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<mods:title id="1DF35DB34E20F110E22B7021CA43DDDA">Revision of Leocrates Kinberg, 1866 and Leocratides Ehlers, 1908 (Annelida, Errantia, Hesionidae)</mods:title>
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<treatment id="2D1987E4FFB6737BFF23FDE2F72B60A5" ID-DOI="http://doi.org/10.5281/zenodo.3680223" ID-GBIF-Taxon="162161375" ID-Zenodo-Dep="3680223" LSID="urn:lsid:plazi:treatment:2D1987E4FFB6737BFF23FDE2F72B60A5" httpUri="http://treatment.plazi.org/id/2D1987E4FFB6737BFF23FDE2F72B60A5" lastPageId="87" lastPageNumber="88" pageId="85" pageNumber="86">
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<heading id="FE47819EFFB67379FF23FDE2F7EA633B" bold="true" box="[151,904,535,562]" fontSize="11" level="1" pageId="85" pageNumber="86" reason="1">
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<taxonomicName id="62B04D71FFB67379FF23FDE2F6E8633B" ID-CoL="a06da30e-be4b-4408-b625-737156a5532d" authority="(Treadwell, 1906)" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Treadwell" baseAuthorityYear="1906" box="[151,650,535,562]" class="Polychaeta" family="Hesionidae" genus="Paradalhousia" kingdom="Animalia" order="Phyllodocida" pageId="85" pageNumber="86" phylum="Annelida" rank="species" species="oculata" status="comb. nov.">
<emphasis id="97C4EAE0FFB67379FF23FDE2F5C86338" bold="true" box="[151,426,535,562]" italics="true" pageId="85" pageNumber="86">Paradalhousia oculata</emphasis>
(
<bibRefCitation id="C1214B03FFB67379FE0DFDE2F6E0633B" author="Treadwell" box="[441,642,535,561]" firstAuthor="Treadwell" pageId="85" pageNumber="86" pagination="1145 - 1181" refId="ref89458" refString="Treadwell, A. L. (1906 [1903]) Polychaetous annelids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission, 23 (3), 1145 - 1181. [http: // docs. lib. noaa. gov / rescue / Fish _ Commission _ Bulletins / BFC 1903 - v 23. pt 3. pdf]" type="journal article" year="1906">Treadwell, 1906</bibRefCitation>
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, reinstated
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<subSubSection id="EDAA6579FFB67379FF23FDCEF576635C" box="[151,276,571,598]" pageId="85" pageNumber="86" type="description">
<paragraph id="A50F36F2FFB67379FF23FDCEF576635C" blockId="85.[151,904,535,598]" box="[151,276,571,598]" pageId="85" pageNumber="86">
<figureCitation id="3D8B2A77FFB67379FF23FDCEF48F635C" box="[151,237,571,598]" captionStart="FIGURE 49" captionStartId="85.[151,250,1859,1884]" captionTargetBox="[173,1417,949,1832]" captionTargetId="figure@85.[170,1418,946,1834]" captionTargetPageId="85" captionText="FIGURE 49. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, holotype (USNM 5200). A. Dorsal view, depressed by glass slide. B. Anterior end, dorsal view (*: denticle). C. Same, MGS (*: denticle). D. Pharynx, frontal view, MGS, denticles slightly darker than surrounding areas. E. Middle chaetiger, left parapodium, anterior view (insets: (left) notacicular and (right) neuracicular lobes tips, and medium- and small neurochaetal blades). Scale bars: A: 1 mm, B, C: 0.3 mm, D: 0.5 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672650" httpUri="https://zenodo.org/record/3672650/files/figure.png" pageId="85" pageNumber="86">Figs 49</figureCitation>
,
<figureCitation id="3D8B2A77FFB67379FF4CFDCEF576635C" box="[248,276,571,598]" captionStart="FIGURE 50" captionStartId="86.[151,250,1314,1339]" captionTargetBox="[172,1415,669,1285]" captionTargetId="figure@86.[170,1418,668,1288]" captionTargetPageId="86" captionText="FIGURE 50. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, non-type specimen (AMNH 476). A. Dorsal view. B. Anterior region, dorsal view (*: left dorsolateral tubercle). C. Anterior region, oblique left lateral view (*: left dorsolateral tubercles). Scale bars: A: 1.7 mm, B, C: 0.4 mm (photos: V.M. Conde-Vela)." figureDoi="http://doi.org/10.5281/zenodo.3672652" httpUri="https://zenodo.org/record/3672652/files/figure.png" pageId="85" pageNumber="86">50</figureCitation>
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<paragraph id="A50F36F2FFB67379FF23FD8CF6ED639B" blockId="85.[151,1436,632,780]" box="[151,655,632,657]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="85A011DCFFB67379FF23FD8CF6ED639B" box="[151,655,632,657]" pageId="85" pageNumber="86">
<treatmentCitation id="241110E3FFB67379FF23FD8CF644639B" author="Treadwell, A. L." box="[151,550,632,657]" page="1148" pageId="85" pageNumber="86" year="1906">
<taxonomicName id="62B04D71FFB67379FF23FD8CF644639B" authority="Treadwell 1906: 1148" authorityName="Treadwell" authorityPageNumber="1148" authorityYear="1906" box="[151,550,632,657]" class="Bivalvia" family="Hyriidae" genus="Castalia" kingdom="Animalia" order="Unionoida" pageId="85" pageNumber="86" phylum="Mollusca" rank="species" species="oculata">
<emphasis id="97C4EAE0FFB67379FF23FD8CF55D639B" box="[151,319,633,657]" italics="true" pageId="85" pageNumber="86">Castalia oculata</emphasis>
<bibRefCitation id="C1214B03FFB67379FEF2FD8DF644639B" author="Treadwell" box="[326,550,632,657]" firstAuthor="Treadwell" pageId="85" pageNumber="86" pagination="1145 - 1181" refId="ref89458" refString="Treadwell, A. L. (1906 [1903]) Polychaetous annelids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission, 23 (3), 1145 - 1181. [http: // docs. lib. noaa. gov / rescue / Fish _ Commission _ Bulletins / BFC 1903 - v 23. pt 3. pdf]" type="journal article" year="1906">Treadwell 1906: 1148</bibRefCitation>
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,
<figureCitation id="3D8B2A77FFB67379FD84FD8DF6E8639B" box="[560,650,632,657]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[151,250,1362,1387]" captionStartId-1="10.[151,250,1806,1831]" captionStartId-2="12.[151,250,1852,1877]" captionTargetBox-0="[151,1436,705,1338]" captionTargetBox-1="[174,1413,418,1781]" captionTargetBox-2="[178,1414,423,1820]" captionTargetId-0="figure@4.[151,1436,705,1338]" captionTargetId-1="figure@10.[170,1418,417,1781]" captionTargetId-2="figure@12.[170,1418,417,1828]" captionTargetPageId-0="4" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="FIGURE 1. Anterior end in dorsal view and exposed pharynx in frontal or oblique ventral view, rearranged after Pettibone (1970). A. Leocrates claparedii (Costa in Claparède, 1868), anterior end. B. Same, exposed pharynx. C. Leocrates oculatus (Treadwell, 1906), anterior end. D. Same, exposed pharynx. E. Leocrates diplognathus (Monro, 1926), anterior end. F. Same, exposed pharynx. G. Leocratides filamentosus Ehlers, 1908. H. Same, exposed pharynx. Scale bars: A, B: 0.4 mm, C, D, F: 0.8 mm, E: 0.3 mm, G, H: 1.0 mm (asterisk indicates nuchal organ; De: denticles; DJ: double jaw; DT: dorsolateral tubercles; LV: lateral vesicle, SJ: single jaw; TJ: transverse jaw) (Images produced by Marian Pettibone; reproduced with permission of the National Museum of Natural History, Smithsonian Institution)." captionText-1="FIGURE 2. A. Leocrates claparedii (Costa in Claparède, 1868), non-type specimens (USNM 5131), anterior end, dorsal view, pharynx everted. B. Another specimen, anterior end, dorsal view, pharynx fully everted. C. Same, pharynx and jaws, left lateral view (1: Pharynx ring I, 2: Pharynx ring II, AM: anterior margin, DT: dorsolateral tubercle, LJ: lower jaw, LV: lateral vesicle, Pe: peristomium, TB: tentacular belt, UJ: upper jaw). D. Lamprophaea poupini n. sp., holotype (UF 858), anterior end, right lateral view, after SAS (18: tentacular cirri, Ch: chaetiger, LA: lateral antenna, NO: nuchal organ lobe, Pa: palp, TB: tentacular belt). Scale bars: A, C: 0.5 mm, B: 0.7 mm, D: 0.3 mm." captionText-2="FIGURE 3. Lamprophaea longicirrata (Treadwell, 1902), new combination, non-type specimens, Florida (ECOSUR 3075), prostomia, dorsal view. A. Smallest specimen, 13x2 mm. B. Medium-sized specimen, 32x3.5 mm. C. Large specimen, pharynx slightly exposed, 32x4 mm. D. Small specimen, pharynx exposed, 20x3 mm. Scale bars: A: 0.25 mm, BC: 0.4 mm, D: 0.30 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3672549" figureDoi-1="http://doi.org/10.5281/zenodo.3672551" figureDoi-2="http://doi.org/10.5281/zenodo.3672553" httpUri-0="https://zenodo.org/record/3672549/files/figure.png" httpUri-1="https://zenodo.org/record/3672551/files/figure.png" httpUri-2="https://zenodo.org/record/3672553/files/figure.png" pageId="85" pageNumber="86">Figs 13</figureCitation>
.
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<treatmentCitationGroup id="85A011DCFFB67379FF23FD6DF65E63BA" box="[151,572,663,688]" pageId="85" pageNumber="86">
?
<treatmentCitation id="241110E3FFB67379FF16FD6CF65A63BA" author="Hartman, O." box="[162,568,663,688]" page="278" pageId="85" pageNumber="86" year="1956">
<taxonomicName id="62B04D71FFB67379FF16FD6CF65A63BA" ID-CoL="6P8X3" authority=": Hartman 1956: 278" authorityName="Hartman" authorityPageNumber="278" authorityYear="1956" box="[162,568,663,688]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="85" pageNumber="86" phylum="Annelida" rank="species" species="oculatus">
<emphasis id="97C4EAE0FFB67379FF16FD6CF50163BA" box="[162,355,664,688]" italics="true" pageId="85" pageNumber="86">Leocrates oculatus</emphasis>
:
<bibRefCitation id="C1214B03FFB67379FEC4FD6DF65A63BA" author="Hartman" box="[368,568,663,688]" firstAuthor="Hartman" pageId="85" pageNumber="86" pagination="245 - 310" refId="ref85359" refString="Hartman, O. (1956) Polychaetous annelids erected by Treadwell, 1891 to 1948, together with a brief chronology. Bulletin of the American Museum of Natural Hislory, 109, 245 - 310. [http: // digitallibrary. amnh. org / handle / 2246 / 1145]" type="journal article" year="1956">Hartman 1956: 278</bibRefCitation>
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.
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</paragraph>
<paragraph id="A50F36F2FFB67379FF23FD42F6B163C5" blockId="85.[151,1436,632,780]" box="[151,723,694,719]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="85A011DCFFB67379FF23FD42F6B163C5" box="[151,723,694,719]" pageId="85" pageNumber="86">
<treatmentCitation id="241110E3FFB67379FF23FD42F63E63C5" author="Hartman, O." box="[151,604,694,719]" page="192" pageId="85" pageNumber="86" year="1966">
<taxonomicName id="62B04D71FFB67379FF23FD42F63E63C5" ID-CoL="6P8X3" authority=": Hartman 1966: 191 - 192" authorityName="Hartman" authorityPageNumber="191 - 192" authorityYear="1966" box="[151,604,694,719]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="85" pageNumber="86" phylum="Annelida" rank="species" species="oculatus">
<emphasis id="97C4EAE0FFB67379FF23FD42F53563C4" box="[151,343,694,718]" italics="true" pageId="85" pageNumber="86">Leocrates oculatus</emphasis>
:
<bibRefCitation id="C1214B03FFB67379FED1FD42F63E63C5" author="Hartman" box="[357,604,694,719]" firstAuthor="Hartman" pageId="85" pageNumber="86" pagination="163 - 252" refId="ref85506" refString="Hartman, O. (1966) Polychaetous annelids of the Hawaiian Islands. Occasional Papers of the Bernice P. Bishop Museum, 23, 163 - 252." type="journal article" year="1966">Hartman 1966: 191192</bibRefCitation>
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(n. comb.).
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</paragraph>
<paragraph id="A50F36F2FFB67379FF23FD23F5F56206" blockId="85.[151,1436,632,780]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="85A011DCFFB67379FF23FD23F5F56206" pageId="85" pageNumber="86">
<treatmentCitation id="241110E3FFB67379FF23FD23F62D63E7" author="Pettibone, M. H." box="[151,591,724,749]" page="221" pageId="85" pageNumber="86" year="1970">
<taxonomicName id="62B04D71FFB67379FF23FD23F62D63E7" ID-CoL="6P8X5" authority=": Pettibone 1970: 219 - 221" authorityName=": Pettibone" authorityPageNumber="219 - 221" authorityYear="1970" box="[151,591,724,749]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="85" pageNumber="86" phylum="Annelida" rank="species" species="giardi">
<emphasis id="97C4EAE0FFB67379FF23FD23F52263E7" box="[151,320,725,749]" italics="true" pageId="85" pageNumber="86">Leocrates giardi</emphasis>
:
<bibRefCitation id="C1214B03FFB67379FEFBFD21F62D63E7" author="Pettibone" box="[335,591,724,749]" firstAuthor="Pettibone" pageId="85" pageNumber="86" pagination="199 - 270" refId="ref88038" refString="Pettibone, M. H. (1970) Polychaeta Errantia of the Siboga Expedition, 4. Some additional polychaetes of the Polynoidae, Hesionidae, Nereidae, Goniadidae, Eunicidae, and Onuphidae, selected as new species by the late Dr. Hermann Augener with remarks on other related species. Siboga Expeditie Monographie, 24 (1 d), 199 - 270." type="journal article" year="1970">Pettibone 1970: 219-221</bibRefCitation>
</taxonomicName>
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,
<figureCitation id="3D8B2A77FFB67379FDEFFD21F6AC63E7" box="[603,718,724,749]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStartId-0="34.[151,250,1685,1710]" captionStartId-1="37.[151,250,1269,1294]" captionStartId-2="38.[151,250,1899,1924]" captionTargetBox-0="[174,1414,459,1655]" captionTargetBox-1="[173,1416,311,1241]" captionTargetBox-2="[175,1417,852,1873]" captionTargetId-0="figure@34.[170,1418,453,1660]" captionTargetId-1="figure@37.[170,1418,308,1244]" captionTargetId-2="figure@38.[170,1418,849,1874]" captionTargetPageId-0="34" captionTargetPageId-1="37" captionTargetPageId-2="38" captionText-0="FIGURE 17. Lamprophaea cuprea Grube, 1867, reinstated, largest non-type specimen (USNM 19194). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetiger 8, right parapodium, anterior view, MGS (insets: notacicular and neuracicular lobes). D. Same, neurochaetae. Scale bars: A: 1.4 mm, B, C: 0.3 mm, D: 80 µm." captionText-1="FIGURE 18. Lamprophaea longicirrata (Treadwell, 1902), new combination, holotype (USNM 15915), MGS. A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, ventral view. D. Chaetiger 10, right parapodium, anterior view (insets: longest and shortest neurochaetal blades, and notacicular lobe). Scale bars: A: 0.7 mm, B, C: 0.6 mm, D: 0.9 mm." captionText-2="FIGURE 19. Lamprophaea ockeri n. sp., holotype (LACM 10143). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetiger 8, left parapodium, anterior view, MGS. D. Same, neurochaetal blades including longest and shortest ones. Scale bars: A: 1.2 mm, B: 0.8 mm, C: 0.5 mm, D: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3672584" figureDoi-1="http://doi.org/10.5281/zenodo.3672586" figureDoi-2="http://doi.org/10.5281/zenodo.3672588" httpUri-0="https://zenodo.org/record/3672584/files/figure.png" httpUri-1="https://zenodo.org/record/3672586/files/figure.png" httpUri-2="https://zenodo.org/record/3672588/files/figure.png" pageId="85" pageNumber="86">Figs 1719</figureCitation>
(only
<figureCitation id="3D8B2A77FFB67379FCA7FD21F73D63E7" box="[787,863,724,749]" captionStart="FIGURE 17" captionStartId="34.[151,250,1685,1710]" captionTargetBox="[174,1414,459,1655]" captionTargetId="figure@34.[170,1418,453,1660]" captionTargetPageId="34" captionText="FIGURE 17. Lamprophaea cuprea Grube, 1867, reinstated, largest non-type specimen (USNM 19194). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetiger 8, right parapodium, anterior view, MGS (insets: notacicular and neuracicular lobes). D. Same, neurochaetae. Scale bars: A: 1.4 mm, B, C: 0.3 mm, D: 80 µm." figureDoi="http://doi.org/10.5281/zenodo.3672584" httpUri="https://zenodo.org/record/3672584/files/figure.png" pageId="85" pageNumber="86">Fig. 17</figureCitation>
);
<treatmentCitation id="241110E3FFB67379FCC0FD21F17F63E7" author="Bailey-Brock, J. H. &amp; Hartman, O." box="[884,1309,724,749]" page="263" pageId="85" pageNumber="86" year="1987">
<bibRefCitation id="C1214B03FFB67379FCC0FD21F17F63E7" author="Bailey-Brock &amp; Hartman" box="[884,1309,724,749]" firstAuthor="Bailey-Brock" pageId="85" pageNumber="86" pagination="216 - 454" refId="ref82163" refString="Bailey-Brock, J. H. &amp; Hartman, O. (1987) Polychaeta. In: Devaney, D. M. &amp; Eldredge, L. G. (Eds.), Reef Shore Fauna of Hawaii. Section 2. Platyhelminthes through Phoronida &amp; Section 3. Sipuncula through Annelida. Bishop Museum Special Publication, 64, pp. 216 - 454." type="journal article" year="1987">Bailey-Brock &amp; Hartman 1987: 263264</bibRefCitation>
</treatmentCitation>
,
<figureCitation id="3D8B2A77FFB67379FA9DFD21F10763E7" box="[1321,1381,724,749]" captionStart="FIGURE 3" captionStartId="12.[151,250,1852,1877]" captionTargetBox="[178,1414,423,1820]" captionTargetId="figure@12.[170,1418,417,1828]" captionTargetPageId="12" captionText="FIGURE 3. Lamprophaea longicirrata (Treadwell, 1902), new combination, non-type specimens, Florida (ECOSUR 3075), prostomia, dorsal view. A. Smallest specimen, 13x2 mm. B. Medium-sized specimen, 32x3.5 mm. C. Large specimen, pharynx slightly exposed, 32x4 mm. D. Small specimen, pharynx exposed, 20x3 mm. Scale bars: A: 0.25 mm, BC: 0.4 mm, D: 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.3672553" httpUri="https://zenodo.org/record/3672553/files/figure.png" pageId="85" pageNumber="86">Fig. 3</figureCitation>
.II.34 (
<emphasis id="97C4EAE0FFB67379FF64FD00F4946206" box="[208,246,757,780]" italics="true" pageId="85" pageNumber="86">non</emphasis>
<bibRefCitation id="C1214B03FFB67379FF48FD06F5EE6206" author="Gravier" box="[252,396,755,780]" firstAuthor="Gravier" pageId="85" pageNumber="86" pagination="137 - 282" refId="ref84500" refString="Gravier, C. (1900) Contribution a l'etude des annelides polychetes de la Mer Rouge. Nouvelles Archives du Museum d'Histoire Naturelle, 4 eme Serie, 2 (2), 137 - 282. [http: // www. biodiversitylibrary. org / item / 113551 # page / 149 / mode / 1 up]" type="journal article" year="1900">Gravier, 1900</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFB6737AFF23FCCFF55961DD" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86" type="materials_examined">
<paragraph id="A50F36F2FFB67379FF23FCCFF5E46296" blockId="85.[151,1436,825,924]" pageId="85" pageNumber="86">
<emphasis id="97C4EAE0FFB67379FF23FCCFF55F625E" bold="true" box="[151,317,825,852]" pageId="85" pageNumber="86">Type material</emphasis>
.
<materialsCitation id="15D83CAFFFB67379FEFEFCCCF5E46296" ID-GBIF-Occurrence="2565941471" collectingDate="1902-07-18" collectionCode="USNM" county="Maui Island" elevation="317" latitude="21.609722" location="Aleunihana Channel" longLatPrecision="21" longitude="-156.0861" municipality="South of Puhielele Point" pageId="85" pageNumber="86" specimenCode="USNM 5200" specimenCount="1" stateProvince="Hawaii" typeStatus="holotype">
<emphasis id="97C4EAE0FFB67379FEFEFCCCF5C2625E" bold="true" box="[330,416,825,852]" pageId="85" pageNumber="86">
<collectingRegion id="6774F810FFB67379FEFEFCCCF5C2625E" box="[330,416,825,852]" country="United States of America" name="Hawaii" pageId="85" pageNumber="86">Hawaii</collectingRegion>
</emphasis>
.
<typeStatus id="7A0B8850FFB67379FE19FCCFF677625E" box="[429,533,826,852]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
(
<specimenCode id="F5169E89FFB67379FD97FCCFF6DA625E" box="[547,696,825,852]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="85" pageNumber="86" type="Museum">USNM 5200</specimenCode>
),
<collectingCounty id="4C6E4E7EFFB67379FD7FFCCFF731625E" box="[715,851,826,852]" pageId="85" pageNumber="86">Maui Island</collectingCounty>
,
<collectingMunicipality id="456BAC88FFB67379FCEAFCCFF00F625E" box="[862,1133,826,852]" pageId="85" pageNumber="86">South of Puhielele Point</collectingMunicipality>
(
<geoCoordinate id="C0845035FFB67379FBC8FCCCF16D625F" box="[1148,1295,825,853]" degrees="21" direction="north" minutes="36" orientation="latitude" pageId="85" pageNumber="86" precision="15" seconds="35" value="21.609722">21°3635” N</geoCoordinate>
,
<geoCoordinate id="C0845035FFB67379FAADFCCCF4D76273" degrees="156" direction="west" minutes="05" orientation="longitude" pageId="85" pageNumber="86" precision="15" seconds="10" value="-156.0861">156°0510” W</geoCoordinate>
),
<location id="A06F6029FFB67379FF73FCABF5CC6272" LSID="urn:lsid:plazi:treatment:2D1987E4FFB6737BFF23FDE2F72B60A5:A06F6029FFB67379FF73FCABF5CC6272" box="[199,430,862,888]" county="Maui Island" latitude="21.609722" longLatPrecision="21" longitude="-156.0861" municipality="South of Puhielele Point" name="Aleunihana Channel" pageId="85" pageNumber="86" stateProvince="Hawaii">Aleunihana Channel</location>
, between
<collectingRegion id="6774F810FFB67379FDA9FCABF60C6272" box="[541,622,862,888]" country="United States of America" name="Hawaii" pageId="85" pageNumber="86">Hawaii</collectingRegion>
and
<location id="A06F6029FFB67379FD17FCABF7556272" LSID="urn:lsid:plazi:treatment:2D1987E4FFB6737BFF23FDE2F72B60A5:A06F6029FFB67379FD17FCABF7556272" box="[675,823,862,888]" county="Maui Island" latitude="21.609722" longLatPrecision="21" longitude="-156.0861" municipality="South of Puhielele Point" name="Maui Islands" pageId="85" pageNumber="86" stateProvince="Hawaii">Maui Islands</location>
, USFC
<location id="A06F6029FFB67379FC3AFCABF03F6272" LSID="urn:lsid:plazi:treatment:2D1987E4FFB6737BFF23FDE2F72B60A5:A06F6029FFB67379FC3AFCABF03F6272" box="[910,1117,862,888]" county="Maui Island" latitude="21.609722" longLatPrecision="21" longitude="-156.0861" municipality="South of Puhielele Point" name="Steamer Albatross" pageId="85" pageNumber="86" stateProvince="Hawaii">Steamer Albatross</location>
, Sta. 4066, 88
<quantity id="62489B17FFB67379FABCFCABF1336273" box="[1288,1361,862,889]" metricMagnitude="2" metricUnit="m" metricValue="3.17" pageId="85" pageNumber="86" unit="m" value="317.0">
<elevation id="2E9DD1C1FFB67379FABCFCABF1336273" box="[1288,1361,862,889]" metricMagnitude="2" metricUnit="m" metricValue="3.17" pageId="85" pageNumber="86" unit="m" value="317.0">317 m</elevation>
</quantity>
, rocky bottom,
<date id="D10E1032FFB67379FF41FC77F5E36296" box="[245,385,898,924]" pageId="85" pageNumber="86" value="1902-07-18">
<collectingDate id="C14AE9DAFFB67379FF41FC77F5E36296" box="[245,385,898,924]" pageId="85" pageNumber="86" value="1902-07-18">18 Jul. 1902</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption id="F1CF667AFFB67379FF23F8B6F55F66E1" ID-DOI="http://doi.org/10.5281/zenodo.3672650" ID-Zenodo-Dep="3672650" httpUri="https://zenodo.org/record/3672650/files/figure.png" pageId="85" pageNumber="86" startId="85.[151,250,1859,1884]" targetBox="[173,1417,949,1832]" targetPageId="85">
<paragraph id="A50F36F2FFB67379FF23F8B6F55F66E1" blockId="85.[151,1437,1859,2028]" pageId="85" pageNumber="86">
<emphasis id="97C4EAE0FFB67379FF23F8B6F57B6656" bold="true" box="[151,281,1859,1884]" pageId="85" pageNumber="86">FIGURE 49</emphasis>
.
<taxonomicName id="62B04D71FFB67379FE92F8B1F6AE6656" authority="(Treadwell, 1906)" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Treadwell" baseAuthorityYear="1906" box="[294,716,1859,1884]" class="Polychaeta" family="Hesionidae" genus="Paradalhousia" kingdom="Animalia" order="Phyllodocida" pageId="85" pageNumber="86" phylum="Annelida" rank="species" species="oculata" status="comb. nov.">
<emphasis id="97C4EAE0FFB67379FE92F8B1F66C6656" box="[294,526,1860,1884]" italics="true" pageId="85" pageNumber="86">Paradalhousia oculata</emphasis>
(
<bibRefCitation id="C1214B03FFB67379FDAFF8B6F6A76656" author="Treadwell" box="[539,709,1859,1884]" firstAuthor="Treadwell" pageId="85" pageNumber="86" pagination="1145 - 1181" refId="ref89458" refString="Treadwell, A. L. (1906 [1903]) Polychaetous annelids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission, 23 (3), 1145 - 1181. [http: // docs. lib. noaa. gov / rescue / Fish _ Commission _ Bulletins / BFC 1903 - v 23. pt 3. pdf]" type="journal article" year="1906">Treadwell, 1906</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="8CF7579BFFB67379FD63F8B1F7EA6656" box="[727,904,1859,1884]" pageId="85" pageNumber="86" rank="species">new combination</taxonomicNameLabel>
, reinstated, holotype (USNM 5200). A. Dorsal view, depressed by glass slide. B. Anterior end, dorsal view (*: denticle). C. Same, MGS (*: denticle). D. Pharynx, frontal view, MGS, denticles slightly darker than surrounding areas. E. Middle chaetiger, left parapodium, anterior view (insets: (left) notacicular and (right) neuracicular lobes tips, and medium- and small neurochaetal blades). Scale bars: A: 1 mm, B, C: 0.3 mm, D: 0.5 mm, E: 0.4 mm.
</paragraph>
</caption>
<paragraph id="A50F36F2FFB5737AFF73FF6DF55961DD" blockId="86.[151,1437,152,646]" pageId="86" pageNumber="87">
<materialsCitation id="15D83CAFFFB5737AFF73FF6DF55561DD" ID-GBIF-Occurrence="2565941449" collectionCode="AMNH" location="One" pageId="86" pageNumber="87" specimenCode="AMNH 476" specimenCount="2" stateProvince="Hawaii" typeStatus="syntype">
<emphasis id="97C4EAE0FFB5737AFF73FF6DF5D061B9" bold="true" box="[199,434,152,179]" pageId="86" pageNumber="87">Additional material</emphasis>
.
<emphasis id="97C4EAE0FFB5737AFE74FF6DF67461B9" bold="true" box="[448,534,152,179]" pageId="86" pageNumber="87">
<collectingRegion id="6774F810FFB5737AFE74FF6DF67461B9" box="[448,534,152,179]" country="United States of America" name="Hawaii" pageId="86" pageNumber="87">Hawaii</collectingRegion>
</emphasis>
.
<location id="A06F6029FFB5737AFD90FF6CF63161B9" LSID="urn:lsid:plazi:treatment:2D1987E4FFB6737BFF23FDE2F72B60A5:A06F6029FFB5737AFD90FF6CF63161B9" box="[548,595,153,179]" name="One" pageId="86" pageNumber="87" stateProvince="Hawaii">One</location>
specimen (
<specimenCode id="F5169E89FFB5737AFD65FF6CF73F61B9" box="[721,861,152,179]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="86" pageNumber="87">AMNH 476</specimenCode>
), no further data [labelled as
<typeStatus id="7A0B8850FFB5737AFB16FF6CF09F61B9" box="[1186,1277,153,179]" pageId="86" pageNumber="87" type="syntype">syntype</typeStatus>
;
<quantity id="62489B17FFB5737AFAB3FF6CF13F61B9" box="[1287,1373,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="86" pageNumber="87" unit="mm" value="7.1">7.1 mm</quantity>
long,
<quantity id="62489B17FFB5737AFF23FF48F48F61DD" box="[151,237,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="86" pageNumber="87" unit="mm" value="2.3">2.3 mm</quantity>
wide]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFB5737AFF73FF15F5AF6639" pageId="86" pageNumber="87" type="description">
<paragraph id="A50F36F2FFB5737AFF73FF15F540606C" blockId="86.[151,1437,152,646]" pageId="86" pageNumber="87">
<emphasis id="97C4EAE0FFB5737AFF73FF15F53361F1" bold="true" box="[199,337,224,251]" pageId="86" pageNumber="87">Description</emphasis>
.
<typeStatus id="7A0B8850FFB5737AFEEBFF15F5A561F0" box="[351,455,224,250]" pageId="86" pageNumber="87" type="holotype">Holotype</typeStatus>
(USNM 5200) complete, bent ventrally, breaking into two (
<figureCitation id="3D8B2A77FFB5737AFBD5FF15F0AB61F1" box="[1121,1225,224,251]" captionStart="FIGURE 49" captionStartId="85.[151,250,1859,1884]" captionTargetBox="[173,1417,949,1832]" captionTargetId="figure@85.[170,1418,946,1834]" captionTargetPageId="85" captionText="FIGURE 49. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, holotype (USNM 5200). A. Dorsal view, depressed by glass slide. B. Anterior end, dorsal view (*: denticle). C. Same, MGS (*: denticle). D. Pharynx, frontal view, MGS, denticles slightly darker than surrounding areas. E. Middle chaetiger, left parapodium, anterior view (insets: (left) notacicular and (right) neuracicular lobes tips, and medium- and small neurochaetal blades). Scale bars: A: 1 mm, B, C: 0.3 mm, D: 0.5 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672650" httpUri="https://zenodo.org/record/3672650/files/figure.png" pageId="86" pageNumber="87">Fig. 49A</figureCitation>
), pharynx fully exposed; left parapodia of chaetigers 69, and right parapodia of chaetigers 67 previously removed (all in container). Body colorless,
<quantity id="62489B17FFB5737AFEFAFEDCF5FF6049" box="[334,413,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="86" pageNumber="87" unit="mm" value="11.0">11 mm</quantity>
long,
<quantity id="62489B17FFB5737AFE50FEDCF6446049" box="[484,550,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="86" pageNumber="87" unit="mm" value="2.0">2 mm</quantity>
wide, 16 chaetigers; eyes brownish, middorsal peristomial tubercle pale; most cirri broken.
</paragraph>
<paragraph id="A50F36F2FFB5737AFF73FE85F76B60D9" blockId="86.[151,1437,152,646]" pageId="86" pageNumber="87">
Prostomium slightly longer than wide, slightly wider anteriorly (
<figureCitation id="3D8B2A77FFB5737AFC18FE85F05A6081" box="[940,1080,368,395]" captionStart="FIGURE 49" captionStartId="85.[151,250,1859,1884]" captionTargetBox="[173,1417,949,1832]" captionTargetId="figure@85.[170,1418,946,1834]" captionTargetPageId="85" captionText="FIGURE 49. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, holotype (USNM 5200). A. Dorsal view, depressed by glass slide. B. Anterior end, dorsal view (*: denticle). C. Same, MGS (*: denticle). D. Pharynx, frontal view, MGS, denticles slightly darker than surrounding areas. E. Middle chaetiger, left parapodium, anterior view (insets: (left) notacicular and (right) neuracicular lobes tips, and medium- and small neurochaetal blades). Scale bars: A: 1 mm, B, C: 0.3 mm, D: 0.5 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672650" httpUri="https://zenodo.org/record/3672650/files/figure.png" pageId="86" pageNumber="87">Fig. 49B, C</figureCitation>
). Lateral antennae with ceratophores distinct, longer than prostomium, slightly longer than palps; palpophores slightly longer than palpostyles. Median antenna broken, inserted between anterior eyes.
</paragraph>
<paragraph id="A50F36F2FFB5737AFF73FE28F0DE6311" blockId="86.[151,1437,152,646]" pageId="86" pageNumber="87">
Eyes brownish, round; anterior eyes emarginate anterolaterally, slightly larger and slightly more distant to each other than posterior round ones; in lateral view anterior and posterior eyes fused (
<figureCitation id="3D8B2A77FFB5737AFB9CFDF5F0CD6311" box="[1064,1199,512,539]" captionStart="FIGURE 49" captionStartId="85.[151,250,1859,1884]" captionTargetBox="[173,1417,949,1832]" captionTargetId="figure@85.[170,1418,946,1834]" captionTargetPageId="85" captionText="FIGURE 49. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, holotype (USNM 5200). A. Dorsal view, depressed by glass slide. B. Anterior end, dorsal view (*: denticle). C. Same, MGS (*: denticle). D. Pharynx, frontal view, MGS, denticles slightly darker than surrounding areas. E. Middle chaetiger, left parapodium, anterior view (insets: (left) notacicular and (right) neuracicular lobes tips, and medium- and small neurochaetal blades). Scale bars: A: 1 mm, B, C: 0.3 mm, D: 0.5 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672650" httpUri="https://zenodo.org/record/3672650/files/figure.png" pageId="86" pageNumber="87">Fig. 49B, C</figureCitation>
).
</paragraph>
<paragraph id="A50F36F2FFB5737AFF73FDD1F6CA638C" blockId="86.[151,1437,152,646]" pageId="86" pageNumber="87">Nuchal organs lobes U-shaped, parallel, prostomial posterior projections pale; middle furrow deep. Lateral ciliated bands visible dorsally. Tentacular cirri mostly broken or without tips, longest one reaching chaetiger 6. Lateral cushions low, entire, longitudinal striae visible.</paragraph>
<caption id="F1CF667AFFB5737AFF23FAD7F0796489" ID-DOI="http://doi.org/10.5281/zenodo.3672652" ID-Zenodo-Dep="3672652" httpUri="https://zenodo.org/record/3672652/files/figure.png" pageId="86" pageNumber="87" startId="86.[151,250,1314,1339]" targetBox="[172,1415,669,1285]" targetPageId="86">
<paragraph id="A50F36F2FFB5737AFF23FAD7F0796489" blockId="86.[151,1437,1314,1411]" pageId="86" pageNumber="87">
<emphasis id="97C4EAE0FFB5737AFF23FAD7F57E6431" bold="true" box="[151,284,1314,1339]" pageId="86" pageNumber="87">FIGURE 50</emphasis>
.
<taxonomicName id="62B04D71FFB5737AFE98FAD7F6B96431" authority="(Treadwell, 1906)" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Treadwell" baseAuthorityYear="1906" box="[300,731,1314,1339]" class="Polychaeta" family="Hesionidae" genus="Paradalhousia" kingdom="Animalia" order="Phyllodocida" pageId="86" pageNumber="87" phylum="Annelida" rank="species" species="oculata" status="comb. nov.">
<emphasis id="97C4EAE0FFB5737AFE98FAD7F6756430" box="[300,535,1314,1338]" italics="true" pageId="86" pageNumber="87">Paradalhousia oculata</emphasis>
(
<bibRefCitation id="C1214B03FFB5737AFD93FAD7F6B66431" author="Treadwell" box="[551,724,1314,1339]" firstAuthor="Treadwell" pageId="86" pageNumber="87" pagination="1145 - 1181" refId="ref89458" refString="Treadwell, A. L. (1906 [1903]) Polychaetous annelids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission, 23 (3), 1145 - 1181. [http: // docs. lib. noaa. gov / rescue / Fish _ Commission _ Bulletins / BFC 1903 - v 23. pt 3. pdf]" type="journal article" year="1906">Treadwell, 1906</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="8CF7579BFFB5737AFD5DFAD6F7FF6431" box="[745,925,1314,1339]" pageId="86" pageNumber="87" rank="species">new combination</taxonomicNameLabel>
, reinstated, non-type specimen (AMNH 476). A. Dorsal view. B. Anterior region, dorsal view (*: left dorsolateral tubercle). C. Anterior region, oblique left lateral view (*: left dorsolateral tubercles). Scale bars: A: 1.7 mm, B, C: 0.4 mm (photos: V.M. Conde-Vela).
</paragraph>
</caption>
<paragraph id="A50F36F2FFB5737AFF73FA20F7C26719" blockId="86.[151,1437,1493,2023]" pageId="86" pageNumber="87">
Pharynx fully everted; anterior margin with 24 denticles, middorsal and midventral areas bare (
<figureCitation id="3D8B2A77FFB5737AFB40FA20F13C64E5" box="[1268,1374,1493,1519]" captionStart="FIGURE 49" captionStartId="85.[151,250,1859,1884]" captionTargetBox="[173,1417,949,1832]" captionTargetId="figure@85.[170,1418,946,1834]" captionTargetPageId="85" captionText="FIGURE 49. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, holotype (USNM 5200). A. Dorsal view, depressed by glass slide. B. Anterior end, dorsal view (*: denticle). C. Same, MGS (*: denticle). D. Pharynx, frontal view, MGS, denticles slightly darker than surrounding areas. E. Middle chaetiger, left parapodium, anterior view (insets: (left) notacicular and (right) neuracicular lobes tips, and medium- and small neurochaetal blades). Scale bars: A: 1 mm, B, C: 0.3 mm, D: 0.5 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672650" httpUri="https://zenodo.org/record/3672650/files/figure.png" pageId="86" pageNumber="87">Fig. 49D</figureCitation>
). Upper and lower jaws single, visible, exposed, brownish in frontal view.
</paragraph>
<paragraph id="A50F36F2FFB5737AFF73F9E8F1E367C2" blockId="86.[151,1437,1493,2023]" pageId="86" pageNumber="87">
Dorsal cirri without tips, about as long as body width (excluding parapodia). Chaetigers 13 without notochaetae, notochaetae present along chaetigers 416, about 50 per bundle, as long as neurochaetae, subdistally denticulate, denticles fine, small. Notacicular and neuracicular lobes tapered, blunt (
<figureCitation id="3D8B2A77FFB5737AFBB8F990F0116775" box="[1036,1139,1637,1663]" captionStart="FIGURE 49" captionStartId="85.[151,250,1859,1884]" captionTargetBox="[173,1417,949,1832]" captionTargetId="figure@85.[170,1418,946,1834]" captionTargetPageId="85" captionText="FIGURE 49. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, holotype (USNM 5200). A. Dorsal view, depressed by glass slide. B. Anterior end, dorsal view (*: denticle). C. Same, MGS (*: denticle). D. Pharynx, frontal view, MGS, denticles slightly darker than surrounding areas. E. Middle chaetiger, left parapodium, anterior view (insets: (left) notacicular and (right) neuracicular lobes tips, and medium- and small neurochaetal blades). Scale bars: A: 1 mm, B, C: 0.3 mm, D: 0.5 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672650" httpUri="https://zenodo.org/record/3672650/files/figure.png" pageId="86" pageNumber="87">Fig. 49E</figureCitation>
, insets); notacicular lobes twice longer than wide, neuracicular ones as long as wide. Neurochaetae about 50 per bundle, blades decreasing in size ventrally, 67 times longer than wide; blades bidentate, guard approaching subdistal tooth (
<figureCitation id="3D8B2A77FFB5737AFB7CF958F14B67CD" box="[1224,1321,1709,1735]" captionStart="FIGURE 49" captionStartId="85.[151,250,1859,1884]" captionTargetBox="[173,1417,949,1832]" captionTargetId="figure@85.[170,1418,946,1834]" captionTargetPageId="85" captionText="FIGURE 49. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, holotype (USNM 5200). A. Dorsal view, depressed by glass slide. B. Anterior end, dorsal view (*: denticle). C. Same, MGS (*: denticle). D. Pharynx, frontal view, MGS, denticles slightly darker than surrounding areas. E. Middle chaetiger, left parapodium, anterior view (insets: (left) notacicular and (right) neuracicular lobes tips, and medium- and small neurochaetal blades). Scale bars: A: 1 mm, B, C: 0.3 mm, D: 0.5 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672650" httpUri="https://zenodo.org/record/3672650/files/figure.png" pageId="86" pageNumber="87">Fig. 49E</figureCitation>
insets).
</paragraph>
<paragraph id="A50F36F2FFB5737AFF73F924F5DF6605" blockId="86.[151,1437,1493,2023]" pageId="86" pageNumber="87">Posterior region tapered. Prepygidial segment without dorsal cirri, ventral cirri medially broken. Pygidium tubular, anal cirri missing.</paragraph>
<paragraph id="A50F36F2FFB5737AFF73F8ECF5AF6639" blockId="86.[151,1437,1493,2023]" box="[199,461,1817,1843]" pageId="86" pageNumber="87">
Oocytes about
<quantity id="62489B17FFB5737AFEDBF8ECF5AB6639" box="[367,457,1817,1843]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="86" pageNumber="87" unit="mm" value="100.0">100 µm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFB5737BFF73F8C8F6306081" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87" type="discussion">
<paragraph id="A50F36F2FFB5737BFF73F8C8F0D161DD" blockId="86.[151,1437,1493,2023]" lastBlockId="87.[151,1437,152,431]" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87">
<emphasis id="97C4EAE0FFB5737AFF73F8C8F556665D" bold="true" box="[199,308,1853,1879]" pageId="86" pageNumber="87">Remarks</emphasis>
.
<taxonomicName id="62B04D71FFB5737AFEF0F8C8F6A0665C" authority="Treadwell, 1906" authorityName="Treadwell" authorityYear="1906" box="[324,706,1852,1879]" class="Bivalvia" family="Hyriidae" genus="Castalia" kingdom="Animalia" order="Unionoida" pageId="86" pageNumber="87" phylum="Mollusca" rank="species" species="oculata">
<emphasis id="97C4EAE0FFB5737AFEF0F8C8F663665D" box="[324,513,1853,1879]" italics="true" pageId="86" pageNumber="87">Castalia oculata</emphasis>
<bibRefCitation id="C1214B03FFB5737AFDBDF8C8F6A0665C" author="Treadwell" box="[521,706,1852,1879]" firstAuthor="Treadwell" pageId="86" pageNumber="87" pagination="1145 - 1181" refId="ref89458" refString="Treadwell, A. L. (1906 [1903]) Polychaetous annelids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission, 23 (3), 1145 - 1181. [http: // docs. lib. noaa. gov / rescue / Fish _ Commission _ Bulletins / BFC 1903 - v 23. pt 3. pdf]" type="journal article" year="1906">Treadwell, 1906</bibRefCitation>
</taxonomicName>
must be reinstated and newly combined.
<bibRefCitation id="C1214B03FFB5737AFB17F8C8F1FE665D" author="Pettibone" box="[1187,1436,1853,1879]" firstAuthor="Pettibone" pageId="86" pageNumber="87" pagination="199 - 270" refId="ref88038" refString="Pettibone, M. H. (1970) Polychaeta Errantia of the Siboga Expedition, 4. Some additional polychaetes of the Polynoidae, Hesionidae, Nereidae, Goniadidae, Eunicidae, and Onuphidae, selected as new species by the late Dr. Hermann Augener with remarks on other related species. Siboga Expeditie Monographie, 24 (1 d), 199 - 270." type="journal article" year="1970">Pettibone (1970: 219)</bibRefCitation>
regarded it as a junior synonym of
<taxonomicName id="62B04D71FFB5737AFD97F894F7E06671" authority="Gravier, 1900" authorityName="Gravier" authorityYear="1900" box="[547,898,1889,1915]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="86" pageNumber="87" phylum="Annelida" rank="species" species="giardi">
<emphasis id="97C4EAE0FFB5737AFD97F894F6BE6671" box="[547,732,1889,1915]" italics="true" pageId="86" pageNumber="87">Leocrates giardi</emphasis>
<bibRefCitation id="C1214B03FFB5737AFD50F894F7E06671" author="Gravier" box="[740,898,1889,1915]" firstAuthor="Gravier" pageId="86" pageNumber="87" pagination="137 - 282" refId="ref84500" refString="Gravier, C. (1900) Contribution a l'etude des annelides polychetes de la Mer Rouge. Nouvelles Archives du Museum d'Histoire Naturelle, 4 eme Serie, 2 (2), 137 - 282. [http: // www. biodiversitylibrary. org / item / 113551 # page / 149 / mode / 1 up]" type="journal article" year="1900">Gravier, 1900</bibRefCitation>
</taxonomicName>
, but they belong to different genera. Further,
<taxonomicName id="62B04D71FFB5737AFA37F894F4896695" authorityName="Treadwell" authorityYear="1906" class="Bivalvia" family="Hyriidae" genus="Castalia" kingdom="Animalia" order="Unionoida" pageId="86" pageNumber="87" phylum="Mollusca" rank="species" species="oculata">
<emphasis id="97C4EAE0FFB5737AFA37F894F4896695" italics="true" pageId="86" pageNumber="87">C. oculata</emphasis>
</taxonomicName>
deserves a new combination into
<taxonomicName id="62B04D71FFB5737AFDDAF870F7766695" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[622,788,1925,1951]" class="Polychaeta" family="Hesionidae" genus="Paradalhousia" kingdom="Animalia" order="Phyllodocida" pageId="86" pageNumber="81" phylum="Annelida" rank="genus" status="gen. nov.">
<emphasis id="97C4EAE0FFB5737AFDDAF870F7766695" box="[622,788,1925,1951]" italics="true" pageId="86" pageNumber="87">Paradalhousia</emphasis>
</taxonomicName>
because it has U-shaped nuchal organs, single upper and lower jaws, and pale neurochaetae. On the other hand,
<taxonomicName id="62B04D71FFB5737AFCB1F85CF02A66C9" authority="(Treadwell, 1906)" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Treadwell" baseAuthorityYear="1906" box="[773,1096,1960,1987]" class="Bivalvia" family="Hyriidae" genus="Castalia" kingdom="Animalia" order="Unionoida" pageId="86" pageNumber="87" phylum="Mollusca" rank="species" species="oculata" status="comb. nov.">
<emphasis id="97C4EAE0FFB5737AFCB1F85CF71466C9" box="[773,886,1961,1987]" italics="true" pageId="86" pageNumber="87">P. oculata</emphasis>
(
<bibRefCitation id="C1214B03FFB5737AFC31F85CF05D66C9" author="Treadwell" box="[901,1087,1960,1987]" firstAuthor="Treadwell" pageId="86" pageNumber="87" pagination="1145 - 1181" refId="ref89458" refString="Treadwell, A. L. (1906 [1903]) Polychaetous annelids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission, 23 (3), 1145 - 1181. [http: // docs. lib. noaa. gov / rescue / Fish _ Commission _ Bulletins / BFC 1903 - v 23. pt 3. pdf]" type="journal article" year="1906">Treadwell, 1906</bibRefCitation>
)
</taxonomicName>
<emphasis id="97C4EAE0FFB5737AFBFBF85CF0D466C9" bold="true" box="[1103,1206,1961,1987]" pageId="86" pageNumber="87">
<taxonomicNameLabel id="8CF7579BFFB5737AFBFBF85CF0D466C9" box="[1103,1206,1961,1987]" pageId="86" pageNumber="87" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
, resembles
<taxonomicName id="62B04D71FFB5737AFA89F85CF51766ED" authority="(Monro, 1926)" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Monro" baseAuthorityYear="1926" class="Polychaeta" family="Hesionidae" genus="Paradalhousia" kingdom="Animalia" order="Phyllodocida" pageId="86" pageNumber="87" phylum="Annelida" rank="species" species="papillosa">
<emphasis id="97C4EAE0FFB5737AFA89F85CF4A466ED" italics="true" pageId="86" pageNumber="87">P. papillosa</emphasis>
(
<bibRefCitation id="C1214B03FFB5737AFF62F838F50E66ED" author="Monro" box="[214,364,1996,2023]" firstAuthor="Monro" pageId="86" pageNumber="87" pagination="311 - 323" refId="ref87314" refString="Monro, C. C. A. (1926) Polychaeta of the H. M. S. Alert Expedition, 1881 - 1882. Families Hesionidae and Nereidae. Journal of the Linnaean Society, Zoology, 36, 311 - 323. [http: // onlinelibrary. wiley. com / doi / 10.1111 / j. 1096 - 3642.1926. tb 02172. x / epdf] https: // doi. org / 10.1111 / j. 1096 - 3642.1926. tb 02172. x" type="journal article" year="1926">Monro, 1926</bibRefCitation>
)
</taxonomicName>
reinst.,
<emphasis id="97C4EAE0FFB5737AFE65F838F65B66ED" bold="true" box="[465,569,1997,2023]" pageId="86" pageNumber="87">n. comb.</emphasis>
but they differ in the number of neurochaetae and in the orientation of their teeth. In
<taxonomicName id="62B04D71FFB4737BFF4DFF6FF50B61B9" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Treadwell" baseAuthorityYear="1906" box="[249,361,153,179]" class="Bivalvia" family="Hyriidae" genus="Castalia" kingdom="Animalia" order="Unionoida" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="oculata">
<emphasis id="97C4EAE0FFB4737BFF4DFF6FF50B61B9" box="[249,361,153,179]" italics="true" pageId="87" pageNumber="88">P. oculata</emphasis>
</taxonomicName>
there are about 50 neurochaetae per bundle, and their distal teeth are slightly falcate, whereas in
<taxonomicName id="62B04D71FFB4737BFF23FF4BF57961DD" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Monro" baseAuthorityYear="1926" box="[151,283,189,215]" class="Polychaeta" family="Hesionidae" genus="Paradalhousia" kingdom="Animalia" order="Phyllodocida" pageId="87" pageNumber="88" phylum="Annelida" rank="species" species="papillosa">
<emphasis id="97C4EAE0FFB4737BFF23FF4BF57961DD" box="[151,283,189,215]" italics="true" pageId="87" pageNumber="88">P. papillosa</emphasis>
</taxonomicName>
neurochaetae number about 20(30) per bundle, and their distal teeth are straight.
</paragraph>
<paragraph id="A50F36F2FFB4737BFF73FF14F6306081" blockId="87.[151,1437,152,431]" pageId="87" pageNumber="88">
The specimen deposited in
<collectingRegion id="6774F810FFB4737BFE4FFF14F60F61F1" box="[507,621,225,251]" country="United States of America" name="New York" pageId="87" pageNumber="88">New York</collectingRegion>
(AMNH 476) is labeled as
<typeStatus id="7A0B8850FFB4737BFC13FF14F06061F1" box="[935,1026,225,251]" pageId="87" pageNumber="88" type="syntype">syntype</typeStatus>
. However, it was not included in the original description, which is the requisite in order to be included as a
<typeStatus id="7A0B8850FFB4737BFC17FEF0F7986015" box="[931,1018,261,287]" pageId="87" pageNumber="88" type="syntype">syntype</typeStatus>
(
<bibRefCitation id="C1214B03FFB4737BFBBCFEF0F0EE6015" author="ICZN" box="[1032,1164,261,287]" firstAuthor="ICZN" pageId="87" pageNumber="88" refId="ref85916" refString="ICZN (International Commission on Zoological Nomenclature). (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature in association with the British Museum (Natural History), London, 306 pp. [http: // www. iczn. org / iczn / index. jsp]" type="book" year="1999">ICZN 1999</bibRefCitation>
,
<collectingRegion id="6774F810FFB4737BFB21FEF0F0DC6015" box="[1173,1214,261,287]" country="Djibouti" name="Arta" pageId="87" pageNumber="88">Art</collectingRegion>
. 72.4.1), and consequently it is herein regarded as an additional specimen. However, it completely matches the
<typeStatus id="7A0B8850FFB4737BFB2DFEDCF0996049" box="[1177,1275,297,323]" pageId="87" pageNumber="88" type="holotype">holotype</typeStatus>
in body shape (
<figureCitation id="3D8B2A77FFB4737BFF14FEB8F56A606D" box="[160,264,332,359]" captionStart="FIGURE 50" captionStartId="86.[151,250,1314,1339]" captionTargetBox="[172,1415,669,1285]" captionTargetId="figure@86.[170,1418,668,1288]" captionTargetPageId="86" captionText="FIGURE 50. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, non-type specimen (AMNH 476). A. Dorsal view. B. Anterior region, dorsal view (*: left dorsolateral tubercle). C. Anterior region, oblique left lateral view (*: left dorsolateral tubercles). Scale bars: A: 1.7 mm, B, C: 0.4 mm (photos: V.M. Conde-Vela)." figureDoi="http://doi.org/10.5281/zenodo.3672652" httpUri="https://zenodo.org/record/3672652/files/figure.png" pageId="87" pageNumber="88">Fig. 50A</figureCitation>
), anterior end appendages (
<figureCitation id="3D8B2A77FFB4737BFD8DFEB8F6FD606D" box="[569,671,332,359]" captionStart="FIGURE 50" captionStartId="86.[151,250,1314,1339]" captionTargetBox="[172,1415,669,1285]" captionTargetId="figure@86.[170,1418,668,1288]" captionTargetPageId="86" captionText="FIGURE 50. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, non-type specimen (AMNH 476). A. Dorsal view. B. Anterior region, dorsal view (*: left dorsolateral tubercle). C. Anterior region, oblique left lateral view (*: left dorsolateral tubercles). Scale bars: A: 1.7 mm, B, C: 0.4 mm (photos: V.M. Conde-Vela)." figureDoi="http://doi.org/10.5281/zenodo.3672652" httpUri="https://zenodo.org/record/3672652/files/figure.png" pageId="87" pageNumber="88">Fig. 50B</figureCitation>
), and pharynx features (
<figureCitation id="3D8B2A77FFB4737BFC18FEB8F071606D" box="[940,1043,332,359]" captionStart="FIGURE 50" captionStartId="86.[151,250,1314,1339]" captionTargetBox="[172,1415,669,1285]" captionTargetId="figure@86.[170,1418,668,1288]" captionTargetPageId="86" captionText="FIGURE 50. Paradalhousia oculata (Treadwell, 1906), new combination, reinstated, non-type specimen (AMNH 476). A. Dorsal view. B. Anterior region, dorsal view (*: left dorsolateral tubercle). C. Anterior region, oblique left lateral view (*: left dorsolateral tubercles). Scale bars: A: 1.7 mm, B, C: 0.4 mm (photos: V.M. Conde-Vela)." figureDoi="http://doi.org/10.5281/zenodo.3672652" httpUri="https://zenodo.org/record/3672652/files/figure.png" pageId="87" pageNumber="88">Fig. 50C</figureCitation>
), rendering it enigmatic why it was not included in the original description.
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFB4737BFF73FE61F72B60A5" box="[199,841,404,431]" pageId="87" pageNumber="88" type="distribution">
<paragraph id="A50F36F2FFB4737BFF73FE61F72B60A5" blockId="87.[151,1437,152,431]" box="[199,841,404,431]" pageId="87" pageNumber="88">
<emphasis id="97C4EAE0FFB4737BFF73FE61F53B60A5" bold="true" box="[199,345,404,431]" pageId="87" pageNumber="88">Distribution</emphasis>
.
<collectingRegion id="6774F810FFB4737BFED3FE60F5D960A5" box="[359,443,405,431]" country="United States of America" name="Hawaii" pageId="87" pageNumber="88">Hawaii</collectingRegion>
, rocky bottoms in
<quantity id="62489B17FFB4737BFD3FFE60F69E60A5" box="[651,764,405,431]" metricMagnitude="2" metricUnit="m" metricValue="2.025" metricValueMax="3.17" metricValueMin="0.88" pageId="87" pageNumber="88" unit="m" value="202.5" valueMax="317.0" valueMin="88.0">88317 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>