743]" captionStart="FIGURE 2" captionStartId="3.[151,250,1463,1486]" captionTargetBox="[153,1429,195,1462]" captionTargetId="figure@3.[151,1436,163,1468]" captionTargetPageId="3" captionText="FIGURE 2. Echiniscus clavispinosus sp. nov. A, anterior portion of the body; arrow a: indicates the pointed clava; arrow b: indicates a small spine on the first pair of legs. B, anterior portion of the body; arrow indicates the pointed clava. C, Posterior portion of the body; arrow a: indicates the papilla on the hind legs; arrow b: indicates the spur of one of the internal claws. D, anterior portion of a two-clawed larva; it is evident that the plate ornamentation lacks dark tubercles. Arrow indicates the pointed clava. Scale bars = 10 µm. A, C, D: PCM images; B: DIC image." httpUri="https://zenodo.org/record/204754/files/figure.png" pageId="3" pageNumber="33">Fig. 2</figureCitation>
|
||
A, B); no other trunk appendages present.
|
||
</paragraph>
|
||
<paragraph id="8D0A36D5B613FFB0FF1829F0FCEAF8F3" blockId="3.[151,1436,1647,2031]" pageId="3" pageNumber="33">
|
||
Claws well developed (
|
||
<figureCitation id="158E2A50B613FFB0FE1029F0FDDBF93B" box="[463,530,1755,1779]" captionStart="FIGURE 2" captionStartId="3.[151,250,1463,1486]" captionTargetBox="[153,1429,195,1462]" captionTargetId="figure@3.[151,1436,163,1468]" captionTargetPageId="3" captionText="FIGURE 2. Echiniscus clavispinosus sp. nov. A, anterior portion of the body; arrow a: indicates the pointed clava; arrow b: indicates a small spine on the first pair of legs. B, anterior portion of the body; arrow indicates the pointed clava. C, Posterior portion of the body; arrow a: indicates the papilla on the hind legs; arrow b: indicates the spur of one of the internal claws. D, anterior portion of a two-clawed larva; it is evident that the plate ornamentation lacks dark tubercles. Arrow indicates the pointed clava. Scale bars = 10 µm. A, C, D: PCM images; B: DIC image." httpUri="https://zenodo.org/record/204754/files/figure.png" pageId="3" pageNumber="33">Fig. 2</figureCitation>
|
||
A, C); internal and external claws on the first pair of legs 13.3 and 11.8 µm long respectively; they were slightly longer on the fourth pair of legs (14.3 µm and 13.3 µm respectively). Internal claws with a well developed spur directed downwards (
|
||
<figureCitation id="158E2A50B613FFB0FD622808FCC9F8F3" box="[701,768,1827,1851]" captionStart="FIGURE 2" captionStartId="3.[151,250,1463,1486]" captionTargetBox="[153,1429,195,1462]" captionTargetId="figure@3.[151,1436,163,1468]" captionTargetPageId="3" captionText="FIGURE 2. Echiniscus clavispinosus sp. nov. A, anterior portion of the body; arrow a: indicates the pointed clava; arrow b: indicates a small spine on the first pair of legs. B, anterior portion of the body; arrow indicates the pointed clava. C, Posterior portion of the body; arrow a: indicates the papilla on the hind legs; arrow b: indicates the spur of one of the internal claws. D, anterior portion of a two-clawed larva; it is evident that the plate ornamentation lacks dark tubercles. Arrow indicates the pointed clava. Scale bars = 10 µm. A, C, D: PCM images; B: DIC image." httpUri="https://zenodo.org/record/204754/files/figure.png" pageId="3" pageNumber="33">Fig. 2</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
<paragraph id="8D0A36D5B613FFB0FF1A2863FEBDF897" blockId="3.[151,1436,1647,2031]" box="[197,372,1863,1888]" pageId="3" pageNumber="33">Eggs unknown.</paragraph>
|
||
<paragraph id="8D0A36D5B613FFB0FF1A2840FAB0F803" blockId="3.[151,1436,1647,2031]" pageId="3" pageNumber="33">
|
||
<emphasis id="BFC1EAC7B613FFB0FF1A2840FE82F84C" bold="true" box="[197,331,1899,1924]" pageId="3" pageNumber="33">Variability.</emphasis>
|
||
The
|
||
<typeStatus id="520E8877B613FFB0FE5C2847FE27F84C" box="[387,494,1900,1924]" pageId="3" pageNumber="33" type="paratype">paratypes</typeStatus>
|
||
(body length 133–241 µm) were similar to the
|
||
<typeStatus id="520E8877B613FFB0FC202840FBA8F84B" box="[1023,1121,1899,1923]" pageId="3" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
in both qualitative and metric characters (
|
||
<tableCitation id="C037036EB613FFB0FEE428A4FE47F86F" box="[315,398,1935,1959]" captionStart="TABLE 1" captionStartId="4.[151,235,506,527]" captionTargetBox="[151,1436,627,1187]" captionTargetPageId="4" captionText="TABLE 1. Measurements of the holotype, the smallest and the largest specimens of Echiniscus clavispinosus sp. nov. (% bo = percentage of body length; % sc = percentage of scapular plate length)." httpUri="http://table.plazi.org/id/D9CA665DB614FFB7FF482ED1FCA1FDE7" pageId="3" pageNumber="33" tableUuid="D9CA665DB614FFB7FF482ED1FCA1FDE7">Table 1</tableCitation>
|
||
). However, we would like to emphasise the curiosity found in two of the
|
||
<typeStatus id="520E8877B613FFB0FB1A28BBFAFDF860" box="[1221,1332,1936,1960]" pageId="3" pageNumber="33" type="paratype">paratypes</typeStatus>
|
||
. One had a deeply forked buccal cirrus, and the other a deeply forked cirrusi
|
||
<emphasis id="BFC1EAC7B613FFB0FC59289FFC5EF804" box="[902,919,1972,1996]" italics="true" pageId="3" pageNumber="33">A</emphasis>
|
||
. The cause of this anomaly was not found.
|
||
</paragraph>
|
||
<paragraph id="8D0A36D5B613FFB7FF1A28F3FEC6FE07" blockId="3.[151,1436,1647,2031]" lastBlockId="4.[151,1437,151,463]" lastPageId="4" lastPageNumber="34" pageId="3" pageNumber="33">
|
||
We consider it important to stress the differences between the two-clawed larvae (body lengths
|
||
<emphasis id="BFC1EAC7B613FFB0FB3528F3FB37F827" box="[1258,1278,2008,2031]" italics="true" pageId="3" pageNumber="33">c.</emphasis>
|
||
142–144 µm) and the adults. In the larvae the cuticular ornamentation lacked the dark tubercles and was instead formed by small, dense dots, sometimes joined together, and by sparse light spots (
|
||
<figureCitation id="158E2A50B614FFB7FC5C2F97FC01FF1C" box="[899,968,188,212]" captionStart="FIGURE 2" captionStartId="3.[151,250,1463,1486]" captionTargetBox="[153,1429,195,1462]" captionTargetId="figure@3.[151,1436,163,1468]" captionTargetPageId="3" captionText="FIGURE 2. Echiniscus clavispinosus sp. nov. A, anterior portion of the body; arrow a: indicates the pointed clava; arrow b: indicates a small spine on the first pair of legs. B, anterior portion of the body; arrow indicates the pointed clava. C, Posterior portion of the body; arrow a: indicates the papilla on the hind legs; arrow b: indicates the spur of one of the internal claws. D, anterior portion of a two-clawed larva; it is evident that the plate ornamentation lacks dark tubercles. Arrow indicates the pointed clava. Scale bars = 10 µm. A, C, D: PCM images; B: DIC image." httpUri="https://zenodo.org/record/204754/files/figure.png" pageId="4" pageNumber="34">Fig. 2</figureCitation>
|
||
D); these light spots, more evident in the posterior margins of the scapular, median and paired plates, were larger than those observed on the adults. We were sure that the two-clawed larvae belong to
|
||
<taxonomicName id="4AB54D56B614FFB7FDAF2E2FFC41FED4" box="[624,904,260,284]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="clavispinosus" status="sp. nov.">
|
||
<emphasis id="BFC1EAC7B614FFB7FDAF2E2FFC41FED4" box="[624,904,260,284]" italics="true" pageId="4" pageNumber="34">Echiniscus clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BFC1EAC7B614FFB7FC4F2E2FFC22FED3" bold="true" box="[912,1003,260,284]" pageId="4" pageNumber="34">
|
||
<taxonomicNameLabel id="A4F257BCB614FFB7FC4F2E2FFC22FED3" box="[912,1003,260,284]" pageId="4" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
due to the distinctive (pointed) clavae shape and because they have the smooth unornamented anterior portion of the median plate 2 and the unornamented area between the paired plates III and the terminal plate. The differences of the cuticular ornamentation seemed to be in relation to the developmental stage and not body length because the four two-clawed larvae were slightly larger (142–144 µm long) than a four-clawed
|
||
<typeStatus id="520E8877B614FFB7FCDC2EBEFCAAFE65" box="[771,867,405,429]" pageId="4" pageNumber="34" type="paratype">paratype</typeStatus>
|
||
(133 µm) which had typical adult cuticular ornamentation.
|
||
</paragraph>
|
||
<caption id="D9CA665DB614FFB7FF482ED1FCA1FDE7" ID-Table-UUID="D9CA665DB614FFB7FF482ED1FCA1FDE7" httpUri="http://table.plazi.org/id/D9CA665DB614FFB7FF482ED1FCA1FDE7" pageId="4" pageNumber="34" targetBox="[151,1436,627,1187]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="8D0A36D5B614FFB7FF482ED1FCA1FDE7" blockId="4.[151,1436,506,608]" pageId="4" pageNumber="34">
|
||
<emphasis id="BFC1EAC7B614FFB7FF482ED1FECDFDD8" bold="true" box="[151,260,506,528]" pageId="4" pageNumber="34">TABLE 1.</emphasis>
|
||
Measurements of the holotype, the smallest and the largest specimens of
|
||
<taxonomicName id="4AB54D56B614FFB7FC2F2ED1FB25FDD8" box="[1008,1260,506,528]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="clavispinosus" status="sp. nov.">
|
||
<emphasis id="BFC1EAC7B614FFB7FC2F2ED1FB25FDD8" box="[1008,1260,506,528]" italics="true" pageId="4" pageNumber="34">Echiniscus clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A4F257BCB614FFB7FB2C2ED0FA8AFDD8" box="[1267,1347,507,528]" pageId="4" pageNumber="34" rank="species">
|
||
<emphasis id="BFC1EAC7B614FFB7FB2C2ED0FAF5FDD8" bold="true" box="[1267,1340,507,528]" pageId="4" pageNumber="34">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="BFC1EAC7B614FFB7FA8D2ED1FA41FDD8" box="[1362,1416,506,528]" italics="true" pageId="4" pageNumber="34">% bo</emphasis>
|
||
= percentage of body length;
|
||
<emphasis id="BFC1EAC7B614FFB7FE732D32FE15FDE7" box="[428,476,537,559]" italics="true" pageId="4" pageNumber="34">% sc</emphasis>
|
||
= percentage of scapular plate length).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8D0A36D5B614FFB7FF432D61FADBFDA8" blockId="4.[151,1436,506,608]" box="[156,1298,586,608]" pageId="4" pageNumber="34">
|
||
Characters Smallest specimen
|
||
<typeStatus id="520E8877B614FFB7FD292D61FC9DFDA8" box="[758,852,586,608]" pageId="4" pageNumber="34" type="holotype">Holotype</typeStatus>
|
||
Largest specimen
|
||
</paragraph>
|
||
</subSubSection>
|
||
<paragraph id="8D0A36D5B614FFB7FF432DB6FA8EFB6B" pageId="4" pageNumber="34">
|
||
<table id="FFB5C475B614004CFF482D58FA55FB6B" box="[151,1436,627,1187]" gridcols="9" gridrows="4" pageId="4" pageNumber="34">
|
||
<tr id="33853497B614004CFF482D58FA55FD14" box="[151,1436,627,732]" gridrow="0" pageId="4" pageNumber="34">
|
||
<th id="70545DEBB614004CFF482D58FEB2FD14" box="[151,379,627,732]" gridcol="0" gridrow="0" pageId="4" pageNumber="34">Body length Scapular plate length</th>
|
||
<th id="70545DEBB614004CFE6E2D58FE14FD14" box="[433,477,627,732]" gridcol="1" gridrow="0" pageId="4" pageNumber="34">µm 133 26.6</th>
|
||
<th id="70545DEBB614004CFDF82D58FD92FD14" box="[551,603,627,732]" gridcol="2" gridrow="0" pageId="4" pageNumber="34">% bo - 20.0</th>
|
||
<th id="70545DEBB614004CFD512D58FD76FD14" box="[654,703,627,732]" gridcol="3" gridrow="0" pageId="4" pageNumber="34">% sc - -</th>
|
||
<th id="70545DEBB614004CFD2A2D58FCEBFD14" box="[757,802,627,732]" gridcol="4" gridrow="0" pageId="4" pageNumber="34">µm 220 43.6</th>
|
||
<th id="70545DEBB614004CFCB22D58FC39FD14" box="[877,1008,627,732]" gridcol="5" gridrow="0" pageId="4" pageNumber="34">% bo % sc - - 19.8 -</th>
|
||
<th id="70545DEBB614004CFBBF2D58FB44FD14" box="[1120,1165,627,732]" gridcol="6" gridrow="0" pageId="4" pageNumber="34">µm 241 48.8</th>
|
||
<th id="70545DEBB614004CFB072D58FAC4FD14" box="[1240,1293,627,732]" gridcol="7" gridrow="0" pageId="4" pageNumber="34">% bo - 20.2</th>
|
||
<th id="70545DEBB614004CFAE02D58FA55FD14" box="[1343,1436,627,732]" gridcol="8" gridrow="0" pageId="4" pageNumber="34">% sc - -</th>
|
||
</tr>
|
||
<tr id="33853497B614004CFF482DDBFA55FC49" box="[151,1436,752,897]" gridrow="1" pageId="4" pageNumber="34">
|
||
<th id="70545DEBB614004CFF482DDBFEB2FC49" box="[151,379,752,897]" gridcol="0" gridrow="1" pageId="4" pageNumber="34">Internal buccal cirrus External buccal cirrus Cephalic papilla Clava</th>
|
||
<td id="70545DEBB614004CFE6E2DDBFE14FC49" box="[433,477,752,897]" gridcol="1" gridrow="1" pageId="4" pageNumber="34">7.1 8.6 4.5 4.8</td>
|
||
<td id="70545DEBB614004CFDF82DDBFD92FC49" box="[551,603,752,897]" gridcol="2" gridrow="1" pageId="4" pageNumber="34">5.3 6.5 3.4 3.6</td>
|
||
<td id="70545DEBB614004CFD512DDBFD76FC49" box="[654,703,752,897]" gridcol="3" gridrow="1" pageId="4" pageNumber="34">26.7 32.3 16.9 18.1</td>
|
||
<td id="70545DEBB614004CFD2A2DDBFCEBFC49" box="[757,802,752,897]" gridcol="4" gridrow="1" pageId="4" pageNumber="34">9.6 11.9 6.8 7.2</td>
|
||
<td id="70545DEBB614004CFCB22DDBFC39FC49" box="[877,1008,752,897]" gridcol="5" gridrow="1" pageId="4" pageNumber="34">4.4 22.0 5.4 27.3 3.1 15.6 3.3 16.5</td>
|
||
<td id="70545DEBB614004CFBBF2DDBFB44FC49" box="[1120,1165,752,897]" gridcol="6" gridrow="1" pageId="4" pageNumber="34">12.8 15.8 7.3 8.1</td>
|
||
<td id="70545DEBB614004CFB072DDBFAC4FC49" box="[1240,1293,752,897]" gridcol="7" gridrow="1" pageId="4" pageNumber="34">5.3 6.6 3.0 3.4</td>
|
||
<td id="70545DEBB614004CFAE02DDBFA55FC49" box="[1343,1436,752,897]" gridcol="8" gridrow="1" pageId="4" pageNumber="34">26.2 32.4 15.0 16.6</td>
|
||
</tr>
|
||
<tr id="33853497B614004CFF482CBEFA55FC1C" box="[151,1436,917,980]" gridrow="2" pageId="4" pageNumber="34">
|
||
<th id="70545DEBB614004CFF482CBEFEB2FC1C" box="[151,379,917,980]" gridcol="0" gridrow="2" pageId="4" pageNumber="34">
|
||
Cirrus
|
||
<emphasis id="BFC1EAC7B614FFB7FF3E2CBDFF39FC63" box="[225,240,918,939]" italics="true" pageId="4" pageNumber="34">A</emphasis>
|
||
Spine on leg I
|
||
</th>
|
||
<td id="70545DEBB614004CFE6E2CBEFE14FC1C" box="[433,477,917,980]" gridcol="1" gridrow="2" pageId="4" pageNumber="34">14.5?</td>
|
||
<td id="70545DEBB614004CFDF82CBEFD92FC1C" box="[551,603,917,980]" gridcol="2" gridrow="2" pageId="4" pageNumber="34">10.9 -</td>
|
||
<td id="70545DEBB614004CFD512CBEFD76FC1C" box="[654,703,917,980]" gridcol="3" gridrow="2" pageId="4" pageNumber="34">54.5 -</td>
|
||
<td id="70545DEBB614004CFD2A2CBEFCEBFC1C" box="[757,802,917,980]" gridcol="4" gridrow="2" pageId="4" pageNumber="34">26.0 2.4</td>
|
||
<td id="70545DEBB614004CFCB22CBEFC39FC1C" box="[877,1008,917,980]" gridcol="5" gridrow="2" pageId="4" pageNumber="34">11.8 59.6 1.1 5.5</td>
|
||
<td id="70545DEBB614004CFBBF2CBEFB44FC1C" box="[1120,1165,917,980]" gridcol="6" gridrow="2" pageId="4" pageNumber="34">32.4 3.4</td>
|
||
<td id="70545DEBB614004CFB072CBEFAC4FC1C" box="[1240,1293,917,980]" gridcol="7" gridrow="2" pageId="4" pageNumber="34">13.4 1.4</td>
|
||
<td id="70545DEBB614004CFAE02CBEFA55FC1C" box="[1343,1436,917,980]" gridcol="8" gridrow="2" pageId="4" pageNumber="34">66.4 7.0</td>
|
||
</tr>
|
||
<tr id="33853497B614004CFF482CC3FA55FB6B" box="[151,1436,1000,1187]" gridrow="3" pageId="4" pageNumber="34">
|
||
<th id="70545DEBB614004CFF482CC3FEB2FB6B" box="[151,379,1000,1187]" gridcol="0" gridrow="3" pageId="4" pageNumber="34">Internal claw leg I External claw leg I Internal claw leg IV External claw leg IV Papilla leg IV</th>
|
||
<td id="70545DEBB614004CFE6E2CC3FE14FB6B" box="[433,477,1000,1187]" gridcol="1" gridrow="3" pageId="4" pageNumber="34">9.2 8.1 9.3 8.1?</td>
|
||
<td id="70545DEBB614004CFDF82CC3FD92FB6B" box="[551,603,1000,1187]" gridcol="2" gridrow="3" pageId="4" pageNumber="34">6.9 6.1 7.0 6.1 -</td>
|
||
<td id="70545DEBB614004CFD512CC3FD76FB6B" box="[654,703,1000,1187]" gridcol="3" gridrow="3" pageId="4" pageNumber="34">34.6 30.5 35.0 30.5 -</td>
|
||
<td id="70545DEBB614004CFD2A2CC3FCEBFB6B" box="[757,802,1000,1187]" gridcol="4" gridrow="3" pageId="4" pageNumber="34">13.3 11.8 14.3 13.3 3.3</td>
|
||
<td id="70545DEBB614004CFCB22CC3FC39FB6B" box="[877,1008,1000,1187]" gridcol="5" gridrow="3" pageId="4" pageNumber="34">6.1 30.5 5.4 27.1 6.5 32.8 6.1 30.5 1.5 7.6</td>
|
||
<td id="70545DEBB614004CFBBF2CC3FB44FB6B" box="[1120,1165,1000,1187]" gridcol="6" gridrow="3" pageId="4" pageNumber="34">15.4 14.4 17.6 15.5?</td>
|
||
<td id="70545DEBB614004CFB072CC3FAC4FB6B" box="[1240,1293,1000,1187]" gridcol="7" gridrow="3" pageId="4" pageNumber="34">6.4 6.0 7.3 6.4 -</td>
|
||
<td id="70545DEBB614004CFAE02CC3FA55FB6B" box="[1343,1436,1000,1187]" gridcol="8" gridrow="3" pageId="4" pageNumber="34">31.6 29.5 36.1 31.8 -</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C5AF655EB614FFB5FF1A2BC3FDDCFC7F" lastPageId="6" lastPageNumber="36" pageId="4" pageNumber="34" type="etymology">
|
||
<paragraph id="8D0A36D5B614FFB7FF1A2BC3FB92FB37" blockId="4.[151,1436,1255,2006]" box="[197,1115,1255,1280]" pageId="4" pageNumber="34">
|
||
<emphasis id="BFC1EAC7B614FFB7FF1A2BC3FE85FAC8" bold="true" box="[197,332,1256,1280]" pageId="4" pageNumber="34">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="4AB54D56B614FFB7FE172BC3FD96FAC8" box="[456,607,1256,1280]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="clavispinosus">
|
||
<emphasis id="BFC1EAC7B614FFB7FE172BC3FD96FAC8" box="[456,607,1256,1280]" italics="true" pageId="4" pageNumber="34">clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
refers to the pointed apex shape of the clava.
|
||
</paragraph>
|
||
<paragraph id="8D0A36D5B614FFB7FF1A2A20FCA0F98F" blockId="4.[151,1436,1255,2006]" pageId="4" pageNumber="34">
|
||
<emphasis id="BFC1EAC7B614FFB7FF1A2A20FE02FAEC" bold="true" box="[197,459,1291,1316]" pageId="4" pageNumber="34">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4AB54D56B614FFB7FE0E2A27FD21FAEC" box="[465,744,1292,1316]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="clavispinosus" status="sp. nov.">
|
||
<emphasis id="BFC1EAC7B614FFB7FE0E2A27FD21FAEC" box="[465,744,1292,1316]" italics="true" pageId="4" pageNumber="34">Echiniscus clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A4F257BCB614FFB7FD302A27FC81FAEB" box="[751,840,1292,1316]" pageId="4" pageNumber="34" rank="species">
|
||
<emphasis id="BFC1EAC7B614FFB7FD302A27FC81FAEB" bold="true" box="[751,840,1292,1316]" pageId="4" pageNumber="34">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
belongs to the
|
||
<taxonomicName id="4AB54D56B614FFB7FC2D2A27FBF0FAEC" box="[1010,1081,1292,1316]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="viridis">
|
||
<emphasis id="BFC1EAC7B614FFB7FC2D2A27FBF0FAEC" box="[1010,1081,1292,1316]" italics="true" pageId="4" pageNumber="34">viridis</emphasis>
|
||
</taxonomicName>
|
||
group, a small group of species within the genus
|
||
<taxonomicName id="4AB54D56B614FFB7FE892A1BFE06FA80" box="[342,463,1328,1352]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="BFC1EAC7B614FFB7FE892A1BFE06FA80" box="[342,463,1328,1352]" italics="true" pageId="4" pageNumber="34">Echiniscus</emphasis>
|
||
</taxonomicName>
|
||
characterised by the green colour, by plate ornamentation comprised of dark tubercles, fine dots and light spots, by the absence of dorsal and lateral trunk appendages (cirrus
|
||
<emphasis id="BFC1EAC7B614FFB7FB832A7FFBA4FAA4" box="[1116,1133,1364,1388]" italics="true" pageId="4" pageNumber="34">A</emphasis>
|
||
excluded), and well developed claws. Traditionally, on the basis of the colour, five species were attributed to this group:
|
||
<taxonomicName id="4AB54D56B614FFB7FB122A52FAFCFA58" box="[1229,1333,1400,1424]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="viridis">
|
||
<emphasis id="BFC1EAC7B614FFB7FB122A52FAFCFA58" box="[1229,1333,1400,1424]" italics="true" pageId="4" pageNumber="34">E. viridis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4AB54D56B614FFB7FA9B2A52FF16FA7C" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="rufoviridis">
|
||
<emphasis id="BFC1EAC7B614FFB7FA9B2A52FF16FA7C" italics="true" pageId="4" pageNumber="34">E. rufoviridis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4AB54D56B614FFB7FF2D2AB7FE5BFA7C" box="[242,402,1436,1460]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="viridissimus">
|
||
<emphasis id="BFC1EAC7B614FFB7FF2D2AB7FE5BFA7C" box="[242,402,1436,1460]" italics="true" pageId="4" pageNumber="34">E.viridissimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4AB54D56B614FFB7FE792AB6FDF1FA7C" box="[422,568,1436,1460]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="perviridis">
|
||
<emphasis id="BFC1EAC7B614FFB7FE792AB6FDF1FA7C" box="[422,568,1436,1460]" italics="true" pageId="4" pageNumber="34">E. perviridis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4AB54D56B614FFB7FDA42AB6FCDBFA7C" box="[635,786,1436,1460]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="viridianus">
|
||
<emphasis id="BFC1EAC7B614FFB7FDA42AB6FCDBFA7C" box="[635,786,1436,1460]" italics="true" pageId="4" pageNumber="34">E. viridianus</emphasis>
|
||
</taxonomicName>
|
||
. However, two of these species,
|
||
<taxonomicName id="4AB54D56B614FFB7FB792AB6FA88FA7C" box="[1190,1345,1436,1460]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="rufoviridis">
|
||
<emphasis id="BFC1EAC7B614FFB7FB792AB6FA88FA7C" box="[1190,1345,1436,1460]" italics="true" pageId="4" pageNumber="34">E. rufoviridis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4AB54D56B614FFB7FA5B2AB6FED7FA13" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="viridissimus">
|
||
<emphasis id="BFC1EAC7B614FFB7FA5B2AB6FED7FA13" italics="true" pageId="4" pageNumber="34">E. viridissimus</emphasis>
|
||
</taxonomicName>
|
||
, have distinctly different cuticular ornamentation (
|
||
<figureCitation id="158E2A50B614FFB7FC842AE9FC54FA12" box="[859,925,1474,1498]" captionStart="FIGURE 3" captionStartId="5.[151,250,1692,1715]" captionTargetBox="[171,1417,393,1649]" captionTargetId="figure@5.[151,1436,373,1671]" captionTargetPageId="5" captionText="FIGURE 3. Details of the cuticular ornamentation (PCM images) in: A, Echinicus clavispinosus sp. nov.; B, Echiniscus viridis; C, Echiniscus viridianus; D, Echnisicus perviridis; E, Echiniscus rufoviridis and, F, Echinisicus viridissimus. The differences between the first three species on one hand and Echniscus rufoviridis and Echiniscus viridissimus on the other hand are very evident. Scale bar: 10 µm." httpUri="https://zenodo.org/record/204755/files/figure.png" pageId="4" pageNumber="34">Fig. 3</figureCitation>
|
||
) (Pilato
|
||
<emphasis id="BFC1EAC7B614FFB7FBDF2AEFFBF0FA13" box="[1024,1081,1475,1499]" italics="true" pageId="4" pageNumber="34">et al.</emphasis>
|
||
, 2007; 2008a) and should probably be excluded from the group. Nevertheless, to avoid misunderstandings and to make the species diagnosis easier, we incorporated all the green coloured
|
||
<taxonomicName id="4AB54D56B614FFB7FD742920FCEDF9EB" box="[683,804,1547,1571]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="BFC1EAC7B614FFB7FD742920FCEDF9EB" box="[683,804,1547,1571]" italics="true" pageId="4" pageNumber="34">Echiniscus</emphasis>
|
||
</taxonomicName>
|
||
species lacking dorsal and lateral trunk appendages (including
|
||
<taxonomicName id="4AB54D56B614FFB7FECE291BFE7EF98F" box="[273,439,1583,1607]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="viridissimus">
|
||
<emphasis id="BFC1EAC7B614FFB7FECE291BFE7EF98F" box="[273,439,1583,1607]" italics="true" pageId="4" pageNumber="34">E. viridissimus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4AB54D56B614FFB7FE31291BFD4AF98F" box="[494,643,1583,1607]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="rufoviridis">
|
||
<emphasis id="BFC1EAC7B614FFB7FE31291BFD4AF98F" box="[494,643,1583,1607]" italics="true" pageId="4" pageNumber="34">E. rufoviridis</emphasis>
|
||
</taxonomicName>
|
||
) into the key below.
|
||
</paragraph>
|
||
<paragraph id="8D0A36D5B614FFB7FF182978FC0CF8EA" blockId="4.[151,1436,1255,2006]" pageId="4" pageNumber="34">
|
||
<taxonomicName id="4AB54D56B614FFB7FF182978FE17F9A3" box="[199,478,1619,1643]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="clavispinosus" status="sp. nov.">
|
||
<emphasis id="BFC1EAC7B614FFB7FF182978FE17F9A3" box="[199,478,1619,1643]" italics="true" pageId="4" pageNumber="34">Echiniscus clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BFC1EAC7B614FFB7FE3A2978FDF6F9A2" bold="true" box="[485,575,1619,1643]" pageId="4" pageNumber="34">
|
||
<taxonomicNameLabel id="A4F257BCB614FFB7FE3A2978FDF6F9A2" box="[485,575,1619,1643]" pageId="4" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differed from the other three true ‘
|
||
<taxonomicName id="4AB54D56B614FFB7FC182978FBC7F9A3" box="[967,1038,1619,1643]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="viridis">
|
||
<emphasis id="BFC1EAC7B614FFB7FC182978FBC7F9A3" box="[967,1038,1619,1643]" italics="true" pageId="4" pageNumber="34">viridis</emphasis>
|
||
</taxonomicName>
|
||
group’ species by the unsculptured area between paired plate III and terminal plate; pointed clavae, instead of with rounded apices; and a stronger spur on internal claws. Differences among these species regarding quantitative characters are illustrated in
|
||
<tableCitation id="C037036EB614FFB7FAD029B1FAACF97A" box="[1295,1381,1690,1714]" captionStart="TABLE 2" captionStartId="6.[151,235,152,173]" captionText="TABLE 2. Measurements of single specimens of Echiniscus clavispinosus sp. nov., E. viridis, E. perviridis and E. viridianus (% bo = percentage of body length; % sc = percentage of scapular plate length)." pageId="4" pageNumber="34">Table 2</tableCitation>
|
||
. It is important to stress that the values relative to the external buccal cirrus, clava and spine on the first pair of legs of
|
||
<taxonomicName id="4AB54D56B614FFB7FA5B29EBFF16F933" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="viridis">
|
||
<emphasis id="BFC1EAC7B614FFB7FA5B29EBFF16F933" italics="true" pageId="4" pageNumber="34">E. viridis</emphasis>
|
||
</taxonomicName>
|
||
and the values relative to the leg IV claws of
|
||
<taxonomicName id="4AB54D56B614FFB7FD3D29CFFCBAF933" box="[738,883,1763,1787]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="viridianus">
|
||
<emphasis id="BFC1EAC7B614FFB7FD3D29CFFCBAF933" box="[738,883,1763,1787]" italics="true" pageId="4" pageNumber="34">E. viridianus</emphasis>
|
||
</taxonomicName>
|
||
are different from those indicated by Pilato
|
||
<emphasis id="BFC1EAC7B614FFB7FABB29CFFA55F933" box="[1380,1436,1763,1787]" italics="true" pageId="4" pageNumber="34">et al.</emphasis>
|
||
(2008a; tables 1 and 2) in which they, for a misunderstanding, are wrong.
|
||
</paragraph>
|
||
<paragraph id="8D0A36D5B614FFB7FF182806FA53F845" blockId="4.[151,1436,1255,2006]" pageId="4" pageNumber="34">
|
||
The new species differed from
|
||
<taxonomicName id="4AB54D56B614FFB7FDF62804FD58F88E" box="[553,657,1838,1862]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="viridis">
|
||
<emphasis id="BFC1EAC7B614FFB7FDF62804FD58F88E" box="[553,657,1838,1862]" italics="true" pageId="4" pageNumber="34">E. viridis</emphasis>
|
||
</taxonomicName>
|
||
in the absence of dark tubercles on the cephalic and neck plates; the smooth anterior portion of the median plate 2; the distance between the dark tubercles much shorter than the diameter of the tubercles (
|
||
<figureCitation id="158E2A50B614FFB7FE5B285EFE01F845" box="[388,456,1909,1933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1692,1715]" captionTargetBox="[171,1417,393,1649]" captionTargetId="figure@5.[151,1436,373,1671]" captionTargetPageId="5" captionText="FIGURE 3. Details of the cuticular ornamentation (PCM images) in: A, Echinicus clavispinosus sp. nov.; B, Echiniscus viridis; C, Echiniscus viridianus; D, Echnisicus perviridis; E, Echiniscus rufoviridis and, F, Echinisicus viridissimus. The differences between the first three species on one hand and Echniscus rufoviridis and Echiniscus viridissimus on the other hand are very evident. Scale bar: 10 µm." httpUri="https://zenodo.org/record/204755/files/figure.png" pageId="4" pageNumber="34">Fig. 3</figureCitation>
|
||
, A and B); more visible fine dots and light spots; and slightly longer cirrus
|
||
<emphasis id="BFC1EAC7B614FFB7FACA285DFAEFF846" box="[1301,1318,1910,1934]" italics="true" pageId="4" pageNumber="34">A</emphasis>
|
||
(
|
||
<tableCitation id="C037036EB614FFB7FAEA285EFA44F845" box="[1333,1421,1909,1933]" captionStart="TABLE 2" captionStartId="6.[151,235,152,173]" captionText="TABLE 2. Measurements of single specimens of Echiniscus clavispinosus sp. nov., E. viridis, E. perviridis and E. viridianus (% bo = percentage of body length; % sc = percentage of scapular plate length)." pageId="4" pageNumber="34">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8D0A36D5B614FFB6FF1828B2FE2FFF1C" blockId="4.[151,1436,1255,2006]" lastBlockId="5.[151,1436,151,320]" lastPageId="5" lastPageNumber="35" pageId="4" pageNumber="34">
|
||
<taxonomicName id="4AB54D56B614FFB7FF1828B2FE17F879" box="[199,478,1945,1969]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="clavispinosus" status="sp. nov.">
|
||
<emphasis id="BFC1EAC7B614FFB7FF1828B2FE17F879" box="[199,478,1945,1969]" italics="true" pageId="4" pageNumber="34">Echiniscus clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BFC1EAC7B614FFB7FE3A28B1FDF6F879" bold="true" box="[485,575,1946,1970]" pageId="4" pageNumber="34">
|
||
<taxonomicNameLabel id="A4F257BCB614FFB7FE3A28B1FDF6F879" box="[485,575,1946,1970]" pageId="4" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also differed from
|
||
<taxonomicName id="4AB54D56B614FFB7FCC828B1FC6AF879" box="[791,931,1945,1969]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="34" phylum="Tardigrada" rank="species" species="perviridis">
|
||
<emphasis id="BFC1EAC7B614FFB7FCC828B1FC6AF879" box="[791,931,1945,1969]" italics="true" pageId="4" pageNumber="34">E. perviridis</emphasis>
|
||
</taxonomicName>
|
||
in lacking tubercles in the cephalic and neck plates, the smooth anterior portion of median plate 2; finer and denser dots and light spots on the plates (
|
||
<figureCitation id="158E2A50B614FFB7FAE92896FAB2F81D" box="[1334,1403,1981,2005]" captionStart="FIGURE 3" captionStartId="5.[151,250,1692,1715]" captionTargetBox="[171,1417,393,1649]" captionTargetId="figure@5.[151,1436,373,1671]" captionTargetPageId="5" captionText="FIGURE 3. Details of the cuticular ornamentation (PCM images) in: A, Echinicus clavispinosus sp. nov.; B, Echiniscus viridis; C, Echiniscus viridianus; D, Echnisicus perviridis; E, Echiniscus rufoviridis and, F, Echinisicus viridissimus. The differences between the first three species on one hand and Echniscus rufoviridis and Echiniscus viridissimus on the other hand are very evident. Scale bar: 10 µm." httpUri="https://zenodo.org/record/204755/files/figure.png" pageId="4" pageNumber="34">Fig. 3</figureCitation>
|
||
, A and D), less visible green area bordering the trunk plates; dentate collar with smaller teeth; and much shorter cirrus
|
||
<emphasis id="BFC1EAC7B615FFB6FF482F97FF61FF1C" box="[151,168,188,212]" italics="true" pageId="5" pageNumber="35">A</emphasis>
|
||
and shorter claws (
|
||
<tableCitation id="C037036EB615FFB6FE5E2F97FE10FF1C" box="[385,473,188,212]" captionStart="TABLE 2" captionStartId="6.[151,235,152,173]" captionText="TABLE 2. Measurements of single specimens of Echiniscus clavispinosus sp. nov., E. viridis, E. perviridis and E. viridianus (% bo = percentage of body length; % sc = percentage of scapular plate length)." pageId="5" pageNumber="35">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8D0A36D5B615FFB6FF182FF4FB0EFEF7" blockId="5.[151,1436,151,320]" pageId="5" pageNumber="35">
|
||
The new species differed from
|
||
<emphasis id="BFC1EAC7B615FFB6FDF92FCAFD77FF30" box="[550,702,224,248]" italics="true" pageId="5" pageNumber="35">
|
||
<taxonomicName id="4AB54D56B615FFB6FDF92FCAFD73FF30" box="[550,698,224,248]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="35" phylum="Tardigrada" rank="species" species="viridianus">E. viridianus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the most similar species, by the absence of dark tubercles on the cephalic and neck plates; less visible and more dense fine dots and light spots on the plates (
|
||
<figureCitation id="158E2A50B615FFB6FB0E2E2FFAD0FED4" box="[1233,1305,260,284]" captionStart="FIGURE 3" captionStartId="5.[151,250,1692,1715]" captionTargetBox="[171,1417,393,1649]" captionTargetId="figure@5.[151,1436,373,1671]" captionTargetPageId="5" captionText="FIGURE 3. Details of the cuticular ornamentation (PCM images) in: A, Echinicus clavispinosus sp. nov.; B, Echiniscus viridis; C, Echiniscus viridianus; D, Echnisicus perviridis; E, Echiniscus rufoviridis and, F, Echinisicus viridissimus. The differences between the first three species on one hand and Echniscus rufoviridis and Echiniscus viridissimus on the other hand are very evident. Scale bar: 10 µm." httpUri="https://zenodo.org/record/204755/files/figure.png" pageId="5" pageNumber="35">Fig. 3</figureCitation>
|
||
, A and C); smooth anterior portion of the median plate 2; less visible green area bordering the trunk plates.
|
||
</paragraph>
|
||
<caption id="D9CA665DB615FFB6FF4829B7FE33F8C6" httpUri="https://zenodo.org/record/204755/files/figure.png" pageId="5" pageNumber="35" targetBox="[171,1417,393,1649]" targetPageId="5">
|
||
<paragraph id="8D0A36D5B615FFB6FF4829B7FE33F8C6" blockId="5.[151,1437,1692,1806]" pageId="5" pageNumber="35">
|
||
<emphasis id="BFC1EAC7B615FFB6FF4829B7FEDFF979" bold="true" box="[151,278,1692,1715]" pageId="5" pageNumber="35">FIGURE 3.</emphasis>
|
||
Details of the cuticular ornamentation (PCM images) in: A,
|
||
<taxonomicName id="4AB54D56B615FFB6FC7B29B7FB55F97A" box="[932,1180,1692,1714]" genus="Echinicus" pageId="5" pageNumber="35" rank="species" species="clavispinosus" status="sp. nov.">
|
||
<emphasis id="BFC1EAC7B615FFB6FC7B29B7FB55F97A" box="[932,1180,1692,1714]" italics="true" pageId="5" pageNumber="35">Echinicus clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BFC1EAC7B615FFB6FB7A29B6FACBF97A" bold="true" box="[1189,1282,1693,1714]" pageId="5" pageNumber="35">
|
||
<taxonomicNameLabel id="A4F257BCB615FFB6FB7A29B6FB34F97A" box="[1189,1277,1693,1714]" pageId="5" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
;
|
||
</emphasis>
|
||
B,
|
||
<taxonomicName id="4AB54D56B615FFB6FAF129B7FF11F919" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="35" phylum="Tardigrada" rank="species" species="viridis">
|
||
<emphasis id="BFC1EAC7B615FFB6FAF129B7FF11F919" italics="true" pageId="5" pageNumber="35">Echiniscus viridis</emphasis>
|
||
</taxonomicName>
|
||
; C,
|
||
<taxonomicName id="4AB54D56B615FFB6FEDB2990FE29F919" box="[260,480,1723,1745]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="35" phylum="Tardigrada" rank="species" species="viridianus">
|
||
<emphasis id="BFC1EAC7B615FFB6FEDB2990FE29F919" box="[260,480,1723,1745]" italics="true" pageId="5" pageNumber="35">Echiniscus viridianus</emphasis>
|
||
</taxonomicName>
|
||
; D,
|
||
<taxonomicName id="4AB54D56B615FFB6FDD12990FD2FF919" box="[526,742,1723,1745]" genus="Echnisicus" pageId="5" pageNumber="35" rank="species" species="perviridis">
|
||
<emphasis id="BFC1EAC7B615FFB6FDD12990FD2FF919" box="[526,742,1723,1745]" italics="true" pageId="5" pageNumber="35">Echnisicus perviridis</emphasis>
|
||
</taxonomicName>
|
||
; E,
|
||
<taxonomicName id="4AB54D56B615FFB6FCCE2990FC38F919" box="[785,1009,1723,1745]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="35" phylum="Tardigrada" rank="species" species="rufoviridis">
|
||
<emphasis id="BFC1EAC7B615FFB6FCCE2990FC38F919" box="[785,1009,1723,1745]" italics="true" pageId="5" pageNumber="35">Echiniscus rufoviridis</emphasis>
|
||
</taxonomicName>
|
||
and, F,
|
||
<emphasis id="BFC1EAC7B615FFB6FB9B2990FAF2F919" box="[1092,1339,1723,1745]" italics="true" pageId="5" pageNumber="35">
|
||
Echinisicus
|
||
<taxonomicName id="4AB54D56B615FFB6FB1E2990FAF2F919" box="[1217,1339,1723,1745]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="35" phylum="Tardigrada" rank="species" species="viridissimus">viridissimus</taxonomicName>
|
||
</emphasis>
|
||
. The differences between the first three species on one hand and
|
||
<taxonomicName id="4AB54D56B615FFB6FD3C29F1FC75F938" box="[739,956,1754,1776]" genus="Echniscus" pageId="5" pageNumber="35" rank="species" species="rufoviridis">
|
||
<emphasis id="BFC1EAC7B615FFB6FD3C29F1FC75F938" box="[739,956,1754,1776]" italics="true" pageId="5" pageNumber="35">Echniscus rufoviridis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4AB54D56B615FFB6FC2F29F1FB16F938" box="[1008,1247,1754,1776]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="35" phylum="Tardigrada" rank="species" species="viridissimus">
|
||
<emphasis id="BFC1EAC7B615FFB6FC2F29F1FB16F938" box="[1008,1247,1754,1776]" italics="true" pageId="5" pageNumber="35">Echiniscus viridissimus</emphasis>
|
||
</taxonomicName>
|
||
on the other hand are very evident. Scale bar: 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D9CA665DB616FFB5FF482FB3FC09FF04" pageId="6" pageNumber="36">
|
||
<paragraph id="8D0A36D5B616FFB5FF482FB3FC09FF04" blockId="6.[151,1436,152,832]" pageId="6" pageNumber="36">
|
||
<emphasis id="BFC1EAC7B616FFB5FF482FB3FECCFF65" bold="true" box="[151,261,152,173]" pageId="6" pageNumber="36">TABLE 2.</emphasis>
|
||
Measurements of single specimens of
|
||
<taxonomicName id="4AB54D56B616FFB5FD4A2FB3FC5BFF66" box="[661,914,152,174]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="36" phylum="Tardigrada" rank="species" species="clavispinosus" status="sp. nov.">
|
||
<emphasis id="BFC1EAC7B616FFB5FD4A2FB3FC5BFF66" box="[661,914,152,174]" italics="true" pageId="6" pageNumber="36">Echiniscus clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BFC1EAC7B616FFB5FC452FB3FC22FF65" bold="true" box="[922,1003,152,173]" pageId="6" pageNumber="36">
|
||
<taxonomicNameLabel id="A4F257BCB616FFB5FC452FB3FC22FF65" box="[922,1003,152,173]" pageId="6" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="BFC1EAC7B616FFB5FC252FB2FB2DFF66" box="[1018,1252,152,174]" italics="true" pageId="6" pageNumber="36">
|
||
<taxonomicName id="4AB54D56B616FFB5FC252FB2FB93FF66" box="[1018,1114,152,174]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="36" phylum="Tardigrada" rank="species" species="viridis">E. viridis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4AB54D56B616FFB5FBBA2FB2FB2DFF66" box="[1125,1252,152,174]" genus="Echnisicus" pageId="6" pageNumber="36" rank="species" species="perviridis">E. perviridis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4AB54D56B616FFB5FAC72FB2FA55FF66" box="[1304,1436,152,174]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="36" phylum="Tardigrada" rank="species" species="viridianus">
|
||
<emphasis id="BFC1EAC7B616FFB5FAC72FB2FAEEFF66" box="[1304,1319,153,174]" italics="true" pageId="6" pageNumber="36">E</emphasis>
|
||
.
|
||
<emphasis id="BFC1EAC7B616FFB5FAEA2FB3FA55FF66" box="[1333,1436,152,174]" italics="true" pageId="6" pageNumber="36">viridianus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="BFC1EAC7B616FFB5FF402F9CFF1CFF05" box="[159,213,183,205]" italics="true" pageId="6" pageNumber="36">% bo</emphasis>
|
||
= percentage of body length;
|
||
<emphasis id="BFC1EAC7B616FFB5FDDB2F9CFDFDFF05" box="[516,564,183,205]" italics="true" pageId="6" pageNumber="36">% sc</emphasis>
|
||
= percentage of scapular plate length).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8D0A36D5B616FFB5FF402FC3FAF8FF36" blockId="6.[151,1436,152,832]" box="[159,1329,232,254]" pageId="6" pageNumber="36">
|
||
Characters
|
||
<taxonomicName id="4AB54D56B616FFB5FE4E2FC2FDFFFF36" box="[401,566,232,254]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="36" phylum="Tardigrada" rank="species" species="clavispinosus" status="sp. nov.">
|
||
<emphasis id="BFC1EAC7B616FFB5FE4E2FC2FDFFFF36" box="[401,566,232,254]" italics="true" pageId="6" pageNumber="36">E. clavispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A4F257BCB616FFB5FDE32FC3FD45FF36" box="[572,652,232,254]" pageId="6" pageNumber="36" rank="species">
|
||
<emphasis id="BFC1EAC7B616FFB5FDE32FC3FD4CFF35" bold="true" box="[572,645,232,253]" pageId="6" pageNumber="36">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<taxonomicName id="4AB54D56B616FFB5FD7E2FC2FD34FF36" box="[673,765,232,254]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="36" phylum="Tardigrada" rank="species" species="viridis">
|
||
<emphasis id="BFC1EAC7B616FFB5FD7E2FC2FD34FF36" box="[673,765,232,254]" italics="true" pageId="6" pageNumber="36">E. viridis</emphasis>
|
||
</taxonomicName>
|
||
*
|
||
<emphasis id="BFC1EAC7B616FFB5FC462FC2FAF8FF36" box="[921,1329,232,254]" italics="true" pageId="6" pageNumber="36">
|
||
<taxonomicName id="4AB54D56B616FFB5FC462FC2FBDEFF36" box="[921,1047,232,254]" genus="Echnisicus" pageId="6" pageNumber="36" rank="species" species="perviridis">E. perviridis</taxonomicName>
|
||
<taxonomicName id="4AB54D56B616FFB5FB7D2FC2FAEEFF36" box="[1186,1319,232,254]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="36" phylum="Tardigrada" rank="species" species="viridianus">E. viridianus</taxonomicName>
|
||
*
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D0A36D5B616FFB5FE4E2E39FA4AFEEF" blockId="6.[151,1436,152,832]" box="[401,1411,273,295]" pageId="6" pageNumber="36">µm % bo % sc µm % bo % sc µm % bo % sc µm % bo % sc</paragraph>
|
||
<paragraph id="8D0A36D5B616FFB5FF402E11FB01FE98" blockId="6.[151,1436,152,832]" box="[159,1224,314,336]" pageId="6" pageNumber="36">Body length 241 - - 190 290 231</paragraph>
|
||
<paragraph id="8D0A36D5B616FFB5FF402E48FAE2FEB1" blockId="6.[151,1436,152,832]" box="[159,1323,355,378]" pageId="6" pageNumber="36">Scapular plate length 48.8 20.2 - 41.8 22.0 56.5 19.5 43.1 18.7</paragraph>
|
||
<paragraph id="8D0A36D5B616FFB5FF402EA6FAB7FC88" blockId="6.[151,1436,152,832]" pageId="6" pageNumber="36">
|
||
Internal buccal cirrus 12.8 5.3 26.2 9.0 4.7 21.5 - - - 13.9 6.0 32.3 External buccal cirrus 15.8 6.6 32.4 10.3 5.4 24.6 - - - 14.7 6.4 34.1 Cephalic papilla 7.3 3.0 15.0 6.4 3.4 15.3 6.0 2.1 10.6 5.8 2.5 13.5 Clava 8.1 3.4 16.6 6.0 3.2 14.4 - - - 5.5 2.4 12.8 Cirrus
|
||
<emphasis id="BFC1EAC7B616FFB5FF3B2D18FF3AFD80" box="[228,243,563,584]" italics="true" pageId="6" pageNumber="36">A</emphasis>
|
||
32.4 13.4 66.4 18.2 9.6 43.5 114.0 39.3 201.8 29.0 12.6 67.3 Spine on leg I 3.4 1.4 7.0 3.3 1.7 7.9 - - - 2.6 1.1 6.0 Internal claw leg I 15.4 6.4 31.6 12.6 6.6 30.1 - - - 15.1 6.5 35.0 External claw leg I 14.4 6.0 29.5 11.9 6.3 28.5 21.4 7.4 37.9 14.5 6.3 33.6 Internal claw leg IV 17.6 7.3 36.1 15.5 8.2 37.1 30.0 10.3 53.1 16.9 7.3 39.2 External claw leg IV 15.5 6.4 31.8 14.7 7.7 35.2 28.5 9.8 50.4 15.2 6.6 35.3 Papilla leg IV - - - 3.0 1.6 7.2 - - - 4.8 2.1 11.1
|
||
</paragraph>
|
||
<paragraph id="8D0A36D5B616FFB5FF482C49FDDCFC7F" blockId="6.[151,1436,865,952]" pageId="6" pageNumber="36">
|
||
* As above mentioned in the text, the
|
||
<emphasis id="BFC1EAC7B616FFB5FDCB2C4AFD8AFCBF" box="[532,579,865,887]" italics="true" pageId="6" pageNumber="36">%bo</emphasis>
|
||
and
|
||
<emphasis id="BFC1EAC7B616FFB5FDAB2C4AFD57FCBF" box="[628,670,865,887]" italics="true" pageId="6" pageNumber="36">%sc</emphasis>
|
||
values relative to the external buccal cirrus, clava and spine on the first pair of legs of
|
||
<taxonomicName id="4AB54D56B616FFB5FF232CA8FE91FC50" box="[252,344,898,920]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="36" phylum="Tardigrada" rank="species" species="viridis">
|
||
<emphasis id="BFC1EAC7B616FFB5FF232CA8FE91FC50" box="[252,344,898,920]" italics="true" pageId="6" pageNumber="36">E. viridis</emphasis>
|
||
</taxonomicName>
|
||
and the values regarding the leg IV claws of
|
||
<taxonomicName id="4AB54D56B616FFB5FCFF2CA8FC6BFC50" box="[800,930,898,920]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="36" phylum="Tardigrada" rank="species" species="viridianus">
|
||
<emphasis id="BFC1EAC7B616FFB5FCFF2CA8FC6BFC50" box="[800,930,898,920]" italics="true" pageId="6" pageNumber="36">E. viridianus</emphasis>
|
||
</taxonomicName>
|
||
differ from those indicated by Pilato
|
||
<emphasis id="BFC1EAC7B616FFB5FAC32CA8FA86FC50" box="[1308,1359,898,920]" italics="true" pageId="6" pageNumber="36">et al.</emphasis>
|
||
(2008a, tables 1 and 2), which were incorrect.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |