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<document id="75208D96AD29034BFB7CB5F00C209D9C" ID-CLB-Dataset="8782" ID-DOI="10.11646/zootaxa.4822.3.2" ID-GBIF-Dataset="3c15254c-086c-448f-bfe2-e3cd208dcc87" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401617" ID-ZooBank="86DC1241-FE11-49CC-9674-ACAB1DF92F62" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609337343636" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="525F87CCFF9DFF8B9BEDFB2CFD32FEB6" docLanguage="en" docName="zootaxa.4822.3.2.pdf" docOrigin="Zootaxa 4822 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Himaloconnus Franz 1979" docType="treatment" docVersion="7" lastPageNumber="336" masterDocId="AE66FFB4FF9CFF889B7AFFE2FFD9FFF7" masterDocTitle="Himaloconnus Franz and Nogunius gen. n. of Japan (Coleoptera: Staphylinidae Scydmaeninae)" masterLastPageNumber="360" masterPageNumber="334" pageNumber="335" updateTime="1698849732245" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3EEF557D8AE471999798AF7D24640F97">Himaloconnus Franz and Nogunius gen. n. of Japan (Coleoptera: Staphylinidae Scydmaeninae)</mods:title>
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<mods:namePart id="8808BB9BCEB97B44D10ED03BEF702075">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="A5F9DA9BF1A1C7EEEE99344791D2934D">2020</mods:date>
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<treatment id="525F87CCFF9DFF8B9BEDFB2CFD32FEB6" ID-DOI="http://doi.org/10.5281/zenodo.4450821" ID-GBIF-Taxon="177009884" ID-Zenodo-Dep="4450821" LSID="urn:lsid:plazi:treatment:525F87CCFF9DFF8B9BEDFB2CFD32FEB6" httpUri="http://treatment.plazi.org/id/525F87CCFF9DFF8B9BEDFB2CFD32FEB6" lastPageId="3" lastPageNumber="336" pageId="1" pageNumber="335">
<subSubSection id="92EC6551FF9DFF899BEDFB2CFE44FB1E" box="[151,413,1230,1257]" pageId="1" pageNumber="335" type="nomenclature">
<paragraph id="DA4936DAFF9DFF899BEDFB2CFE44FB1E" blockId="1.[151,413,1230,1257]" box="[151,413,1230,1257]" pageId="1" pageNumber="335">
<heading id="810181B6FF9DFF899BEDFB2CFE44FB1E" bold="true" box="[151,413,1230,1257]" fontSize="11" level="1" pageId="1" pageNumber="335" reason="1">
<taxonomicName id="1DF64D59FF9DFF899BEDFB2CFE44FB1E" ID-CoL="8XLT8" authority="Franz" authorityName="Franz" authorityYear="1979" box="[151,413,1230,1257]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF9DFF899BEDFB2CFE44FB1E" bold="true" box="[151,413,1230,1257]" pageId="1" pageNumber="335">
<emphasis id="E882EAC8FF9DFF899BEDFB2CFE93FB1E" bold="true" box="[151,330,1230,1257]" italics="true" pageId="1" pageNumber="335">Himaloconnus</emphasis>
Franz
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF9DFF899BEDFAF5FB4DF96D" pageId="1" pageNumber="335" type="reference_group">
<paragraph id="DA4936DAFF9DFF899BEDFAF5FD65FAA1" blockId="1.[151,1437,1303,1437]" pageId="1" pageNumber="335">
<treatmentCitationGroup id="FAE611F4FF9DFF899BEDFAF5FD65FAA1" pageId="1" pageNumber="335">
<treatmentCitation id="5B5710CBFF9DFF899BEDFAF5FDDAFAC6" author="Franz, H." box="[151,515,1303,1329]" page="244" pageId="1" pageNumber="335" year="1979">
<taxonomicName id="1DF64D59FF9DFF899BEDFAF5FDDAFAC6" ID-CoL="8XLT8" authority="Franz, 1979: 244" authorityName="Franz" authorityPageNumber="244" authorityYear="1979" box="[151,515,1303,1329]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF9DFF899BEDFAF5FEE3FAC6" box="[151,314,1303,1329]" italics="true" pageId="1" pageNumber="335">Himaloconnus</emphasis>
<bibRefCitation id="BE674B2BFF9DFF899A39FAF5FDDAFAC6" author="Franz, H." box="[323,515,1303,1329]" pageId="1" pageNumber="335" pagination="235 - 274" refId="ref16048" refString="Franz, H. (1979) Weitere Beitrage zur Kenntnis der Scydmaenidenfauna des Himalaja und seiner Grenzgebiete (Coleoptera: Scydmaenidae). Entomologica Basiliensia, 4, 235 - 274." type="journal article" year="1979">Franz, 1979: 244</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(as subgenus of
<taxonomicName id="1DF64D59FF9DFF8999BEFAF5FCEBFAC7" box="[708,818,1303,1328]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF9DFF8999BEFAF5FCEBFAC7" box="[708,818,1303,1328]" italics="true" pageId="1" pageNumber="335">Euconnus</emphasis>
</taxonomicName>
).
<typeStatus id="054D8878FF9DFF899830FAF5FC5BFAC6" box="[842,898,1303,1329]" pageId="1" pageNumber="335">Type</typeStatus>
species:
<taxonomicName id="1DF64D59FF9DFF899891FAF5FA87FAC6" authority="Franz, 1979" authorityName="Franz" authorityYear="1979" box="[1003,1374,1303,1329]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="shutjensis">
<emphasis id="E882EAC8FF9DFF899891FAF5FB16FAC6" box="[1003,1231,1303,1329]" italics="true" pageId="1" pageNumber="335">Euconnus shutjensis</emphasis>
<bibRefCitation id="BE674B2BFF9DFF899FA2FAF5FA87FAC6" author="Franz, H." box="[1240,1374,1303,1329]" pageId="1" pageNumber="335" pagination="235 - 274" refId="ref16048" refString="Franz, H. (1979) Weitere Beitrage zur Kenntnis der Scydmaenidenfauna des Himalaja und seiner Grenzgebiete (Coleoptera: Scydmaenidae). Entomologica Basiliensia, 4, 235 - 274." type="journal article" year="1979">Franz, 1979</bibRefCitation>
</taxonomicName>
(des. orig.). Elevated to genus by
<bibRefCitation id="BE674B2BFF9DFF899AAEFAD9FD61FAA1" author="Jaloszynski, P." box="[468,696,1339,1366]" pageId="1" pageNumber="335" pagination="463 - 472" refId="ref17112" refString="Jaloszynski, P. (2016 b) Taxonomy of &quot; Euconnus complex &quot;. Part VIII. Subgenera Napoconnus, Himaloconnus and Nepalocon- nus removed from Euconnus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4103 (5), 463 - 472. https: // doi. org / 10.11646 / zootaxa. 4103.5.4" type="journal article" year="2016">Jałoszyński (2016b)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="DA4936DAFF9DFF899BEDFABDFCA2FA6A" blockId="1.[151,1437,1303,1437]" pageId="1" pageNumber="335">
<treatmentCitationGroup id="FAE611F4FF9DFF899BEDFABDFCA2FA6A" pageId="1" pageNumber="335">
<treatmentCitation id="5B5710CBFF9DFF899BEDFABDFDC4FA8E" author="Franz, H." box="[151,541,1375,1401]" page="247" pageId="1" pageNumber="335" year="1979">
<taxonomicName id="1DF64D59FF9DFF899BEDFABDFDC4FA8E" authority="Franz, 1979: 247" authorityName="Franz" authorityPageNumber="247" authorityYear="1979" box="[151,541,1375,1401]" class="Insecta" family="Scydmaenidae" genus="Himalayaconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF9DFF899BEDFABDFE8CFA8E" box="[151,341,1375,1401]" italics="true" pageId="1" pageNumber="335">Himalayaconnus</emphasis>
<bibRefCitation id="BE674B2BFF9DFF899A26FABDFDC4FA8E" author="Franz, H." box="[348,541,1375,1401]" pageId="1" pageNumber="335" pagination="235 - 274" refId="ref16048" refString="Franz, H. (1979) Weitere Beitrage zur Kenntnis der Scydmaenidenfauna des Himalaja und seiner Grenzgebiete (Coleoptera: Scydmaenidae). Entomologica Basiliensia, 4, 235 - 274." type="journal article" year="1979">Franz, 1979: 247</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Alternate original spelling for
<taxonomicName id="1DF64D59FF9DFF899801FABDFBC7FA8E" ID-CoL="8XLT8" authorityName="Franz" authorityYear="1979" box="[891,1054,1375,1401]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF9DFF899801FABDFBC7FA8E" box="[891,1054,1375,1401]" italics="true" pageId="1" pageNumber="335">Himaloconnus</emphasis>
</taxonomicName>
; presumably lapsus calami.
<emphasis id="E882EAC8FF9DFF899BEDFA61FE8CFA6A" box="[151,341,1411,1437]" italics="true" pageId="1" pageNumber="335">Himalayoconnus</emphasis>
<treatmentCitation id="5B5710CBFF9DFF899A26FA61FDC4FA6A" author="Franz, H." box="[348,541,1411,1437]" page="48" pageId="1" pageNumber="335" year="1981">
<bibRefCitation id="BE674B2BFF9DFF899A26FA61FDC4FA6A" author="Franz, H." box="[348,541,1411,1437]" pageId="1" pageNumber="335" pagination="45 - 50" refId="ref16112" refString="Franz, H. (1981 b) Weitere Scydmaeniden aus dem Himalaya (Scydmaenidae, Col.). Koleopterologische Rundschau, 55, 45 - 50." type="journal article" year="1981">Franz, 1981b: 48</bibRefCitation>
</treatmentCitation>
. Subsequent incorrect spelling.
</treatmentCitationGroup>
</paragraph>
<paragraph id="DA4936DAFF9DFF899BEDFA28FB4DF96D" blockId="1.[151,1437,1482,2013]" pageId="1" pageNumber="335">
<emphasis id="E882EAC8FF9DFF899BEDFA28FE22FA13" bold="true" box="[151,507,1482,1509]" pageId="1" pageNumber="335">Composition and distribution.</emphasis>
Nine nominal species are known (in alphabetical order):
<taxonomicName id="1DF64D59FF9DFF899F10FA29FE44F9FD" ID-CoL="3LXSK" authority="(Franz, 1985) (Taiwan)" baseAuthorityName="Franz" baseAuthorityYear="1985" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="alishanensis">
<emphasis id="E882EAC8FF9DFF899F10FA29FA45FA12" box="[1130,1436,1483,1509]" italics="true" pageId="1" pageNumber="335">Himaloconnus alishanensis</emphasis>
(
<bibRefCitation id="BE674B2BFF9DFF899BE5FA0DFEFFF9FE" author="Franz, H." box="[159,294,1518,1545]" pageId="1" pageNumber="335" pagination="91 - 128" refId="ref16141" refString="Franz, H. (1985) Neue und ungenugend bekannte Scydmaeniden (Coleoptera) aus Taiwan, Fukien und Thailand. Mitteilungen der Munchner Entomologischen Gesellschaft, 74, 91 - 128. [1984]" type="journal article" year="1985">Franz, 1985</bibRefCitation>
) (Taiwan)
</taxonomicName>
;
<taxonomicName id="1DF64D59FF9DFF899ADCFA0DFC41F9FD" ID-CoL="3LXSL" authority="(Franz, 1981a) (Nepal)" baseAuthorityName="Franz" baseAuthorityYear="1981" box="[422,920,1519,1546]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="hindu">
<emphasis id="E882EAC8FF9DFF899ADCFA0DFD49F9FE" box="[422,656,1519,1545]" italics="true" pageId="1" pageNumber="335">Himaloconnus hindu</emphasis>
(
<bibRefCitation id="BE674B2BFF9DFF8999E7FA0DFCE8F9FE" author="Franz, H." box="[669,817,1519,1545]" pageId="1" pageNumber="335" pagination="295 - 316" refId="ref16081" refString="Franz, H. (1981 a) 10. Beitrag zur Kenntnis der Scydmaenidenfauna des Himalaja (Col.). Entomologica Basiliensia, 6, 295 - 316." type="journal article" year="1981">Franz, 1981a</bibRefCitation>
) (
<collectingCountry id="A2E1764AFF9DFF89983DFA0DFC49F9FD" box="[839,912,1519,1546]" name="Nepal" pageId="1" pageNumber="335">Nepal</collectingCountry>
)
</taxonomicName>
;
<taxonomicName id="1DF64D59FF9DFF8998D8FA0DFED9F9D9" ID-CoL="3LXSM" authority="(Franz, 1985) (Taiwan)" baseAuthorityName="Franz" baseAuthorityYear="1985" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="klapperichianus">
<emphasis id="E882EAC8FF9DFF8998D8FA0DFB26F9FE" box="[930,1279,1519,1545]" italics="true" pageId="1" pageNumber="335">Himaloconnus klapperichianus</emphasis>
(
<bibRefCitation id="BE674B2BFF9DFF899E77FA0DFA4DF9FE" author="Franz, H." box="[1293,1428,1518,1545]" pageId="1" pageNumber="335" pagination="91 - 128" refId="ref16141" refString="Franz, H. (1985) Neue und ungenugend bekannte Scydmaeniden (Coleoptera) aus Taiwan, Fukien und Thailand. Mitteilungen der Munchner Entomologischen Gesellschaft, 74, 91 - 128. [1984]" type="journal article" year="1985">Franz, 1985</bibRefCitation>
) (Taiwan)
</taxonomicName>
;
<taxonomicName id="1DF64D59FF9DFF899A75F9F1FCBFF9D9" ID-CoL="3LXSN" authority="(Franz, 1979) (Nepal)" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[271,870,1555,1582]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="nagarkotensis">
<emphasis id="E882EAC8FF9DFF899A75F9F1FD83F9DA" box="[271,602,1555,1581]" italics="true" pageId="1" pageNumber="335">Himaloconnus nagarkotensis</emphasis>
(
<bibRefCitation id="BE674B2BFF9DFF899917F9F1FD23F9DA" author="Franz, H." box="[621,762,1555,1581]" pageId="1" pageNumber="335" pagination="235 - 274" refId="ref16048" refString="Franz, H. (1979) Weitere Beitrage zur Kenntnis der Scydmaenidenfauna des Himalaja und seiner Grenzgebiete (Coleoptera: Scydmaenidae). Entomologica Basiliensia, 4, 235 - 274." type="journal article" year="1979">Franz, 1979</bibRefCitation>
) (
<collectingCountry id="A2E1764AFF9DFF89986FF9F1FC87F9D9" box="[789,862,1555,1582]" name="Nepal" pageId="1" pageNumber="335">Nepal</collectingCountry>
)
</taxonomicName>
;
<taxonomicName id="1DF64D59FF9DFF89980FF9F1FF1FF9A5" ID-CoL="3LXSP" authority="(Franz, 1981b) (Nepal)" baseAuthorityName="Franz" baseAuthorityYear="1981" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="pokharanus">
<emphasis id="E882EAC8FF9DFF89980FF9F1FB71F9DA" box="[885,1192,1555,1581]" italics="true" pageId="1" pageNumber="335">Himaloconnus pokharanus</emphasis>
(
<bibRefCitation id="BE674B2BFF9DFF899FC1F9F1FA8FF9D9" author="Franz, H." box="[1211,1366,1555,1582]" pageId="1" pageNumber="335" pagination="45 - 50" refId="ref16112" refString="Franz, H. (1981 b) Weitere Scydmaeniden aus dem Himalaya (Scydmaenidae, Col.). Koleopterologische Rundschau, 55, 45 - 50." type="journal article" year="1981">Franz, 1981b</bibRefCitation>
) (
<collectingCountry id="A2E1764AFF9DFF899E0BF9F1FF64F9A5" name="Nepal" pageId="1" pageNumber="335">Nepal</collectingCountry>
)
</taxonomicName>
;
<taxonomicName id="1DF64D59FF9DFF899BABF9D5FD79F9A6" ID-CoL="3LXSQ" authority="(Franz, 1979)" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[209,672,1591,1617]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="ramamensis">
<emphasis id="E882EAC8FF9DFF899BABF9D5FDDBF9A7" box="[209,514,1591,1617]" italics="true" pageId="1" pageNumber="335">Himaloconnus ramamensis</emphasis>
(
<bibRefCitation id="BE674B2BFF9DFF89996AF9D5FD41F9A6" author="Franz, H." box="[528,664,1591,1617]" pageId="1" pageNumber="335" pagination="235 - 274" refId="ref16048" refString="Franz, H. (1979) Weitere Beitrage zur Kenntnis der Scydmaenidenfauna des Himalaja und seiner Grenzgebiete (Coleoptera: Scydmaenidae). Entomologica Basiliensia, 4, 235 - 274." type="journal article" year="1979">Franz, 1979</bibRefCitation>
)
</taxonomicName>
(
<collectingCountry id="A2E1764AFF9DFF8999D4F9D5FD34F9A6" box="[686,749,1591,1617]" name="India" pageId="1" pageNumber="335">India</collectingCountry>
:
<collectingRegion id="1832F838FF9DFF89998DF9D5FC93F9A5" box="[759,842,1591,1618]" country="India" name="Sikkim" pageId="1" pageNumber="335">Sikkim</collectingRegion>
/Darjiling);
<taxonomicName id="1DF64D59FF9DFF8998B7F9D5FE83F981" ID-CoL="3LXSR" authority="(Newton, 2017) (Thailand)" baseAuthorityName="Newton" baseAuthorityYear="2017" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="reductipenisides">
<emphasis id="E882EAC8FF9DFF8998B7F9D5FAF4F9A6" box="[973,1325,1591,1617]" italics="true" pageId="1" pageNumber="335">Himaloconnus reductipenisides</emphasis>
(
<bibRefCitation id="BE674B2BFF9DFF899E41F9D5FF08F982" author="Newton, A. F." pageId="1" pageNumber="335" pagination="1 - 52" refId="ref17327" refString="Newton, A. F. (2017) Nomenclatural and taxonomic changes in Staphyliniformia (Coleoptera). Insecta Mundi, 0595, 1 - 52." type="book chapter" year="2017">Newton, 2017</bibRefCitation>
) (
<collectingCountry id="A2E1764AFF9DFF899B92F9B9FE8BF981" box="[232,338,1627,1654]" name="Thailand" pageId="1" pageNumber="335">Thailand</collectingCountry>
)
</taxonomicName>
;
<taxonomicName id="1DF64D59FF9DFF899A1CF9B9FC58F981" ID-CoL="3LXSS" authority="(Franz, 1979) (Nepal)" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[358,897,1627,1654]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="shutjensis">
<emphasis id="E882EAC8FF9DFF899A1CF9B9FDA6F982" box="[358,639,1627,1653]" italics="true" pageId="1" pageNumber="335">Himaloconnus shutjensis</emphasis>
(
<bibRefCitation id="BE674B2BFF9DFF8999F5F9B9FCC1F982" author="Franz, H." box="[655,792,1627,1653]" pageId="1" pageNumber="335" pagination="235 - 274" refId="ref16048" refString="Franz, H. (1979) Weitere Beitrage zur Kenntnis der Scydmaenidenfauna des Himalaja und seiner Grenzgebiete (Coleoptera: Scydmaenidae). Entomologica Basiliensia, 4, 235 - 274." type="journal article" year="1979">Franz, 1979</bibRefCitation>
) (
<collectingCountry id="A2E1764AFF9DFF89984AF9B9FCA0F981" box="[816,889,1627,1654]" name="Nepal" pageId="1" pageNumber="335">Nepal</collectingCountry>
)
</taxonomicName>
; and
<taxonomicName id="1DF64D59FF9DFF8998C7F9B9FA93F982" ID-CoL="3LXST" authority="(Franz, 1979)" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[957,1354,1627,1653]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="topali">
<emphasis id="E882EAC8FF9DFF8998C7F9B9FB70F982" box="[957,1193,1627,1653]" italics="true" pageId="1" pageNumber="335">Himaloconnus topali</emphasis>
(
<bibRefCitation id="BE674B2BFF9DFF899FC3F9B9FA9BF982" author="Franz, H." box="[1209,1346,1627,1653]" pageId="1" pageNumber="335" pagination="235 - 274" refId="ref16048" refString="Franz, H. (1979) Weitere Beitrage zur Kenntnis der Scydmaenidenfauna des Himalaja und seiner Grenzgebiete (Coleoptera: Scydmaenidae). Entomologica Basiliensia, 4, 235 - 274." type="journal article" year="1979">Franz, 1979</bibRefCitation>
)
</taxonomicName>
(
<collectingCountry id="A2E1764AFF9DFF899E20F9B9FA40F982" box="[1370,1433,1627,1653]" name="India" pageId="1" pageNumber="335">India</collectingCountry>
:
<collectingRegion id="1832F838FF9DFF899BEDF99DFF33F96D" box="[151,234,1663,1690]" country="India" name="Sikkim" pageId="1" pageNumber="335">Sikkim</collectingRegion>
/Darjiling). Many undescribed species occur in Southeast and
<collectingCountry id="A2E1764AFF9DFF8998DAF99DFBD0F96E" box="[928,1033,1663,1689]" name="China" pageId="1" pageNumber="335">East Asia</collectingCountry>
(pers. obs.).
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF9DFF8B9BBDF941FD32FEB6" lastPageId="3" lastPageNumber="337" pageId="1" pageNumber="335" type="discussion">
<paragraph id="DA4936DAFF9DFF8A9BBDF941FEA4FF0E" blockId="1.[151,1437,1482,2013]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="336" pageId="1" pageNumber="335">
<emphasis id="E882EAC8FF9DFF899BBDF941FEE2F94A" bold="true" box="[199,315,1699,1725]" pageId="1" pageNumber="335">Remarks.</emphasis>
Morphological structures were illustrated and emended diagnosis was given by
<bibRefCitation id="BE674B2BFF9DFF899FCFF941FA40F949" author="Jaloszynski, P." box="[1205,1433,1699,1726]" pageId="1" pageNumber="335" pagination="463 - 472" refId="ref17112" refString="Jaloszynski, P. (2016 b) Taxonomy of &quot; Euconnus complex &quot;. Part VIII. Subgenera Napoconnus, Himaloconnus and Nepalocon- nus removed from Euconnus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4103 (5), 463 - 472. https: // doi. org / 10.11646 / zootaxa. 4103.5.4" type="journal article" year="2016">Jałoszyński (2016b)</bibRefCitation>
. The diagnosis of
<taxonomicName id="1DF64D59FF9DFF899A21F925FE27F916" authorityName="Franz" authorityYear="1979" box="[347,510,1735,1761]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF9DFF899A21F925FE27F916" box="[347,510,1735,1761]" italics="true" pageId="1" pageNumber="335">Himaloconnus</emphasis>
</taxonomicName>
was based on structures of the
<typeStatus id="054D8878FF9DFF899818F925FC4BF916" box="[866,914,1735,1761]" pageId="1" pageNumber="335">type</typeStatus>
species,
<taxonomicName id="1DF64D59FF9DFF899883F925FB04F916" authorityName="Franz" authorityYear="1979" box="[1017,1245,1735,1761]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="shutjensis">
<emphasis id="E882EAC8FF9DFF899883F925FB04F916" box="[1017,1245,1735,1761]" italics="true" pageId="1" pageNumber="335">Euconnus shutjensis</emphasis>
</taxonomicName>
, known to occur in
<collectingCountry id="A2E1764AFF9DFF899BCFF909FF21F8F2" box="[181,248,1771,1797]" name="Nepal" pageId="1" pageNumber="335">Nepal</collectingCountry>
(
<bibRefCitation id="BE674B2BFF9DFF899A72F909FE04F8F1" author="Jaloszynski, P." box="[264,477,1771,1798]" pageId="1" pageNumber="335" pagination="463 - 472" refId="ref17112" refString="Jaloszynski, P. (2016 b) Taxonomy of &quot; Euconnus complex &quot;. Part VIII. Subgenera Napoconnus, Himaloconnus and Nepalocon- nus removed from Euconnus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4103 (5), 463 - 472. https: // doi. org / 10.11646 / zootaxa. 4103.5.4" type="journal article" year="2016">Jałoszyński 2016b</bibRefCitation>
), and included the following characters: head with long tempora; pronotum subtrapezoidal, broadest near base and strongly narrowing anteriorly; head and pronotum with thick bristles; antenna with distinct trimerous club; hypostomal ridges extending to the level of posterior tentorial pits; mouthparts with “rounded lateral lobes flanking prementum, which are presumably associated with hypopharynx” (see remarks below); pronotum with one pair of small lateral pits, lacking sublateral carinae; hypomeral ridges incomplete, obliterated anteriorly and posteriorly; prosternum with rudimentary prosternal carina; mesoventral intercoxal process carinate, T-shaped in cross-section, broad and strongly elevated, broadening posteriorly, anteriorly not connected to anterior ridge of mesoventrite; metaventral intercoxal process narrowly separating metacoxae, with a pair of posteriorly directed, pointed spines adjacent at middle; each elytron with two distinct but small and setose basal foveae; aedeagus with free parameres.
</paragraph>
<paragraph id="DA4936DAFF9EFF8A9BBDFEE1FD69FB32" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="336">
Characters found in the Japanese species (illustrated in
<figureCitation id="42CD2A5FFF9EFF8A9841FEE1FCA5FEEA" box="[827,892,259,285]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[161,1426,932,1936]" captionTargetId="figure-414@4.[151,1436,921,1940]" captionTargetPageId="4" captionText="FIGURES 1–3. Dorsal habitus of males. Himaloconnus klapperichianus yaeyamanus ssp. n. (1); H. klapperichianus amami- anus ssp. n. (2); and H. klapperichianus okinawanus ssp. n. (3)." figureDoi="http://doi.org/10.5281/zenodo.4401619" httpUri="https://zenodo.org/record/4401619/files/figure.png" pageId="2" pageNumber="336">Figs 1</figureCitation>
–43) largely agree with those given above, with few notable differences. The hypostomal ridges are nearly complete, obliterated slightly in front of posterior tentorial pits (
<figureCitation id="42CD2A5FFF9EFF8A9A6EFEA9FEB0FE92" box="[276,361,331,357]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Fig. 12</figureCitation>
); the pronotum lacks antebasal pits; the hypomeral ridges are complete, although indeed more distinct near middle and less so in their anterior and posterior portions (
<figureCitation id="42CD2A5FFF9EFF8A98D2FE8DFC23FE7E" box="[936,1018,367,393]" captionStart="FIGURES 18–21" captionStartId="8.[151,264,1417,1441]" captionTargetBox="[158,1430,390,1382]" captionTargetId="figure-105@8.[158,1430,381,1394]" captionTargetPageId="8" captionText="FIGURES 18–21. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Prothorax in ventral view (18–19), mesonotum (Fig. 20), and metanotum (Fig. 21; postnotum omitted). Abbreviations: alc, alacrista; ap, apodeme; hyr, hypomeral ridge; mma, median membranous area; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pst+pn, prosternum fused with pronotum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; scl3, metascutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401625" httpUri="https://zenodo.org/record/4401625/files/figure.png" pageId="2" pageNumber="336">Fig. 18</figureCitation>
); the prosternal carina (
<figureCitation id="42CD2A5FFF9EFF8A9F87FE8DFAA5FE7E" box="[1277,1404,367,393]" captionStart="FIGURES 18–21" captionStartId="8.[151,264,1417,1441]" captionTargetBox="[158,1430,390,1382]" captionTargetId="figure-105@8.[158,1430,381,1394]" captionTargetPageId="8" captionText="FIGURES 18–21. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Prothorax in ventral view (18–19), mesonotum (Fig. 20), and metanotum (Fig. 21; postnotum omitted). Abbreviations: alc, alacrista; ap, apodeme; hyr, hypomeral ridge; mma, median membranous area; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pst+pn, prosternum fused with pronotum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; scl3, metascutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401625" httpUri="https://zenodo.org/record/4401625/files/figure.png" pageId="2" pageNumber="336">Figs 18–19</figureCitation>
) is entirely lacking; the mesoventral intercoxal process (
<figureCitation id="42CD2A5FFF9EFF8A9999FE71FCBBFE5A" box="[739,866,403,429]" captionStart="FIGURES 22–26" captionStartId="9.[151,264,1818,1842]" captionTargetBox="[157,1430,496,1785]" captionTargetId="figure-152@9.[151,1436,488,1794]" captionTargetPageId="9" captionText="FIGURES 22–26. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Pterothorax in ventral (22–23) and lateral (24) views; elytral base (25); and abdomen in ventral view (26). Abbreviations: aest3, mesanepisternum; ar, anterior ridge; bef, basal elytral foveae; dlf, dorsolateral fovea; epm3, metepimeron; lmc, lateral metaventral carina; lmfa, lateral metafurcal arm; mcp, mesocoxal process; mcr, mesocoxal rest; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401627" httpUri="https://zenodo.org/record/4401627/files/figure.png" pageId="2" pageNumber="336">Figs 22–23</figureCitation>
) is parallel-sided between mesocoxae; and the basal elytral foveae are asetose (
<figureCitation id="42CD2A5FFF9EFF8A9AA7FE55FDEBFE26" box="[477,562,438,465]" captionStart="FIGURES 22–26" captionStartId="9.[151,264,1818,1842]" captionTargetBox="[157,1430,496,1785]" captionTargetId="figure-152@9.[151,1436,488,1794]" captionTargetPageId="9" captionText="FIGURES 22–26. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Pterothorax in ventral (22–23) and lateral (24) views; elytral base (25); and abdomen in ventral view (26). Abbreviations: aest3, mesanepisternum; ar, anterior ridge; bef, basal elytral foveae; dlf, dorsolateral fovea; epm3, metepimeron; lmc, lateral metaventral carina; lmfa, lateral metafurcal arm; mcp, mesocoxal process; mcr, mesocoxal rest; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401627" httpUri="https://zenodo.org/record/4401627/files/figure.png" pageId="2" pageNumber="336">Fig. 25</figureCitation>
). Examination of a disarticulated specimen of a
<taxonomicName id="1DF64D59FF9EFF8A9F2AFE55FB2AFE26" authorityName="Franz" authorityYear="1979" box="[1104,1267,439,465]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF9EFF8A9F2AFE55FB2AFE26" box="[1104,1267,439,465]" italics="true" pageId="2" pageNumber="336">Himaloconnus</emphasis>
</taxonomicName>
from Ishigakijima allowed for identifying the “rounded lateral lobes flanking prementum, which are presumably associated with hypopharynx” clearly as lateral lobes of hypopharynx (
<figureCitation id="42CD2A5FFF9EFF8A987CFE1DFC50FDEE" box="[774,905,511,537]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Figs 12, 17</figureCitation>
), which are unique in being strongly expanded laterally and forming large lateral ‘flaps’, in
<figureCitation id="42CD2A5FFF9EFF8A99FFFDC1FCF1FDCA" box="[645,808,547,573]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Figs 12 and 17</figureCitation>
oriented in such a way that only their dorsal margins are visible and their anterior surface is directed anterad; in fig. 14 of
<bibRefCitation id="BE674B2BFF9EFF8A980EFDA5FB81FD95" author="Jaloszynski, P." box="[884,1112,583,610]" pageId="2" pageNumber="336" pagination="463 - 472" refId="ref17112" refString="Jaloszynski, P. (2016 b) Taxonomy of &quot; Euconnus complex &quot;. Part VIII. Subgenera Napoconnus, Himaloconnus and Nepalocon- nus removed from Euconnus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4103 (5), 463 - 472. https: // doi. org / 10.11646 / zootaxa. 4103.5.4" type="journal article" year="2016">Jałoszyński (2016b)</bibRefCitation>
, this structure was annotated with a question mark and is visible at each side of the prementum as an oval lobe, but it is clear that the lobe is flexible and can bend, as the left and right lobe are in different orientations. This hypertrophied lateral hypopharyngeal lobe seems unique for
<taxonomicName id="1DF64D59FF9EFF8A9AE2FD51FDE2FD3A" authorityName="Franz" authorityYear="1979" box="[408,571,691,717]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF9EFF8A9AE2FD51FDE2FD3A" box="[408,571,691,717]" italics="true" pageId="2" pageNumber="336">Himaloconnus</emphasis>
</taxonomicName>
and should be included in the diagnosis. Similarly, the U-shaped impression on the submentum found in
<taxonomicName id="1DF64D59FF9EFF8A9AB7FD35FD3CFD06" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[461,741,727,753]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="species" species="shutjensis">
<emphasis id="E882EAC8FF9EFF8A9AB7FD35FD3CFD06" box="[461,741,727,753]" italics="true" pageId="2" pageNumber="336">Himaloconnus shutjensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="BE674B2BFF9EFF8A9989FD35FC1CFD06" author="Jaloszynski, P." box="[755,965,727,753]" pageId="2" pageNumber="336" pagination="463 - 472" refId="ref17112" refString="Jaloszynski, P. (2016 b) Taxonomy of &quot; Euconnus complex &quot;. Part VIII. Subgenera Napoconnus, Himaloconnus and Nepalocon- nus removed from Euconnus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4103 (5), 463 - 472. https: // doi. org / 10.11646 / zootaxa. 4103.5.4" type="journal article" year="2016">Jałoszyński 2016b</bibRefCitation>
; figs 14–15) covered with numerous pores and what seems to be remains of a secretion, was found in all (male and female) specimens collected in
<collectingCountry id="A2E1764AFF9EFF8A9E64FD19FAB8FCE2" box="[1310,1377,763,789]" name="Japan" pageId="2" pageNumber="336">Japan</collectingCountry>
. This median impression (
<figureCitation id="42CD2A5FFF9EFF8A9A01FCFDFE14FCCE" box="[379,461,799,825]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Fig. 12</figureCitation>
) can also be added to the genus diagnosis, especially that it is well-visible also under a stereomicroscope. The complete or slightly incomplete hypostomal ridges seem a variable character, and while they are incomplete in specimens from the Yaeyama Archipelago, they are complete in
<taxonomicName id="1DF64D59FF9EFF8A9F45FC85FB09FC76" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[1087,1232,871,897]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="species" species="shutjensis">
<emphasis id="E882EAC8FF9EFF8A9F45FC85FB09FC76" box="[1087,1232,871,897]" italics="true" pageId="2" pageNumber="336">H. shutjensis</emphasis>
</taxonomicName>
and in specimens from Amami-Ôshima. The prosternal process or carina in
<taxonomicName id="1DF64D59FF9EFF8A9850FC69FC62FC52" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[810,955,907,933]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="species" species="shutjensis">
<emphasis id="E882EAC8FF9EFF8A9850FC69FC62FC52" box="[810,955,907,933]" italics="true" pageId="2" pageNumber="336">H. shutjensis</emphasis>
</taxonomicName>
is vestigial, barely discernable, so its lack in some species of the genus is not surprising. The number and arrangement of antebasal pronotal pits may vary among species within a genus, and the lack of pits in the Japanese specimens (as well as in
<taxonomicName id="1DF64D59FF9EFF8A9FFAFC31FA8CFC1A" baseAuthorityName="Franz" baseAuthorityYear="1985" box="[1152,1365,979,1005]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="species" species="klapperichianus">
<emphasis id="E882EAC8FF9EFF8A9FFAFC31FA8CFC1A" box="[1152,1365,979,1005]" italics="true" pageId="2" pageNumber="336">H. klapperichianus</emphasis>
</taxonomicName>
found in geographically close Taiwan) is not a character of great taxonomic value. The shape of the mesoventral carina (parallel-sided vs. slightly broadening posterad) is another character clearly variable within genus. The basal elytral foveae in
<taxonomicName id="1DF64D59FF9EFF8A9A7DFBDDFE41FBAE" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[263,408,1087,1113]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="species" species="shutjensis">
<emphasis id="E882EAC8FF9EFF8A9A7DFBDDFE41FBAE" box="[263,408,1087,1113]" italics="true" pageId="2" pageNumber="336">H. shutjensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="BE674B2BFF9EFF8A9AD2FBDDFDA5FBAE" author="Jaloszynski, P." box="[424,636,1087,1113]" pageId="2" pageNumber="336" pagination="463 - 472" refId="ref17112" refString="Jaloszynski, P. (2016 b) Taxonomy of &quot; Euconnus complex &quot;. Part VIII. Subgenera Napoconnus, Himaloconnus and Nepalocon- nus removed from Euconnus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4103 (5), 463 - 472. https: // doi. org / 10.11646 / zootaxa. 4103.5.4" type="journal article" year="2016">Jałoszyński 2016b</bibRefCitation>
; fig. 13) are filled with a few setae only (in contrast to many genera of
<taxonomicName id="1DF64D59FF9EFF8A9BEDFB81FECEFB8A" authorityName="Ganglbauer" authorityYear="1899" box="[151,279,1123,1149]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="tribe" tribe="Stenichnini">Stenichnini</taxonomicName>
that have densely setose foveae, with numerous ‘wooly’ setae on their circumference directed toward the center), and again, reduction of this character is not of great taxonomic value. This variability, however, must be included in an emended diagnosis, given below.
</paragraph>
<paragraph id="DA4936DAFF9EFF8A9BBDFB2DFEB3F8F2" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="336">
In addition to previously known characters, a more detailed examination of disarticulated specimens allowed for a better characterization of
<taxonomicName id="1DF64D59FF9EFF8A9A81FB11FD47FAFA" authorityName="Franz" authorityYear="1979" box="[507,670,1267,1293]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF9EFF8A9A81FB11FD47FAFA" box="[507,670,1267,1293]" italics="true" pageId="2" pageNumber="336">Himaloconnus</emphasis>
</taxonomicName>
. The following structures, poorly visible in the unique
<typeStatus id="054D8878FF9EFF8A9E60FB11FAA5FAFA" box="[1306,1404,1267,1293]" pageId="2" pageNumber="336" type="holotype">holotype</typeStatus>
of
<taxonomicName id="1DF64D59FF9EFF8A9BEDFAF5FEFEFAC6" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[151,295,1303,1329]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="species" species="shutjensis">
<emphasis id="E882EAC8FF9EFF8A9BEDFAF5FEFEFAC6" box="[151,295,1303,1329]" italics="true" pageId="2" pageNumber="336">H. shutjensis</emphasis>
</taxonomicName>
and studied only by scanning electron microscopy, are well-visible in transparent slides: labrum (
<figureCitation id="42CD2A5FFF9EFF8A9E15FAF5FF6DFAA2" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Fig. 15</figureCitation>
) transverse, with slightly concave anterior margin and sparse, nearly symmetrical dorsal setae; mandibles (
<figureCitation id="42CD2A5FFF9EFF8A9E15FAD9FF6DFA8D" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Fig. 16</figureCitation>
) symmetrical, each with broad base, distinct setose prostheca and a small preapical tooth; prementum (
<figureCitation id="42CD2A5FFF9EFF8A9E38FABDFA4DFA8E" box="[1346,1428,1375,1401]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Fig. 17</figureCitation>
) with a small papillate ligula (
<figureCitation id="42CD2A5FFF9EFF8A9A9AFA61FDBAFA6A" box="[480,611,1411,1437]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Figs 12, 17</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9915FA61FD54FA6A" box="[623,653,1411,1437]" italics="true" pageId="2" pageNumber="336">lig</emphasis>
) bearing a pair of small setae, and with labial palps with palpomere I (
<figureCitation id="42CD2A5FFF9EFF8A9BDAFA45FF2AFA36" box="[160,243,1447,1473]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Fig. 17</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9B85FA45FEFAFA36" box="[255,291,1447,1473]" italics="true" pageId="2" pageNumber="336">lp1</emphasis>
) short, ring-like, palpomere II (
<figureCitation id="42CD2A5FFF9EFF8A99F2FA45FD05FA36" box="[648,732,1447,1473]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Fig. 17</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9992FA45FCD5FA36" box="[744,780,1447,1473]" italics="true" pageId="2" pageNumber="336">lp2</emphasis>
) elongate, subcylindrical, and palpomere III (
<figureCitation id="42CD2A5FFF9EFF8A9E75FA45FABAFA36" box="[1295,1379,1447,1473]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Fig. 17</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9E15FA45FA4AFA36" box="[1391,1427,1447,1473]" italics="true" pageId="2" pageNumber="336">lp3</emphasis>
) narrow, elongate, tapering distad and slightly curved; tentorium (
<figureCitation id="42CD2A5FFF9EFF8A9813FA29FC3EFA12" box="[873,999,1483,1509]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Figs 11, 14</figureCitation>
) with small corporatentorium (
<figureCitation id="42CD2A5FFF9EFF8A9E38FA29FF6FF9FE" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Figs 11, 14</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9BB8FA0DFF0FF9FF" box="[194,214,1519,1544]" italics="true" pageId="2" pageNumber="336">ct</emphasis>
), bisinuate tentorial bridge (
<figureCitation id="42CD2A5FFF9EFF8A996FFA0DFD4EF9FE" box="[533,663,1519,1545]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Figs 11, 14</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A99D8FA0DFD61F9FE" box="[674,696,1519,1545]" italics="true" pageId="2" pageNumber="336">tb</emphasis>
) with anteromedian projection, long gular ridges (
<figureCitation id="42CD2A5FFF9EFF8A9F94FA0DFAA9F9FE" box="[1262,1392,1519,1545]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Figs 11, 14</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9E06FA0DFA4CF9FF" box="[1404,1429,1519,1544]" italics="true" pageId="2" pageNumber="336">gr</emphasis>
; with membranous ventral portions and sclerotized dorsal portions, the latter visible in lateral and dorsal view as long ‘arms’), short dorsal tentorial arms (
<figureCitation id="42CD2A5FFF9EFF8A9955F9D5FD5AF9A6" box="[559,643,1591,1617]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Fig. 14</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A99F5F9D5FD6AF9A6" box="[655,691,1591,1617]" italics="true" pageId="2" pageNumber="336">dta</emphasis>
) not reaching vertex, short and stout posterior tentorial arms (
<figureCitation id="42CD2A5FFF9EFF8A9E15F9D5FF6FF982" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Fig. 14</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9BB9F9B9FF3EF983" box="[195,231,1627,1652]" italics="true" pageId="2" pageNumber="336">pta</emphasis>
), and long, slender anterior tentorial arms (
<figureCitation id="42CD2A5FFF9EFF8A99AFF9B9FC81F982" box="[725,856,1627,1653]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1751,1775]" captionTargetBox="[151,1432,493,1717]" captionTargetId="figure-112@7.[151,1436,489,1728]" captionTargetPageId="7" captionText="FIGURES 10–17. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Head in dorsal (10–11), ventral (12) and lateral (13–14) views; labrum in dorsal view (15); right mandible in dorsal view (16); prementum in ventral view (17). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; dta, dorsal tentorial arm; fo, foramen occipitale; gal, galea; gr, gular ridge; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1–3, labial palpomere I–III; mi, median impression; mn, mentum; mxp1–4, maxillary palpomere I–IV; occ, occipital constriction; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401623" httpUri="https://zenodo.org/record/4401623/files/figure.png" pageId="2" pageNumber="336">Figs 11, 14</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A981FF9B9FC50F983" box="[869,905,1627,1652]" italics="true" pageId="2" pageNumber="336">ata</emphasis>
); mesonotum (
<figureCitation id="42CD2A5FFF9EFF8A9F4FF9B9FB51F982" box="[1077,1160,1627,1653]" captionStart="FIGURES 18–21" captionStartId="8.[151,264,1417,1441]" captionTargetBox="[158,1430,390,1382]" captionTargetId="figure-105@8.[158,1430,381,1394]" captionTargetPageId="8" captionText="FIGURES 18–21. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Prothorax in ventral view (18–19), mesonotum (Fig. 20), and metanotum (Fig. 21; postnotum omitted). Abbreviations: alc, alacrista; ap, apodeme; hyr, hypomeral ridge; mma, median membranous area; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pst+pn, prosternum fused with pronotum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; scl3, metascutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401625" httpUri="https://zenodo.org/record/4401625/files/figure.png" pageId="2" pageNumber="336">Fig. 20</figureCitation>
) with mesoscutum (
<figureCitation id="42CD2A5FFF9EFF8A9E15F9B9FF6FF96E" captionStart="FIGURES 18–21" captionStartId="8.[151,264,1417,1441]" captionTargetBox="[158,1430,390,1382]" captionTargetId="figure-105@8.[158,1430,381,1394]" captionTargetPageId="8" captionText="FIGURES 18–21. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Prothorax in ventral view (18–19), mesonotum (Fig. 20), and metanotum (Fig. 21; postnotum omitted). Abbreviations: alc, alacrista; ap, apodeme; hyr, hypomeral ridge; mma, median membranous area; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pst+pn, prosternum fused with pronotum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; scl3, metascutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401625" httpUri="https://zenodo.org/record/4401625/files/figure.png" pageId="2" pageNumber="336">Fig. 20</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9BB8F99DFF31F96E" box="[194,232,1663,1689]" italics="true" pageId="2" pageNumber="336">sc2</emphasis>
) and mesoscutellum (
<figureCitation id="42CD2A5FFF9EFF8A9A9AF99DFDEAF96E" box="[480,563,1663,1689]" captionStart="FIGURES 18–21" captionStartId="8.[151,264,1417,1441]" captionTargetBox="[158,1430,390,1382]" captionTargetId="figure-105@8.[158,1430,381,1394]" captionTargetPageId="8" captionText="FIGURES 18–21. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Prothorax in ventral view (18–19), mesonotum (Fig. 20), and metanotum (Fig. 21; postnotum omitted). Abbreviations: alc, alacrista; ap, apodeme; hyr, hypomeral ridge; mma, median membranous area; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pst+pn, prosternum fused with pronotum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; scl3, metascutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401625" httpUri="https://zenodo.org/record/4401625/files/figure.png" pageId="2" pageNumber="336">Fig. 20</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9945F99DFDB4F96E" box="[575,621,1663,1689]" italics="true" pageId="2" pageNumber="336">scl2</emphasis>
) demarcated by scutoscutellar suture (
<figureCitation id="42CD2A5FFF9EFF8A9F64F99DFBABF96E" box="[1054,1138,1663,1689]" captionStart="FIGURES 18–21" captionStartId="8.[151,264,1417,1441]" captionTargetBox="[158,1430,390,1382]" captionTargetId="figure-105@8.[158,1430,381,1394]" captionTargetPageId="8" captionText="FIGURES 18–21. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Prothorax in ventral view (18–19), mesonotum (Fig. 20), and metanotum (Fig. 21; postnotum omitted). Abbreviations: alc, alacrista; ap, apodeme; hyr, hypomeral ridge; mma, median membranous area; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pst+pn, prosternum fused with pronotum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; scl3, metascutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401625" httpUri="https://zenodo.org/record/4401625/files/figure.png" pageId="2" pageNumber="336">Fig. 20</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9F04F99DFB46F96F" box="[1150,1183,1663,1688]" italics="true" pageId="2" pageNumber="336">sss</emphasis>
); metanotum (
<figureCitation id="42CD2A5FFF9EFF8A9E3FF99DFA4EF96E" box="[1349,1431,1663,1689]" captionStart="FIGURES 18–21" captionStartId="8.[151,264,1417,1441]" captionTargetBox="[158,1430,390,1382]" captionTargetId="figure-105@8.[158,1430,381,1394]" captionTargetPageId="8" captionText="FIGURES 18–21. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Prothorax in ventral view (18–19), mesonotum (Fig. 20), and metanotum (Fig. 21; postnotum omitted). Abbreviations: alc, alacrista; ap, apodeme; hyr, hypomeral ridge; mma, median membranous area; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pst+pn, prosternum fused with pronotum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; scl3, metascutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401625" httpUri="https://zenodo.org/record/4401625/files/figure.png" pageId="2" pageNumber="336">Fig. 21</figureCitation>
, postnotum omitted) developed as in all winged
<taxonomicName id="1DF64D59FF9EFF8A99CFF941FCE1F94A" authorityName="Ganglbauer" authorityYear="1899" box="[693,824,1699,1725]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="tribe" tribe="Stenichnini">Stenichnini</taxonomicName>
; deep dorsolateral foveae of mesothorax (
<figureCitation id="42CD2A5FFF9EFF8A9E6EF941FA41F94A" box="[1300,1432,1699,1725]" captionStart="FIGURES 22–26" captionStartId="9.[151,264,1818,1842]" captionTargetBox="[157,1430,496,1785]" captionTargetId="figure-152@9.[151,1436,488,1794]" captionTargetPageId="9" captionText="FIGURES 22–26. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Pterothorax in ventral (22–23) and lateral (24) views; elytral base (25); and abdomen in ventral view (26). Abbreviations: aest3, mesanepisternum; ar, anterior ridge; bef, basal elytral foveae; dlf, dorsolateral fovea; epm3, metepimeron; lmc, lateral metaventral carina; lmfa, lateral metafurcal arm; mcp, mesocoxal process; mcr, mesocoxal rest; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401627" httpUri="https://zenodo.org/record/4401627/files/figure.png" pageId="2" pageNumber="336">Figs 22–24</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9BEDF925FF6CF916" box="[151,181,1735,1761]" italics="true" pageId="2" pageNumber="336">dlf</emphasis>
); pleural meso- and metaventral structures largely fused together (
<figureCitation id="42CD2A5FFF9EFF8A98D4F925FBDDF916" box="[942,1028,1735,1761]" captionStart="FIGURES 22–26" captionStartId="9.[151,264,1818,1842]" captionTargetBox="[157,1430,496,1785]" captionTargetId="figure-152@9.[151,1436,488,1794]" captionTargetPageId="9" captionText="FIGURES 22–26. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Pterothorax in ventral (22–23) and lateral (24) views; elytral base (25); and abdomen in ventral view (26). Abbreviations: aest3, mesanepisternum; ar, anterior ridge; bef, basal elytral foveae; dlf, dorsolateral fovea; epm3, metepimeron; lmc, lateral metaventral carina; lmfa, lateral metafurcal arm; mcp, mesocoxal process; mcr, mesocoxal rest; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401627" httpUri="https://zenodo.org/record/4401627/files/figure.png" pageId="2" pageNumber="336">Fig. 24</figureCitation>
); and elongate, thin-walled spermatheca (
<figureCitation id="42CD2A5FFF9EFF8A9B98F909FEECF8F2" box="[226,309,1771,1797]" captionStart="FIGURES 22–26" captionStartId="9.[151,264,1818,1842]" captionTargetBox="[157,1430,496,1785]" captionTargetId="figure-152@9.[151,1436,488,1794]" captionTargetPageId="9" captionText="FIGURES 22–26. Morphological structures of Himaloconnus klapperichianus yaeyamanus ssp. n., female. Pterothorax in ventral (22–23) and lateral (24) views; elytral base (25); and abdomen in ventral view (26). Abbreviations: aest3, mesanepisternum; ar, anterior ridge; bef, basal elytral foveae; dlf, dorsolateral fovea; epm3, metepimeron; lmc, lateral metaventral carina; lmfa, lateral metafurcal arm; mcp, mesocoxal process; mcr, mesocoxal rest; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401627" httpUri="https://zenodo.org/record/4401627/files/figure.png" pageId="2" pageNumber="336">Fig. 26</figureCitation>
;
<emphasis id="E882EAC8FF9EFF8A9A3BF909FE83F8F3" box="[321,346,1771,1796]" italics="true" pageId="2" pageNumber="336">sp</emphasis>
).
</paragraph>
<paragraph id="DA4936DAFF9EFF8B9BBDF8EDFD32FEB6" blockId="2.[151,1437,151,2013]" lastBlockId="3.[151,1436,151,321]" lastPageId="3" lastPageNumber="337" pageId="2" pageNumber="336">
<emphasis id="E882EAC8FF9EFF8A9BBDF8EDFD5FF8DE" bold="true" box="[199,646,1806,1833]" pageId="2" pageNumber="336">
Emended diagnosis of
<taxonomicName id="1DF64D59FF9EFF8A9AAEF8ECFDA6F8DE" authorityName="Franz" authorityYear="1979" box="[468,639,1806,1833]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF9EFF8A9AAEF8ECFDA6F8DE" bold="true" box="[468,639,1806,1833]" italics="true" pageId="2" pageNumber="336">Himaloconnus</emphasis>
</taxonomicName>
.
</emphasis>
Head with long tempora (i.e., eyes situated anteriorly); pronotum subtrapezoidal, broadest near base and strongly narrowing anteriorly; head and pronotum with thick bristles; antennal fossae narrowly separated at middle; antenna with distinct trimerous club; hypostomal ridges complete or almost complete (i.e., posteriorly reaching or almost reaching the level of posterior tentorial pits); lateral lobes of hypopharynx conspicuously large, forming oval flexible ‘flaps’; submentum with deep U-shaped impression with sharply marked lateral and posterior margins; pronotum with one pair of lateral pits or lacking pits, lacking sublateral carinae; notosternal sutures obliterated; hypomeral ridges complete or partly obliterated; prosternal process lacking or vestigial; mesoventral intercoxal process carinate, broad and strongly elevated, anteriorly not connected to anterior ridge of mesoventrite; metaventral intercoxal process narrowly separating metacoxae, composed of a pair of pointed spines adjacent at middle; each elytron with two distinct but small basal foveae (asetose or filled with sparse setae); aedeagus with free parameres; spermatheca elongate.
</paragraph>
</subSubSection>
</treatment>
</document>
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