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<document ID-CLB-Dataset="6049" ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.63775" ID-GBIF-Dataset="e6957b3b-f228-4d6a-b430-35d41cdb2caa" ID-PMC="PMC8222308" ID-Pensoft-Pub="1313-2970-1044-375" ID-Pensoft-UUID="EBD9E81E5EC55AA491952682787793A4" ID-PubMed="34183881" ID-ZooBank="D50CC77C2E6941FDA9BD395B025C43AE" ModsDocID="1313-2970-1044-375" checkinTime="1623875406767" checkinUser="pensoft" docAuthor="Will, Kipling &amp; Gueorguiev, Borislav" docDate="2021" docId="8FD37B09F28452A9B8457C6BBA2ADD1B" docLanguage="en" docName="ZooKeys 1044: 375-425" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.63775" docTitle="Coptocarpus amieuensis Will &amp; Guéorguiev 2021, sp. nov." docType="treatment" docUuid="6FDFFBC6-C84B-4070-AEC5-D4D39F730356" docUuidSource="ZooBank" docVersion="6" id="EBD9E81E5EC55AA491952682787793A4" lastPageNumber="375" masterDocId="EBD9E81E5EC55AA491952682787793A4" masterDocTitle="Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)" masterLastPageNumber="425" masterPageNumber="375" pageNumber="375" updateTime="1732931690188" updateUser="ExternalLinkService">
<mods:mods id="8FED048FC8227F2F2242C623F566153B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4AC7584450A1800C79369688F2E78BA4">
<mods:title id="F7C81C0501D2D1268623D42D646064F7">Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)</mods:title>
</mods:titleInfo>
<mods:name id="78F88A97A257029436A2524FC70A7BD5" type="personal">
<mods:role id="7CF927C31FB5E75D61F92635A39019F9">
<mods:roleTerm id="D667189E5102B87CAB01C8D035AD3FE5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD43CC2AA731E04A3B276615364ED86E">Will, Kipling</mods:namePart>
<mods:nameIdentifier id="8D5E8BA07E170C66D03BE36D52FB0571" type="ORCID">https://orcid.org/0000-0002-7056-9011</mods:nameIdentifier>
<mods:affiliation id="2C186FD34A92146661E930E7BA73670D">University of California Berkeley, Essig Museum of Entomology. Berkeley, California, USA</mods:affiliation>
<mods:nameIdentifier id="7F00A7815358FBEB97058AFF6BCC3B1D" type="email">kipwill@berkeley.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="4414F084FB54D6E55E2C6D2CA27876ED" type="personal">
<mods:role id="2BB8A42CE5AC9F174041178CCEF98947">
<mods:roleTerm id="4BA7BD4864BF1A376ECDB4530939475E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E164CC98B6C6C82A235F5BE3CEC0A33F">Gueorguiev, Borislav</mods:namePart>
<mods:affiliation id="E1FEE6AC8BDD7149A05170A62C422EBF">National Museum of Natural History, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:typeOfResource id="042E347932A4549D06AF0E1927401A0B">text</mods:typeOfResource>
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<mods:title id="A1FB9D6ED51561BFB0D5F08F5EBB0F78">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="E965A5B069C40C8E89164A1FB6AB39B4">
<mods:date id="90638F7EF026957FE5F2DBD3C39CF2DE">2021</mods:date>
<mods:detail id="4D45F634C36C69F6995C4570DC93CAF4" type="pubDate">
<mods:number id="FA4413075009AD9809C3463362512628">2021-06-16</mods:number>
</mods:detail>
<mods:detail id="7DCEAE63326519750A847CE319AB49E5" type="volume">
<mods:number id="D8C5208C623FB71B14B29ED424C2BD7B">1044</mods:number>
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<mods:start id="4F5B1C6B95A1FB3D599B5D7AF78A5A8C">375</mods:start>
<mods:end id="4DEF001382C87F23E06F498E06401D65">425</mods:end>
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<mods:location id="7E147E1BAB9BC5EE4CBAAB2E3E52F492">
<mods:url id="746EEA916EBB452B3C2872990298507E">http://dx.doi.org/10.3897/zookeys.1044.63775</mods:url>
</mods:location>
<mods:classification id="C419F58316BF514209FCC75056C3CD43">journal article</mods:classification>
<mods:identifier id="D7AADDAE8177C200649E81C7A8F72A91" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.63775</mods:identifier>
<mods:identifier id="E766DF7B63AF8E975FCA0A02F708DBD4" type="Pensoft-Pub">1313-2970-1044-375</mods:identifier>
<mods:identifier id="444B820E55224DB6E98756FD5AA7144A" type="ZooBank">D50CC77C2E6941FDA9BD395B025C43AE</mods:identifier>
<mods:identifier id="D6B31A71E955C67B634EA50374E736FB" type="Pensoft-UUID">EBD9E81E5EC55AA491952682787793A4</mods:identifier>
</mods:mods>
<treatment id="8FD37B09F28452A9B8457C6BBA2ADD1B" ID-GBIF-Taxon="182754028" LSID="urn:lsid:zoobank.org:act:6FDFFBC6-C84B-4070-AEC5-D4D39F730356" httpUri="http://treatment.plazi.org/id/8FD37B09F28452A9B8457C6BBA2ADD1B" lastPageNumber="375" pageId="0" pageNumber="375" scope_family="Carabidae" scope_order="Coleoptera" scope_subTribe="Thryptocerina" scope_tribe="Oodini">
<subSubSection id="D66C3914DE16F63CC257E1E45CF7248A" pageId="0" pageNumber="375" type="nomenclature">
<paragraph id="B16A3BA3676EF5A612060AF444E01BFA" pageId="0" pageNumber="375">
<taxonomicName id="9BDBB880C34487079A241E504B09701C" ID-CoL="8M8N2" LSID="http://zoobank.org/6FDFFBC6-C84B-4070-AEC5-D4D39F730356" authority="Will &amp; Guéorguiev, 2021" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus amieuensis" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="amieuensis" status="sp. nov.">Coptocarpus amieuensis</taxonomicName>
<taxonomicNameLabel id="373D1638DE49E4ECB05706DD1E2269DB" pageId="0" pageNumber="375">sp. nov.</taxonomicNameLabel>
<figureCitation id="837DA14DC92D5EA2D047DEA17CD1A696" captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Dorsal habitus of 18 C. erwini sp. nov. 19 holotype of Coptocarpus cyllodinus, with labels 20 C. amieuensis sp. nov. 21 C. lescheni sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.63775.figures18-21" httpUri="https://binary.pensoft.net/fig/554053" pageId="0" pageNumber="375">Figs 20</figureCitation>
<figureCitation id="57340CE1FABA230792C80056C0E0109D" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">, 25</figureCitation>
<figureCitation id="A962D36192FE5082C5A8202768B91D29" captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">, 35</figureCitation>
<figureCitation id="859CBBADA7B27A4E79F0187B7A43BC60" captionStart="Figures 52, 53" captionStartId="F15" captionText="Figures 52, 53. Maps of localities of New Caledonian Oodini." figureDoi="10.3897/zookeys.1044.63775.figures52-53" httpUri="https://binary.pensoft.net/fig/554062" pageId="0" pageNumber="375">, 53</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="EE17E9068B6A9F98D8068527D2365A0B" pageId="0" pageNumber="375" type="materials_examined">
<paragraph id="330332C143A0D971A5714E56DCBF6400" pageId="0" pageNumber="375">Material examined.</paragraph>
<paragraph id="046E8CF329AA3F2C93A523E722BAE69C" pageId="0" pageNumber="375">
<materialsCitation id="AF1B595AB9635E5CE003A5761760E28A" collectingDate="2007-03-04" collectingDateMax="2007-03-16" collectingDateMin="2007-03-04" country="NEW CALEDONIA" elevation="440" latitude="-21.56" location="New Caledonia" longLatPrecision="21" longitude="165.75555" specimenCount="♂" typeStatus="Holotype">
<emphasis id="2A43945B62EC0C24397C87318AE4CEB9" bold="true" italics="true" pageId="0" pageNumber="375">
<typeStatus id="21D55D541D5A3E25C3E0216E6CE51698">Holotype</typeStatus>
</emphasis>
:
<location id="EBDA7C800FB390BF335803A8294A14E3" LSID="urn:lsid:plazi:treatment:8FD37B09F28452A9B8457C6BBA2ADD1B:EBDA7C800FB390BF335803A8294A14E3" country="NEW CALEDONIA" latitude="-21.56" longLatPrecision="21" longitude="165.75555" name="New Caledonia">New Caledonia</location>
•
<specimenCount id="3261EFB449A3B89798FC719B7C2FBEE8">♂</specimenCount>
; &quot;
<geoCoordinate id="B69EACB00215B85C96802B9FF057CA5B" degrees="21" direction="south" minutes="33" orientation="latitude" precision="15" seconds="36" value="-21.56">21°33'36&quot;S</geoCoordinate>
/
<geoCoordinate id="682B55BAB753CA16CEB34AF6E595D191" degrees="165" direction="east" minutes="45" orientation="longitude" precision="15" seconds="20" value="165.75555">165°45'20&quot;E</geoCoordinate>
<collectingCountry id="40D3AF7D7F97D2CCE3A18FEE3DAFEC9F" name="New Caledonia">NEW CALEDONIA</collectingCountry>
: Prov Sud. Col
<normalizedToken id="28E7D036101F54BC5ED88259292F6948" originalValue="d’Amieu">d'Amieu</normalizedToken>
,
<quantity id="3DFFA6DD062964EE658FDC600F690D03" metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">
<elevation id="8CAADD4F1C612165A92B86C9AF7DBF4A" metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">440m</elevation>
</quantity>
el,
<collectingDate id="10C2EF1BA25E4E409D8692259A0400A1" value="2007-03-04" valueMax="2007-03-16" valueMin="2007-03-04">4-16.iii.2007</collectingDate>
headlamp search&quot;//
<normalizedToken id="EA58C56E1ED12DA8BD5539F9365DF4AF" originalValue="“EMEC1139863”">&quot;EMEC1139863&quot;</normalizedToken>
</materialsCitation>
<typeStatus id="D907607ECFFBC2EB8C921E2C0D7B3CBA">Holotype</typeStatus>
pinned, with genitalia in a separate microvial. Specimen relatively well-preserved, but with missing palpomeres, left antennomeres 2-11, right tarsomeres 1-5, left and right mesotibiae and mesotarsomeres 1-5, and left and right metatarsomeres 1-5. Source collection EMEC, deposited MNHN. Type locality as given on label.
</paragraph>
</subSubSection>
<subSubSection id="68E3DDCC799FF6187F63BA698727FDD2" pageId="0" pageNumber="375" type="diagnosis">
<paragraph id="9C955EC6026A270DF03CEBA67087FB39" pageId="0" pageNumber="375">Diagnosis.</paragraph>
<paragraph id="189C24C984D2CC74B59002DF3BA187F0" pageId="0" pageNumber="375">
Most similar to
<taxonomicName id="3B0A06092CE4D2B80FE722F342E4BB28" lsidName="C. magnus" pageId="0" pageNumber="375" rank="species" species="magnus">
<emphasis id="61827FC988C651CCA29E8C31731624C1" italics="true" pageId="0" pageNumber="375">C. magnus</emphasis>
</taxonomicName>
. However, the latter species has striae 2-5 shallowly impressed from near the elytra base to the apex (vs. 2-4 impressed only in the apical 1/3, and 5 in apical 1/2) and apical lamella of the median lobe more widely rounded at tip (Figs
<figureCitation id="EAC58CD944A662F088A6A68DD1CB29F5" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">25</figureCitation>
,
<figureCitation id="C2DBC6746FEE325C68DD1B092A280D85" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">26</figureCitation>
). Also resembles
<taxonomicName id="3DDC9C7DB66E52FD9755696942F0D238" lsidName="C. cyllodinus" pageId="0" pageNumber="375" rank="species" species="cyllodinus">
<emphasis id="76BF8C1283FBC00CD89D5DFEB819851B" italics="true" pageId="0" pageNumber="375">C. cyllodinus</emphasis>
</taxonomicName>
but in
<taxonomicName id="29B71C211EECE92452923D7625E18BCC" lsidName="C. amieuensis" pageId="0" pageNumber="375" rank="species" species="amieuensis">
<emphasis id="D284DDE040C4B77F94D8D7E338BDD18F" italics="true" pageId="0" pageNumber="375">C. amieuensis</emphasis>
</taxonomicName>
the eyes are conical, head lacking a notch deep enough to receive the antenna ventral to the eye and the male metafemur does not have a prominent, ventral tooth. The male aedeagus is distinctly different (Fig.
<figureCitation id="6CEE844C0F34EDA96AD1CB5558CE09A0" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D3AB967317CBCCF3243072531810DDDB" pageId="0" pageNumber="375" type="description">
<paragraph id="2F6FFC46DA218A263687B07FA8506AC3" pageId="0" pageNumber="375">Description.</paragraph>
<paragraph id="5626BC2362D9403B5D14EF1E7722B44B" pageId="0" pageNumber="375">
<emphasis id="3AA940799F4427DAB457E041DF4EE7EE" bold="true" pageId="0" pageNumber="375">
<emphasis id="1CD155E5ED661A4EA774C5AFBD6EEB9A" italics="true" pageId="0" pageNumber="375">Habitus</emphasis>
.
</emphasis>
Small sized for oodines BL: 7.20 mm, BW: 3.50 mm, with ovate, moderately convex body (Fig.
<figureCitation id="568DDD93407736F5BC5B9DE3C7E72AE0" captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Dorsal habitus of 18 C. erwini sp. nov. 19 holotype of Coptocarpus cyllodinus, with labels 20 C. amieuensis sp. nov. 21 C. lescheni sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.63775.figures18-21" httpUri="https://binary.pensoft.net/fig/554053" pageId="0" pageNumber="375">20</figureCitation>
).
<emphasis id="0BF9CED4A4BBB299A2CB6597103D7672" bold="true" pageId="0" pageNumber="375">
<emphasis id="02CAE3F5B5B59B4B8C9831A31FE58148" italics="true" pageId="0" pageNumber="375">Color and luster</emphasis>
.
</emphasis>
Dorsal and ventral surface primarily black, with pronotum sides brownish black, fore part of head and femora dark reddish brown, antennomeres 1-3 and tibiae reddish brown, palpi, antennomeres 4-12 and tarsomeres yellowish brown. Integument moderately glossy, without spectral iridescence.
<emphasis id="6B76F7B52A1746C17FD4E2107711E5D7" bold="true" pageId="0" pageNumber="375">
<emphasis id="74F5A9F8F0F7344F05BD94BCD01EA54B" italics="true" pageId="0" pageNumber="375">Microsculpture and punctation</emphasis>
.
</emphasis>
Head, pronotum and elytra with isodiametric meshes well impressed, prosternum in front and abdominal ventrites in middle with slightly transverse meshes, prosternal process with isodiametric meshes less impressed than on dorsal surface, rest of ventral surface with scarcely-visible sculpticells or sculpticells not apparent. Micropunctation evident on head, pronotum, prosternum medially (including prosternal process) and medial parts of abdominal ventrites, more or less uniform, lacking on remaining part of ventral surface; abdominal ventrite 1 with some large punctures at sides, ventrites 2-5 at sides moderately rugose.
<emphasis id="1A095B8B7C1666F15396CAE96EA4C1A6" bold="true" pageId="0" pageNumber="375">
<emphasis id="B559F212BB5E9F06CCC26A81ED4CEB36" italics="true" pageId="0" pageNumber="375">Chaetotaxy</emphasis>
.
</emphasis>
Labrum with six setae distributed more or less evenly across width each in its own socket. Clypeal setae present. Supraorbital setae absent. Elytron without discal setiferous punctures. Abdominal ventrites 1-5 without ambulatory setae.
<emphasis id="8FDE1E82FDD166AE87D6610C71E738C4" bold="true" pageId="0" pageNumber="375">Head.</emphasis>
Less than 2 1/2
<normalizedToken id="393F002E1E661FE58EFA5770DFCE6A8D" originalValue="×">x</normalizedToken>
as wide as pronotum (Suppl. material 2: Table S1). Eye relatively large, moderately protruded, EyW/HW: 1.47. Labrum anterior margin straight. Frontoclypeal suture present medially, almost obliterated at sides. Antenna moderately long, with last segment not exceeding pronotal base and pubescence starting from segment 4. Mentum tooth with paramedial border distinct.
<emphasis id="CB6611BD649496EE526AB386EB73896C" bold="true" pageId="0" pageNumber="375">
<emphasis id="C12EA425D4CDD70F89EA339802057E05" italics="true" pageId="0" pageNumber="375">Thorax</emphasis>
.
</emphasis>
Pronotum ~ 1 2/3 wider than long (PW/PL: 1.70); width at apex less than 2
<normalizedToken id="8501C752CE461B788D92810C76E90FA1" originalValue="×">x</normalizedToken>
at widest point (PW/PA: 2.16). Disc with middle line very fine, without apical transverse impression; anterior angles slightly, widely rounded at tips, anterior margin concave, anterior submarginal sulcus present, interrupted in middle. Prosternal process rhomboidal, finely bordered at apex, not bordered between coxae. Mesosternum anteriorly with two prominent, symmetrical tubercles separated by a space smaller or equal to diameter of one of them. Metepisternum wider than long, with lateral margin barely convex, coadunation with epipleuron long, extended along most of length.
<emphasis id="2ACDF6445D809AD201AAE7E8ACC4EF82" bold="true" pageId="0" pageNumber="375">Elytra.</emphasis>
Slightly longer than wide (EL/EW: 1.11). Basal margin forming distinct denticle at shoulder, ended medially just before level of parascutellar puncture. Humeral submarginal carina present. Apical sinuation shallow. Parascutellar striole absent, striae 1-5 present at apex, ended well before basal margin; striae 6 and 7 well impressed throughout, 7 reaching basal margin, 6 ended near but not reaching margin. Elytral intervals 1-6 where evident at apex, convex, interval 7 slightly convex in anterior 1/2, clearly convex in posterior 1/2, interval 8 convex throughout, from 2-3
<normalizedToken id="D77C989F266E7C92B14202311BFBAF29" originalValue="×">x</normalizedToken>
narrower than interval 7. Granulation in marginal furrow evident in first 1/5 and last 3/5, lacking in second 1/5.
<emphasis id="320879EDD6AA4F7518F67286F66B66C0" bold="true" pageId="0" pageNumber="375">
<emphasis id="D23324571733BA9E15F4659D55066194" italics="true" pageId="0" pageNumber="375">Legs</emphasis>
.
</emphasis>
Male protarsomeres 1-3 asymmetrically expanded, beneath with adhesive setae (tarsomeres 2 and 3 fully covered, tarsomere 1 covered only on apical 1/4), each of them transverse, wider than long.
<emphasis id="CC3C43287F9109B0D48383F21DD32021" bold="true" pageId="0" pageNumber="375">
<emphasis id="CC930F78BF1BB31BDA685FE30A864CC6" italics="true" pageId="0" pageNumber="375">Female genitalia</emphasis>
.
</emphasis>
Female specimens unknown.
<emphasis id="486EA32CAA71169DEC85A8E679541022" bold="true" pageId="0" pageNumber="375">
<emphasis id="06406640B17FBCA652E3BCB90D860A04" italics="true" pageId="0" pageNumber="375">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view long, curved ventrally, with rather thin apex (Fig.
<figureCitation id="3B4662E303F4303213C0BEC89A753747" captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">35</figureCitation>
); lobe in dorsal view, with apical lamella well bent to left, with left side clearly concave and right side slightly sinuate (Fig.
<figureCitation id="A68B306EBB39B1DB8FFC8D76CF3403EE" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">25</figureCitation>
); basal bulb rounded dorsally; sclerotized portion of the endophallus with three sclerites, most clearly visible from ventral side.
</paragraph>
</subSubSection>
<subSubSection id="17CA4EA971416E01158E6A52ED47A801" pageId="0" pageNumber="375" type="etymology">
<paragraph id="5F0ADBD583B973302ED90F68F12574FA" pageId="0" pageNumber="375">Etymology.</paragraph>
<paragraph id="B7DB8980E37C6136B00D3926F77BE8CC" pageId="0" pageNumber="375">
The specific epithet
<taxonomicName id="3BD4529851105CF793B386ECD58A5601" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus amieuensis" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="amieuensis">
<emphasis id="618D9514B847AC89C5F16197564F32CB" italics="true" pageId="0" pageNumber="375">Coptocarpus amieuensis</emphasis>
</taxonomicName>
is based on the type locality Col
<normalizedToken id="34A6F6201A851DB9986410548F85FDD6" originalValue="d’Amieu">d'Amieu</normalizedToken>
and is treated as an adjective.
</paragraph>
</subSubSection>
</treatment>
</document>
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