338 lines
23 KiB
XML
338 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.72.56223" ID-GBIF-Dataset="f61aa496-eaf5-4d9a-bcc3-e93e4f71bbd8" ID-PMC="PMC7481264" ID-Pensoft-Pub="1314-4049-72-43" ID-Pensoft-UUID="5243D130A7EF55F3856E1E3722F204D5" ID-PubMed="32963488" ModsDocID="1314-4049-72-43" checkinTime="1599126124911" checkinUser="pensoft" docAuthor="Pykaelae, Juha, Kantelinen, Annina & Myllys, Leena" docDate="2020" docId="FAEF26ED6270581B8803B6DAFF75F133" docLanguage="en" docName="MycoKeys 72: 43-92" docOrigin="MycoKeys 72" docSource="http://dx.doi.org/10.3897/mycokeys.72.56223" docTitle="Verrucaria vacillans Pykaelae & Myllys 2020, sp. nov." docType="treatment" docVersion="4" id="5243D130A7EF55F3856E1E3722F204D5" lastPageNumber="43" masterDocId="5243D130A7EF55F3856E1E3722F204D5" masterDocTitle="Taxonomy of Verrucaria species characterised by large spores, perithecia leaving pits in the rock and a pale thin thallus in Finland" masterLastPageNumber="92" masterPageNumber="43" pageNumber="43" updateTime="1668136534071" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomy of Verrucaria species characterised by large spores, perithecia leaving pits in the rock and a pale thin thallus in Finland</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pykaelae, Juha</mods:namePart>
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<mods:affiliation>Biodiversity Centre, Finnish Environment Institute, Latokartanonkaari 11, 00790 Helsinki, Finland</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7566-9310</mods:nameIdentifier>
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<mods:nameIdentifier type="email">juha.pykala@ymparisto.fi</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kantelinen, Annina</mods:namePart>
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<mods:affiliation>Botanical Museum, Finnish Museum of Natural History, P. O. Box 7, FI- 00014 University of Helsinki, Finland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Myllys, Leena</mods:namePart>
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<mods:affiliation>Botanical Museum, Finnish Museum of Natural History, P. O. Box 7, FI- 00014 University of Helsinki, Finland</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-9566-9473</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>72</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>43</mods:start>
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<mods:end>92</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.72.56223</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.72.56223</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-72-43</mods:identifier>
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<mods:identifier type="Pensoft-UUID">5243D130A7EF55F3856E1E3722F204D5</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="167465739" LSID="urn:lsid:plazi:treatment:FAEF26ED6270581B8803B6DAFF75F133" httpUri="http://treatment.plazi.org/id/FAEF26ED6270581B8803B6DAFF75F133" lastPageNumber="43" pageId="0" pageNumber="43">
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<subSubSection pageId="0" pageNumber="43" type="nomenclature">
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<paragraph pageId="0" pageNumber="43">
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<taxonomicName LSID="FAEF26ED-6270-581B-8803-B6DAFF75F133" authority="Pykälä & Myllys" authorityName="Pykaelae & Myllys" authorityYear="2020" class="Eurotiomycetes" family="Verrucariaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria vacillans" order="Verrucariales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="vacillans" status="sp. nov.">
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Verrucaria vacillans
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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& Myllys
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F3" captionText="Figure 2. Habitus of the new Verrucaria species A V. bifurcata (holotype) B V. cavernarum (holotype) C V. difficilis (holotype) D V. fuscozonata (holotype) E V. kuusamoensis (holotype) F V. subdevergens (holotype) G V. vacillans (holotype). Scale bars: 1 mm (A-D), 0.5 mm (E-G)." figureDoi="10.3897/mycokeys.72.56223.figure2" httpUri="https://binary.pensoft.net/fig/448799" pageId="0" pageNumber="43">Fig. 2G</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="diagnosis">
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<paragraph pageId="0" pageNumber="43">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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Species characterised by dark lines between contiguous conspecific thalli, pale usually endolithic thallus, perithecia leaving shallow to deep pits in the rock, very variable involucrellum, ascospores (18-)23-28(-32)
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<normalizedToken originalValue="×">x</normalizedToken>
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(8-)11-13(-15) mm, morphologically rather similar to the Finnish species of the
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<taxonomicName lsidName="V. subtilis" pageId="0" pageNumber="43" rank="species" species="subtilis">
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<emphasis italics="true" pageId="0" pageNumber="43">V. subtilis</emphasis>
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</taxonomicName>
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complex, but the sequence divergence in ITS 4.5-6.8%.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="holotype">
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<paragraph pageId="0" pageNumber="43">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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Finland.
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<normalizedToken originalValue="Enontekiön">Enontekioen</normalizedToken>
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Lappi,
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<normalizedToken originalValue="Enontekiö">Enontekioe</normalizedToken>
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,
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<normalizedToken originalValue="Porojärvet">Porojaervet</normalizedToken>
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, Toskalharji, Toskalpahta, fell, SW-slope, scree, on dolomite boulder, 795 m alt.,
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<geoCoordinate degrees="69" direction="north" minutes="11" orientation="latitude" precision="925" value="69.183334">69°11'N</geoCoordinate>
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,
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<geoCoordinate degrees="21" direction="east" minutes="29" orientation="longitude" precision="925" value="21.483334">21°29'E</geoCoordinate>
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, 1 Aug 2011, J.
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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43118 (H9205851, GenBank accession number: MT229829).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="description">
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<paragraph pageId="0" pageNumber="43">Description.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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Prothallus absent. Thallus white, whitish grey or pale brownish, mainly endolithic to thinly epilithic, 20-170 mm thick, algal cells 5-10 mm, contiguous conspecific thalli separated by dark lines, 0.21-0.41 mm wide. Perithecia 0.15-0.47 mm in diam., 1/4-3/4-immersed, usually leaving shallow to fairly deep pits in the rock, rarely few perithecia not leaving pits, often surrounded by a thalline collar, 60-160(-200) perithecia/cm2. Ostiole tiny or conspicuous, pale to dark, plane or depressed, ca. 20-40(-60) mm wide, wider ostiolar depression occasionally present, up to 160 mm wide. Involucrellum apical, covering half of the exciple, exceeding half of the exciple or rarely to the exciple base, 30-70(-90) mm thick, appressed to the exciple, moderately diverging from the exciple, strongly diverging from the exciple or even spreading outwards away from the exciple. Exciple 0.15-0.26 mm, wall dark brown or black, 17-35 mm thick. Periphysoids ca. 25-40(-50)
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<normalizedToken originalValue="×">x</normalizedToken>
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1.5-2.5 mm, branching. Asci 67-84
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<normalizedToken originalValue="×">x</normalizedToken>
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27-28 mm, 8-spored. Ascospores 0-septate, (18.1-)22.7-25.3-28.0(-31.7)
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<normalizedToken originalValue="×">x</normalizedToken>
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(8.3-)10.8-11.9-13.1(-15.2) mm (n = 228), perispore 1-1.5 mm thick.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="habitat">
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<paragraph pageId="0" pageNumber="43">Habitat and distribution.</paragraph>
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<paragraph pageId="0" pageNumber="43">The species is restricted in Finland to the calcareous mountains (Scandes) in NW Finland above the tree level. It always grows on dolomite. It grows on rock outcrops, boulders, stones and pebbles.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="etymology">
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<paragraph pageId="0" pageNumber="43">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="43">The specific epithet refers to the high morphological variation in the involucrellum from apical to (rarely) reaching the exciple base level, from being appressed to the exciple to spreading outwards away from the exciple and from fairly thin to thick.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="materials_examined">
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<paragraph pageId="0" pageNumber="43">Other specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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Finland.
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<normalizedToken originalValue="Enontekiön">Enontekioen</normalizedToken>
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Lappi,
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<normalizedToken originalValue="Enontekiö">Enontekioe</normalizedToken>
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,
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<normalizedToken originalValue="Porojärvet">Porojaervet</normalizedToken>
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, Toskalharji, Toskalpahta, fell, SW-slope, scree, on dolomite pebbles, 785 m alt.,
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<geoCoordinate degrees="69" direction="north" minutes="11" orientation="latitude" precision="925" value="69.183334">69°11'N</geoCoordinate>
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,
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<geoCoordinate degrees="21" direction="east" minutes="29" orientation="longitude" precision="925" value="21.483334">21°29'E</geoCoordinate>
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, 1 Aug 2011, J.
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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43058 (H);
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<normalizedToken originalValue="Enontekiö">Enontekioe</normalizedToken>
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,
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<normalizedToken originalValue="Porojärvet">Porojaervet</normalizedToken>
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, Toskalharji,
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<normalizedToken originalValue="Toskaljärvi">Toskaljaervi</normalizedToken>
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N, fell, gentle SE-slope, dolomite rock outcrop, on dolomite stones, with
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<taxonomicName lsidName="V. foveolata" pageId="0" pageNumber="43" rank="species" species="foveolata">
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<emphasis italics="true" pageId="0" pageNumber="43">V. foveolata</emphasis>
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</taxonomicName>
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, 730 m alt.,
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<geoCoordinate degrees="69" direction="north" minutes="12" orientation="latitude" precision="925" value="69.2">69°12'N</geoCoordinate>
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,
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<geoCoordinate degrees="21" direction="east" minutes="26" orientation="longitude" precision="925" value="21.433332">21°26'E</geoCoordinate>
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, 2 Aug 2011, J.
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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43232 (H);
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<normalizedToken originalValue="Enontekiö">Enontekioe</normalizedToken>
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,
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<normalizedToken originalValue="Porojärvet">Porojaervet</normalizedToken>
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, Toskalharji,
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<normalizedToken originalValue="Toskaljärvi">Toskaljaervi</normalizedToken>
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N, fell, dolomite rock, gentle S-slope, on dolomite stone, 720 m alt.,
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<geoCoordinate degrees="69" direction="north" minutes="12" orientation="latitude" precision="925" value="69.2">69°12'N</geoCoordinate>
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,
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<geoCoordinate degrees="21" direction="east" minutes="26" orientation="longitude" precision="925" value="21.433332">21°26'E</geoCoordinate>
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, 2 Aug 2011, J.
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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43272 (H);
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<normalizedToken originalValue="Enontekiö">Enontekioe</normalizedToken>
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,
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<normalizedToken originalValue="Porojärvet">Porojaervet</normalizedToken>
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, Toskalharji,
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<normalizedToken originalValue="Toskaljärvi">Toskaljaervi</normalizedToken>
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N, fell, dolomite rock, on SE-facing wall, 720 m alt.,
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<geoCoordinate degrees="69" direction="north" minutes="12" orientation="latitude" precision="925" value="69.2">69°12'N</geoCoordinate>
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,
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<geoCoordinate degrees="21" direction="east" minutes="26" orientation="longitude" precision="925" value="21.433332">21°26'E</geoCoordinate>
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, 2 Aug 2011, J.
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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43296 (H);
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<normalizedToken originalValue="Enontekiö">Enontekioe</normalizedToken>
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,
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<normalizedToken originalValue="Porojärvet">Porojaervet</normalizedToken>
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, Toskalharji,
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<normalizedToken originalValue="Toskaljärvi">Toskaljaervi</normalizedToken>
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N, fell, dolomite rock, gentle SE-slope, on dolomite pebbles, 730 m alt.,
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<geoCoordinate degrees="69" direction="north" minutes="12" orientation="latitude" precision="925" value="69.2">69°12'N</geoCoordinate>
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,
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<geoCoordinate degrees="21" direction="east" minutes="26" orientation="longitude" precision="925" value="21.433332">21°26'E</geoCoordinate>
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, 2 Aug 2011, J.
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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43302 (H);
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<normalizedToken originalValue="Enontekiö">Enontekioe</normalizedToken>
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,
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<normalizedToken originalValue="Porojärvet">Porojaervet</normalizedToken>
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, Toskalharji,
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<normalizedToken originalValue="Toskaljärvi">Toskaljaervi</normalizedToken>
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N, fell, dolomite scree, on dolomite boulder, rather abundant, 710 m alt.,
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<geoCoordinate degrees="69" direction="north" minutes="11" orientation="latitude" precision="925" value="69.183334">69°11'N</geoCoordinate>
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,
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<geoCoordinate degrees="21" direction="east" minutes="26" orientation="longitude" precision="925" value="21.433332">21°26'E</geoCoordinate>
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, 2 Aug 2011, J.
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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43384 (H);
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<normalizedToken originalValue="Enontekiö">Enontekioe</normalizedToken>
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,
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<normalizedToken originalValue="Kilpisjärvi">Kilpisjaervi</normalizedToken>
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, Saana, nature reserve, E-part, fell, dolomite rock, on SW-facing wall, 880 m alt.,
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<geoCoordinate degrees="69" direction="north" minutes="02" orientation="latitude" precision="925" value="69.03333">69°02'N</geoCoordinate>
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,
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<geoCoordinate degrees="20" direction="east" minutes="51" orientation="longitude" precision="925" value="20.85">20°51'E</geoCoordinate>
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, 10 Aug 2011, J.
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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44075, 44081b (H);
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<normalizedToken originalValue="Enontekiö">Enontekioe</normalizedToken>
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,
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<normalizedToken originalValue="Kilpisjärvi">Kilpisjaervi</normalizedToken>
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, Saana, fell, steep NE-slope, dolomite rock, on NE-facing wall, 820 m alt.,
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<geoCoordinate degrees="69" direction="north" minutes="02" orientation="latitude" precision="925" value="69.03333">69°02'N</geoCoordinate>
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,
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<geoCoordinate degrees="20" direction="east" minutes="51" orientation="longitude" precision="925" value="20.85">20°51'E</geoCoordinate>
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, 11 Aug 2011, J.
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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44142, 44162 (H);
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<normalizedToken originalValue="Enontekiö">Enontekioe</normalizedToken>
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,
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<normalizedToken originalValue="Kilpisjärvi">Kilpisjaervi</normalizedToken>
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, Saana, nature reserve, E-part, fell, steep SW-slope, dolomite rock, on SW-facing wall, 730 m alt.,
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<geoCoordinate degrees="69" direction="north" minutes="02" orientation="latitude" precision="925" value="69.03333">69°02'N</geoCoordinate>
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,
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<geoCoordinate degrees="20" direction="east" minutes="51" orientation="longitude" precision="925" value="20.85">20°51'E</geoCoordinate>
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, 12 Aug 2011, J.
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<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
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44255 (H).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="notes">
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<paragraph pageId="0" pageNumber="43">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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Based on ITS sequences,
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<taxonomicName lsidName="V. vacillans" pageId="0" pageNumber="43" rank="species" species="vacillans">
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<emphasis italics="true" pageId="0" pageNumber="43">V. vacillans</emphasis>
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</taxonomicName>
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is genetically well distinct from other
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<taxonomicName class="Arthoniomycetes" family="Arthoniaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria" order="Arthoniales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria</emphasis>
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</taxonomicName>
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species. However, it may be confused with several other species.
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<taxonomicName authorityName="Pykaelae & Myllys" authorityYear="2020" class="Arthoniomycetes" family="Arthoniaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria vacillans" order="Arthoniales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="vacillans">
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<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria vacillans</emphasis>
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</taxonomicName>
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is most difficult to separate from
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<taxonomicName lsidName="V. cavernarum" pageId="0" pageNumber="43" rank="species" species="cavernarum">
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<emphasis italics="true" pageId="0" pageNumber="43">V. cavernarum</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="V. difficilis" pageId="0" pageNumber="43" rank="species" species="difficilis">
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<emphasis italics="true" pageId="0" pageNumber="43">V. difficilis</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="V. subtilis" pageId="0" pageNumber="43" rank="species" species="subtilis">
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<emphasis italics="true" pageId="0" pageNumber="43">V. subtilis</emphasis>
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</taxonomicName>
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. In these three species, dark lines between contiguous conspecific thalli are never present.
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<taxonomicName authorityName="Pykaelae & Myllys" authorityYear="2020" class="Arthoniomycetes" family="Arthoniaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria cavernarum" order="Arthoniales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="cavernarum">
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<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria cavernarum</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="V. subtilis" pageId="0" pageNumber="43" rank="species" species="subtilis">
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<emphasis italics="true" pageId="0" pageNumber="43">V. subtilis</emphasis>
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</taxonomicName>
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have an involucrellum seldom exceeding half of the exciple (and then only in a minority of perithecia). The exciple of
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<taxonomicName lsidName="V. subtilis" pageId="0" pageNumber="43" rank="species" species="subtilis">
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<emphasis italics="true" pageId="0" pageNumber="43">V. subtilis</emphasis>
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</taxonomicName>
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is sometimes pale (although usually dark). The spores tend to be slightly broader in
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<taxonomicName lsidName="V. vacillans" pageId="0" pageNumber="43" rank="species" species="vacillans">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. vacillans</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName lsidName="V. subtilis" pageId="0" pageNumber="43" rank="species" species="subtilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. subtilis</emphasis>
|
||
</taxonomicName>
|
||
. However, specimens of
|
||
<taxonomicName lsidName="V. vacillans" pageId="0" pageNumber="43" rank="species" species="vacillans">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. vacillans</emphasis>
|
||
</taxonomicName>
|
||
without dark lines and with a short involucrellum may not be possible to separate from
|
||
<taxonomicName lsidName="V. cavernarum" pageId="0" pageNumber="43" rank="species" species="cavernarum">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. cavernarum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="V. subtilis" pageId="0" pageNumber="43" rank="species" species="subtilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. subtilis</emphasis>
|
||
</taxonomicName>
|
||
by morphology. Specimens of
|
||
<taxonomicName lsidName="V. vacillans" pageId="0" pageNumber="43" rank="species" species="vacillans">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. vacillans</emphasis>
|
||
</taxonomicName>
|
||
with a deep reaching involucrellum may not be separable from
|
||
<taxonomicName lsidName="V. difficilis" pageId="0" pageNumber="43" rank="species" species="difficilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. difficilis</emphasis>
|
||
</taxonomicName>
|
||
if dark lines are absent.
|
||
<taxonomicName authorityName="Pykaelae & Myllys" authorityYear="2020" class="Arthoniomycetes" family="Arthoniaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria vacillans" order="Arthoniales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="vacillans">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria vacillans</emphasis>
|
||
</taxonomicName>
|
||
may also be confused with
|
||
<taxonomicName lsidName="V. devergens" pageId="0" pageNumber="43" rank="species" species="devergens">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. devergens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="V. kuusamoensis" pageId="0" pageNumber="43" rank="species" species="kuusamoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. kuusamoensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="V. epilithea" pageId="0" pageNumber="43" rank="species" species="epilithea">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. epilithea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="V. muralis" pageId="0" pageNumber="43" rank="species" species="muralis">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. muralis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName authorityName="Pykaelae, Kantelinen & Myllys" authorityYear="2020" class="Arthoniomycetes" family="Arthoniaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria kuusamoensis" order="Arthoniales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="kuusamoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria kuusamoensis</emphasis>
|
||
</taxonomicName>
|
||
has an involucrellum usually exceeding half of the exciple, larger spores and dark lines are rather rare.
|
||
<taxonomicName authorityName="Nyl., Flora 55: 362" authorityYear="1872" class="Arthoniomycetes" family="Arthoniaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria devergens" order="Arthoniales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="devergens">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria devergens</emphasis>
|
||
</taxonomicName>
|
||
has larger spores and the involucrellum is usually absent or sometimes apical.
|
||
<taxonomicName class="Arthoniomycetes" family="Arthoniaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria epilithea" order="Arthoniales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="epilithea">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria epilithea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="V. muralis" pageId="0" pageNumber="43" rank="species" species="muralis">
|
||
<emphasis italics="true" pageId="0" pageNumber="43">V. muralis</emphasis>
|
||
</taxonomicName>
|
||
have perithecia not leaving pits or the pits are shallow, the spores do not exceed 26 mm in length and dark lines are absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="43" type="names considered inapplicable to the species treated above">
|
||
<paragraph pageId="0" pageNumber="43">Names considered inapplicable to the species treated above</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |