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<document id="F4B15609BE99CD1818EB34BF39B39836" ID-DOI="10.11646/zootaxa.3640.1.6" ID-GBIF-Dataset="85bebfe2-a082-48ef-b84f-67bdf1b068ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="283657" ID-ZooBank="2392C313-68F6-4FB0-B798-6D8CAA4BA04C" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460592819764" checkinUser="plazi" docAuthor="Silveira, Luiz Felipe Lima Da &amp; Mermudes, José Ricardo M." docDate="2013" docId="5C29879AFFD7FFD8FF3B5AA0CC30FD01" docLanguage="en" docName="zt03640p087.pdf" docOrigin="Zootaxa 3640 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Memoan ciceroi Silveira &amp; Mermudes, sp. nov." docType="treatment" docVersion="6" lastPageNumber="86" masterDocId="A010FFE2FFD2FFDFFFAC5C29C83BFFA6" masterDocTitle="Memoan ciceroi gen. et sp. nov., a remarkable new firefly genus and species from the Atlantic Rainforest (Coleoptera: Lampyridae)" masterLastPageNumber="87" masterPageNumber="79" pageNumber="84" updateTime="1698326268522" updateUser="plazi">
<mods:mods id="B28F878C1F8A5AB95C27F3F50F5DDD7D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5BFAC443DC8983331AB6BB75BB6422C0">Memoan ciceroi gen. et sp. nov., a remarkable new firefly genus and species from the Atlantic Rainforest (Coleoptera: Lampyridae)</mods:title>
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<mods:name id="86C817967302D729901633F0A3AFCAC3" type="personal">
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<mods:namePart id="C4AEACFA2144200CAFD2FB7BD051FFB5">Silveira, Luiz Felipe Lima Da</mods:namePart>
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<mods:namePart id="68BE63B2CB2ACAA3B90EFF1F818E7512">Mermudes, José Ricardo M.</mods:namePart>
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<mods:title id="61C210BB603CEC4AC25F5A2D5FCC9541">Zootaxa</mods:title>
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<mods:date id="BA6447C2E511E6C8D88227A24458C28B">2013</mods:date>
<mods:detail id="AB86BE58F08E9D2B84A1E84F8CFC8F42" type="volume">
<mods:number id="0673F1DE4E4106B77C6C348F60ECE6F4">3640</mods:number>
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<mods:identifier id="8608196D929289B3D62C17A34FE57E66" type="DOI">10.11646/zootaxa.3640.1.6</mods:identifier>
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<treatment id="5C29879AFFD7FFD8FF3B5AA0CC30FD01" ID-DOI="http://doi.org/10.5281/zenodo.5670104" ID-GBIF-Taxon="127663844" ID-Zenodo-Dep="5670104" LSID="urn:lsid:plazi:treatment:5C29879AFFD7FFD8FF3B5AA0CC30FD01" httpUri="http://treatment.plazi.org/id/5C29879AFFD7FFD8FF3B5AA0CC30FD01" lastPageId="7" lastPageNumber="86" pageId="5" pageNumber="84">
<subSubSection id="9C9A6507FFD7FFDAFF3B5AA0C922F965" pageId="5" pageNumber="84" type="nomenclature">
<paragraph id="D43F368CFFD7FFDAFF3B5AA0CAEEF904" blockId="5.[151,725,1672,1731]" box="[151,725,1672,1699]" pageId="5" pageNumber="84">
<heading id="8F7781E0FFD7FFDAFF3B5AA0CAEEF904" bold="true" box="[151,725,1672,1699]" fontSize="11" level="1" pageId="5" pageNumber="84" reason="1">
<emphasis id="E6F4EA9EFFD7FFDAFF3B5AA0CAEEF904" bold="true" box="[151,725,1672,1699]" pageId="5" pageNumber="84">
<taxonomicName id="13804D0FFFD7FFDAFF3B5AA0CA4BF904" ID-CoL="3ZP9B" authority="Silveira &amp; Mermudes" authorityName="Silveira &amp; Mermudes" box="[151,624,1672,1699]" class="Insecta" family="Lampyridae" genus="Memoan" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="ciceroi" status="sp. nov.">
<emphasis id="E6F4EA9EFFD7FFDAFF3B5AA0C960F905" bold="true" box="[151,347,1673,1699]" italics="true" pageId="5" pageNumber="84">Memoan ciceroi</emphasis>
Silveira &amp; Mermudes
</taxonomicName>
<taxonomicNameLabel id="FDC757E5FFD7FFDAFDD45AA1CAEEF904" box="[632,725,1672,1698]" pageId="5" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D43F368CFFD7FFDAFF3B5A82C922F965" blockId="5.[151,725,1672,1731]" box="[151,281,1707,1731]" pageId="5" pageNumber="84">
(
<figureCitation id="4CBB2A09FFD7FFDAFF335A82C92BF965" box="[159,272,1707,1731]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 5" captionStart-2="FIGURES 6 – 8" captionStart-3="FIGURES 9 – 13" captionStartId-0="3.[151,264,1823,1846]" captionStartId-1="4.[151,264,1934,1957]" captionStartId-2="5.[151,264,1514,1537]" captionStartId-3="6.[151,264,1877,1900]" captionTargetBox-0="[174,1391,223,1780]" captionTargetBox-1="[200,1390,230,1909]" captionTargetBox-2="[187,1397,226,1452]" captionTargetBox-3="[201,1366,307,1828]" captionTargetId-0="figure@3.[151,1436,195,1830]" captionTargetId-1="figure@4.[179,1408,181,1935]" captionTargetId-2="figure@5.[151,1436,193,1493]" captionTargetId-3="figure@6.[183,1410,269,1864]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 1 – 3. Memoan ciceroi gen. et sp. nov., holotype male from Estação Biológica Santa Lúcia, Santa Teresa, Espírito Santo, Brazil: 1, habitus dorsal; 2, habitus ventral; 3, head, frontal view, which shows the fusion of the fronto-clypeus and the labrum." captionText-1="FIGURES 4 – 5. Memoan ciceroi gen. nov. et sp. nov.: 4, head and thorax, ventral. Note the subcontiguous eyes and the conspicuous sutures between mesosternum and mesanepisternum, and between the mesanepisternum and mesanepimerum. 5, Mouthparts. Note the one-segmented labial palps." captionText-2="FIGURES 6 – 8. Memoan ciceroi gen. nov. et sp. nov.: 6, apical antennomeres. Note the fusion between the two apical-most antennomeres. 7, pronotum and elytral base, dorsal. Note the deep punctuation all along the pronotum, scutellum and elytra; and the pronotal parassagital, posterior elevations. 8, right wing." captionText-3="FIGURES 9 – 13. Memoan ciceroi gen. nov. et sp. nov.: 9, abdomen, left lateral; 10 abdomen, ventral (anterior end to left of page). 11, Detail of the abdomen, dorsal (arrows shows the dorsal spiracles); 12, aedeagal sheath: sintergite, dorsal and sternite IX, ventral (anterior ends to bottom of page). 13, aedeagus, dorsal." httpUri-0="https://zenodo.org/record/283658/files/figure.png" httpUri-1="https://zenodo.org/record/283659/files/figure.png" httpUri-2="https://zenodo.org/record/283660/files/figure.png" httpUri-3="https://zenodo.org/record/283661/files/figure.png" pageId="5" pageNumber="84">Figs 1–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9C9A6507FFD7FFDAFF3B5ADDC9B6F889" pageId="5" pageNumber="84" type="etymology">
<paragraph id="D43F368CFFD7FFDAFF3B5ADDC9B6F889" blockId="5.[151,1436,1779,2020]" pageId="5" pageNumber="84">
<emphasis id="E6F4EA9EFFD7FFDAFF3B5ADDC925F8AA" bold="true" box="[151,286,1780,1804]" pageId="5" pageNumber="84">Etymology.</emphasis>
The species is named after the american entomologist Joseph Cicero, whose contribution to firefly research is admirable.
</paragraph>
</subSubSection>
<subSubSection id="9C9A6507FFD7FFD8FF6B5B12CCA5FDB1" lastPageId="7" lastPageNumber="86" pageId="5" pageNumber="84" type="description">
<paragraph id="D43F368CFFD7FFDAFF6B5B12CC56F83D" blockId="5.[151,1436,1779,2020]" pageId="5" pageNumber="84">
<emphasis id="E6F4EA9EFFD7FFDAFF6B5B12CA10F8F2" bold="true" box="[199,555,1851,1876]" pageId="5" pageNumber="84">Description (adult, male, n=2)</emphasis>
. Body length:
<quantity id="13789B69FFD7FFDAFD7C5B12CB4DF8F5" box="[720,886,1851,1875]" metricMagnitude="-3" metricUnit="m" metricValue="5.08" metricValueMax="5.2" metricValueMin="4.96" pageId="5" pageNumber="84" unit="mm" value="5.08" valueMax="5.2" valueMin="4.96">4.96–5.20 mm</quantity>
; pronotal length:
<quantity id="13789B69FFD7FFDAFB955B12CCE4F8F5" box="[1081,1247,1851,1875]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="9.0" metricValueMin="7.5" pageId="5" pageNumber="84" unit="mm" value="0.825" valueMax="0.9" valueMin="0.75">0.75–0.90 mm</quantity>
; greater pronotal width:
<quantity id="13789B69FFD7FFDAFF455B76C9A8F8D1" box="[233,403,1887,1911]" metricMagnitude="-3" metricUnit="m" metricValue="1.245" metricValueMax="1.3" metricValueMin="1.19" pageId="5" pageNumber="84" unit="mm" value="1.245" valueMax="1.3" valueMin="1.19">1.19–1.30 mm</quantity>
; elytral length
<quantity id="13789B69FFD7FFDAFDE95B76CAD5F8D1" box="[581,750,1887,1911]" metricMagnitude="-3" metricUnit="m" metricValue="4.23" metricValueMax="4.3" metricValueMin="4.16" pageId="5" pageNumber="84" unit="mm" value="4.23" valueMax="4.3" valueMin="4.16">4.16–4.30 mm</quantity>
; elytral width
<quantity id="13789B69FFD7FFDAFC365B76CC78F8D1" box="[922,1091,1887,1911]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="5" pageNumber="84" unit="mm" value="1.275" valueMax="1.3" valueMin="1.25">1.25–1.30 mm</quantity>
. Integument overall brown to pale-yellow, including appendices, abdominal segments translucent from eight forward.
</paragraph>
<paragraph id="D43F368CFFD7FFD9FF6B5B81CD2AFF51" blockId="5.[151,1436,1779,2020]" lastBlockId="6.[151,1436,151,247]" lastPageId="6" lastPageNumber="85" pageId="5" pageNumber="84">
<emphasis id="E6F4EA9EFFD7FFDAFF6B5B81C93CF866" bold="true" box="[199,263,1960,1984]" pageId="5" pageNumber="84">Head</emphasis>
. (
<figureCitation id="4CBB2A09FFD7FFDAFE8F5B8EC9A1F819" box="[291,410,1959,1983]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 5" captionStart-2="FIGURES 6 – 8" captionStartId-0="3.[151,264,1823,1846]" captionStartId-1="4.[151,264,1934,1957]" captionStartId-2="5.[151,264,1514,1537]" captionTargetBox-0="[174,1391,223,1780]" captionTargetBox-1="[200,1390,230,1909]" captionTargetBox-2="[187,1397,226,1452]" captionTargetId-0="figure@3.[151,1436,195,1830]" captionTargetId-1="figure@4.[179,1408,181,1935]" captionTargetId-2="figure@5.[151,1436,193,1493]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1 – 3. Memoan ciceroi gen. et sp. nov., holotype male from Estação Biológica Santa Lúcia, Santa Teresa, Espírito Santo, Brazil: 1, habitus dorsal; 2, habitus ventral; 3, head, frontal view, which shows the fusion of the fronto-clypeus and the labrum." captionText-1="FIGURES 4 – 5. Memoan ciceroi gen. nov. et sp. nov.: 4, head and thorax, ventral. Note the subcontiguous eyes and the conspicuous sutures between mesosternum and mesanepisternum, and between the mesanepisternum and mesanepimerum. 5, Mouthparts. Note the one-segmented labial palps." captionText-2="FIGURES 6 – 8. Memoan ciceroi gen. nov. et sp. nov.: 6, apical antennomeres. Note the fusion between the two apical-most antennomeres. 7, pronotum and elytral base, dorsal. Note the deep punctuation all along the pronotum, scutellum and elytra; and the pronotal parassagital, posterior elevations. 8, right wing." httpUri-0="https://zenodo.org/record/283658/files/figure.png" httpUri-1="https://zenodo.org/record/283659/files/figure.png" httpUri-2="https://zenodo.org/record/283660/files/figure.png" pageId="5" pageNumber="84">FIGS 1–6</figureCitation>
) Antennomeres densely and uniformly hairy. Scape nearly twice as long as pedicel, constrained on basal half, enlarging to apex. Pedicel as long as wide. Antennomere III longer than pedicel; III–IX laterally compressed, enlarging towards their apices; III–VI slightly decrease in length; VII–IX subequal in length to VI; X and XI connate; X + XI longer than IX. Maxillary palpomeres cylindrical, I–III transverse, I and III subequal and shorter than II; IV acuminate and membranous at apex, longer than the sum of II and III.
</paragraph>
<caption id="80FF6604FFD4FFD9FF3B5B7CCB01F80F" httpUri="https://zenodo.org/record/283661/files/figure.png" pageId="6" pageNumber="85" targetBox="[201,1366,307,1828]" targetPageId="6">
<paragraph id="D43F368CFFD4FFD9FF3B5B7CCB01F80F" blockId="6.[151,1436,1877,1961]" pageId="6" pageNumber="85">
<emphasis id="E6F4EA9EFFD4FFD9FF3B5B7CC973F8CD" bold="true" box="[151,328,1877,1900]" pageId="6" pageNumber="85">FIGURES 9–13.</emphasis>
<taxonomicName id="13804D0FFFD4FFD9FEFC5B7EC9CEF8CA" box="[336,501,1879,1900]" class="Insecta" family="Lampyridae" genus="Memoan" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="ciceroi" status="gen. nov.">
<emphasis id="E6F4EA9EFFD4FFD9FEFC5B7EC9CEF8CA" box="[336,501,1879,1900]" italics="true" pageId="6" pageNumber="85">Memoan ciceroi</emphasis>
</taxonomicName>
<emphasis id="E6F4EA9EFFD4FFD9FE515B7FCAEAF8CD" bold="true" box="[509,721,1878,1900]" pageId="6" pageNumber="85">
<taxonomicNameLabel id="FDC757E5FFD4FFD9FE515B7FCA66F8CD" box="[509,605,1878,1899]" pageId="6" pageNumber="85" rank="genus">gen. nov.</taxonomicNameLabel>
<emphasis id="E6F4EA9EFFD4FFD9FDC95B7ECA4CF8CA" bold="true" box="[613,631,1879,1900]" italics="true" pageId="6" pageNumber="85">et</emphasis>
sp. nov.
</emphasis>
: 9, abdomen, left lateral; 10 abdomen, ventral (anterior end to left of page). 11, Detail of the abdomen, dorsal (arrows shows the dorsal spiracles); 12, aedeagal sheath: sintergite, dorsal and sternite IX, ventral (anterior ends to bottom of page). 13, aedeagus, dorsal.
</paragraph>
</caption>
<paragraph id="D43F368CFFD5FFD8FF6B5CB1C93EFE99" blockId="7.[151,1436,151,680]" pageId="7" pageNumber="86">
<emphasis id="E6F4EA9EFFD5FFD8FF6B5CB1C91CFF16" bold="true" box="[199,295,152,176]" pageId="7" pageNumber="86">Thorax.</emphasis>
(FIGS 4,7) Anterior margin of pronotum with two feebly para-sagital sinuosities. Prosternum medially glabrous, with many lateral bristles. Meso-scutellum sub-pentagonal, with dense and deep punctures; lateral margins convergent posteriad, posterior margin emarginate; with sparse bristles in the anterior region of the punctures. Hypomeron with deep punctures, except for the central region. Mesosternum projecting and rounded posteriad.
</paragraph>
<paragraph id="D43F368CFFD5FFD8FF6B5D62CCA5FDB1" blockId="7.[151,1436,151,680]" pageId="7" pageNumber="86">
<emphasis id="E6F4EA9EFFD5FFD8FF6B5D62C978FEC2" bold="true" box="[199,323,331,356]" pageId="7" pageNumber="86">Abdomen.</emphasis>
(
<figureCitation id="4CBB2A09FFD5FFD8FEFE5D65C9EAFEC2" box="[338,465,332,356]" captionStart="FIGURES 9 – 13" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[201,1366,307,1828]" captionTargetId="figure@6.[183,1410,269,1864]" captionTargetPageId="6" captionText="FIGURES 9 – 13. Memoan ciceroi gen. nov. et sp. nov.: 9, abdomen, left lateral; 10 abdomen, ventral (anterior end to left of page). 11, Detail of the abdomen, dorsal (arrows shows the dorsal spiracles); 12, aedeagal sheath: sintergite, dorsal and sternite IX, ventral (anterior ends to bottom of page). 13, aedeagus, dorsal." httpUri="https://zenodo.org/record/283661/files/figure.png" pageId="7" pageNumber="86">FIGS 9–13</figureCitation>
) Abdominal segments translucent from eight forward; lanterns occupying most of the venrite area. Pygidium with median lobe slightly acuminate, lateral lobes acute and projected.
<emphasis id="E6F4EA9EFFD5FFD8FB4D5D46C936FE0A" bold="true" pageId="7" pageNumber="86">Terminalia and aedeagus:</emphasis>
(
<figureCitation id="4CBB2A09FFD5FFD8FE8F5DBDC98CFE0A" box="[291,439,404,428]" captionStart="FIGURES 9 – 13" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[201,1366,307,1828]" captionTargetId="figure@6.[183,1410,269,1864]" captionTargetPageId="6" captionText="FIGURES 9 – 13. Memoan ciceroi gen. nov. et sp. nov.: 9, abdomen, left lateral; 10 abdomen, ventral (anterior end to left of page). 11, Detail of the abdomen, dorsal (arrows shows the dorsal spiracles); 12, aedeagal sheath: sintergite, dorsal and sternite IX, ventral (anterior ends to bottom of page). 13, aedeagus, dorsal." httpUri="https://zenodo.org/record/283661/files/figure.png" pageId="7" pageNumber="86">FIGS 12–13</figureCitation>
) sintergite half the sternite IX length, slightly asymmetrical; sternite IX narrowly prolonged anteriorly. Aedeagus almost entirely membranous. Phallobase symmetrical. Parameres basally constricted, with two thirds of phallus length. Apex of parameres and phallus rounded. Phallus slightly wider than a paramere. Ventral plate of phallus present, sides subparallel, slightly constrained medially.
</paragraph>
</subSubSection>
<subSubSection id="9C9A6507FFD5FFD8FF6B5E0DCC30FD01" pageId="7" pageNumber="86" type="materials_examined">
<paragraph id="D43F368CFFD5FFD8FF6B5E0DCC30FD01" blockId="7.[151,1436,151,680]" pageId="7" pageNumber="86">
<emphasis id="E6F4EA9EFFD5FFD8FF6B5E0DC906FD9A" bold="true" box="[199,317,548,572]" pageId="7" pageNumber="86">
<typeStatus id="0B3B882EFFD5FFD8FF6B5E0DC903FD9A" box="[199,312,548,572]" pageId="7" pageNumber="86" type="holotype">Holotype</typeStatus>
:
</emphasis>
adult, male from
<collectingCountry id="AC97761CFFD5FFD8FDB45E0DCA64FD9A" box="[536,607,548,572]" name="Brazil" pageId="7" pageNumber="86">Brazil</collectingCountry>
, Espírito Santo State, Estação Biológica da
<collectingCountry id="AC97761CFFD5FFD8FBC15E0DCCC2FD9A" box="[1133,1273,548,572]" name="Saint Lucia" pageId="7" pageNumber="86">Santa Lúcia</collectingCountry>
, Santa Teresa Municipality.
<date id="A03E104CFFD5FFD8FE945E6EC980FDF9" box="[312,443,583,607]" pageId="7" pageNumber="86" value="2011-04-02">02/IV/2011</date>
, Silveira, L; Dumas, L. Nessimian, J.
<emphasis id="E6F4EA9EFFD5FFD8FC2A5E61CB8DFDC6" box="[902,950,584,608]" italics="true" pageId="7" pageNumber="86">leg..</emphasis>
<emphasis id="E6F4EA9EFFD5FFD8FC6E5E61CC14FDC6" bold="true" box="[962,1071,584,608]" pageId="7" pageNumber="86">
<typeStatus id="0B3B882EFFD5FFD8FC6E5E61CC14FDC6" box="[962,1071,584,608]" pageId="7" pageNumber="86" type="paratype">Paratype</typeStatus>
</emphasis>
: adult, male; same data of the
<typeStatus id="0B3B882EFFD5FFD8FF3B5E45C8C6FD22" box="[151,253,620,644]" pageId="7" pageNumber="86" type="holotype">holotype</typeStatus>
. Abdomen of
<typeStatus id="0B3B882EFFD5FFD8FE305E44C9C7FD23" box="[412,508,621,645]" pageId="7" pageNumber="86" type="paratype">paratype</typeStatus>
is missing. Genitalia of both specimens were deformed during the study due to its membranous nature, and we only got presentable illustration of its dorsal view.
</paragraph>
</subSubSection>
</treatment>
</document>
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