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<document id="54DFDB46304BF6382CFF87A2D94EA202" ID-DOI="10.5281/zenodo.186895" ID-GBIF-Dataset="be950fd9-a476-4479-b6ec-a085df0ff9cb" ID-ISSN="1175-5326" ID-Zenodo-Dep="186895" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461059228515" checkinUser="plazi" docAuthor="Christidis, Faye" docDate="2009" docId="0511E618924BEF1EDE83A1B9FE7D67E4" docLanguage="en" docName="zt02063p068.pdf" docOrigin="Zootaxa 2063" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Riekophlebia Christidis, 2009, gen. n" docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="F9289E60924AEF1CDE14A12EFF98673C" masterDocTitle="Riekophlebia crocina, a new genus and species of Atalophlebiinae (Ephemeroptera: Leptophlebiidae) from the Wet Tropics bioregion of north-eastern Australia" masterLastPageNumber="68" masterPageNumber="64" pageNumber="65" updateTime="1698581785873" updateUser="plazi">
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<mods:title id="1A61F54905D8E42174AE0852DA29C6BF">Riekophlebia crocina, a new genus and species of Atalophlebiinae (Ephemeroptera: Leptophlebiidae) from the Wet Tropics bioregion of north-eastern Australia</mods:title>
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<mods:namePart id="FDBD0790E0D32385519D82FD8B256108">Christidis, Faye</mods:namePart>
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<mods:date id="F015260FD27181B281D254775AA16CB5">2009</mods:date>
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<treatment id="0511E618924BEF1EDE83A1B9FE7D67E4" ID-DOI="http://doi.org/10.5281/zenodo.6214492" ID-GBIF-Taxon="119613781" ID-Zenodo-Dep="6214492" LSID="urn:lsid:plazi:treatment:0511E618924BEF1EDE83A1B9FE7D67E4" httpUri="http://treatment.plazi.org/id/0511E618924BEF1EDE83A1B9FE7D67E4" lastPageId="2" lastPageNumber="66" pageId="1" pageNumber="65">
<subSubSection id="C5A20485924BEF1DDE83A1B9FE10678D" box="[151,392,151,177]" pageId="1" pageNumber="65" type="nomenclature">
<paragraph id="8D07570E924BEF1DDE83A1B9FE10678D" blockId="1.[151,392,151,177]" box="[151,392,151,177]" pageId="1" pageNumber="65">
<heading id="D64FE062924BEF1DDE83A1B9FE10678D" bold="true" box="[151,392,151,177]" fontSize="11" level="1" pageId="1" pageNumber="65" reason="1">
<emphasis id="BFCC8B1C924BEF1DDE83A1B9FE10678D" bold="true" box="[151,392,151,177]" pageId="1" pageNumber="65">
<taxonomicName id="4AB82C8D924BEF1DDE83A1B9FEAF678D" ID-CoL="79Z6" box="[151,311,151,177]" class="Insecta" family="Leptophlebiidae" genus="Riekophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BFCC8B1C924BEF1DDE83A1B9FEAF678D" bold="true" box="[151,311,151,177]" italics="true" pageId="1" pageNumber="65">Riekophlebia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4FF3667924BEF1DDF2AA1B9FE10678D" box="[318,392,151,177]" pageId="1" pageNumber="65" rank="genus">gen. n</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C5A20485924BEF1DDE83A1CFFDD567C4" box="[151,589,225,249]" pageId="1" pageNumber="65" type="materials_examined">
<paragraph id="8D07570E924BEF1DDE83A1CFFDD567C4" blockId="1.[151,1437,225,2029]" box="[151,589,225,249]" pageId="1" pageNumber="65">
<emphasis id="BFCC8B1C924BEF1DDE83A1CFFEBB67C5" bold="true" box="[151,291,225,249]" pageId="1" pageNumber="65">
<typeStatus id="5203E9AC924BEF1DDE83A1CFFF5767C5" box="[151,207,225,249]" pageId="1" pageNumber="65">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4AB82C8D924BEF1DDF25A1CFFD8D67C4" box="[305,533,225,248]" class="Insecta" family="Leptophlebiidae" genus="Riekophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="crocina" status="sp. nov.">
<emphasis id="BFCC8B1C924BEF1DDF25A1CFFD8D67C4" box="[305,533,225,248]" italics="true" pageId="1" pageNumber="65">Riekophlebia crocina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4FF3667924BEF1DDC08A1CFFDD567C4" box="[540,589,225,248]" pageId="1" pageNumber="65" rank="species">sp. n</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C5A20485924BEF1DDE83A02EFED464C3" pageId="1" pageNumber="65" type="diagnosis">
<paragraph id="8D07570E924BEF1DDE83A02EFBF66535" blockId="1.[151,1437,225,2029]" pageId="1" pageNumber="65">
<emphasis id="BFCC8B1C924BEF1DDE83A02EFE886626" bold="true" box="[151,272,256,282]" pageId="1" pageNumber="65">Diagnosis</emphasis>
.
<emphasis id="BFCC8B1C924BEF1DDF0BA02EFEF76626" bold="true" box="[287,367,256,282]" pageId="1" pageNumber="65">Imago</emphasis>
. Wings hyaline (
<figureCitation id="15834B8B924BEF1DDC2DA02FFDE76627" box="[569,639,257,283]" captionStart="FIGURES 1 5" captionStartId="2.[151,271,1630,1654]" captionTargetBox="[186,1401,271,1596]" captionTargetId="figure@2.[151,1436,256,1622]" captionTargetPageId="2" captionText="FIGURES 1 5. Riekophlebia crocina. Male imago: (1) wings; (2) hind wing; (3) abdominal terga 3 5; (4) penes and claspers, ventral view. Female imago: (5) ninth abdominal sternum." httpUri="https://zenodo.org/record/186896/files/figure.png" pageId="1" pageNumber="65">Fig. 1</figureCitation>
). Forewing length-width ratio 3.03.2; MA forked at just over half distance from base of vein to margin of wing, fork symmetrical; MP2 attached to MP1 at about 0.24 length of MP1; ICu1 not attached at base to CuA-CuP cross vein.
<collectingCountry id="F5AF179E924BEF1DDD30A07FFCC76657" box="[804,863,337,363]" name="India" pageId="1" pageNumber="65">Hind</collectingCountry>
wing 0.180.20 times length of forewing; costal margin convex; vein Sc about 0.8 length of hind wing (
<figureCitation id="15834B8B924BEF1DDD23A055FC1B66A9" box="[823,899,379,405]" captionStart="FIGURES 1 5" captionStartId="2.[151,271,1630,1654]" captionTargetBox="[186,1401,271,1596]" captionTargetId="figure@2.[151,1436,256,1622]" captionTargetPageId="2" captionText="FIGURES 1 5. Riekophlebia crocina. Male imago: (1) wings; (2) hind wing; (3) abdominal terga 3 5; (4) penes and claspers, ventral view. Female imago: (5) ninth abdominal sternum." httpUri="https://zenodo.org/record/186896/files/figure.png" pageId="1" pageNumber="65">Fig. 2</figureCitation>
). Tarsal claws dissimilar, one with an apical hook, the other pad-like. Male genitalia with claspers 3 segmented, segment 1 of claspers narrowing at about 0.4 of length; penes fused along most of their length, each lobe with a large ventral spine near apex; styliger plate with deep cleft (
<figureCitation id="15834B8B924BEF1DDF81A0C1FE466535" box="[405,478,495,521]" captionStart="FIGURES 1 5" captionStartId="2.[151,271,1630,1654]" captionTargetBox="[186,1401,271,1596]" captionTargetId="figure@2.[151,1436,256,1622]" captionTargetPageId="2" captionText="FIGURES 1 5. Riekophlebia crocina. Male imago: (1) wings; (2) hind wing; (3) abdominal terga 3 5; (4) penes and claspers, ventral view. Female imago: (5) ninth abdominal sternum." httpUri="https://zenodo.org/record/186896/files/figure.png" pageId="1" pageNumber="65">Fig. 4</figureCitation>
). Female ninth sternum entire, pointed apically (
<figureCitation id="15834B8B924BEF1DDA03A0C1FBC76535" box="[1047,1119,495,521]" captionStart="FIGURES 1 5" captionStartId="2.[151,271,1630,1654]" captionTargetBox="[186,1401,271,1596]" captionTargetId="figure@2.[151,1436,256,1622]" captionTargetPageId="2" captionText="FIGURES 1 5. Riekophlebia crocina. Male imago: (1) wings; (2) hind wing; (3) abdominal terga 3 5; (4) penes and claspers, ventral view. Female imago: (5) ninth abdominal sternum." httpUri="https://zenodo.org/record/186896/files/figure.png" pageId="1" pageNumber="65">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8D07570E924BEF1DDED2A33BFED464C3" blockId="1.[151,1437,225,2029]" pageId="1" pageNumber="65">
<emphasis id="BFCC8B1C924BEF1DDED2A33BFEB26513" bold="true" box="[198,298,533,559]" pageId="1" pageNumber="65">Nymph.</emphasis>
Head prognathus. Antennae about two times length of head. Mouthparts as in
<figureCitation id="15834B8B924BEF1DDAECA33BFA006513" box="[1272,1432,533,559]" captionStart="FIGURES 6 13" captionStartId="3.[151,270,1682,1706]" captionTargetBox="[154,1421,356,1638]" captionTargetId="figure@3.[151,1436,351,1674]" captionTargetPageId="3" captionText="FIGURES 6 13. Riekophlebia crocina. Nymph: (6) labrum and clypeus; (7) right mandible; (8) maxilla, ventral view; (9) hypopharynx; (10) labium, dorsal view left of midline and ventral view right of midline; (11) segment 3 of labial palp, dorsal view; (12) foreleg; (13) abdominal gill." httpUri="https://zenodo.org/record/186897/files/figure.png" pageId="1" pageNumber="65">Figures 611</figureCitation>
. Labrum broader than clypeus; length 1.181.20 times maximum length of clypeus; labrum broadest beyond midlength; anterior margin with median U-shaped notch; dorsal surface with two setal fringes (
<figureCitation id="15834B8B924BEF1DDB2BA34DFA156541" box="[1343,1421,611,637]" captionStart="FIGURES 6 13" captionStartId="3.[151,270,1682,1706]" captionTargetBox="[154,1421,356,1638]" captionTargetId="figure@3.[151,1436,351,1674]" captionTargetPageId="3" captionText="FIGURES 6 13. Riekophlebia crocina. Nymph: (6) labrum and clypeus; (7) right mandible; (8) maxilla, ventral view; (9) hypopharynx; (10) labium, dorsal view left of midline and ventral view right of midline; (11) segment 3 of labial palp, dorsal view; (12) foreleg; (13) abdominal gill." httpUri="https://zenodo.org/record/186897/files/figure.png" pageId="1" pageNumber="65">Fig. 6</figureCitation>
). Clypeus with lateral margins slightly divergent apically. Outer margin of mandible with small median setal tuft and series of fine setae between tuft and outer incisor (
<figureCitation id="15834B8B924BEF1DDD52A39EFC1665F6" box="[838,910,688,714]" captionStart="FIGURES 6 13" captionStartId="3.[151,270,1682,1706]" captionTargetBox="[154,1421,356,1638]" captionTargetId="figure@3.[151,1436,351,1674]" captionTargetPageId="3" captionText="FIGURES 6 13. Riekophlebia crocina. Nymph: (6) labrum and clypeus; (7) right mandible; (8) maxilla, ventral view; (9) hypopharynx; (10) labium, dorsal view left of midline and ventral view right of midline; (11) segment 3 of labial palp, dorsal view; (12) foreleg; (13) abdominal gill." httpUri="https://zenodo.org/record/186897/files/figure.png" pageId="1" pageNumber="65">Fig. 7</figureCitation>
); incisors slender; prostheca well developed. Maxilla broad apically, with 2527 subapical pectinate setae (
<figureCitation id="15834B8B924BEF1DDD8FA3F9FC7F65CD" box="[923,999,727,753]" captionStart="FIGURES 6 13" captionStartId="3.[151,270,1682,1706]" captionTargetBox="[154,1421,356,1638]" captionTargetId="figure@3.[151,1436,351,1674]" captionTargetPageId="3" captionText="FIGURES 6 13. Riekophlebia crocina. Nymph: (6) labrum and clypeus; (7) right mandible; (8) maxilla, ventral view; (9) hypopharynx; (10) labium, dorsal view left of midline and ventral view right of midline; (11) segment 3 of labial palp, dorsal view; (12) foreleg; (13) abdominal gill." httpUri="https://zenodo.org/record/186897/files/figure.png" pageId="1" pageNumber="65">Fig. 8</figureCitation>
). Ligula of hypopharynx with well developed lateral processes, anterior margin cleft (
<figureCitation id="15834B8B924BEF1DDCE1A3D3FCA6642B" box="[757,830,765,791]" captionStart="FIGURES 6 13" captionStartId="3.[151,270,1682,1706]" captionTargetBox="[154,1421,356,1638]" captionTargetId="figure@3.[151,1436,351,1674]" captionTargetPageId="3" captionText="FIGURES 6 13. Riekophlebia crocina. Nymph: (6) labrum and clypeus; (7) right mandible; (8) maxilla, ventral view; (9) hypopharynx; (10) labium, dorsal view left of midline and ventral view right of midline; (11) segment 3 of labial palp, dorsal view; (12) foreleg; (13) abdominal gill." httpUri="https://zenodo.org/record/186897/files/figure.png" pageId="1" pageNumber="65">Fig. 9</figureCitation>
). Labium with glossae on about the same plane as paraglossae. Lateral margins of submentum with a few basal setae (
<figureCitation id="15834B8B924BEF1DDDA5A20AFB916402" box="[945,1033,804,830]" captionStart="FIGURES 6 13" captionStartId="3.[151,270,1682,1706]" captionTargetBox="[154,1421,356,1638]" captionTargetId="figure@3.[151,1436,351,1674]" captionTargetPageId="3" captionText="FIGURES 6 13. Riekophlebia crocina. Nymph: (6) labrum and clypeus; (7) right mandible; (8) maxilla, ventral view; (9) hypopharynx; (10) labium, dorsal view left of midline and ventral view right of midline; (11) segment 3 of labial palp, dorsal view; (12) foreleg; (13) abdominal gill." httpUri="https://zenodo.org/record/186897/files/figure.png" pageId="1" pageNumber="65">Fig. 10</figureCitation>
). Labial palp with segments 1 and 2 subequal in length, segment 3 about half length of segment 2. Tarsal claws with ventral teeth, increasing in size apically. Abdomen with posterolateral spines present on segments 29; lateral margins of abdominal segments bare, without setae. Gills present on segments 17; each gill lanceolate, lateral tracheae moderately developed (
<figureCitation id="15834B8B924BEF1DDF0BA291FEEE64E5" box="[287,374,959,985]" captionStart="FIGURES 6 13" captionStartId="3.[151,270,1682,1706]" captionTargetBox="[154,1421,356,1638]" captionTargetId="figure@3.[151,1436,351,1674]" captionTargetPageId="3" captionText="FIGURES 6 13. Riekophlebia crocina. Nymph: (6) labrum and clypeus; (7) right mandible; (8) maxilla, ventral view; (9) hypopharynx; (10) labium, dorsal view left of midline and ventral view right of midline; (11) segment 3 of labial palp, dorsal view; (12) foreleg; (13) abdominal gill." httpUri="https://zenodo.org/record/186897/files/figure.png" pageId="1" pageNumber="65">Fig. 13</figureCitation>
). Caudal filaments 1.5 to 2 times length of body; each segment with apical whorl of spines and short setae.
</paragraph>
</subSubSection>
<subSubSection id="C5A20485924BEF1DDED2A522FED560D1" pageId="1" pageNumber="65" type="discussion">
<paragraph id="8D07570E924BEF1DDED2A522FDAA6121" blockId="1.[151,1437,225,2029]" pageId="1" pageNumber="65">
<emphasis id="BFCC8B1C924BEF1DDED2A522FED8631A" bold="true" box="[198,320,1036,1062]" pageId="1" pageNumber="65">Remarks.</emphasis>
<taxonomicName id="4AB82C8D924BEF1DDF5AA522FE716319" box="[334,489,1036,1061]" class="Insecta" family="Leptophlebiidae" genus="Riekophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDF5AA522FE716319" box="[334,489,1036,1061]" italics="true" pageId="1" pageNumber="65">Riekophlebia</emphasis>
</taxonomicName>
can be distinguished from all other leptophlebiid genera by the following combination of characters. Imago: (1) penes fused along most of their length, each lobe with large ventral spine near apex; (2) segment 1 of claspers narrowing at about 0.4 of length; (3) styliger plate with deep cleft; (4) tarsal claws dissimilar; (5) female ninth sternum entire and pointed apically. Nymph: (1) anterior margin of labrum with median U-shaped notch; (2) labrum length 1.181.20 times maximum length of clypeus (3) labrum broadest beyond midlength; (4) maxillae broad apically, with 2527 subapical pectinate setae; (5) lateral margins of submentum with a few basal setae; (6) abdomen with posterolateral spines present on segments 29; (7) lateral margins of abdominal segments without setal fringe; (8) gills with lateral tracheae.
<taxonomicName id="4AB82C8D924BEF1DDE83A46CFEAA6267" box="[151,306,1346,1371]" class="Insecta" family="Leptophlebiidae" genus="Riekophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDE83A46CFEAA6267" box="[151,306,1346,1371]" italics="true" pageId="1" pageNumber="65">Riekophlebia</emphasis>
</taxonomicName>
is a member of the
<taxonomicName id="4AB82C8D924BEF1DDC0DA46CFD356267" box="[537,685,1346,1371]" class="Insecta" family="Leptophlebiidae" genus="Meridialaris" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDC0DA46CFD356267" box="[537,685,1346,1371]" italics="true" pageId="1" pageNumber="65">Meridialaris</emphasis>
</taxonomicName>
lineage as defined by
<bibRefCitation id="E9292AFF924BEF1DDDA6A46CFB696260" author="Pescador" box="[946,1265,1346,1372]" pageId="1" pageNumber="68" refString="Pescador, M. L. &amp; Peters, W. L. (1980) Phylogenetic relationships and zoogeography of cool-adapted Leptophlebiidae (Ephemeroptera) in Southern South America. In Flannagan, J. F. &amp; Marshall, K. E. (Eds.), Advances in Ephemeroptera Biology. Plenum Press, New York, pp. 43 - 56." type="book chapter" year="1980">Pescador and Peters (1980)</bibRefCitation>
and Christidis (2005), based on the possession of the following synapomorphic character states: adult tarsal claws dissimilar; ninth abdominal sternum of female entire; labrum of nymph with U-shaped notch on anterior margin; labrum broader than clypeus; maxilla broad apically; lateral margins of submentum with few basal setae. Other Australian members of the
<taxonomicName id="4AB82C8D924BEF1DDC13A4F2FD2962C9" box="[519,689,1500,1525]" class="Insecta" family="Leptophlebiidae" genus="Meridialaris" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDC13A4F2FD2962C9" box="[519,689,1500,1525]" italics="true" pageId="1" pageNumber="65">Meridialaris</emphasis>
</taxonomicName>
lineage include the genera
<taxonomicName id="4AB82C8D924BEF1DDA33A4F2FA8262C9" box="[1063,1306,1500,1525]" class="Insecta" family="Leptophlebiidae" genus="Austrophlebioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDA33A4F2FA8262C9" box="[1063,1306,1500,1525]" italics="true" pageId="1" pageNumber="65">Austrophlebioides</emphasis>
</taxonomicName>
,
<taxonomicName id="4AB82C8D924BEF1DDB39A4F2FA0D62C9" box="[1325,1429,1500,1525]" class="Insecta" family="Leptophlebiidae" genus="Kirrara" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDB39A4F2FA0D62C9" box="[1325,1429,1500,1525]" italics="true" pageId="1" pageNumber="65">Kirrara</emphasis>
</taxonomicName>
,
<taxonomicName id="4AB82C8D924BEF1DDE83A72DFEC26120" box="[151,346,1539,1564]" class="Insecta" family="Leptophlebiidae" genus="Tillyardophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDE83A72DFEC26120" box="[151,346,1539,1564]" italics="true" pageId="1" pageNumber="65">Tillyardophlebia</emphasis>
</taxonomicName>
and
<taxonomicName id="4AB82C8D924BEF1DDF86A72DFDB36120" box="[402,555,1539,1564]" class="Insecta" family="Leptophlebiidae" genus="Manggabora" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDF86A72DFDB36120" box="[402,555,1539,1564]" italics="true" pageId="1" pageNumber="65">Manggabora</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8D07570E924BEF1DDED2A704FED560D1" blockId="1.[151,1437,225,2029]" pageId="1" pageNumber="65">
<taxonomicName id="4AB82C8D924BEF1DDED2A704FEF9617F" box="[198,353,1578,1603]" class="Insecta" family="Leptophlebiidae" genus="Riekophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDED2A704FEF9617F" box="[198,353,1578,1603]" italics="true" pageId="1" pageNumber="65">Riekophlebia</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4AB82C8D924BEF1DDC9EA704FCD5617F" box="[650,845,1578,1603]" class="Insecta" family="Leptophlebiidae" genus="Tillyardophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDC9EA704FCD5617F" box="[650,845,1578,1603]" italics="true" pageId="1" pageNumber="65">Tillyardophlebia</emphasis>
</taxonomicName>
and
<emphasis id="BFCC8B1C924BEF1DDD9FA704FBB3617F" box="[907,1067,1578,1603]" italics="true" pageId="1" pageNumber="65">
<taxonomicName id="4AB82C8D924BEF1DDD9FA704FBBF617F" box="[907,1063,1578,1603]" class="Insecta" family="Leptophlebiidae" genus="Manggabora" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">Manggabora</taxonomicName>
,
</emphasis>
but can be distinguished from these genera as follows.
<taxonomicName id="4AB82C8D924BEF1DDFABA77FFDC46156" box="[447,604,1617,1642]" class="Insecta" family="Leptophlebiidae" genus="Riekophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDFABA77FFDC46156" box="[447,604,1617,1642]" italics="true" pageId="1" pageNumber="65">Riekophlebia</emphasis>
</taxonomicName>
is separated from
<taxonomicName id="4AB82C8D924BEF1DDD29A77FFC416156" box="[829,985,1617,1642]" class="Insecta" family="Leptophlebiidae" genus="Manggabora" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDD29A77FFC416156" box="[829,985,1617,1642]" italics="true" pageId="1" pageNumber="65">Manggabora</emphasis>
</taxonomicName>
in the imago by the deep cleft on the posterior margin of the styliger plate, and absence of a large ventral projection on the penes; and in the nymph by the presence of posterolateral spines on abdominal segments 29, a maxilla with 2527 subapical pectinate setae, and presence of setae on the outer margin of mandible between the median setal tuft and the outer incisor.
<taxonomicName id="4AB82C8D924BEF1DDEE0A7C5FE176038" box="[244,399,1771,1796]" class="Insecta" family="Leptophlebiidae" genus="Riekophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDEE0A7C5FE176038" box="[244,399,1771,1796]" italics="true" pageId="1" pageNumber="65">Riekophlebia</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4AB82C8D924BEF1DDC80A7C5FCCE6038" box="[660,854,1771,1796]" class="Insecta" family="Leptophlebiidae" genus="Tillyardophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDC80A7C5FCCE6038" box="[660,854,1771,1796]" italics="true" pageId="1" pageNumber="65">Tillyardophlebia</emphasis>
</taxonomicName>
by the form of the male genitalia including penes that are fused along most of their length, presence of large ventral spine near apex of each penes lobe, absence of a pair of large curved ventral spines near base of penes, and a styliger plate with a deep cleft. The nymph of
<taxonomicName id="4AB82C8D924BEF1DDE83A671FEAE6044" box="[151,310,1887,1912]" class="Insecta" family="Leptophlebiidae" genus="Riekophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDE83A671FEAE6044" box="[151,310,1887,1912]" italics="true" pageId="1" pageNumber="65">Riekophlebia</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4AB82C8D924BEF1DDC53A671FD5E6044" box="[583,710,1887,1913]" class="Insecta" family="Leptophlebiidae" genus="Tillyardophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="dostinei">
<emphasis id="BFCC8B1C924BEF1DDC53A671FDCE6044" box="[583,598,1887,1912]" italics="true" pageId="1" pageNumber="65">T</emphasis>
.
<emphasis id="BFCC8B1C924BEF1DDC72A671FD5E6044" box="[614,710,1887,1912]" italics="true" pageId="1" pageNumber="65">dostinei</emphasis>
</taxonomicName>
by the presence of lateral tracheae on the gills; and from
<taxonomicName id="4AB82C8D924BEF1DDB92A671FF7B60A3" class="Insecta" family="Leptophlebiidae" genus="Tillyardophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="alpina">
<emphasis id="BFCC8B1C924BEF1DDB92A671FF7B60A3" italics="true" pageId="1" pageNumber="65">T. alpina</emphasis>
</taxonomicName>
and
<taxonomicName id="4AB82C8D924BEF1DDF09A6A8FE1E60A3" box="[285,390,1926,1951]" class="Insecta" family="Leptophlebiidae" genus="Tillyardophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="rufosa">
<emphasis id="BFCC8B1C924BEF1DDF09A6A8FE1E60A3" box="[285,390,1926,1951]" italics="true" pageId="1" pageNumber="65">T. rufosa</emphasis>
</taxonomicName>
by the form of the labrum which in
<taxonomicName id="4AB82C8D924BEF1DDD2CA6A8FC4C60A3" box="[824,980,1926,1951]" class="Insecta" family="Leptophlebiidae" genus="Riekophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="BFCC8B1C924BEF1DDD2CA6A8FC4C60A3" box="[824,980,1926,1951]" italics="true" pageId="1" pageNumber="65">Riekophlebia</emphasis>
</taxonomicName>
is broadest beyond midlength and has angular lateral margins whereas in
<taxonomicName id="4AB82C8D924BEF1DDC56A682FD3560F9" box="[578,685,1964,1989]" class="Insecta" family="Leptophlebiidae" genus="Tillyardophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="alpina">
<emphasis id="BFCC8B1C924BEF1DDC56A682FD3560F9" box="[578,685,1964,1989]" italics="true" pageId="1" pageNumber="65">T. alpina</emphasis>
</taxonomicName>
and
<taxonomicName id="4AB82C8D924BEF1DDCFEA682FCCB60F9" box="[746,851,1964,1989]" class="Insecta" family="Leptophlebiidae" genus="Tillyardophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="rufosa">
<emphasis id="BFCC8B1C924BEF1DDCFEA682FCCB60F9" box="[746,851,1964,1989]" italics="true" pageId="1" pageNumber="65">T. rufosa</emphasis>
</taxonomicName>
is broadest at about midlength and has rounded lateral margins.
</paragraph>
</subSubSection>
<subSubSection id="C5A204859248EF1EDED2A1B9FE7D67E4" pageId="2" pageNumber="66" type="etymology">
<paragraph id="8D07570E9248EF1EDED2A1B9FE7D67E4" blockId="2.[151,1436,151,216]" pageId="2" pageNumber="66">
<emphasis id="BFCC8B1C9248EF1EDED2A1B9FECB678D" bold="true" box="[198,339,151,177]" pageId="2" pageNumber="66">Etymology.</emphasis>
This genus is named for Edgar Riek, in recognition of his contribution to mayfly research in
<collectingCountry id="F5AF179E9248EF1EDE83A190FE9E67E4" box="[151,262,190,216]" name="Australia" pageId="2" pageNumber="66">Australia</collectingCountry>
. Gender, feminine.
</paragraph>
</subSubSection>
</treatment>
</document>