293 lines
52 KiB
XML
293 lines
52 KiB
XML
<document id="CD1BA8FE53D62D9EA393813BA45B64B3" ID-ISSN="11755334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629706483752" checkinUser="felipe" docAuthor="Martynov, Alexander" docDate="2010" docId="BC5D5914FFD05223FF48F9DE85DCFE62" docLanguage="en" docName="zt02697p154.pdf" docOrigin="Zootaxa 2697" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Ophiolycus Mortensen 1933" docType="treatment" docVersion="2" lastPageNumber="106" masterDocId="4064216CFFB6524AFFDFFFEC840FFFBE" masterDocTitle="Reassessment of the classification of the Ophiuroidea (Echinodermata), based on morphological characters. I. General character evaluation and delineation of the families Ophiomyxidae and Ophiacanthidae 2697" masterLastPageNumber="154" masterPageNumber="1" pageNumber="103" updateTime="1699098856182" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="527DF02D37FF3769DE8C563AE9E8508F">Reassessment of the classification of the Ophiuroidea (Echinodermata), based on morphological characters. I. General character evaluation and delineation of the families Ophiomyxidae and Ophiacanthidae 2697</mods:title>
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<mods:namePart id="99150923125259E3607AFEC81234598B">Martynov, Alexander</mods:namePart>
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<mods:title id="BF3AF7CA17ACE3B5B98D8872067D0915">Zootaxa</mods:title>
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<mods:date id="AB68C736436CF743059DD3231BFC2F7F">2010</mods:date>
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<mods:number id="4E312EC3EC0C5C1ED76A154466683018">2697</mods:number>
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<mods:identifier id="83C433F57A67EDFE23D0760FECE43B35" type="ISSN">11755334</mods:identifier>
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<subSubSection id="7CEEBB89FFD0522CFF48F9DE8640F9F2" box="[151,591,1586,1612]" pageId="102" pageNumber="103" type="nomenclature">
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<paragraph id="344BE802FFD0522CFF48F9DE8640F9F2" blockId="102.[151,591,1586,1647]" box="[151,591,1586,1612]" pageId="102" pageNumber="103">
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<heading id="6F035F6EFFD0522CFF48F9DE8640F9F2" bold="true" box="[151,591,1586,1612]" fontSize="11" level="1" pageId="102" pageNumber="103" reason="1">
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<emphasis id="06803410FFD0522CFF48F9DE8640F9F2" bold="true" box="[151,591,1586,1612]" pageId="102" pageNumber="103">
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Genus
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<taxonomicName id="F3F49381FFD0522CFF31F9DE8640F9F2" ID-CoL="7PQY8" authority="Mortensen, 1933" authorityName="Mortensen" authorityYear="1933" box="[238,591,1586,1612]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="102" pageNumber="103" phylum="Echinodermata" rank="genus">
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<emphasis id="06803410FFD0522CFF31F9DE8578F9F2" bold="true" box="[238,375,1586,1612]" italics="true" pageId="102" pageNumber="103">Ophiolycus</emphasis>
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Mortensen, 1933
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7CEEBB89FFD0522CFF48F9B985AAF9D1" box="[151,421,1621,1647]" pageId="102" pageNumber="103" type="description">
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<paragraph id="344BE802FFD0522CFF48F9B985AAF9D1" blockId="102.[151,591,1586,1647]" box="[151,421,1621,1647]" pageId="102" pageNumber="103">
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<figureCitation id="ACCFF487FFD0522CFF48F9B98540F9D1" box="[151,335,1621,1647]" captionStart="FIGURE 27" captionStartId="38.[151,256,1544,1568]" captionTargetBox="[158,1428,194,1521]" captionTargetId="figure-295@38.[158,1428,194,1523]" captionTargetPageId="38" captionText="FIGURE 27. Arm spine articulations and spines of the various taxa of the family Ophiomyxidae. A–C, Ophioscolex glacialis Müller & Troschel, 1842, ZMMU D-817, 27 mm dd; spine articulations of the proximal segments; D. spine, lateral view, E. spine, ventral view showing large nerve opening; F. Ophiomyxa brevirima H.L. Clark, 1915, ZMMU D- 725, 19 mm dd, distal segments; G. Ophiomyxa brevirima, proximal segments; H. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, proximal segments; I. Ophiocymbium cavernosum, ZMMU D-795, 10.5 mm, proximal segment; J. Ophiocymbium tanyae sp. nov., holotype, ZMMU D-799, 9.5 mm dd, proximal segments; K. same, middle segments; L. Ophiologimus cf. secundus Koehler, 1914, ZMMU D-815, 4 mm dd, proximal segments; M. same a proximal segment with spines; N. same, details of the articulations of the proximal segments; O. Ophiobyrsa rudis Lyman, 1878, ZMMU D-868, 12.5 mm dd; P. Ophiologimus cf. secundus, most proximal segments; Q. Ophiolycus purpureus (Düben & Koren, 1846), ZMMU D-845, 5.5 mm dd, most proximal segments; R. same, proximal segments; S. Ophiolycus nutrix Mortensen, 1936, ZMMU D-818, 8 mm dd, proximal segments with spines; T. same, other proximal segments; U. same, spine base, ventral view; Scale bars: A-B, 300 µ; C, 200 µ; D, 400 µ; E, 150 µ; F, 100 µ; G, 150 µ; H, 100 µ; I, 100 µ; J, 100 µ; K, 200 µ; L, 100 µ; M, 100 µ; N, 100 µ; O, 250 µ; P, 100 µ; Q, 100 µ; R, 100 µ; S–T, 120 µ; U, 150 µ. Photos: Alexander Martynov." pageId="102" pageNumber="103">Figures 27Q–U</figureCitation>
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;
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<figureCitation id="ACCFF487FFD0522CFE83F9B985AAF9D1" box="[348,421,1621,1647]" captionStart-0="FIGURE 71" captionStart-1="FIGURE 72" captionStart-2="FIGURE 73" captionStart-3="FIGURE 74" captionStartId-0="103.[151,256,1758,1782]" captionStartId-1="104.[151,255,1858,1882]" captionStartId-2="105.[151,255,1724,1748]" captionStartId-3="106.[151,255,1650,1674]" captionTargetBox-0="[158,1428,866,1736]" captionTargetBox-1="[158,1428,538,1828]" captionTargetBox-2="[158,1428,530,1703]" captionTargetBox-3="[159,1427,194,1628]" captionTargetId-0="figure-414@103.[158,1428,866,1736]" captionTargetId-1="figure-249@104.[158,1428,538,1828]" captionTargetId-2="figure-260@105.[158,1428,530,1703]" captionTargetId-3="figure-143@106.[159,1427,194,1628]" captionTargetPageId-0="103" captionTargetPageId-1="104" captionTargetPageId-2="105" captionTargetPageId-3="106" captionText-0="FIGURE 71. Ophiolycus dentatus (Lyman, 1878), ZMUC, 19.5 mm dd, external views and details (A–H). A. dorsal view; B. disk details showing small radial shields concealed under the skin; C. ventral view; D. ventral view showing genital slits; E. proximal arm segments in situ; F. same, details showing fragmented dorsal arm plates concealed under the skin; G. distal arm segments in situ showing entire dorsal arm plates; H. distal segments, hooks; lateral plate and spine articulations, proximal segments; proximal vertebra, proximal view; I. Ophioscolex glacialis, distal segments, hook-shaped spines. Scale bars: E, 1 mm; F, 300 µ; G, H–I, 100 µ; Photos: A–C Tatiana Korshunova; E–I Alexander Martynov." captionText-1="FIGURE 72. Ophiolycus dentatus (Lyman, 1878), ZMUC, details. A–B, arm spine articulations, proximal segments; C. same, dorsalmost articulation details; D. same, ventralmost articulation details; E. arm spine articulation, distal segments; F. dorslamost spine; G. spine; H. proximal vertebra, dorsal view; I. same, ventral view; J. same, proximal view; K. same, distal view; L. spine, ventral view; M. solid proximal spine, sectioned transversally showing only small cavities. Scale bars: A, F, G–K, 300 µ; B–E, L, M, 100 µ; Photos: Alexander Martynov" captionText-2="FIGURE 73. Ophiolycus purpureus (Düben & Koren, 1846), Mareano program, 8 mm dd, external views and details. A; dorsal view; B. ventral view; C. dorsal view showing distal arm segments with hooks; D. dorsal disk view, details showing small radial shields; E. interradial view showing long genital slits; F. arm spine articulations, proximal segments; G. arm spine articulation bear hooks, distal segments; H. proximalmost segments showing large distinct dorsalmost articulations; I. arm spine articulations, proximal segments; J. distal segments, hooks; K. spine; L. same, ventral view; M. spine section showing only small cavities. Scale bars: F-H, J-M, 100 µ; I, 300 µ. Photos: A–E Tatiana Korshunova; F–M Alexander Martynov." captionText-3="FIGURE 74. Ophiolycus purpureus (Düben & Koren, 1846), Mareano program, 8 mm dd, details. A. proximal vertebra, dorsal view; B. same, ventral view; C. same, proximal view; D. same, distal view; E. radial shield, ventral view; F. dental plate; G. adradial genital plate, articulation surface; H. jaw, adradial view; I. adradial and abradial genital plates in situ. Scale bars: A–B, E, H, I, 300 µ; C, D, F–G, 100 µ. Photos: Alexander Martynov." pageId="102" pageNumber="103">71–74</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="7CEEBB89FFD0522CFF48F94C8757F909" box="[151,856,1696,1719]" pageId="102" pageNumber="103" type="reference_group">
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<paragraph id="344BE802FFD0522CFF48F94C8757F909" blockId="102.[151,938,1696,1751]" box="[151,856,1696,1719]" pageId="102" pageNumber="103">
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<treatmentCitationGroup id="14E4CF2CFFD0522CFF48F94C8757F909" box="[151,856,1696,1719]" pageId="102" pageNumber="103">
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<taxonomicName id="F3F49381FFD0522CFF48F94C851FF909" ID-CoL="7PQY8" authorityName="Mortensen" authorityYear="1933" box="[151,272,1696,1719]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="102" pageNumber="103" phylum="Echinodermata" rank="genus">
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<emphasis id="06803410FFD0522CFF48F94C851FF909" box="[151,272,1696,1719]" italics="true" pageId="102" pageNumber="103">Ophiolycus</emphasis>
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</taxonomicName>
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(as subgenus of
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<taxonomicName id="F3F49381FFD0522CFE1CF94C8647F909" ID-CoL="7PNFH" authorityName=": Muller & Troschel" authorityYear="1842" box="[451,584,1696,1719]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioscolex" kingdom="Animalia" order="Ophioscolecida" pageId="102" pageNumber="103" phylum="Echinodermata" rank="genus">
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<emphasis id="06803410FFD0522CFE1CF94C8647F909" box="[451,584,1696,1719]" italics="true" pageId="102" pageNumber="103">Ophioscolex</emphasis>
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</taxonomicName>
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):
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<treatmentCitation id="B555CE13FFD0522CFD80F94C8757F909" author="Mortensen, T." box="[607,856,1696,1719]" page="315" pageId="102" pageNumber="103" year="1933">
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<bibRefCitation id="506595F3FFD0522CFD80F94C8757F909" author="Mortensen, T." box="[607,856,1696,1719]" pageId="102" pageNumber="103" pagination="215 - 400" refId="ref90767" refString="Mortensen, T. (1933 b). Echinoderms of South Africa. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 93, 215 - 400." type="journal article" year="1933">Mortensen, 1933b: 315</bibRefCitation>
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</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="7CEEBB89FFD0522CFF48F92C87A5F969" box="[151,938,1728,1751]" pageId="102" pageNumber="103" type="materials_examined">
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||
<paragraph id="344BE802FFD0522CFF48F92C87A5F969" blockId="102.[151,938,1696,1751]" box="[151,938,1728,1751]" pageId="102" pageNumber="103">
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<typeStatus id="EB4F56A0FFD0522CFF48F92C84C3F969" box="[151,204,1728,1751]" pageId="102" pageNumber="103">Type</typeStatus>
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species:
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<taxonomicName id="F3F49381FFD0522CFEF2F92C86A5F969" authority="Lyman, 1878" authorityName="Lyman" authorityYear="1878" box="[301,682,1728,1751]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioscolex" kingdom="Animalia" order="Ophioscolecida" pageId="102" pageNumber="103" phylum="Echinodermata" rank="species" species="dentatus">
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<emphasis id="06803410FFD0522CFEF2F92C861CF969" box="[301,531,1728,1751]" italics="true" pageId="102" pageNumber="103">Ophioscolex dentatus</emphasis>
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<bibRefCitation id="506595F3FFD0522CFDC5F92C86A5F969" author="Lyman, T." box="[538,682,1728,1751]" pageId="102" pageNumber="103" pagination="65 - 168" refId="ref90096" refString="Lyman, T. (1878) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger, under Prof. Sir Wyville Thomson, F. R. S. Part 1. Bulletin of the Museum of Comparative Zoology, Harvard 5, 65 - 168." type="journal article" year="1878">Lyman, 1878</bibRefCitation>
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</taxonomicName>
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, by original designation
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</paragraph>
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</subSubSection>
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||
<subSubSection id="7CEEBB89FFD0522DFF48F8E98741FD02" lastPageId="103" lastPageNumber="104" pageId="102" pageNumber="103" type="diagnosis">
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<paragraph id="344BE802FFD0522DFF48F8E98741FD02" blockId="102.[151,1436,1797,2010]" lastBlockId="103.[151,1436,152,812]" lastPageId="103" lastPageNumber="104" pageId="102" pageNumber="103">
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<emphasis id="06803410FFD0522CFF48F8E98518F8A1" bold="true" box="[151,279,1797,1823]" pageId="102" pageNumber="103">Diagnosis.</emphasis>
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The disk is covered with numerous small, uniform scales, conspicuous when dried, covered by a thin skin layer. The radial shields are moderately developed, elongated, visible in dried specimens. The adradial genital plate is well developed, tightly articulating with the radial shield, proximally slightly curved. The abradial genital plate is short and externally only slightly conspicuous. The articulation surface of the adradial genital plate is an elongate elevation with a small pit dorsally. The articulation surface of the radial shield is a slightly prominent elevation with distinct proximal groove. The genital slits are long; genital plates bordering about two thirds of the slits. The jaw bears numerous spiniform oral papillae, similar in shape to the cluster of the 3–4 ventralmost teeth. Two adoral shield papillae and one adjacent oral papilla of each jaw are spiniform, approximately 2 times as long as the oral papillae, placed around the second tentacle pore. The half-jaws are slightly elongated. The adradial sides of the jaws bear a few sharp straight folds distally. The dental plate is small, elongated, entire, without folds and with few rounded sockets, alternately placed. The teeth are spiniform and few in number (besides the apical papillae there are 4–5 teeth). All teeth are of similar shape. The oral shield is rhomboidal, almost devoid of a distinct distal lobe. The adoral shields have distal bilobed wings and are proximally very narrow bars. Dorsal arm plates are moderately developed in some segments but fragmented proximally. Ventral arm plate is well defined. The arm spine articulations are placed almost parallel in relation to the lateral plate. The muscle and nerve openings are similar in size. A low perforated lobe occupies the dorsal part of the articulations. The sigmoidal fold is absent. The spines are relatively long, flattened, solid over almost their entire length, containing only a small lumen. Distally the dorsal and middle arm spines transform into hooks. One small spiniform tentacle scale is placed on the lateral arm plate. The vertebrae have a broad dorsal keel, with a blunt end; the dorsal medial suture is indistinct. The articulation is zygospondylous. The podial basins are large.
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</paragraph>
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</subSubSection>
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<subSubSection id="7CEEBB89FFD1522DFF19FD2B86D6FC92" pageId="103" pageNumber="104" type="materials_examined">
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<paragraph id="344BE802FFD1522DFF19FD2B86D6FC92" blockId="103.[151,1436,152,812]" pageId="103" pageNumber="104">
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<emphasis id="06803410FFD1522DFF19FD2B85AFFD5F" bold="true" box="[198,416,711,737]" pageId="103" pageNumber="104">Material studied.</emphasis>
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<taxonomicName id="F3F49381FFD1522DFE75FD248755FD5C" authority="(Lyman, 1878)" baseAuthorityName="Lyman" baseAuthorityYear="1878" box="[426,858,712,738]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="103" pageNumber="104" phylum="Echinodermata" rank="species" species="dentatus">
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<emphasis id="06803410FFD1522DFE75FD248693FD5F" box="[426,668,712,737]" italics="true" pageId="103" pageNumber="104">Ophiolycus dentatus</emphasis>
|
||
(
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<bibRefCitation id="506595F3FFD1522DFD71FD24875EFD5C" author="Lyman, T." box="[686,849,712,738]" pageId="103" pageNumber="104" pagination="65 - 168" refId="ref90096" refString="Lyman, T. (1878) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger, under Prof. Sir Wyville Thomson, F. R. S. Part 1. Bulletin of the Museum of Comparative Zoology, Harvard 5, 65 - 168." type="journal article" year="1878">Lyman, 1878</bibRefCitation>
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||
)
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</taxonomicName>
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,
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<materialsCitation id="849CE25FFFD1522DFCB6FD2486D9FC92" collectionCode="ZMUC, ZMMU" pageId="103" pageNumber="104" specimenCount="19">
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||
<collectionCode id="52E570C7FFD1522DFCB6FD2487CEFD5C" box="[873,961,712,738]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="103" pageNumber="104">ZMUC</collectionCode>
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||
,
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<specimenCount id="22F2238BFFD1522DFC10FD24806EFD5C" box="[975,1121,712,738]" pageId="103" pageNumber="104" type="generic">3 specimens</specimenCount>
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||
(
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||
<figureCitation id="ACCFF487FFD1522DFBABFD2480F0FD5C" box="[1140,1279,712,738]" captionStart-0="FIGURE 71" captionStart-1="FIGURE 72" captionStartId-0="103.[151,256,1758,1782]" captionStartId-1="104.[151,255,1858,1882]" captionTargetBox-0="[158,1428,866,1736]" captionTargetBox-1="[158,1428,538,1828]" captionTargetId-0="figure-414@103.[158,1428,866,1736]" captionTargetId-1="figure-249@104.[158,1428,538,1828]" captionTargetPageId-0="103" captionTargetPageId-1="104" captionText-0="FIGURE 71. Ophiolycus dentatus (Lyman, 1878), ZMUC, 19.5 mm dd, external views and details (A–H). A. dorsal view; B. disk details showing small radial shields concealed under the skin; C. ventral view; D. ventral view showing genital slits; E. proximal arm segments in situ; F. same, details showing fragmented dorsal arm plates concealed under the skin; G. distal arm segments in situ showing entire dorsal arm plates; H. distal segments, hooks; lateral plate and spine articulations, proximal segments; proximal vertebra, proximal view; I. Ophioscolex glacialis, distal segments, hook-shaped spines. Scale bars: E, 1 mm; F, 300 µ; G, H–I, 100 µ; Photos: A–C Tatiana Korshunova; E–I Alexander Martynov." captionText-1="FIGURE 72. Ophiolycus dentatus (Lyman, 1878), ZMUC, details. A–B, arm spine articulations, proximal segments; C. same, dorsalmost articulation details; D. same, ventralmost articulation details; E. arm spine articulation, distal segments; F. dorslamost spine; G. spine; H. proximal vertebra, dorsal view; I. same, ventral view; J. same, proximal view; K. same, distal view; L. spine, ventral view; M. solid proximal spine, sectioned transversally showing only small cavities. Scale bars: A, F, G–K, 300 µ; B–E, L, M, 100 µ; Photos: Alexander Martynov" pageId="103" pageNumber="104">Figs 71–72</figureCitation>
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);
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<taxonomicName id="F3F49381FFD1522DFAC9FD248577FCB9" authority=", ZMMU" authorityName="ZMMU" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="103" pageNumber="104" phylum="Echinodermata" rank="species" species="purpureus">
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<emphasis id="06803410FFD1522DFAC9FD248501FCB8" italics="true" pageId="103" pageNumber="104">Ophiolycus purpureus</emphasis>
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, ZMMU
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</taxonomicName>
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D-845,
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<specimenCount id="22F2238BFFD1522DFE05FD018673FCB9" box="[474,636,749,775]" pageId="103" pageNumber="104" type="generic">11 specimens</specimenCount>
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||
; Mareano,
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||
<specimenCount id="22F2238BFFD1522DFD21FD018780FCB9" box="[766,911,749,775]" pageId="103" pageNumber="104" type="generic">3 specimens</specimenCount>
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||
(
|
||
<figureCitation id="ACCFF487FFD1522DFC41FD018028FCB9" box="[926,1063,749,775]" captionStart-0="FIGURE 73" captionStart-1="FIGURE 74" captionStartId-0="105.[151,255,1724,1748]" captionStartId-1="106.[151,255,1650,1674]" captionTargetBox-0="[158,1428,530,1703]" captionTargetBox-1="[159,1427,194,1628]" captionTargetId-0="figure-260@105.[158,1428,530,1703]" captionTargetId-1="figure-143@106.[159,1427,194,1628]" captionTargetPageId-0="105" captionTargetPageId-1="106" captionText-0="FIGURE 73. Ophiolycus purpureus (Düben & Koren, 1846), Mareano program, 8 mm dd, external views and details. A; dorsal view; B. ventral view; C. dorsal view showing distal arm segments with hooks; D. dorsal disk view, details showing small radial shields; E. interradial view showing long genital slits; F. arm spine articulations, proximal segments; G. arm spine articulation bear hooks, distal segments; H. proximalmost segments showing large distinct dorsalmost articulations; I. arm spine articulations, proximal segments; J. distal segments, hooks; K. spine; L. same, ventral view; M. spine section showing only small cavities. Scale bars: F-H, J-M, 100 µ; I, 300 µ. Photos: A–E Tatiana Korshunova; F–M Alexander Martynov." captionText-1="FIGURE 74. Ophiolycus purpureus (Düben & Koren, 1846), Mareano program, 8 mm dd, details. A. proximal vertebra, dorsal view; B. same, ventral view; C. same, proximal view; D. same, distal view; E. radial shield, ventral view; F. dental plate; G. adradial genital plate, articulation surface; H. jaw, adradial view; I. adradial and abradial genital plates in situ. Scale bars: A–B, E, H, I, 300 µ; C, D, F–G, 100 µ. Photos: Alexander Martynov." pageId="103" pageNumber="104">Figs 73–74</figureCitation>
|
||
);
|
||
<taxonomicName id="F3F49381FFD1522DFBE3FD0184DBFC92" authority="Mortensen, 1936" authorityName="Mortensen" authorityYear="1936" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="103" pageNumber="104" phylum="Echinodermata" rank="species" species="nutrix">
|
||
<emphasis id="06803410FFD1522DFBE3FD01811FFCB8" box="[1084,1296,749,774]" italics="true" pageId="103" pageNumber="104">Ophiolycus nutrix</emphasis>
|
||
<bibRefCitation id="506595F3FFD1522DFAC8FD0184DBFC92" author="Mortensen, T." pageId="103" pageNumber="104" pagination="199 - 348" refId="ref90871" refString="Mortensen, T. (1936) Echinoidea and Ophiuroidea. Discovery Reports, 12, 199 - 348." type="journal article" year="1936">Mortensen, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<collectionCode id="52E570C7FFD1522DFF3FFCFE8534FC92" box="[224,315,786,812]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="103" pageNumber="104" type="Museum">ZMMU</collectionCode>
|
||
D-818,
|
||
<specimenCount id="22F2238BFFD1522DFE43FCFE8623FC92" box="[412,556,786,812]" pageId="103" pageNumber="104" type="generic">2 specimens</specimenCount>
|
||
(
|
||
<figureCitation id="ACCFF487FFD1522DFDE3FCFE86C2FC92" box="[572,717,786,812]" captionStart="FIGURE 27" captionStartId="38.[151,256,1544,1568]" captionTargetBox="[158,1428,194,1521]" captionTargetId="figure-295@38.[158,1428,194,1523]" captionTargetPageId="38" captionText="FIGURE 27. Arm spine articulations and spines of the various taxa of the family Ophiomyxidae. A–C, Ophioscolex glacialis Müller & Troschel, 1842, ZMMU D-817, 27 mm dd; spine articulations of the proximal segments; D. spine, lateral view, E. spine, ventral view showing large nerve opening; F. Ophiomyxa brevirima H.L. Clark, 1915, ZMMU D- 725, 19 mm dd, distal segments; G. Ophiomyxa brevirima, proximal segments; H. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, proximal segments; I. Ophiocymbium cavernosum, ZMMU D-795, 10.5 mm, proximal segment; J. Ophiocymbium tanyae sp. nov., holotype, ZMMU D-799, 9.5 mm dd, proximal segments; K. same, middle segments; L. Ophiologimus cf. secundus Koehler, 1914, ZMMU D-815, 4 mm dd, proximal segments; M. same a proximal segment with spines; N. same, details of the articulations of the proximal segments; O. Ophiobyrsa rudis Lyman, 1878, ZMMU D-868, 12.5 mm dd; P. Ophiologimus cf. secundus, most proximal segments; Q. Ophiolycus purpureus (Düben & Koren, 1846), ZMMU D-845, 5.5 mm dd, most proximal segments; R. same, proximal segments; S. Ophiolycus nutrix Mortensen, 1936, ZMMU D-818, 8 mm dd, proximal segments with spines; T. same, other proximal segments; U. same, spine base, ventral view; Scale bars: A-B, 300 µ; C, 200 µ; D, 400 µ; E, 150 µ; F, 100 µ; G, 150 µ; H, 100 µ; I, 100 µ; J, 100 µ; K, 200 µ; L, 100 µ; M, 100 µ; N, 100 µ; O, 250 µ; P, 100 µ; Q, 100 µ; R, 100 µ; S–T, 120 µ; U, 150 µ. Photos: Alexander Martynov." pageId="103" pageNumber="104">Figs 27S–U</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="608BB88AFFD1522DFF48F9328509F80B" pageId="103" pageNumber="104" startId="103.[151,256,1758,1782]" targetBox="[158,1428,866,1736]" targetPageId="103">
|
||
<paragraph id="344BE802FFD1522DFF48F9328509F80B" blockId="103.[151,1436,1758,1973]" pageId="103" pageNumber="104">
|
||
<emphasis id="06803410FFD1522DFF48F9328525F948" bold="true" box="[151,298,1758,1782]" pageId="103" pageNumber="104">FIGURE 71.</emphasis>
|
||
<taxonomicName id="F3F49381FFD1522DFEEDF93286B0F94B" authority="(Lyman, 1878)" baseAuthorityName="Lyman" baseAuthorityYear="1878" box="[306,703,1758,1781]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="103" pageNumber="104" phylum="Echinodermata" rank="species" species="dentatus">
|
||
<emphasis id="06803410FFD1522DFEEDF932861DF94B" box="[306,530,1758,1781]" italics="true" pageId="103" pageNumber="104">Ophiolycus dentatus</emphasis>
|
||
(
|
||
<bibRefCitation id="506595F3FFD1522DFDFDF93286B7F94B" author="Lyman, T." box="[546,696,1758,1781]" pageId="103" pageNumber="104" pagination="65 - 168" refId="ref90096" refString="Lyman, T. (1878) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger, under Prof. Sir Wyville Thomson, F. R. S. Part 1. Bulletin of the Museum of Comparative Zoology, Harvard 5, 65 - 168." type="journal article" year="1878">Lyman, 1878</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ZMUC, 19.5 mm dd, external views and details (A–H). A. dorsal view; B. disk details showing small radial shields concealed under the skin; C. ventral view; D. ventral view showing genital slits; E. proximal arm segments in situ; F. same, details showing fragmented dorsal arm plates concealed under the skin; G. distal arm segments in situ showing entire dorsal arm plates; H. distal segments, hooks; lateral plate and spine articulations, proximal segments; proximal vertebra, proximal view; I.
|
||
<taxonomicName id="F3F49381FFD1522DFC33F8B280D3F8CB" authorityName="Muller & Troschel" authorityYear="1842" box="[1004,1244,1886,1909]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioscolex" kingdom="Animalia" order="Ophioscolecida" pageId="103" pageNumber="104" phylum="Echinodermata" rank="species" species="glacialis">
|
||
<emphasis id="06803410FFD1522DFC33F8B280D3F8CB" box="[1004,1244,1886,1909]" italics="true" pageId="103" pageNumber="104">Ophioscolex glacialis</emphasis>
|
||
</taxonomicName>
|
||
, distal segments, hook-shaped spines. Scale bars: E, 1 mm; F, 300 µ; G, H–I, 100 µ; Photos: A–C Tatiana Korshunova; E–I Alexander Martynov.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="7CEEBB89FFDE5223FF19FF7B85DCFE62" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105" type="discussion">
|
||
<paragraph id="344BE802FFDE5222FF19FF7B85E6FE54" blockId="104.[151,1436,151,490]" pageId="104" pageNumber="105">
|
||
<emphasis id="06803410FFDE5222FF19FF7B8530FF0F" bold="true" box="[198,319,151,177]" pageId="104" pageNumber="105">Remarks.</emphasis>
|
||
One of the characteristic external features of the genus
|
||
<taxonomicName id="F3F49381FFDE5222FC08FF748053FF0F" authorityName="Mortensen" authorityYear="1933" box="[983,1116,152,177]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDE5222FC08FF748053FF0F" box="[983,1116,152,177]" italics="true" pageId="104" pageNumber="105">Ophiolycus</emphasis>
|
||
</taxonomicName>
|
||
was the presence of hooks on the distal arm segments (
|
||
<bibRefCitation id="506595F3FFDE5222FE3BFF5686B5FF6A" author="Mortensen, T." box="[484,698,186,212]" pageId="104" pageNumber="105" pagination="215 - 400" refId="ref90767" refString="Mortensen, T. (1933 b). Echinoderms of South Africa. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 93, 215 - 400." type="journal article" year="1933">Mortensen 1933b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="506595F3FFDE5222FD18FF56870CFF6A" author="Mortensen, T." box="[711,771,186,212]" pageId="104" pageNumber="105" pagination="199 - 348" refId="ref90871" refString="Mortensen, T. (1936) Echinoidea and Ophiuroidea. Discovery Reports, 12, 199 - 348." type="journal article" year="1936">1936</bibRefCitation>
|
||
) (
|
||
<figureCitation id="ACCFF487FFDE5222FCC3FF568781FF6A" box="[796,910,186,212]" captionStart="FIGURE 71" captionStartId="103.[151,256,1758,1782]" captionTargetBox="[158,1428,866,1736]" captionTargetId="figure-414@103.[158,1428,866,1736]" captionTargetPageId="103" captionText="FIGURE 71. Ophiolycus dentatus (Lyman, 1878), ZMUC, 19.5 mm dd, external views and details (A–H). A. dorsal view; B. disk details showing small radial shields concealed under the skin; C. ventral view; D. ventral view showing genital slits; E. proximal arm segments in situ; F. same, details showing fragmented dorsal arm plates concealed under the skin; G. distal arm segments in situ showing entire dorsal arm plates; H. distal segments, hooks; lateral plate and spine articulations, proximal segments; proximal vertebra, proximal view; I. Ophioscolex glacialis, distal segments, hook-shaped spines. Scale bars: E, 1 mm; F, 300 µ; G, H–I, 100 µ; Photos: A–C Tatiana Korshunova; E–I Alexander Martynov." pageId="104" pageNumber="105">Figs 71H</figureCitation>
|
||
;
|
||
<figureCitation id="ACCFF487FFDE5222FC44FF5687C7FF6A" box="[923,968,186,212]" captionStart="FIGURE 73" captionStartId="105.[151,255,1724,1748]" captionTargetBox="[158,1428,530,1703]" captionTargetId="figure-260@105.[158,1428,530,1703]" captionTargetPageId="105" captionText="FIGURE 73. Ophiolycus purpureus (Düben & Koren, 1846), Mareano program, 8 mm dd, external views and details. A; dorsal view; B. ventral view; C. dorsal view showing distal arm segments with hooks; D. dorsal disk view, details showing small radial shields; E. interradial view showing long genital slits; F. arm spine articulations, proximal segments; G. arm spine articulation bear hooks, distal segments; H. proximalmost segments showing large distinct dorsalmost articulations; I. arm spine articulations, proximal segments; J. distal segments, hooks; K. spine; L. same, ventral view; M. spine section showing only small cavities. Scale bars: F-H, J-M, 100 µ; I, 300 µ. Photos: A–E Tatiana Korshunova; F–M Alexander Martynov." pageId="104" pageNumber="105">73J</figureCitation>
|
||
), whereas the
|
||
<typeStatus id="EB4F56A0FFDE5222FBACFF5680AAFF6A" box="[1139,1189,186,212]" pageId="104" pageNumber="105">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="F3F49381FFDE5222FF48FF318526FF48" authorityName=": Muller & Troschel" authorityYear="1842" box="[151,297,221,246]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioscolex" kingdom="Animalia" order="Ophioscolecida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDE5222FF48FF318526FF48" box="[151,297,221,246]" italics="true" pageId="104" pageNumber="105">Ophioscolex</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3F49381FFDE5222FEE7FF3185CEFF48" authorityName="Muller & Troschel" authorityYear="1842" box="[312,449,221,246]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioscolex" kingdom="Animalia" order="Ophioscolecida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="species" species="glacialis">
|
||
<emphasis id="06803410FFDE5222FEE7FF3185CEFF48" box="[312,449,221,246]" italics="true" pageId="104" pageNumber="105">O. glacialis</emphasis>
|
||
</taxonomicName>
|
||
does not possess hook-shaped spines on the distal segments. However, well-defined hooks on the distal arm segments are apparently widely distributed within various distantly-related taxa of the family
|
||
<taxonomicName id="F3F49381FFDE5222FF31FECE85AAFE82" authorityName="Ljungman" authorityYear="1867" box="[238,421,290,316]" class="Ophiuroidea" family="Ophiomyxidae" kingdom="Animalia" order="Ophiacanthida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="family">Ophiomyxidae</taxonomicName>
|
||
. In the present study, true distal hooks were detected in the genera
|
||
<taxonomicName id="F3F49381FFDE5222FB02FECE8167FE85" authorityName="Muller & Troschel" authorityYear="1842" box="[1245,1384,290,315]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiomyxa" kingdom="Animalia" order="Ophiacanthida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDE5222FB02FECE8167FE85" box="[1245,1384,290,315]" italics="true" pageId="104" pageNumber="105">Ophiomyxa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3F49381FFDE5222FF48FEA9852EFEE0" authorityName="Lyman" authorityYear="1878" box="[151,289,325,350]" class="Ophiuroidea" family="Ophiobyrsidae" genus="Ophiobyrsa" kingdom="Animalia" order="Ophiacanthida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDE5222FF48FEA9852EFEE0" box="[151,289,325,350]" italics="true" pageId="104" pageNumber="105">Ophiobyrsa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="ACCFF487FFDE5222FEEEFEA9859FFEE1" box="[305,400,325,351]" captionStart="FIGURE 1" captionStartId="10.[151,255,1651,1675]" captionTargetBox="[158,1428,194,1630]" captionTargetId="figure-186@10.[158,1428,194,1630]" captionTargetPageId="10" captionText="FIGURE 1. Ophiomyxa pentagona (Lamarck, 1816), ZMMU D-553, 21 mm dd, type species of the genus Ophiomyxa, the family Ophiomyxidae. External morphology and microstructural details. A. dorsal view; B. ventral view; C–H, proximal segments: C. lateral arm plate; D, details of arm spine articulations; E. spine; F. solid spine sectioned transversally; G. spine, ventral view; H. vertebra, dorsal view; I–L. distal segments: I. lateral arm plate; J. details of arm spine articulations; K. vertebra, dorsal view; L. distal arm segments in situ showing hooks; M. articulation surface of the adradial genital plate; N. radial shield, ventral view; O. articulating adradial genital plate and radial shield; P. general overview of the oral structures; Q. teeth in situ; R. dental plate; S. jaw, abradial view; T. jaw, adradial view; Scale bars: C, E, I, L–N, Q, 300 µ; D, F G, K, 100 µ; J, 30 µ; H, O, R–S, 1 mm; P, 600 µ. Photos: A–B Tatiana Korshunova; C–T Alexander Martynov." pageId="104" pageNumber="105">Figs 1L</figureCitation>
|
||
;
|
||
<figureCitation id="ACCFF487FFDE5222FE42FEA985C6FEE1" box="[413,457,325,351]" captionStart="FIGURE 79" captionStartId="113.[151,255,1646,1670]" captionTargetBox="[158,1428,194,1625]" captionTargetId="figure-166@113.[158,1428,194,1625]" captionTargetPageId="113" captionText="FIGURE 79. Ophiobyrsa rudis Lyman, 1878, ZMMU D-868, 12.5 mm dd, external views and details. A. dorsal view; B. ventral view; C. disk, dorsal view, details showing moderately defined radial shields; D. ventral view, details, E. interradial view, showing long genital slits; F. spine articulations, distal segments; G–H, spine articulations, most proximal segments; I. spine articulation, proximal segments, note massive curved lobe of the dorsalmost articulation; J. distal segments showing hooks; K. soild proximal spine, sectioned transversally; L. proximal spine; M. same, ventral view. Scale bars: F–M, 100 µ. Photos: A–E Tatiana Korshunova; F–M Alexander Martynov." pageId="104" pageNumber="105">79J</figureCitation>
|
||
).
|
||
<taxonomicName id="F3F49381FFDE5222FE01FEA987E9FEE1" authority="H. L. Clark, 1911" authorityName="H. L. Clark" authorityYear="1911" box="[478,998,325,351]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiosyzygus" kingdom="Animalia" order="Ophioscolecida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="species" species="disacanthus">
|
||
<emphasis id="06803410FFDE5222FE01FEA9871BFEE0" box="[478,788,325,350]" italics="true" pageId="104" pageNumber="105">Ophiosyzygus disacanthus</emphasis>
|
||
H.L.
|
||
<bibRefCitation id="506595F3FFDE5222FC86FEA987E9FEE1" author="Clark, H. L." box="[857,998,325,351]" pageId="104" pageNumber="105" pagination="1 - 302" refId="ref85928" refString="Clark, H. L. (1911) North Pacific Ophiurans in the collection of the United States National Museum. Bulletin of the United States National Museum 75, 1 - 302." type="journal article" year="1911">Clark, 1911</bibRefCitation>
|
||
</taxonomicName>
|
||
, also has on the distalmost segments small hook-shaped spines (
|
||
<figureCitation id="ACCFF487FFDE5222FE3BFE848640FE3C" box="[484,591,360,386]" captionStart="FIGURE 71" captionStartId="103.[151,256,1758,1782]" captionTargetBox="[158,1428,866,1736]" captionTargetId="figure-414@103.[158,1428,866,1736]" captionTargetPageId="103" captionText="FIGURE 71. Ophiolycus dentatus (Lyman, 1878), ZMUC, 19.5 mm dd, external views and details (A–H). A. dorsal view; B. disk details showing small radial shields concealed under the skin; C. ventral view; D. ventral view showing genital slits; E. proximal arm segments in situ; F. same, details showing fragmented dorsal arm plates concealed under the skin; G. distal arm segments in situ showing entire dorsal arm plates; H. distal segments, hooks; lateral plate and spine articulations, proximal segments; proximal vertebra, proximal view; I. Ophioscolex glacialis, distal segments, hook-shaped spines. Scale bars: E, 1 mm; F, 300 µ; G, H–I, 100 µ; Photos: A–C Tatiana Korshunova; E–I Alexander Martynov." pageId="104" pageNumber="105">Figs 71I</figureCitation>
|
||
;
|
||
<figureCitation id="ACCFF487FFDE5222FD81FE848683FE3C" box="[606,652,360,386]" captionStart="FIGURE 75" captionStartId="108.[151,257,1815,1839]" captionTargetBox="[158,1428,373,1809]" captionTargetId="figure-197@108.[158,1428,373,1809]" captionTargetPageId="108" captionText="FIGURE 75. Ophiosyzygus disacanthus H.L. Clark, 1911, ZMMU D-875, 12 mm dd, external view and details. A. dorsal view; B, same, details showing small radial shields and proximal arm segments; C. ventral view; D. lateral plate and spine articulations, proximal segments; E. spine articulations, distal segments; F. ventral view, details showing mouth parts; G. ventralmost spine articulation, mostproximal segments; H. spine articulation, details, proximal segments; I. distal segments with the spines in situ; J. further distal segments showing modification of the dorsal spine into a hookshaped structure; K. proximal spine, ventral view; L. proximal spine. Scale bars: D, L, 300 µ; E-K, 100 µ. Photos: A–C, F, Tatiana Korshunova; D–E; G–L Alexander Martynov." pageId="104" pageNumber="105">75J</figureCitation>
|
||
), somewhat different from the typical hooks of
|
||
<taxonomicName id="F3F49381FFDE5222FB03FE848167FE3F" authorityName="Muller & Troschel" authorityYear="1842" box="[1244,1384,360,385]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiomyxa" kingdom="Animalia" order="Ophiacanthida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDE5222FB03FE848167FE3F" box="[1244,1384,360,385]" italics="true" pageId="104" pageNumber="105">Ophiomyxa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3F49381FFDE5222FF48FE66852BFE1D" authorityName="Lyman" authorityYear="1878" box="[151,292,394,419]" class="Ophiuroidea" family="Ophiobyrsidae" genus="Ophiobyrsa" kingdom="Animalia" order="Ophiacanthida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDE5222FF48FE66852BFE1D" box="[151,292,394,419]" italics="true" pageId="104" pageNumber="105">Ophiobyrsa</emphasis>
|
||
</taxonomicName>
|
||
. Thus, the hooks or hook-shaped spines are clearly a convergent feature. Besides hook-shaped spines, the genus
|
||
<taxonomicName id="F3F49381FFDE5222FEBBFE4185E5FE78" authorityName="Mortensen" authorityYear="1933" box="[356,490,429,454]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDE5222FEBBFE4185E5FE78" box="[356,490,429,454]" italics="true" pageId="104" pageNumber="105">Ophiolycus</emphasis>
|
||
</taxonomicName>
|
||
is well distinguished from the genus
|
||
<taxonomicName id="F3F49381FFDE5222FC42FE418020FE78" authorityName=": Muller & Troschel" authorityYear="1842" box="[925,1071,429,454]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioscolex" kingdom="Animalia" order="Ophioscolecida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDE5222FC42FE418020FE78" box="[925,1071,429,454]" italics="true" pageId="104" pageNumber="105">Ophioscolex</emphasis>
|
||
</taxonomicName>
|
||
, and the restricted diagnosis of both genera are given above.
|
||
</paragraph>
|
||
<caption id="608BB88AFFDE5222FF48F8AE8036F867" pageId="104" pageNumber="105" startId="104.[151,255,1858,1882]" targetBox="[158,1428,538,1828]" targetPageId="104">
|
||
<paragraph id="344BE802FFDE5222FF48F8AE8036F867" blockId="104.[151,1436,1858,2010]" pageId="104" pageNumber="105">
|
||
<emphasis id="06803410FFDE5222FF48F8AE8528F8E4" bold="true" box="[151,295,1858,1882]" pageId="104" pageNumber="105">FIGURE 72.</emphasis>
|
||
<taxonomicName id="F3F49381FFDE5222FEF1F8AE86BFF8E7" authority="(Lyman, 1878)" baseAuthorityName="Lyman" baseAuthorityYear="1878" box="[302,688,1858,1881]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="104" pageNumber="105" phylum="Echinodermata" rank="species" species="dentatus">
|
||
<emphasis id="06803410FFDE5222FEF1F8AE8607F8E7" box="[302,520,1858,1881]" italics="true" pageId="104" pageNumber="105">Ophiolycus dentatus</emphasis>
|
||
(
|
||
<bibRefCitation id="506595F3FFDE5222FDC8F8AE86A6F8E7" author="Lyman, T." box="[535,681,1858,1881]" pageId="104" pageNumber="105" pagination="65 - 168" refId="ref90096" refString="Lyman, T. (1878) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger, under Prof. Sir Wyville Thomson, F. R. S. Part 1. Bulletin of the Museum of Comparative Zoology, Harvard 5, 65 - 168." type="journal article" year="1878">Lyman, 1878</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ZMUC, details. A–B, arm spine articulations, proximal segments; C. same, dorsalmost articulation details; D. same, ventralmost articulation details; E. arm spine articulation, distal segments; F. dorslamost spine; G. spine; H. proximal vertebra, dorsal view; I. same, ventral view; J. same, proximal view; K. same, distal view; L. spine, ventral view; M. solid proximal spine, sectioned transversally showing only small cavities. Scale bars: A, F, G–K, 300 µ; B–E, L, M, 100 µ; Photos: Alexander Martynov
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="344BE802FFDF5223FF19FF748193FE09" blockId="105.[151,1436,152,476]" pageId="105" pageNumber="106">
|
||
O’Hara (pers. comm.) suggested that
|
||
<taxonomicName id="F3F49381FFDF5223FD40FF748078FF0C" authority="McKnight, 2003" authorityName="McKnight" authorityYear="2003" box="[671,1143,152,178]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="species" species="farquhari">
|
||
<emphasis id="06803410FFDF5223FD40FF7487AEFF0F" box="[671,929,152,177]" italics="true" pageId="105" pageNumber="106">Ophiolycus farquhari</emphasis>
|
||
<bibRefCitation id="506595F3FFDF5223FC70FF748078FF0C" author="McKnight, D. G." box="[943,1143,152,178]" pageId="105" pageNumber="106" pagination="1 - 36" refId="ref90600" refString="McKnight, D. G. (2003) New brittle-stars (Echinodermata: Ophiuroidea) from New Zealand waters. Zootaxa 352, 1 - 36." type="journal article" year="2003">McKnight, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
may be a synonym of
|
||
<taxonomicName id="F3F49381FFDF5223FF48FF5186CFFF69" authority="H. L. Clark, 1925" authorityName="H. L. Clark" authorityYear="1925" box="[151,704,189,215]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiologimus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="species" species="quadrispinus">
|
||
<emphasis id="06803410FFDF5223FF48FF5185E9FF68" box="[151,486,189,214]" italics="true" pageId="105" pageNumber="106">Ophiologimus quadrispinus</emphasis>
|
||
H.L.
|
||
<bibRefCitation id="506595F3FFDF5223FDF1FF5186CFFF69" author="Clark, H. L." box="[558,704,189,215]" pageId="105" pageNumber="106" pagination="89 - 111" refId="ref86065" refString="Clark, H. L. (1925) Marine zoology of tropical central Pacific: Echinoderms other than sea-stars. Bulletin of the Bernice Pauahi Bishop Museum 27, 89 - 111." type="journal article" year="1925">Clark, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="F3F49381FFDF5223FD12FF5187DFFF68" authorityName="McKnight" authorityYear="2003" box="[717,976,189,214]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="species" species="farquhari">
|
||
<emphasis id="06803410FFDF5223FD12FF5187DFFF68" box="[717,976,189,214]" italics="true" pageId="105" pageNumber="106">Ophiolycus farquhari</emphasis>
|
||
</taxonomicName>
|
||
is externally similar to
|
||
<taxonomicName id="F3F49381FFDF5223FB2DFF518539FF45" authorityName="H. L. Clark" authorityYear="1925" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiologimus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="species" species="quadrispinus">
|
||
<emphasis id="06803410FFDF5223FB2DFF518539FF45" italics="true" pageId="105" pageNumber="106">Ophiologimus quadrispinus</emphasis>
|
||
</taxonomicName>
|
||
in general appearance of the disk and arms, elongate radial shields and absence of the long adoral shield papillae. The latter feature is diagnostic for all species of the genus
|
||
<taxonomicName id="F3F49381FFDF5223FB5BFEE48101FE9F" authorityName="Mortensen" authorityYear="1933" box="[1156,1294,264,289]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDF5223FB5BFEE48101FE9F" box="[1156,1294,264,289]" italics="true" pageId="105" pageNumber="106">Ophiolycus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="506595F3FFDF5223FAC0FEE484E9FEF9" author="McKnight, D. G." pageId="105" pageNumber="106" pagination="1 - 36" refId="ref90600" refString="McKnight, D. G. (2003) New brittle-stars (Echinodermata: Ophiuroidea) from New Zealand waters. Zootaxa 352, 1 - 36." type="journal article" year="2003">McKnight (2003)</bibRefCitation>
|
||
mentioned the distalmost spines as “weakly hooked at the tip”. Both the
|
||
<typeStatus id="EB4F56A0FFDF5223FBB5FEC18091FEF9" box="[1130,1182,301,327]" pageId="105" pageNumber="106">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="F3F49381FFDF5223FF48FEBE8530FED5" authorityName="H. L. Clark" authorityYear="1911" box="[151,319,338,363]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiologimus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDF5223FF48FEBE8530FED5" box="[151,319,338,363]" italics="true" pageId="105" pageNumber="106">Ophiologimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3F49381FFDF5223FE90FEBE85D9FED5" authorityName="H. L. Clark" authorityYear="1911" box="[335,470,338,363]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiologimus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="species" species="hexactis">
|
||
<emphasis id="06803410FFDF5223FE90FEBE85D9FED5" box="[335,470,338,363]" italics="true" pageId="105" pageNumber="106">O. hexactis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="ACCFF487FFDF5223FE37FEBE865DFED2" box="[488,594,338,364]" captionStart="FIGURE 48" captionStartId="71.[151,255,1689,1713]" captionTargetBox="[151,1436,194,1655]" captionTargetId="figure-108@71.[146,1438,194,1655]" captionTargetPageId="71" captionText="FIGURE 48. Ophiologimus hexactis H.L. Clark, 1911, holotype USNM 25724, 5 mm dd, external views and details. A. dorsal view; B. ventral view; C–D, arm spine articulations, distal segments; E. arm spine articulations, proximal segments; F. distal segments, hooks; G. spine, proximal segments; H. hollow spine, transversally sectioned; I. spine, ventral view; J. proximal vertebra, dorsal view; K. same, ventral view; L. same, proximal view; M. same, distal view. Scale bars: C–H, J, L–M, 100 µ; I, 30 µ.; K, 300 µ. Photos: A–B Tatiana Korshunova; C–M Alexander Martynov." pageId="105" pageNumber="106">Fig. 48F</figureCitation>
|
||
), and other studied material of
|
||
<taxonomicName id="F3F49381FFDF5223FC14FEBE8083FED5" authorityName="H. L. Clark" authorityYear="1925" box="[971,1164,338,363]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiologimus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="species" species="quadrispinus">
|
||
<emphasis id="06803410FFDF5223FC14FEBE8083FED5" box="[971,1164,338,363]" italics="true" pageId="105" pageNumber="106">O. quadrispinus</emphasis>
|
||
</taxonomicName>
|
||
reveal the presence of well-defined hooks. “
|
||
<taxonomicName id="F3F49381FFDF5223FE4CFE948617FE2F" authorityName="Mortensen" authorityYear="1933" box="[403,536,376,401]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDF5223FE4CFE948617FE2F" box="[403,536,376,401]" italics="true" pageId="105" pageNumber="106">Ophiolycus</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="F3F49381FFDF5223FDF3FE948694FE2F" authorityName="McKnight" authorityYear="2003" box="[556,667,376,401]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="species" species="farquhari">
|
||
<emphasis id="06803410FFDF5223FDF3FE948694FE2F" box="[556,667,376,401]" italics="true" pageId="105" pageNumber="106">farquhari</emphasis>
|
||
</taxonomicName>
|
||
therefore is not included into the genus
|
||
<taxonomicName id="F3F49381FFDF5223FBB2FE9480FCFE2F" authorityName="Mortensen" authorityYear="1933" box="[1133,1267,376,401]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiolycus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDF5223FBB2FE9480FCFE2F" box="[1133,1267,376,401]" italics="true" pageId="105" pageNumber="106">Ophiolycus</emphasis>
|
||
</taxonomicName>
|
||
here, but until a complete revision of the genus
|
||
<taxonomicName id="F3F49381FFDF5223FDC8FE7186B2FE08" authorityName="H. L. Clark" authorityYear="1911" box="[535,701,413,438]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiologimus" kingdom="Animalia" order="Ophioscolecida" pageId="105" pageNumber="106" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFDF5223FDC8FE7186B2FE08" box="[535,701,413,438]" italics="true" pageId="105" pageNumber="106">Ophiologimus</emphasis>
|
||
</taxonomicName>
|
||
will be performed, it is retained as a species of doubtful status.
|
||
</paragraph>
|
||
<paragraph id="344BE802FFDF5223FF19FE2E85DCFE62" blockId="105.[151,1436,152,476]" box="[198,467,450,476]" pageId="105" pageNumber="106">
|
||
<emphasis id="06803410FFDF5223FF19FE2E85BAFE62" bold="true" box="[198,437,450,476]" pageId="105" pageNumber="106">Number of species:</emphasis>
|
||
3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |