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<document id="98DE60D6487B275D987C7EDC3AB7EC53" ID-DOI="10.11646/zootaxa.3077.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5243536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629750005925" checkinUser="felipe" docAuthor="Peck, Stewart B. &amp; Cook, Joyce" docDate="2011" docId="03B375533D62A1659DC0F902FADB0B3E" docLanguage="en" docName="zt03077p118.pdf" docOrigin="Zootaxa 3077 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3077.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Pinodytes contortus Peck &amp; Cook 2011, new species" docType="treatment" docVersion="3" lastPageNumber="24" masterDocId="FF8A0D2B3D74A1729D57FF87FFB30F0A" masterDocTitle="Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077" masterLastPageNumber="118" masterPageNumber="1" pageNumber="23" updateTime="1699105988326" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="B8B8829307C3F20C412B69D3B18CBA9B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="CF19A98676F3343008E78EF15659825F">Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077</mods:title>
</mods:titleInfo>
<mods:name id="3B8C3E8F1D73FEE0B87CBC9E88B1B6F0" type="personal">
<mods:role id="8BD04A7D5ADC39C211BE6B4EBB3FBE36">
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<mods:namePart id="348E7346672EDB5C293811417FAB051D">Peck, Stewart B.</mods:namePart>
</mods:name>
<mods:name id="518CEFCE28D2043C68032B54A34B24CC" type="personal">
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<mods:namePart id="E9142DA690216FAB378262F9F0B15F47">Cook, Joyce</mods:namePart>
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<mods:typeOfResource id="31D0BE79271D48367A7DE10EFC2DD5FA">text</mods:typeOfResource>
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<mods:title id="CC1DAB56449A14D612DEDCC65A2FB629">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4E5A32B213251F089A7991C927ADE49D">
<mods:date id="677FAFAA15E97C31B717DCB98C8E77EE">2011</mods:date>
<mods:detail id="6AF19CCF1DE8A7BB62B25260BCD724AA" type="pubDate">
<mods:number id="66EAD1CB1D3CBD6BFCC198E2D459955C">2011-10-28</mods:number>
</mods:detail>
<mods:detail id="69863F56E55975B04992C211F7C5FD54" type="volume">
<mods:number id="0201B3B69243B9250C2F69E078D22E5E">3077</mods:number>
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<mods:url id="F1BC6D05014C4935B94E450346CED5B4">https://biotaxa.org/Zootaxa/article/view/zootaxa.3077.1.1</mods:url>
</mods:location>
<mods:classification id="0FCAAA79025323B9FE0B0D498C0C9C76">journal article</mods:classification>
<mods:identifier id="25620D8A737917DFF26903799E67ED21" type="DOI">10.11646/zootaxa.3077.1.1</mods:identifier>
<mods:identifier id="5878A91A69E2AA4F017064A6E4C8CB5A" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="4ADDD804D03E89C405A71939D6C27523" type="Zenodo-Dep">5243536</mods:identifier>
</mods:mods>
<treatment id="03B375533D62A1659DC0F902FADB0B3E" LSID="urn:lsid:plazi:treatment:03B375533D62A1659DC0F902FADB0B3E" httpUri="http://treatment.plazi.org/id/03B375533D62A1659DC0F902FADB0B3E" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<subSubSection id="C30097CE3D62A1649DC0F902FD620995" box="[151,721,1669,1695]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BA5C4453D62A1649DC0F902FD620995" blockId="22.[151,721,1669,1728]" box="[151,721,1669,1695]" pageId="22" pageNumber="23">
<heading id="D0ED73293D62A1649DC0F902FD620995" bold="true" box="[151,721,1669,1695]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B96E18573D62A1649DC0F902FD620995" bold="true" box="[151,721,1669,1695]" pageId="22" pageNumber="23">
<taxonomicName id="4C1ABFC63D62A1649DC0F902FD870995" ID-CoL="4HZCC" authority="Peck &amp; Cook" authorityName="Peck &amp; Cook" authorityYear="2011" box="[151,564,1669,1695]" class="Insecta" family="Leiodidae" genus="Pinodytes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="contortus" status="sp. nov.">
<emphasis id="B96E18573D62A1649DC0F902FE360995" bold="true" box="[151,389,1669,1695]" italics="true" pageId="22" pageNumber="23">Pinodytes contortus</emphasis>
Peck &amp; Cook
</taxonomicName>
,
<taxonomicNameLabel id="A25DA52C3D62A1649F16F902FD620995" box="[577,721,1669,1695]" pageId="22" pageNumber="23" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D62A1649DC0F92FFE2F09CA" box="[151,412,1704,1728]" pageId="22" pageNumber="23" type="description">
<paragraph id="8BA5C4453D62A1649DC0F92FFE2F09CA" blockId="22.[151,721,1669,1728]" box="[151,412,1704,1728]" pageId="22" pageNumber="23">
(
<figureCitation id="1321D8C03D62A1649DC8F92FFF4E09CA" box="[159,253,1704,1728]" captionStart="FIGURES 2–10" captionStartId="53.[151,264,1927,1950]" captionTargetBox="[191,1402,155,1906]" captionTargetId="figure-93@53.[185,1402,153,1906]" captionTargetPageId="53" captionText="FIGURES 2–10. Pinodytes habitus photos. 2. P.newtoni, total length 1.38 mm. 3. P. borealis, total length 2.10 mm. 4. P. punctatus, total length 1.36 mm. 5. P.colorado, total length 1.66 mm. 6. P.pusio, total length 1.54 mm. 7. P. ovatus, total length 1.34 mm. 8. P. chandleri, total length 1.54 mm. 9. P. constrictus, total length 1.48 mm. 10. P.contortus, total length 1.16 mm." figureDoi="http://doi.org/10.5281/zenodo.5243541" httpUri="https://zenodo.org/record/5243541/files/figure.png" pageId="22" pageNumber="23">Figs. 10</figureCitation>
,
<figureCitation id="1321D8C03D62A1649C5FF92FFE9509CA" box="[264,294,1704,1728]" captionStart="FIGURE 87" captionStartId="64.[151,250,1637,1660]" captionTargetBox="[345,1242,193,1616]" captionTargetId="figure-120@64.[345,1242,193,1616]" captionTargetPageId="64" captionText="FIGURE 87. Distributions of species in the Pinodytes pusio species group in the southwestern United States. P.pusio, closed circles in the Sierra Nevada and Coast Range mountains of central and southern California. P. ovatus, closed triangles in the Coast Range mountains of southwestern Oregon. P. chandleri, open triangles in the Coast Range mountains of northern California. P. constrictus, open circle in the transverse mountain ranges north of Los Angeles, California. P. contortus, closed square in the northern Sierra Nevada mountains of California. P. eldorado, open squares in the the northern Sierra Nevada mountains of California." figureDoi="http://doi.org/10.5281/zenodo.5243571" httpUri="https://zenodo.org/record/5243571/files/figure.png" pageId="22" pageNumber="23">87</figureCitation>
,
<figureCitation id="1321D8C03D62A1649C65F92FFE2709CA" box="[306,404,1704,1728]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="22" pageNumber="23">112–119</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D62A1659DC0F977FC830E16" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BA5C4453D62A1649DC0F977FA2808EA" blockId="22.[151,1436,1775,2017]" pageId="22" pageNumber="23">
<emphasis id="B96E18573D62A1649DC0F977FE730802" bold="true" box="[151,448,1775,1800]" pageId="22" pageNumber="23">
<emphasis id="B96E18573D62A1649DC0F977FEFB0802" box="[151,328,1775,1800]" inLineHeading="true" pageId="22" pageNumber="23" reason="1">Type material.</emphasis>
<typeStatus id="54A17AE73D62A1649C04F977FE730802" box="[339,448,1776,1800]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="ED0B5C803D62A1649F44F977FDD10802" box="[531,610,1776,1800]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="22" pageNumber="23" type="Museum">CSCA</collectionCode>
).
<emphasis id="B96E18573D62A1649F2FF977FC5F0802" bold="true" box="[632,1004,1775,1800]" pageId="22" pageNumber="23">
<collectingCountry id="F30D84D53D62A1649F2FF977FCEB0802" box="[632,856,1775,1800]" name="United States of America" pageId="22" pageNumber="23">UNITED STATES</collectingCountry>
.
<collectingRegion id="49DE0AA73D62A1649E3FF968FC540802" box="[872,999,1775,1800]" country="United States of America" name="California" pageId="22" pageNumber="23">California</collectingRegion>
:
</emphasis>
El Dorado Co., 5miE Placerville off Hwy. 50,
<date id="FFA4E2853D62A1649C53F893FE380826" box="[260,395,1812,1836]" pageId="22" pageNumber="23" value="1980-06-26">VI-26-1980</date>
, S.
<collectingCountry id="F30D84D53D62A1649CE3F893FE430826" box="[436,496,1812,1836]" name="Cuba" pageId="22" pageNumber="23">Kuba</collectingCountry>
and T. Eichlin, F80-19, Berlesed from
<taxonomicName id="4C1ABFC63D62A1649EFEF893FBA30821" authorityName="Say &amp; Ord" authorityYear="1825" box="[937,1040,1812,1835]" class="Mammalia" family="Cricetidae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">Neotoma</taxonomicName>
nest at base of
<taxonomicName id="4C1ABFC63D62A16499E9F893FF05085A" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="taxifolia">Pseudotsuga taxifolia</taxonomicName>
.
<emphasis id="B96E18573D62A1649D93F8BFFE8F085A" bold="true" box="[196,316,1848,1872]" pageId="22" pageNumber="23">
<typeStatus id="54A17AE73D62A1649D93F8BFFE8F085A" box="[196,316,1848,1872]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
</emphasis>
(10).
<emphasis id="B96E18573D62A1649CD0F8BFFD48085A" bold="true" box="[391,763,1847,1872]" pageId="22" pageNumber="23">
<collectingCountry id="F30D84D53D62A1649CD0F8BFFDD4085A" box="[391,615,1847,1872]" name="United States of America" pageId="22" pageNumber="23">UNITED STATES</collectingCountry>
.
<collectingRegion id="49DE0AA73D62A1649F20F8B0FD45085A" box="[631,758,1847,1872]" country="United States of America" name="California" pageId="22" pageNumber="23">California</collectingRegion>
:
</emphasis>
with same data as
<typeStatus id="54A17AE73D62A1649EB6F8BFFBF5085A" box="[993,1094,1848,1872]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
, 1 (
<collectionCode id="ED0B5C803D62A1649922F8BFFB77085A" box="[1141,1220,1848,1872]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="22" pageNumber="23" type="Museum">CSCA</collectionCode>
); Butte Co., 2miN Hurleton, Stringtown Road,
<date id="FFA4E2853D62A1649CB6F8DBFDD1087E" box="[481,610,1884,1908]" pageId="22" pageNumber="23" value="1983-03-22">22.III.1983</date>
, D.A. Chandler, sift maple &amp; grape litter, 1 (
<collectionCode id="ED0B5C803D62A164992AF8DBFB60087E" box="[1149,1235,1884,1908]" pageId="22" pageNumber="23">UNHC</collectionCode>
); El Dorado Co., Blodgett Forest,
<date id="FFA4E2853D62A1649C00F807FE5F0892" box="[343,492,1920,1944]" pageId="22" pageNumber="23" value="1975-08-27">27.VIII.1975</date>
, F.G. Andrews, M.S. Wasbauer, Ber.
<taxonomicName id="4C1ABFC63D62A1649ECDF806FBEB0892" box="[922,1112,1920,1944]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="ponderosa">
<emphasis id="B96E18573D62A1649ECDF806FBEB0892" box="[922,1112,1920,1944]" italics="true" pageId="22" pageNumber="23">Pinus ponderosa</emphasis>
</taxonomicName>
, 1 (
<collectionCode id="ED0B5C803D62A16499DFF807FB640892" box="[1160,1239,1920,1944]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="22" pageNumber="23" type="Museum">CSCA</collectionCode>
); El Dorado Co., Blodgett Forest,
<date id="FFA4E2853D62A1649C0AF823FE4108B6" box="[349,498,1956,1980]" pageId="22" pageNumber="23" value="1975-08-27">27.VIII.1975</date>
, F.G. Andrews, M.S. Wasbauer, Ber.
<taxonomicName id="4C1ABFC63D62A1649EE6F823FB1908B6" box="[945,1194,1956,1980]" class="Pinopsida" family="Cupressaceae" genus="Libocedrus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="decurrens">
<emphasis id="B96E18573D62A1649EE6F823FB1908B6" box="[945,1194,1956,1980]" italics="true" pageId="22" pageNumber="23">Libocedrus decurrens</emphasis>
</taxonomicName>
duff, 2 (
<collectionCode id="ED0B5C803D62A164984CF823FAD908B6" box="[1307,1386,1956,1980]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="22" pageNumber="23" type="Museum">CSCA</collectionCode>
); El Dorado Co., Blodgett Forest, 13miE Georgetown,
<date id="FFA4E2853D62A1649F83F84FFCE008EA" box="[724,851,1992,2016]" pageId="22" pageNumber="23" value="1976-04-28">28.IV.1976</date>
, J. Doyen, Ber. litter
<taxonomicName id="4C1ABFC63D62A1649911F84FFABD08EA" box="[1094,1294,1992,2016]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="kelloggii">
<emphasis id="B96E18573D62A1649911F84FFABD08EA" box="[1094,1294,1992,2016]" italics="true" pageId="22" pageNumber="23">Quercus kelloggii</emphasis>
</taxonomicName>
, 1 (
<collectionCode id="ED0B5C803D62A164986CF84FFA3C08EA" box="[1339,1423,1992,2016]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="22" pageNumber="23" type="University or college">EMEC</collectionCode>
);
</paragraph>
<paragraph id="8BA5C4453D63A1659DC0FF10FC400FFD" blockId="23.[151,1436,151,1076]" pageId="23" pageNumber="24">
<materialsCitation id="3B72CE183D63A1659DC0FF10FE8B0FDE" collectingDate="1992-04-16" collectionCode="CSCA" collectorName="F. G. Andrews &amp; Ber" county="El Dorado Co." location="Peavine Ridge Rd." municipality="Ice House" pageId="23" pageNumber="24" specimenCount="1">
<collectingCounty id="62C4BCC93D63A1659DC0FF10FE8E0FA5" box="[151,317,151,175]" pageId="23" pageNumber="24">El Dorado Co.</collectingCounty>
,
<location id="8EC5929E3D63A1659C1DFF10FDAC0FA5" LSID="urn:lsid:plazi:treatment:03B375533D62A1659DC0F902FADB0B3E:8EC5929E3D63A1659C1DFF10FDAC0FA5" box="[330,543,151,175]" county="El Dorado Co." municipality="Ice House" name="Peavine Ridge Rd." pageId="23" pageNumber="24">Peavine Ridge Rd.</location>
, 7miSW
<collectingMunicipality id="6BC15E3F3D63A1659FDFFF1FFD4E0FA5" box="[648,765,152,175]" pageId="23" pageNumber="24">Ice House</collectingMunicipality>
,
<date id="FFA4E2853D63A1659E5EFF10FC340FA5" box="[777,903,151,175]" pageId="23" pageNumber="24" value="1992-04-16">
<collectingDate id="EFE01B6D3D63A1659E5EFF10FC340FA5" box="[777,903,151,175]" pageId="23" pageNumber="24" value="1992-04-16">16.IV.1992</collectingDate>
</date>
,
<collectorName id="26EFA1933D63A1659EC4FF10FB9C0FA5" box="[915,1071,151,175]" pageId="23" pageNumber="24">F.G. Andrews</collectorName>
,
<collectorName id="26EFA1933D63A165996CFF1FFBD70FA5" box="[1083,1124,152,175]" pageId="23" pageNumber="24">Ber</collectorName>
duff under
<taxonomicName id="4C1ABFC63D63A16599BEFF1FFF070FDE" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="kelloggii">
<emphasis id="B96E18573D63A16599BEFF1FFF070FDE" italics="true" pageId="23" pageNumber="24">Quercus kelloggii</emphasis>
</taxonomicName>
, 1 (
<collectionCode id="ED0B5C803D63A1659DB6FF3BFE830FDE" box="[225,304,188,212]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="23" pageNumber="24" type="Museum">CSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72CE183D63A1659C13FF3BFB210FDE" box="[324,1170,188,212]" collectingDate="1971-10-14" collectionCode="CSCA" collectorName="E. A. Kane &amp; Ber." county="Placer Co." location="Colfax" pageId="23" pageNumber="24" specimenCount="1">
<collectingCounty id="62C4BCC93D63A1659C13FF3BFE0F0FDE" box="[324,444,188,212]" pageId="23" pageNumber="24">Placer Co.</collectingCounty>
, 2miW
<location id="8EC5929E3D63A1659F41FF3BFDD60FDE" LSID="urn:lsid:plazi:treatment:03B375533D62A1659DC0F902FADB0B3E:8EC5929E3D63A1659F41FF3BFDD60FDE" box="[534,613,188,212]" county="Placer Co." name="Colfax" pageId="23" pageNumber="24">Colfax</location>
,
<date id="FFA4E2853D63A1659F26FF3BFD5A0FDE" box="[625,745,188,212]" pageId="23" pageNumber="24" value="1971-10-14">
<collectingDate id="EFE01B6D3D63A1659F26FF3BFD5A0FDE" box="[625,745,188,212]" pageId="23" pageNumber="24" value="1971-10-14">14.X.1971</collectingDate>
</date>
,
<collectorName id="26EFA1933D63A1659FA1FF3BFCDC0FD9" box="[758,879,188,212]" pageId="23" pageNumber="24">E.A. Kane</collectorName>
,
<collectorName id="26EFA1933D63A1659E2CFF3BFC1A0FD9" box="[891,937,188,211]" pageId="23" pageNumber="24">Ber.</collectorName>
oak duff, 2 (
<collectionCode id="ED0B5C803D63A165996CFF3BFB390FDE" box="[1083,1162,188,212]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="23" pageNumber="24" type="Museum">CSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72CE183D63A16599C9FF3BFC5C0FFD" collectingDate="1971-01-26" collectionCode="CSCA" collectorName="F. G. Andrews &amp; Ber." county="Placer Co." location="Colfax" municipality="Rollins Lake" pageId="23" pageNumber="24" specimenCount="1">
<collectingCounty id="62C4BCC93D63A16599C9FF3BFAA50FDE" box="[1182,1302,188,212]" pageId="23" pageNumber="24">Placer Co.</collectingCounty>
, 3miN
<location id="8EC5929E3D63A165983DFF3BFF0D0FFD" LSID="urn:lsid:plazi:treatment:03B375533D62A1659DC0F902FADB0B3E:8EC5929E3D63A165983DFF3BFF0D0FFD" county="Placer Co." municipality="Rollins Lake" name="Colfax" pageId="23" pageNumber="24">Colfax</location>
,
<collectingMunicipality id="6BC15E3F3D63A1659D9EFF58FEE80FFD" box="[201,347,223,247]" pageId="23" pageNumber="24">Rollins Lake</collectingMunicipality>
,
<date id="FFA4E2853D63A1659C31FF58FE600FFD" box="[358,467,223,247]" pageId="23" pageNumber="24" value="1971-01-26">
<collectingDate id="EFE01B6D3D63A1659C31FF58FE600FFD" box="[358,467,223,247]" pageId="23" pageNumber="24" value="1971-01-26">26.I.1971</collectingDate>
</date>
,
<collectorName id="26EFA1933D63A1659C88FF58FDC80FFD" box="[479,635,223,247]" pageId="23" pageNumber="24">F.G. Andrews</collectorName>
,
<collectorName id="26EFA1933D63A1659FD1FF67FD070FFD" box="[646,692,224,247]" pageId="23" pageNumber="24">Ber.</collectorName>
pine &amp; oak duff, 1 (
<collectionCode id="ED0B5C803D63A1659EC0FF58FC550FFD" box="[919,998,223,247]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="23" pageNumber="24" type="Museum">CSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA5C4453D63A1659D90FE84FC830E16" blockId="23.[151,1436,151,1076]" box="[199,816,259,284]" pageId="23" pageNumber="24">
<emphasis id="B96E18573D63A1659D90FE84FE030E11" box="[199,432,259,284]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">
<emphasis id="B96E18573D63A1659D90FE84FE1A0E16" bold="true" box="[199,425,259,284]" pageId="23" pageNumber="24">Material examined</emphasis>
.
</emphasis>
We have examined
<specimenCount id="9D1C0FCC3D63A1659FC5FE83FC9F0E16" box="[658,812,260,284]" pageId="23" pageNumber="24" type="generic">11 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D63A1659D90FEA0FD2E0B05" pageId="23" pageNumber="24" type="distribution">
<paragraph id="8BA5C4453D63A1659D90FEA0FD110E6E" blockId="23.[151,1436,151,1076]" pageId="23" pageNumber="24">
<emphasis id="B96E18573D63A1659D90FEA0FED30E4A" bold="true" box="[199,352,295,320]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">Distribution.</emphasis>
Specimens (
<figureCitation id="1321D8C03D63A1659CA7FEA0FDF10E35" box="[496,578,295,319]" captionStart="FIGURE 87" captionStartId="64.[151,250,1637,1660]" captionTargetBox="[345,1242,193,1616]" captionTargetId="figure-120@64.[345,1242,193,1616]" captionTargetPageId="64" captionText="FIGURE 87. Distributions of species in the Pinodytes pusio species group in the southwestern United States. P.pusio, closed circles in the Sierra Nevada and Coast Range mountains of central and southern California. P. ovatus, closed triangles in the Coast Range mountains of southwestern Oregon. P. chandleri, open triangles in the Coast Range mountains of northern California. P. constrictus, open circle in the transverse mountain ranges north of Los Angeles, California. P. contortus, closed square in the northern Sierra Nevada mountains of California. P. eldorado, open squares in the the northern Sierra Nevada mountains of California." figureDoi="http://doi.org/10.5281/zenodo.5243571" httpUri="https://zenodo.org/record/5243571/files/figure.png" pageId="23" pageNumber="24">Fig. 87</figureCitation>
) are known from Butte, El Dorado, and Placer counties, on the western side of the Sierra
<collectingRegion id="49DE0AA73D63A1659C7FFECBFECE0E6E" box="[296,381,332,356]" country="United States of America" name="Nevada" pageId="23" pageNumber="24">Nevada</collectingRegion>
Mountains, of
<collectingRegion id="49DE0AA73D63A1659F7EFECBFD2D0E6E" box="[553,670,332,356]" country="United States of America" name="California" pageId="23" pageNumber="24">California</collectingRegion>
.
</paragraph>
<paragraph id="8BA5C4453D63A1659D90FEE8FD2E0B05" blockId="23.[151,1436,151,1076]" pageId="23" pageNumber="24">
<emphasis id="B96E18573D63A1659D90FEE8FE680E8D" box="[199,475,367,392]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">
<emphasis id="B96E18573D63A1659D90FEE8FE670E82" bold="true" box="[199,468,367,392]" pageId="23" pageNumber="24">Diagnostic description</emphasis>
.
</emphasis>
Total length
<quantity id="4CE269A03D63A1659F20FEE8FC930E8D" box="[631,800,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.34" metricValueMin="1.16" pageId="23" pageNumber="24" unit="mm" value="1.25" valueMax="1.34" valueMin="1.16">1.16–1.34 mm</quantity>
; greatest width
<quantity id="4CE269A03D63A1659E8FFEE8FB330E8D" box="[984,1152,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="23" pageNumber="24" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60–0.70 mm</quantity>
; Reddish brown; oval in shape (
<figureCitation id="1321D8C03D63A1659DB0FE13FE890EA6" box="[231,314,404,428]" captionStart="FIGURES 2–10" captionStartId="53.[151,264,1927,1950]" captionTargetBox="[191,1402,155,1906]" captionTargetId="figure-93@53.[185,1402,153,1906]" captionTargetPageId="53" captionText="FIGURES 2–10. Pinodytes habitus photos. 2. P.newtoni, total length 1.38 mm. 3. P. borealis, total length 2.10 mm. 4. P. punctatus, total length 1.36 mm. 5. P.colorado, total length 1.66 mm. 6. P.pusio, total length 1.54 mm. 7. P. ovatus, total length 1.34 mm. 8. P. chandleri, total length 1.54 mm. 9. P. constrictus, total length 1.48 mm. 10. P.contortus, total length 1.16 mm." figureDoi="http://doi.org/10.5281/zenodo.5243541" httpUri="https://zenodo.org/record/5243541/files/figure.png" pageId="23" pageNumber="24">Fig. 10</figureCitation>
).
<emphasis id="B96E18573D63A1659C07FE13FE230EA6" bold="true" box="[336,400,404,428]" pageId="23" pageNumber="24">Head</emphasis>
. Finely, sparsely punctate; shining; vertex with faint reticulate microsculpture. Eyes absent. Antenna (
<figureCitation id="1321D8C03D63A1659C52FE30FED00EC5" box="[261,355,439,463]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Fig. 112</figureCitation>
) with antennomere 2 longer and wider than 3; antennomere 5 wider than 4, longer than 6; antennomere 7 longer but not wider than 8; antennomeres 9 and 10 each with a single sensory vesicle indicated apically by a protruding flange.
<emphasis id="B96E18573D63A1659CC9FD87FDA50D12" bold="true" box="[414,534,512,536]" pageId="23" pageNumber="24">Pronotum</emphasis>
. Finely punctate, punctures separated by 3–4 diameters; shining, microsculpture faintly indicated. Widest near base, weakly narrowing to apex; apical margin weakly emarginate, basal margin nearly straight; apical angles obtuse, basal angles about rectangular.
<emphasis id="B96E18573D63A1659EC8FDCFFC580D6A" bold="true" box="[927,1003,584,608]" pageId="23" pageNumber="24">Elytra</emphasis>
. Finely punctate, in longitudinal rows only basally near suture; punctures joined by fine transverse strioles. Joined elytra slightly wider than pronotum; widest in basal one-third, roundly narrowing to apex.
<emphasis id="B96E18573D63A1659E50FD17FC8C0DA2" bold="true" box="[775,831,656,680]" pageId="23" pageNumber="24">Legs</emphasis>
. Protibia (
<figureCitation id="1321D8C03D63A1659EEDFD08FBA80DAD" box="[954,1051,655,679]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Fig. 113</figureCitation>
) slender, width increases slightly from base to apex; spinose on outer margin near apex and apically; finely spinose on apical one-half of inner margin. Mesotibia (
<figureCitation id="1321D8C03D63A1659C1CFD50FE190DE5" box="[331,426,727,751]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Fig. 114</figureCitation>
) moderately slender; strongly spinose on outer margin and apically. Metatibia (
<figureCitation id="1321D8C03D63A1659863FD50FA270DE5" box="[1332,1428,727,751]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Fig. 115</figureCitation>
) slender, weakly curved; spinose apically and near apex; scattered small spines on outer margin. Metafemur (
<figureCitation id="1321D8C03D63A1659839FD7BFF720C3D" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Fig. 115</figureCitation>
) slender. Male protarsomeres (
<figureCitation id="1321D8C03D63A1659F4CFC98FDCA0C3D" box="[539,633,799,823]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Fig. 113</figureCitation>
) weakly expanded, bearing elongate setae laterally and broad, thin, colorless phanerae ventrally. Mesotarsomeres without phanerae.
<emphasis id="B96E18573D63A1659E05FCC3FC130C56" bold="true" box="[850,928,836,860]" pageId="23" pageNumber="24">Venter</emphasis>
. Mesoventrite (
<figureCitation id="1321D8C03D63A1659903FCC3FB070C56" box="[1108,1204,836,860]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Fig. 119</figureCitation>
) carinate; longitudinal carina with toothlike expansion near middle; not excavated behind transverse carina.
<emphasis id="B96E18573D63A16599DCFCEFFA840C8A" bold="true" box="[1163,1335,871,896]" pageId="23" pageNumber="24">Male genitalia</emphasis>
. Median lobe of aedeagus (
<figureCitation id="1321D8C03D63A1659C3DFC0BFDBC0CAE" box="[362,527,908,932]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Figs. 116, 117</figureCitation>
) broad, with narrow, flattened apex; inverted internal sac (
<figureCitation id="1321D8C03D63A16599E1FC0BFAA40CAE" box="[1206,1303,908,932]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Fig. 117</figureCitation>
) with three clusters of elongate spines and clusters of short spines basally. Parameres (
<figureCitation id="1321D8C03D63A1659954FC28FB190CCD" box="[1027,1194,943,967]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Figs. 116, 117</figureCitation>
) elongate, extending beyond apex of median lobe; apices deflexed, flattened and contorted; each bearing two setae before apex.
<emphasis id="B96E18573D63A165980BFC54FF4F0B1A" bold="true" pageId="23" pageNumber="24">Spermatheca</emphasis>
. Elongate (
<figureCitation id="1321D8C03D63A1659C2AFC70FE6E0B05" box="[381,477,1015,1039]" captionStart="FIGURES 112–119" captionStartId="68.[151,265,1372,1395]" captionTargetBox="[187,1398,197,1345]" captionTargetId="figure-66@68.[180,1406,193,1351]" captionTargetPageId="68" captionText="FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119)." figureDoi="http://doi.org/10.5281/zenodo.5243587" httpUri="https://zenodo.org/record/5243587/files/figure.png" pageId="23" pageNumber="24">Fig. 118</figureCitation>
), tubular, ribbed.
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D63A1659D90FB9BFADB0B3E" box="[199,1384,1052,1076]" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BA5C4453D63A1659D90FB9BFADB0B3E" blockId="23.[151,1436,151,1076]" box="[199,1384,1052,1076]" pageId="23" pageNumber="24">
<emphasis id="B96E18573D63A1659D90FB9BFEFE0B3E" bold="true" box="[199,333,1052,1076]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">Etymology.</emphasis>
The name
<taxonomicName id="4C1ABFC63D63A1659C9CFB9AFD330B3E" authority=", Latin" authorityName="Latin" box="[459,640,1052,1076]" class="Insecta" family="Leiodidae" genus="Pinodytes" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="contortus">
<emphasis id="B96E18573D63A1659C9CFB9AFD860B3E" box="[459,565,1053,1076]" italics="true" pageId="23" pageNumber="24">contortus</emphasis>
, Latin
</taxonomicName>
, twisted, refers to the shape of the paramere apices of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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