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<mods:title id="B43222315FEDA3F0A2A94D180A0B55E8">Revision of Leocrates Kinberg, 1866 and Leocratides Ehlers, 1908 (Annelida, Errantia, Hesionidae)</mods:title>
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<taxonomicName id="62B04D71FFAB7364FF23F926F5A467E7" ID-CoL="84R2X" ID-ENA="1979131" authority="Ehlers, 1908" authorityName="Ehlers" authorityYear="1908" box="[151,454,1747,1774]" class="Polychaeta" family="Hesionidae" genus="Leocratides" kingdom="Animalia" order="Phyllodocida" pageId="72" pageNumber="73" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAB7364FF23F926F5A467E7" bold="true" box="[151,454,1747,1774]" pageId="72" pageNumber="73">
<emphasis id="97C4EAE0FFAB7364FF23F926F54167E4" bold="true" box="[151,291,1747,1774]" italics="true" pageId="72" pageNumber="73">Leocratides</emphasis>
<bibRefCitation id="C1214B03FFAB7364FE9FF921F5A467E7" author="Ehlers" box="[299,454,1747,1774]" firstAuthor="Ehlers" pageId="72" pageNumber="73" pagination="1 - 167" refId="ref83389" refString="Ehlers, E. (1908) Die bodensassigen Anneliden aus den Sammlungen der deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition aud dem Dampfer, Valdivia &quot; 1898 - 1899, 2 (1), 1 - 167, pls. 1 - 23. [https: // www. biodiversitylibrary. org / item / 18687 # page / 3 / mode / 1 up]" type="journal article" year="1908">Ehlers, 1908</bibRefCitation>
</emphasis>
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<subSubSection id="EDAA6579FFAB7364FF23F8EDF5FB6645" pageId="72" pageNumber="73" type="reference_group">
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<treatmentCitation id="241110E3FFAB7364FF23F8EDF5DF663A" author="Ehlers, E." box="[151,445,1815,1840]" page="63" pageId="72" pageNumber="73" year="1908">
<taxonomicName id="62B04D71FFAB7364FF23F8EDF5DF663A" ID-CoL="84R2X" authority="Ehlers, 1908: 63" authorityName="Ehlers" authorityPageNumber="63" authorityYear="1908" box="[151,445,1815,1840]" class="Polychaeta" family="Hesionidae" genus="Leocratides" kingdom="Animalia" order="Phyllodocida" pageId="72" pageNumber="73" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAB7364FF23F8EDF56D663A" box="[151,271,1816,1840]" italics="true" pageId="72" pageNumber="73">Leocratides</emphasis>
<bibRefCitation id="C1214B03FFAB7364FEA1F8E2F5DF663A" author="Ehlers" box="[277,445,1815,1840]" firstAuthor="Ehlers" pageId="72" pageNumber="73" pagination="1 - 167" refId="ref83389" refString="Ehlers, E. (1908) Die bodensassigen Anneliden aus den Sammlungen der deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition aud dem Dampfer, Valdivia &quot; 1898 - 1899, 2 (1), 1 - 167, pls. 1 - 23. [https: // www. biodiversitylibrary. org / item / 18687 # page / 3 / mode / 1 up]" type="journal article" year="1908">Ehlers, 1908: 63</bibRefCitation>
</taxonomicName>
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.
<treatmentCitation id="241110E3FFAB7364FE73F8E2F6CF663A" author="Chamberlin, R. V." box="[455,685,1815,1840]" page="185" pageId="72" pageNumber="73" year="1919">
<bibRefCitation id="C1214B03FFAB7364FE73F8E2F6CF663A" author="Chamberlin" box="[455,685,1815,1840]" firstAuthor="Chamberlin" pageId="72" pageNumber="73" pagination="1 - 514" refId="ref82463" refString="Chamberlin, R. V. (1919) The Annelida Polychaeta of the Albatross Tropical Pacific Expedition, 1891 - 1905. Memoirs of the Museum of Comparative Zoology, Harvard, 31, 1 - 514. [http: // www. biodiversitylibrary. org / bibliography / 49195 # / summary]" type="journal article" year="1919">Chamberlin 1919: 185</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="241110E3FFAB7364FD03F8E2F715663A" author="Fauvel, P." box="[695,887,1815,1840]" page="107" pageId="72" pageNumber="73" year="1953">
<bibRefCitation id="C1214B03FFAB7364FD03F8E2F715663A" author="Fauvel" box="[695,887,1815,1840]" firstAuthor="Fauvel" pageId="72" pageNumber="73" refId="ref84117" refString="Fauvel, P. (1953 b) The Fauna of India including Pakistan, Ceylon, Burma and Malaya: Annelida Polychaeta. Indian Press, Allahabad, 507 pp. [https: // archive. org / details / FBIPolychaeta]" type="book" year="1953" yearSuffix="b">Fauvel 1953b: 107</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="241110E3FFAB7364FC35F8E2F0E0663A" author="Pettibone, M. H." box="[897,1154,1815,1840]" page="229" pageId="72" pageNumber="73" year="1970">
<bibRefCitation id="C1214B03FFAB7364FC35F8E2F0E0663A" author="Pettibone" box="[897,1154,1815,1840]" firstAuthor="Pettibone" pageId="72" pageNumber="73" pagination="199 - 270" refId="ref88038" refString="Pettibone, M. H. (1970) Polychaeta Errantia of the Siboga Expedition, 4. Some additional polychaetes of the Polynoidae, Hesionidae, Nereidae, Goniadidae, Eunicidae, and Onuphidae, selected as new species by the late Dr. Hermann Augener with remarks on other related species. Siboga Expeditie Monographie, 24 (1 d), 199 - 270." type="journal article" year="1970">Pettibone 1970: 229230</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="241110E3FFAB7364FB38F8E2F129663A" author="Fauchald, K." box="[1164,1355,1815,1840]" page="76" pageId="72" pageNumber="73" year="1977">
<bibRefCitation id="C1214B03FFAB7364FB38F8E2F129663A" author="Fauchald" box="[1164,1355,1815,1840]" firstAuthor="Fauchald" pageId="72" pageNumber="73" pagination="1 - 188" refId="ref83459" refString="Fauchald, K. (1977) The polychaete worms. Definitions and keys to the orders, families and genera. Natural History Museum of Los Angeles County, Science Series, 28, 1 - 188. [www. vliz. be / imisdocs / publications / 123110. pdf]" type="journal article" year="1977">Fauchald 1977: 76</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="241110E3FFAB7364FAE1F8E2F5036645" author="Pleijel, F." page="109" pageId="72" pageNumber="73" year="1998">
<bibRefCitation id="C1214B03FFAB7364FAE1F8E2F5036645" author="Pleijel" firstAuthor="Pleijel" pageId="72" pageNumber="73" pagination="89 - 163" refId="ref88106" refString="Pleijel, F. (1998) Phylogeny and classification of Hesionidae (Polychaeta). Zoologica Scripta, 27, 89 - 163. [http: // onlinelibrary. wiley. com / doi / 10.1111 / j. 1463 - 6409.1998. tb 00433. x / pdf] https: // doi. org / 10.1111 / j. 1463 - 6409.1998. tb 00433. x" type="journal article" year="1998">Pleijel, 1998: 109110</bibRefCitation>
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, 160.
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</paragraph>
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<emphasis id="97C4EAE0FFAB7364FF23F8A0F57C6667" bold="true" box="[151,286,1876,1901]" pageId="72" pageNumber="73">
<typeStatus id="7A0B8850FFAB7364FF23F8A0F4AE6667" box="[151,204,1877,1901]" pageId="72" pageNumber="73">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="62B04D71FFAB7364FE9EF8A0F6CD6667" ID-CoL="6P98P" authority="Ehlers, 1908" authorityName="Ehlers" authorityYear="1908" box="[298,687,1876,1901]" class="Polychaeta" family="Hesionidae" genus="Leocratides" kingdom="Animalia" order="Phyllodocida" pageId="72" pageNumber="73" phylum="Annelida" rank="species" species="filamentosus">
<emphasis id="97C4EAE0FFAB7364FE9EF8A0F64A6667" box="[298,552,1877,1901]" italics="true" pageId="72" pageNumber="73">Leocratides filamentosus</emphasis>
<bibRefCitation id="C1214B03FFAB7364FD9AF8A1F6CD6667" author="Ehlers" box="[558,687,1876,1901]" firstAuthor="Ehlers" pageId="72" pageNumber="73" pagination="1 - 167" refId="ref83389" refString="Ehlers, E. (1908) Die bodensassigen Anneliden aus den Sammlungen der deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition aud dem Dampfer, Valdivia &quot; 1898 - 1899, 2 (1), 1 - 167, pls. 1 - 23. [https: // www. biodiversitylibrary. org / item / 18687 # page / 3 / mode / 1 up]" type="journal article" year="1908">Ehlers, 1908</bibRefCitation>
</taxonomicName>
by monotypy.
</paragraph>
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<subSubSection id="EDAA6579FFAB7365FF23F86EF05E6049" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73" type="diagnosis">
<paragraph id="A50F36F2FFAB7365FF23F86EF05E6049" blockId="72.[151,1436,1947,2010]" lastBlockId="73.[151,1437,153,755]" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
<emphasis id="97C4EAE0FFAB7364FF23F86EF56866BC" bold="true" box="[151,266,1947,1974]" pageId="72" pageNumber="73">Diagnosis</emphasis>
.
<taxonomicName id="62B04D71FFAB7364FEACF86EF5F466BF" authorityName="Grube" authorityYear="1850" box="[280,406,1947,1973]" class="Polychaeta" family="Hesionidae" kingdom="Animalia" order="Phyllodocida" pageId="72" pageNumber="73" phylum="Annelida" rank="subFamily" subFamily="Hesioninae">Hesioninae</taxonomicName>
with two lateral antennae, and one median antenna on dorsal prostomial surface. Palps biarticulate, palpophores large, massive, palpostyles smaller, blunt. Eyes dark, black or brown, anterior ones larger than posterior ones, sometimes approaching each other in lateral view. Nuchal organs lobes round, projected posteriorly, parallel to divergent, posterior ciliated bands close to each other. Peristomium with dorsolateral tubercles, often lobate, and ventrolateral ones projected, with a ventral ridge usually with large round papillae. Pharynx with upper jaw double, T-shaped, lower jaw single, fang-shaped; neurochaetal blades bidentate, without guards. Parapodia all sesquiramous. Neurochaetae compound falcigers, blades bidentate, without guards.
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFAA7365FF73FEB8F52263F9" pageId="73" pageNumber="74" type="discussion">
<paragraph id="A50F36F2FFAA7365FF73FEB8F64B6311" blockId="73.[151,1437,153,755]" pageId="73" pageNumber="74">
<emphasis id="97C4EAE0FFAA7365FF73FEB8F556606D" bold="true" box="[199,308,333,359]" pageId="73" pageNumber="74">Remarks</emphasis>
.
<taxonomicName id="62B04D71FFAA7365FEF1FEB8F600606D" authority="Ehlers, 1908" authorityName="Ehlers" authorityYear="1908" box="[325,610,333,359]" class="Polychaeta" family="Hesionidae" genus="Leocratides" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FEF1FEB8F5AB606D" box="[325,457,333,359]" italics="true" pageId="73" pageNumber="74">Leocratides</emphasis>
<bibRefCitation id="C1214B03FFAA7365FE66FEB8F600606D" author="Ehlers" box="[466,610,333,359]" firstAuthor="Ehlers" pageId="73" pageNumber="74" pagination="1 - 167" refId="ref83389" refString="Ehlers, E. (1908) Die bodensassigen Anneliden aus den Sammlungen der deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition aud dem Dampfer, Valdivia &quot; 1898 - 1899, 2 (1), 1 - 167, pls. 1 - 23. [https: // www. biodiversitylibrary. org / item / 18687 # page / 3 / mode / 1 up]" type="journal article" year="1908">Ehlers, 1908</bibRefCitation>
</taxonomicName>
resembles three other
<taxonomicName id="62B04D71FFAA7365FCDEFEB8F78A606D" authorityName="Grube" authorityYear="1850" box="[874,1000,333,359]" class="Polychaeta" family="Hesionidae" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="subFamily" subFamily="Hesioninae">Hesioninae</taxonomicName>
genera by having sesquiramous parapodia:
<taxonomicName id="62B04D71FFAA7365FF53FE84F6586081" authority="McIntosh, 1908" authorityName="McIntosh" authorityYear="1908" box="[231,570,369,395]" class="Polychaeta" family="Hesionidae" genus="Dalhousiella" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FF53FE84F51A6081" box="[231,376,369,395]" italics="true" pageId="73" pageNumber="74">Dalhousiella</emphasis>
<bibRefCitation id="C1214B03FFAA7365FE36FE84F6586081" author="McIntosh" box="[386,570,369,395]" firstAuthor="McIntosh" pageId="73" pageNumber="74" pagination="43 - 70" refId="ref87016" refString="McIntosh, W. C. (1908) A monograph of the British Annelids. Vol. 2. Part 1. Polychaeta: Nephthydidae to Syllidae. Ray Society, London, 233 pp., pls. 43 - 70. [https: // www. biodiversitylibrary. org / item / 116428 # page / 9 / mode / 1 up]" type="book chapter" year="1908">McIntosh, 1908</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="62B04D71FFAA7365FDFCFE84F7DE6081" authority="Salazar-Vallejo, 2016" authorityName="Salazar-Vallejo" authorityYear="2016" box="[584,956,369,395]" class="Polychaeta" family="Hesionidae" genus="Elisesione" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FDFCFE84F6D96081" box="[584,699,369,395]" italics="true" pageId="73" pageNumber="74">Elisesione</emphasis>
<bibRefCitation id="C1214B03FFAA7365FD71FE84F7DE6081" author="Salazar-Vallejo" box="[709,956,369,395]" firstAuthor="Salazar-Vallejo" pageId="73" pageNumber="74" pagination="1 - 12" refId="ref88944" refString="Salazar-Vallejo, S. I. (2016) Elisesione, a new name for Wesenbergia Hartman, 1955, and the description of a new species (Annelida, Hesionidae). ZooKeys, 632, 1 - 12. https: // doi. org / 10.3897 / zookeys. 632.9652" type="journal article" year="2016">Salazar-Vallejo, 2016</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="62B04D71FFAA7365FC49FE87F1FA6081" authority="Savigny in Lamarck, 1818" authorityName="Savigny in Lamarck" authorityYear="1818" box="[1021,1432,369,395]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FC49FE87F03B6081" box="[1021,1113,370,395]" italics="true" pageId="73" pageNumber="74">Hesione</emphasis>
Savigny
<emphasis id="97C4EAE0FFAA7365FB7DFE87F0BD6081" box="[1225,1247,370,395]" italics="true" pageId="73" pageNumber="74">in</emphasis>
<bibRefCitation id="C1214B03FFAA7365FB5DFE84F1FA6081" author="Lamarck" box="[1257,1432,369,395]" firstAuthor="Lamarck" pageId="73" pageNumber="74" refId="ref86554" refString="Lamarck, J. B. (1818) Histoire Naturelle des Animaux sans Vertebres, presentant les Caracteres Generaux et Particuliers de ces Animaux, leur Distribution, leurs Classes, leurs Familles, leurs Genres, et la Citation des principales Especes qui s'y rapportent; Precedes d'une Introduction offrant la Determination des Caracteres Essentiels de l'Animal, sa distinction du Vegetal et des autres Corps Naturels, en fin, l'Exposition des Principes fondamentaux de la Zoologie. Vol. 5. Deterville &amp; Verdiere, Paris, 612 pp. [http: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 64280058]" type="book" year="1818">Lamarck, 1818</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="62B04D71FFAA7365FF23FE60F57960A5" authorityName="Ehlers" authorityYear="1908" box="[151,283,405,431]" class="Polychaeta" family="Hesionidae" genus="Leocratides" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FF23FE60F57960A5" box="[151,283,405,431]" italics="true" pageId="73" pageNumber="74">Leocratides</emphasis>
</taxonomicName>
differs from these genera because it has biarticulate palps (missing in
<taxonomicName id="62B04D71FFAA7365FBF1FE63F0C360A5" authorityName="Savigny in Lamarck" authorityYear="1818" box="[1093,1185,406,431]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FBF1FE63F0C360A5" box="[1093,1185,406,431]" italics="true" pageId="73" pageNumber="74">Hesione</emphasis>
</taxonomicName>
, simple in
<taxonomicName id="62B04D71FFAA7365FA96FE60F1F760A5" authorityName="Salazar-Vallejo" authorityYear="2016" box="[1314,1429,405,431]" class="Polychaeta" family="Hesionidae" genus="Elisesione" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FA96FE60F1F760A5" box="[1314,1429,405,431]" italics="true" pageId="73" pageNumber="74">Elisesione</emphasis>
</taxonomicName>
, biarticulate in
<taxonomicName id="62B04D71FFAA7365FE8BFE4CF5B260D9" authorityName="McIntosh" authorityYear="1908" box="[319,464,441,467]" class="Polychaeta" family="Hesionidae" genus="Dalhousiella" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FE8BFE4CF5B260D9" box="[319,464,441,467]" italics="true" pageId="73" pageNumber="74">Dalhousiella</emphasis>
</taxonomicName>
), median antenna (missing in
<taxonomicName id="62B04D71FFAA7365FC86FE4CF7C760D9" authorityName="Salazar-Vallejo" authorityYear="2016" box="[818,933,441,467]" class="Polychaeta" family="Hesionidae" genus="Elisesione" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FC86FE4CF7C760D9" box="[818,933,441,467]" italics="true" pageId="73" pageNumber="74">Elisesione</emphasis>
</taxonomicName>
and
<taxonomicName id="62B04D71FFAA7365FC55FE4FF05F60D9" authorityName="Savigny in Lamarck" authorityYear="1818" box="[993,1085,442,467]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FC55FE4FF05F60D9" box="[993,1085,442,467]" italics="true" pageId="73" pageNumber="74">Hesione</emphasis>
</taxonomicName>
, present in
<taxonomicName id="62B04D71FFAA7365FB7DFE4CF13860D9" authorityName="McIntosh" authorityYear="1908" box="[1225,1370,441,467]" class="Polychaeta" family="Hesionidae" genus="Dalhousiella" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FFAA7365FB7DFE4CF13860D9" box="[1225,1370,441,467]" italics="true" pageId="73" pageNumber="74">Dalhousiella</emphasis>
</taxonomicName>
), and especially because its pharynx is provided with jaws (missing in the three other genera), being double upper jaws, and single, fang-shaped lower ones.
</paragraph>
<paragraph id="A50F36F2FFAA7365FF73FDD0F52263F9" blockId="73.[151,1437,153,755]" pageId="73" pageNumber="74">
Further, the nuchal organs lobes, as observed in a
<typeStatus id="7A0B8850FFAA7365FD47FDD0F7286335" box="[755,842,549,575]" pageId="73" pageNumber="74" type="syntype">syntype</typeStatus>
of
<taxonomicName id="62B04D71FFAA7365FCDBFDD3F0D36335" authority="Ehlers, 1908" authorityName="Ehlers" authorityYear="1908" box="[879,1201,549,575]" class="Polychaeta" family="Hesionidae" genus="Leocratides" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="species" species="filamentosus">
<emphasis id="97C4EAE0FFAA7365FCDBFDD3F0796335" box="[879,1051,549,575]" italics="true" pageId="73" pageNumber="74">L. filamentosus</emphasis>
<bibRefCitation id="C1214B03FFAA7365FB95FDD0F0D36335" author="Ehlers" box="[1057,1201,549,575]" firstAuthor="Ehlers" pageId="73" pageNumber="74" pagination="1 - 167" refId="ref83389" refString="Ehlers, E. (1908) Die bodensassigen Anneliden aus den Sammlungen der deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition aud dem Dampfer, Valdivia &quot; 1898 - 1899, 2 (1), 1 - 167, pls. 1 - 23. [https: // www. biodiversitylibrary. org / item / 18687 # page / 3 / mode / 1 up]" type="journal article" year="1908">Ehlers, 1908</bibRefCitation>
</taxonomicName>
, the type species, are depressed, divergent, markedly separated middorsally (
<figureCitation id="3D8B2A77FFAA7365FCB7FDBCF7EC6369" box="[771,910,585,611]" captionStart="FIGURE 44" captionStartId="77.[151,250,957,982]" captionTargetBox="[173,1416,183,931]" captionTargetId="figure@77.[170,1418,181,932]" captionTargetPageId="77" captionText="FIGURE 44. Leocratides filamentosus Ehlers, 1908, syntypes (ZMB 4431). A. Smallest syntype, anterior region, dorsal view. B. Anterior end, oblique frontal view. C. Chaetiger 10, left parapodium, anterior view (inset: neuracicular lobe). D. Largest syntype, anterior region, dorsal view. E. Anterior end, oblique frontal view. F. Chaetiger 10, right parapodium, anterior view (inset: neuracicular lobe). Scale bars: A: 1.5 mm; B: 1 mm; C: 0.4 mm; D: 0.9 mm; E: 1.2 mm; F: 0.6 mm." figureDoi="http://doi.org/10.5281/zenodo.3672640" httpUri="https://zenodo.org/record/3672640/files/figure.png" pageId="73" pageNumber="74">Fig. 44A, D</figureCitation>
). There are two other interesting differences in comparison to all other
<taxonomicName id="62B04D71FFAA7365FE16FD98F642638D" authorityName="Grube" authorityYear="1850" box="[418,544,621,647]" class="Polychaeta" family="Hesionidae" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="subFamily" subFamily="Hesioninae">Hesioninae</taxonomicName>
genera, and both might be related to the peristomium. First, there are two complex, dorsolateral tubercles resembling hands by having 34 thick digitate lobes as in
<taxonomicName id="62B04D71FFAA7365FBE2FD67F1FA63A1" authority="Ehlers, 1908" authorityName="Ehlers" authorityYear="1908" box="[1110,1432,657,683]" class="Polychaeta" family="Hesionidae" genus="Leocratides" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="species" species="filamentosus">
<emphasis id="97C4EAE0FFAA7365FBE2FD67F16363A1" box="[1110,1281,657,683]" italics="true" pageId="73" pageNumber="74">L. filamentosus</emphasis>
<bibRefCitation id="C1214B03FFAA7365FABCFD64F1FA63A1" author="Ehlers" box="[1288,1432,657,683]" firstAuthor="Ehlers" pageId="73" pageNumber="74" pagination="1 - 167" refId="ref83389" refString="Ehlers, E. (1908) Die bodensassigen Anneliden aus den Sammlungen der deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition aud dem Dampfer, Valdivia &quot; 1898 - 1899, 2 (1), 1 - 167, pls. 1 - 23. [https: // www. biodiversitylibrary. org / item / 18687 # page / 3 / mode / 1 up]" type="journal article" year="1908">Ehlers, 1908</bibRefCitation>
</taxonomicName>
, or 12 lobes as in
<taxonomicName id="62B04D71FFAA7365FED0FD43F61263C5" authority="(Horst, 1921)" baseAuthorityName="Horst" baseAuthorityYear="1921" box="[356,624,693,719]" class="Polychaeta" family="Hesionidae" genus="Leocratides" kingdom="Animalia" order="Phyllodocida" pageId="73" pageNumber="74" phylum="Annelida" rank="species" species="ehlersi">
<emphasis id="97C4EAE0FFAA7365FED0FD43F5AC63C5" box="[356,462,693,719]" italics="true" pageId="73" pageNumber="74">L. ehlersi</emphasis>
(
<bibRefCitation id="C1214B03FFAA7365FE69FD40F60563C5" author="Horst" box="[477,615,693,719]" firstAuthor="Horst" pageId="73" pageNumber="74" pagination="73 - 83" refId="ref85723" refString="Horst, R. (1921) A review of the family of Hesionidae with a description of two new species. Zoologische Mededeelingen, 6, 73 - 83. [http: // www. repository. naturalis. nl / document / 149445]" type="journal article" year="1921">Horst, 1921</bibRefCitation>
)
</taxonomicName>
. Second, midventrally there is a continuous ridge with 810 large, thick, round papillae.
</paragraph>
</subSubSection>
</treatment>
</document>