treatments-xml/data/BC/5D/59/BC5D5914FF8F5271FF48FB038122FBE9.xml

<document id="7065593A2CA0788FEB2C3FB6BF8799CE" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629706483752" checkinUser="felipe" docAuthor="Martynov, Alexander" docDate="2010" docId="BC5D5914FF8F5271FF48FB038122FBE9" docLanguage="en" docName="zt02697p154.pdf" docOrigin="Zootaxa 2697" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Ophiocymbium ninae Martynov 2010, sp. nov." docType="treatment" docVersion="2" lastPageNumber="60" masterDocId="4064216CFFB6524AFFDFFFEC840FFFBE" masterDocTitle="Reassessment of the classification of the Ophiuroidea (Echinodermata), based on morphological characters. I. General character evaluation and delineation of the families Ophiomyxidae and Ophiacanthidae 2697" masterLastPageNumber="154" masterPageNumber="1" pageNumber="58" updateTime="1699098856182" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C76CEDEEED3725A7A238FC3313C216DF">Reassessment of the classification of the Ophiuroidea (Echinodermata), based on morphological characters. I. General character evaluation and delineation of the families Ophiomyxidae and Ophiacanthidae 2697</mods:title>
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<heading id="6F035F6EFF8F5273FF48FB03860FFAB7" bold="true" box="[151,512,1263,1289]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<emphasis id="06803410FF8F5273FF48FB03860FFAB7" bold="true" box="[151,512,1263,1289]" pageId="57" pageNumber="58">
<taxonomicName id="F3F49381FF8F5273FF48FB038593FAB7" ID-CoL="49WVL" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[151,412,1263,1289]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiocymbium" kingdom="Animalia" order="Ophioscolecida" pageId="57" pageNumber="58" phylum="Echinodermata" rank="species" species="ninae" status="sp. nov.">
<emphasis id="06803410FF8F5273FF48FB038593FAB7" bold="true" box="[151,412,1263,1289]" italics="true" pageId="57" pageNumber="58">Ophiocymbium ninae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1DB3896BFF8F5273FE7CFB03860FFAB7" box="[419,512,1263,1289]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="7CEEBB89FF8F5273FF48FAFE86DBFA92" box="[151,724,1298,1324]" pageId="57" pageNumber="58" type="description">
<paragraph id="344BE802FF8F5273FF48FAFE86DBFA92" blockId="57.[151,724,1263,1324]" box="[151,724,1298,1324]" pageId="57" pageNumber="58">
<figureCitation id="ACCFF487FF8F5273FF48FAFE853EFA92" box="[151,305,1298,1324]" captionStart="FIGURE 4" captionStartId="13.[151,256,1339,1363]" captionTargetBox="[282,1312,194,1330]" captionTargetId="figure-434@13.[282,1312,194,1330]" captionTargetPageId="13" captionText="FIGURE 4. Genital plates and radial shields of various taxa of the families Asteronychidae, Gorgonocephalidae and Ophiomyxidae. A. Gorgonocephalus arcticus Leach, 1819, ZMMU D-613, 39 mm dd, adradial and abradial genital plates in situ, showing articulation surface, dorsal view; B. same, articulation surface of the radial shield, ventral view; C. Asteronyx loveni Müller &amp; Troschel, 1842, ZMMU D-833, 17.5 mm dd, articulation surface of the adradial genital plate, dorsal view; D. same, articulation surface of the radial shield, ventral view. E. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, articulation surface of the adradial genital plate, dorsal view; F. same, radial shield showing articulation surface, ventral view; G. same, articulating adradial genital plate and radial shield, dorsal view; H. Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, dissected disk showing most proximal vertebrae and absence of the both genital plates and radial shield, dorsal view; I. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, adradial and abradial genital plates in situ, ventral view; J. Ophioplexa condita gen. et sp. nov., paratype ca. 17 mm dd, ZMMU D-792, dissected disk showing articulating adradial genital plate and radial shield in situ, dorsal view; K. same, separate articulating adradial genital plate and radial shield, dorsal view; L. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, adradial genital plate, dorsal view; M. Ophiologimus cf. secundus Koehler, 1914, 4 mm dd, ZMMU D-814, adradial genital plate, dorsal view; N. Ophioplexa condita gen. et sp. nov., separate adradial genital plate, dorsal view; O. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, adradial and abradial genital plates, dorsal view; P. Ophioplexa condita gen. et sp. nov., separate articulating adradial genital plate and radial shield, dorsal view; Q. articulation surface of the adradial genital plate, dorsal view; R. radial shield, ventral view, showing distally the distinctive excavated articulation surface; S. articulation surface of the radial shield, ventral view; T. Ophiocymbium ninae sp. nov., paratype 5 mm dd, radial shield, ventral view; U. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, same, articulation surface of the radial shield, proximal view; Scale bars: A– B,1 mm; C–D, 300 µ; E–F, 100 µ; G, 200 µ; H, 300 µ; I, 200 µ; J, 1000 µ; K, 300 µ; L, 100 µ; M, 300 µ; N, 300 µ; O, 200 µ; P, 100 µ; Q–T, 100 µ; U, 200 µ. Photos: Alexander Martynov." pageId="57" pageNumber="58">Figures 4L,T</figureCitation>
; 
<figureCitation id="ACCFF487FF8F5273FEE1FAFE85DCFA92" box="[318,467,1298,1324]" captionStart="FIGURE 12" captionStartId="21.[151,255,1049,1073]" captionTargetBox="[158,1428,194,1027]" captionTargetId="figure-465@21.[158,1428,194,1027]" captionTargetPageId="21" captionText="FIGURE 12. Dental plates and jaws of the genera Ophioscolex, Ophiocymbium and Ophiologimus. A–B, Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, dental plates; C–D, Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, abradial and adradial views of the jaws respectively; E. same, dental plate; F. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-803, dental plate; G. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-805, dental plate; H. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, dental plate; I–J, Ophiologimus cf. secundus Koehler, 1914, ZMMU D-815, 4 mm dd, dental plates; K–L, Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-805, abradial and adradial views of the jaws respectively; M. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, jaw, adradial view; N–O, Ophiologimus cf. secundus Koehler, 1914, ZMMU D-815, 4 mm dd, abradial and adradial views of the jaws respectively; Scale bars: A, 200 µ; B, 200 µ; C–D, 300 µ; E, 100 µ; F, 300 µ; G, 100 µ; H, 300 µ; I–J, 100 µ; K–L, 300 µ; M, 100 µ; N–O, 300 µ. Photos: Alexander Martynov." pageId="57" pageNumber="58">12F–G, K–L</figureCitation>
; 
<figureCitation id="ACCFF487FF8F5273FE00FAFE865FFA92" box="[479,592,1298,1324]" captionStart="FIGURE 17" captionStartId="27.[151,255,1036,1060]" captionTargetBox="[161,1426,186,1015]" captionTargetId="figure-399@27.[161,1426,186,1015]" captionTargetPageId="27" captionText="FIGURE 17. Vertebrae of various taxa of the family Ophiomyxidae. A. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, proximal segments, dorsal and ventral views; B. same, distal segments, dorsal and ventral views; C. same, proximal segments, proximal view; D. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, 10.5 mm dd, proximal segments, dorsal view; E. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, proximal segments, dorsal view; F. Ophioscolex glacialis, proximal segments, distal view; G. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, proximal segments, proximal view; H. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, ventral view; I. same, proximal view; J. same, distal view; K. Ophiomyxa pentagona, ZMMU D-553, 21 mm dd, distal view; L. Ophioplexa condita gen. et sp. nov., holotype ca. 18 mm dd, ZMMU D-788, proximal segments, dorsal view; M. same, ventral view; N. same, proximal view; O. same, distal view; P. Ophiocymbium tanyae sp. nov., holotype 9.5 mm dd, ZMMU D-799, proximal segments, dorsal view; Q. same, ventral view; R. same, distal view; S. same, proximal view. Scale bars: A, 1000 µ; B, C, 200 µ, D, 200 µ; E, 300 µ; F, 200 µ; G, 200 µ; H, 300 µ; I, 100 µ; J, 100 µ; K, 1000 µ; L–S, 300 µ. Photos: Alexander Martynov." pageId="57" pageNumber="58">17E, H–J</figureCitation>
; 
<figureCitation id="ACCFF487FF8F5273FD83FAFE8673FA92" box="[604,636,1298,1324]" captionStart="FIGURE 41" captionStartId="53.[151,255,1505,1529]" captionTargetBox="[158,1428,194,1484]" captionTargetId="figure-189@53.[158,1428,194,1484]" captionTargetPageId="53" captionText="FIGURE 41. External views and vertebrae of Ophiocymbium ninae sp. nov. A. paratype, ZMMU D-804, 5.5 mm dd, dorsal view; B. holotype 6.8 mm dd, ZMMU D-802, 6.8 mm dd, ventral view; C. paratype 6.5 mm dd, ZMMU D-803, ventral view; D. paratype, ZMMU D-803, 6.1 mm dd, ventral view; E. holotype, dorsal view, ZMMU D-802, 6.8 mm dd, F. vertebra, ventral view; G. vertebra, dorsal view; Scale bars: F,G, 300 µ. Photos: A–F Tatiana Korshunova; F–G Alexander Martynov." pageId="57" pageNumber="58">41</figureCitation>
; 
<figureCitation id="ACCFF487FF8F5273FD57FAFE86DBFA92" box="[648,724,1298,1324]" captionStart="FIGURE 55" captionStartId="80.[151,258,1764,1788]" captionTargetBox="[209,1364,194,1740]" captionTargetId="figure-174@80.[209,1364,194,1740]" captionTargetPageId="80" captionText="FIGURE 55. Ophioplexa condita gen. nov., sp. nov. and Ophiocymbium ninae sp. nov., details. A. Ophioplexa condita gen. nov., sp. nov., paratype 19 mm dd, ZMMU D-791, disk ventral view; B. same oral frame showing clusters of the ventral teeth (apical papillae); C. same, one interradius showing apical, oral and adoral shield papillae; D-F, same, most proximal arm segments showing tentacle scales and spines; G-H, arm spine articulations, distal segments; I. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-803, details, part of the oral frame and ventral arm side; J. same, jaw and oral papillae, one interradius; K. ventral arm view showing spines and tentacle scales; L. most proximal segments showing details of tentacle scales and spines; M-N, arm spine articulations, proximal segments; Scale bars: A– B, 1 mm; C–D, 500 µ; E–F, 200 µ; G–H, 100 µ; I, 1 mm; J–N, 300 µ. Photos: Alexander Martynov." pageId="57" pageNumber="58">55I–N</figureCitation>
</paragraph>
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<emphasis id="06803410FF8F5273FF48FAB08504FAC8" bold="true" box="[151,267,1372,1398]" pageId="57" pageNumber="58">Material.</emphasis>
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, 
<collectionCode id="52E570C7FF8F5273FE4FFAB085E4FAC8" box="[400,491,1372,1398]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="57" pageNumber="58" type="Museum">ZMMU</collectionCode>
D-802, 
<collectionCode id="52E570C7FF8F5273FD92FAB0866FFAC8" box="[589,608,1372,1398]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="57" pageNumber="58">R</collectionCode>
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<collectionCode id="52E570C7FF8F5273FDB6FAB08671FAC8" box="[617,638,1372,1398]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="57" pageNumber="58" type="Museum">V</collectionCode>
“Vityaz”, cruise 45, sta. 6109, 
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<collectingDate id="500E372AFF8F5273FC2FFAB0809FFAC8" box="[1008,1168,1372,1398]" pageId="57" pageNumber="58" value="1969-05-15">15 May 1969</collectingDate>
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, 
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<geoCoordinate id="51C08EC5FF8F5273FAFCFAB084BCFA22" degrees="139" direction="west" minutes="44,0" orientation="longitude" pageId="57" pageNumber="58" precision="92" value="-139.73334">139° 44,0' W</geoCoordinate>
– 
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<geoCoordinate id="51C08EC5FF8F5273FE88FA6E85FCFA22" box="[343,499,1410,1436]" degrees="139" direction="west" minutes="43,4" orientation="longitude" pageId="57" pageNumber="58" precision="92" value="-139.72333">139° 43,4' W</geoCoordinate>
, depth 
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, grab dredge &quot;Ocean&quot;. 13 dried 
<typeStatus id="EB4F56A0FF8F5273FBB8FA6E80D5FA22" box="[1127,1242,1410,1436]" pageId="57" pageNumber="58" type="paratype">paratypes</typeStatus>
, 
<collectionCode id="52E570C7FF8F5273FB38FA6E814DFA22" box="[1255,1346,1410,1436]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="57" pageNumber="58" type="Museum">ZMMU</collectionCode>
D-803, same sta. as the 
<typeStatus id="EB4F56A0FF8F5273FE8BFA4B85B1FA7F" box="[340,446,1447,1473]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
. 9 dried 
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, 
<collectionCode id="52E570C7FF8F5273FD7FFA4B86F4FA7F" box="[672,763,1447,1473]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="57" pageNumber="58" type="Museum">ZMMU</collectionCode>
D-804, 
<collectionCode id="52E570C7FF8F5273FC82FA4B877FFA7F" box="[861,880,1447,1473]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="57" pageNumber="58">R</collectionCode>
/ 
<collectionCode id="52E570C7FF8F5273FCA6FA4B8782FA7F" box="[889,909,1447,1473]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="57" pageNumber="58" type="Museum">V</collectionCode>
“Vityaz, cruise 29, sta. 4213, 
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<geoCoordinate id="51C08EC5FF8F5273FABBFA4B84D1FA58" degrees="123" direction="west" minutes="55" orientation="longitude" pageId="57" pageNumber="58" precision="925" value="-123.916664">123° 55' W</geoCoordinate>
, depth 4231– 
<quantity id="F30C45E7FF8F5273FEA1FA2085D2FA58" box="[382,477,1484,1510]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="57" pageNumber="58" unit="m" value="4200.0">
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, Sigsbee trawl. 3 ethanol 
<typeStatus id="EB4F56A0FF8F5273FCCDFA208788FA58" box="[786,903,1484,1510]" pageId="57" pageNumber="58" type="paratype">paratypes</typeStatus>
, 
<collectionCode id="52E570C7FF8F5273FC4CFA2087E0FA58" box="[915,1007,1484,1510]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="57" pageNumber="58" type="Museum">ZMMU</collectionCode>
D-805, 
<collectionCode id="52E570C7FF8F5273FB8CFA208068FA58" box="[1107,1127,1484,1510]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="57" pageNumber="58">R</collectionCode>
/ 
<collectionCode id="52E570C7FF8F5273FBB0FA20808BFA58" box="[1135,1156,1484,1510]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="57" pageNumber="58" type="Museum">V</collectionCode>
“Vityaz, cruise 29, sta. 4213, 
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<geoCoordinate id="51C08EC5FF8F5273FE88FA1E85D3F9B2" box="[343,476,1522,1548]" degrees="123" direction="west" minutes="55" orientation="longitude" pageId="57" pageNumber="58" precision="925" value="-123.916664">123° 55' W</geoCoordinate>
, depth 4231– 
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, Sigsbee trawl.
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</paragraph>
</subSubSection>
<subSubSection id="7CEEBB89FF8F5273FF19F9FB86B7F9C2" pageId="57" pageNumber="58" type="etymology">
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<emphasis id="06803410FF8F5273FF19F9FB855CF98F" bold="true" box="[198,339,1559,1585]" pageId="57" pageNumber="58">Etymology.</emphasis>
This species is named in honour of ophiuroid taxonomist Nina Litvinova (
<collectingRegion id="F63026E0FF8F5273FB24F9FB816DF98F" box="[1275,1378,1559,1585]" country="Russia" name="Moscow" pageId="57" pageNumber="58">Moscow</collectingRegion>
, P.P. Shirshov Institute of Oceanology), whose efforts in describing the deep-water 
<taxonomicName id="F3F49381FF8F5273FB8EF9D080E8F9E8" box="[1105,1255,1596,1622]" class="Ophiuroidea" kingdom="Animalia" pageId="57" pageNumber="58" phylum="Echinodermata" rank="class">Ophiuroidea</taxonomicName>
has elucidated their importance for taxonomy and phylogeny.
</paragraph>
</subSubSection>
<subSubSection id="7CEEBB89FF8F5271FF18F96B8720FC7C" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58" type="description">
<paragraph id="344BE802FF8F5270FF18F96B85EAFDBC" blockId="57.[151,1437,1372,2033]" lastBlockId="58.[151,1436,152,2007]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<emphasis id="06803410FF8F5273FF18F96B863AF91F" bold="true" box="[199,565,1671,1697]" pageId="57" pageNumber="58">
Description of the 
<typeStatus id="EB4F56A0FF8F5273FE1CF96B863FF91F" box="[451,560,1671,1697]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
.
</emphasis>
The disk, 6.8 mm dd, is pentagonal. The disk plates are numerous, somewhat obscured by thin wrinkled skin, when dried. Radial shields are very small irregular plates, similar in size to disk scales, placed at the edge of the disk and scarcely visible (see a detailed radial shield description under 
<typeStatus id="EB4F56A0FF8F5273FF3DF91B8548F8AF" box="[226,327,1783,1809]" pageId="57" pageNumber="58" type="paratype">paratype</typeStatus>
variations). The whole dorsal side of the disk is covered with short cylindrical or subconical, sparsely placed spinelets. The interradii are slightly swollen, ventrally covered with few spinelets. Each jaw bears apically a cluster of 3–4 spiniform teeth or apical papillae. The 3–6 lateral papillae together with the adoral shield papillae are placed along both edges of each jaw in a characteristic, slightly curved row around the second tentacle pore. The adoral shield papillae are long, flattened, similar in size and shape to the arm spines, 
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number in most interradii, five in one interradius. The paired proximalmost oral papillae are placed laterally. Between lateral oral papillae and second tentacle pore the jaws form a rectangular border, which is superficially similar to the block-shaped oral papillae. Numerous spiniform teeth are distributed in a cone-shape, tapering dorsalwards. There are approximately 10 teeth (including apical papillae) on each dental plate. The oral shield is wide, triangular with straight or slightly convex, rather long distal edge, as long as wide, completely separated from the first lateral arm plate by the adoral shields. Distally the oral shield adjoins a supplementary narrow elongated plate or just an elevation formed by adjacent interradial scales. The madreporite has a conspicuous swollen round area occupying a considerable part of its surface. The adoral shield is wing-shaped laterally, widely adjoining the arm, rapidly tapering towards to the midline of the jaws, retaining thus only a narrow bar between jaws and oral shield. True genital slits are barely conspicuous, small openings under the adoral shields, but a long furrow between the most proximal arm segments and the ventral interradius forms a false slit.
</paragraph>
<paragraph id="344BE802FF8C5270FF18FDE185CFFB79" blockId="58.[151,1436,152,2007]" pageId="58" pageNumber="59">Arm length is about 1.7 times the disk diameter. On all arms the most proximal dorsal plates are absent for two-three segments, replaced by a covering of small disc scales. The dorsal arm plates are well developed but rather small, triangular, tapered and increasingly pentagonal towards the middle of the arm, well separated on all segments (for about half of the length of the dorsal arm plate proximally and length of the dorsal plate distally). All dorsal arm plates have pointed proximal and convex distal edges. Arms have weakly developed nodes, but arm spine articulations are placed on lateral plates without forming an elevation. Laterally between two adjacent lateral plates there is a narrow space of uncovered soft tissue. There are two spines on the first arm segment, three on proximal segments under the disk, afterwards four to five and then three distally. The first dorsal spine is slightly longer than the other ones proximally and medially. Spines are flattened, pointed apically. There are no hook-shaped spines. The first ventral arm plate is broadly triangular, the second has straight distal and proximal edges; further ventral arm plates have a characteristic pole-axe shape throughout most of the arm length, with a triangular proximal edge, whereas distally they vary from straight to slightly concave or convex with a small elevation. Ventral plates throughout the length of the arm are contiguous, except for the first two segments. The tentacle pores are large throughout the entire arm length. Basally on the parts of the lateral plate encircling the tentacle pores there are one, or rarely two larger, long, flattened, pointed tentacle scales. On most arm segments there is one tentacle scale. Distally the tentacle scale becomes small and spiniform. The tentacle scales are shorter than arm spines, but conspicuous and capable of covering the whole tentacle pore. Many of the tentacle scales are broken or have fallen off. There are no tentacle scales on the ventral arm plates.</paragraph>
<paragraph id="344BE802FF8C5270FF18FB3E8734F96C" blockId="58.[151,1436,152,2007]" pageId="58" pageNumber="59">
<emphasis id="06803410FF8C5270FF18FB3E85C7FB52" bold="true" box="[199,456,1234,1260]" pageId="58" pageNumber="59">
<typeStatus id="EB4F56A0FF8C5270FF18FB3E8535FB52" box="[199,314,1234,1260]" pageId="58" pageNumber="59" type="paratype">Paratype</typeStatus>
variations.
</emphasis>
A 
<typeStatus id="EB4F56A0FF8C5270FE31FB3E8658FB52" box="[494,599,1234,1260]" pageId="58" pageNumber="59" type="paratype">paratype</typeStatus>
, from the same station as the 
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, 7.0 mm dd, is similar with the 
<typeStatus id="EB4F56A0FF8C5270FF48FB14850CFAAC" box="[151,259,1272,1298]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
, but has a markedly pentagonal disk and some other minor differences. Numerous disk scales are clearly embedded within a thin skin. Apical papillae (teeth) are less numerous and thinner, up to 
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number. A single spiniform tooth in the dorsal part of the dental plate is markedly longer than (up to twice) the apical papillae. The distalmost oral papillae are block-shaped, but short. The following two oral papillae are short and spiniform. The first 1–2 proximal segments near the disk have no dorsal plates and are covered with small scales and skin. The madreporic oral shield is similar in shape to the others but with a swollen area in the middle. Genital slits are very short, placed just below adoral shields, a narrow adradial genital plate is evident in the distal part of the ventral side of the disk, but does not support the walls of the genital slits. The abradial genital plate is absent. Radial shields are very small and inconspicuous. On the 1–2 proximal lateral arm plates under the disk there are two flattened, apically pointed spines, on the proximal and middle lateral plates there are 4–5 spines. Although many tentacle scales are missing, three are present on the most proximal arm segments and some second and third segments. The ventralmost of these three scales is placed at the border of lateral and ventral arm plates, but still on the lateral plate.
</paragraph>
<paragraph id="344BE802FF8C5271FF18F93186EBFDF2" blockId="58.[151,1436,152,2007]" lastBlockId="59.[151,1436,152,1111]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
A small 
<typeStatus id="EB4F56A0FF8C5270FEF4F931859CF949" box="[299,403,1757,1783]" pageId="58" pageNumber="59" type="paratype">paratype</typeStatus>
, from the same station as the 
<typeStatus id="EB4F56A0FF8C5270FD30F9318757F949" box="[751,856,1757,1783]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
, 4.0 mm dd, is similar to the 
<typeStatus id="EB4F56A0FF8C5270FB6FF9318116F949" box="[1200,1305,1757,1783]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
, except for the pentagonal disk and some other differences. There are obvious primary plates joined in a flower-shaped structure. Few disk spines are visible. The radial shields are more evident than in the 
<typeStatus id="EB4F56A0FF8C5270FB69F8C4812FF8FC" box="[1206,1312,1832,1858]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
and other 
<typeStatus id="EB4F56A0FF8C5270FF48F8A18500F8D9" box="[151,271,1869,1895]" pageId="58" pageNumber="59" type="paratype">paratypes</typeStatus>
. Dorsal arm plates proximally are almost rhomboid with rounded angles. Medially and distally dorsal arm plates become almost ovoid. The distance between two adjacent dorsal arm plates proximally is about equal to the dorsal plate length, whereas distally it is up to twice the length of the dorsal plate. Apical papillae (teeth) are considerably fewer than in the 
<typeStatus id="EB4F56A0FF8C5270FD01F851874AF869" box="[734,837,1981,2007]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
and there is an obvious larger middle apical papilla and 1–3 smaller and thinner lateral apical papillae. A single spiniform tooth in the dorsal part of the dental plate is present only in some interradii and is similar in size to the middle apical papilla. The distalmost oral papillae are block-shaped relatively long elevations. The single pair of oral papillae are rectangular and pointed, somewhat similar in shape to the buccal scales of 
<taxonomicName id="F3F49381FF8D5271FCAAFEE48019FE9C" baseAuthorityName="Matsumoto" baseAuthorityYear="1917" box="[885,1046,264,290]" class="Ophiuroidea" family="Amphiuridae" kingdom="Animalia" order="Amphilepidida" pageId="59" pageNumber="60" phylum="Echinodermata" rank="family">Amphiuridae</taxonomicName>
and 
<taxonomicName id="F3F49381FF8D5271FB8AFEE480E1FE9C" baseAuthorityName="Martynov" baseAuthorityYear="2010" box="[1109,1262,264,290]" class="Ophiuroidea" family="Ophiactidae" kingdom="Animalia" order="Amphilepidida" pageId="59" pageNumber="60" phylum="Echinodermata" rank="family">Ophiactidae</taxonomicName>
. The first 1–2 proximal segments near the disk lack dorsal plates and are covered with small scales or almost naked skin exposing the vertebrae. The madreporic oral shield does not differ from the remaining oral shields. Genital slits in most interradii are longer than in the 
<typeStatus id="EB4F56A0FF8D5271FD6DFE948711FE2C" box="[690,798,376,402]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
, whereas in one interradius they are small. A narrow adradial genital plate is obvious in the distal part of the ventral side of the disk, but does not support the walls of the genital slits. The abradial genital plate is absent. On the 1–2 proximal lateral arm plates under the disk there are 2–3 flattened, apically pointed spines, on proximal and middle lateral plates four, and distally three spines. Proximally there are 1–2 large, flattened tentacle scales, which become smaller medially and disappear entirely distally. There are no hook-shaped spines.
</paragraph>
<paragraph id="344BE802FF8D5271FF18FDBB8720FC7C" blockId="59.[151,1436,152,1111]" pageId="59" pageNumber="60">
<emphasis id="06803410FF8D5271FF18FDBB86C9FDCF" bold="true" box="[199,710,599,625]" pageId="59" pageNumber="60">Internal and microstructural characters.</emphasis>
The radial shield is a very small irregular plate, with slightly elevated articulation surface (
<figureCitation id="ACCFF487FF8D5271FE2DFD918642FD29" box="[498,589,637,663]" captionStart="FIGURE 4" captionStartId="13.[151,256,1339,1363]" captionTargetBox="[282,1312,194,1330]" captionTargetId="figure-434@13.[282,1312,194,1330]" captionTargetPageId="13" captionText="FIGURE 4. Genital plates and radial shields of various taxa of the families Asteronychidae, Gorgonocephalidae and Ophiomyxidae. A. Gorgonocephalus arcticus Leach, 1819, ZMMU D-613, 39 mm dd, adradial and abradial genital plates in situ, showing articulation surface, dorsal view; B. same, articulation surface of the radial shield, ventral view; C. Asteronyx loveni Müller &amp; Troschel, 1842, ZMMU D-833, 17.5 mm dd, articulation surface of the adradial genital plate, dorsal view; D. same, articulation surface of the radial shield, ventral view. E. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, articulation surface of the adradial genital plate, dorsal view; F. same, radial shield showing articulation surface, ventral view; G. same, articulating adradial genital plate and radial shield, dorsal view; H. Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, dissected disk showing most proximal vertebrae and absence of the both genital plates and radial shield, dorsal view; I. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, adradial and abradial genital plates in situ, ventral view; J. Ophioplexa condita gen. et sp. nov., paratype ca. 17 mm dd, ZMMU D-792, dissected disk showing articulating adradial genital plate and radial shield in situ, dorsal view; K. same, separate articulating adradial genital plate and radial shield, dorsal view; L. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, adradial genital plate, dorsal view; M. Ophiologimus cf. secundus Koehler, 1914, 4 mm dd, ZMMU D-814, adradial genital plate, dorsal view; N. Ophioplexa condita gen. et sp. nov., separate adradial genital plate, dorsal view; O. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, adradial and abradial genital plates, dorsal view; P. Ophioplexa condita gen. et sp. nov., separate articulating adradial genital plate and radial shield, dorsal view; Q. articulation surface of the adradial genital plate, dorsal view; R. radial shield, ventral view, showing distally the distinctive excavated articulation surface; S. articulation surface of the radial shield, ventral view; T. Ophiocymbium ninae sp. nov., paratype 5 mm dd, radial shield, ventral view; U. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, same, articulation surface of the radial shield, proximal view; Scale bars: A– B,1 mm; C–D, 300 µ; E–F, 100 µ; G, 200 µ; H, 300 µ; I, 200 µ; J, 1000 µ; K, 300 µ; L, 100 µ; M, 300 µ; N, 300 µ; O, 200 µ; P, 100 µ; Q–T, 100 µ; U, 200 µ. Photos: Alexander Martynov." pageId="59" pageNumber="60">Fig. 4T</figureCitation>
). The articulation surface of the abradial genital plate does not have any distinct condyles (
<figureCitation id="ACCFF487FF8D5271FEABFD4E85DCFD02" box="[372,467,674,700]" captionStart="FIGURE 4" captionStartId="13.[151,256,1339,1363]" captionTargetBox="[282,1312,194,1330]" captionTargetId="figure-434@13.[282,1312,194,1330]" captionTargetPageId="13" captionText="FIGURE 4. Genital plates and radial shields of various taxa of the families Asteronychidae, Gorgonocephalidae and Ophiomyxidae. A. Gorgonocephalus arcticus Leach, 1819, ZMMU D-613, 39 mm dd, adradial and abradial genital plates in situ, showing articulation surface, dorsal view; B. same, articulation surface of the radial shield, ventral view; C. Asteronyx loveni Müller &amp; Troschel, 1842, ZMMU D-833, 17.5 mm dd, articulation surface of the adradial genital plate, dorsal view; D. same, articulation surface of the radial shield, ventral view. E. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, articulation surface of the adradial genital plate, dorsal view; F. same, radial shield showing articulation surface, ventral view; G. same, articulating adradial genital plate and radial shield, dorsal view; H. Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, dissected disk showing most proximal vertebrae and absence of the both genital plates and radial shield, dorsal view; I. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, adradial and abradial genital plates in situ, ventral view; J. Ophioplexa condita gen. et sp. nov., paratype ca. 17 mm dd, ZMMU D-792, dissected disk showing articulating adradial genital plate and radial shield in situ, dorsal view; K. same, separate articulating adradial genital plate and radial shield, dorsal view; L. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, adradial genital plate, dorsal view; M. Ophiologimus cf. secundus Koehler, 1914, 4 mm dd, ZMMU D-814, adradial genital plate, dorsal view; N. Ophioplexa condita gen. et sp. nov., separate adradial genital plate, dorsal view; O. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, adradial and abradial genital plates, dorsal view; P. Ophioplexa condita gen. et sp. nov., separate articulating adradial genital plate and radial shield, dorsal view; Q. articulation surface of the adradial genital plate, dorsal view; R. radial shield, ventral view, showing distally the distinctive excavated articulation surface; S. articulation surface of the radial shield, ventral view; T. Ophiocymbium ninae sp. nov., paratype 5 mm dd, radial shield, ventral view; U. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, same, articulation surface of the radial shield, proximal view; Scale bars: A– B,1 mm; C–D, 300 µ; E–F, 100 µ; G, 200 µ; H, 300 µ; I, 200 µ; J, 1000 µ; K, 300 µ; L, 100 µ; M, 300 µ; N, 300 µ; O, 200 µ; P, 100 µ; Q–T, 100 µ; U, 200 µ. Photos: Alexander Martynov." pageId="59" pageNumber="60">Fig. 4L</figureCitation>
). The abradial genital plate is absent. Jaws are slightly elongated (
<figureCitation id="ACCFF487FF8D5271FB28FD4E8181FD02" box="[1271,1422,674,700]" captionStart="FIGURE 12" captionStartId="21.[151,255,1049,1073]" captionTargetBox="[158,1428,194,1027]" captionTargetId="figure-465@21.[158,1428,194,1027]" captionTargetPageId="21" captionText="FIGURE 12. Dental plates and jaws of the genera Ophioscolex, Ophiocymbium and Ophiologimus. A–B, Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, dental plates; C–D, Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, abradial and adradial views of the jaws respectively; E. same, dental plate; F. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-803, dental plate; G. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-805, dental plate; H. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, dental plate; I–J, Ophiologimus cf. secundus Koehler, 1914, ZMMU D-815, 4 mm dd, dental plates; K–L, Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-805, abradial and adradial views of the jaws respectively; M. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, jaw, adradial view; N–O, Ophiologimus cf. secundus Koehler, 1914, ZMMU D-815, 4 mm dd, abradial and adradial views of the jaws respectively; Scale bars: A, 200 µ; B, 200 µ; C–D, 300 µ; E, 100 µ; F, 300 µ; G, 100 µ; H, 300 µ; I–J, 100 µ; K–L, 300 µ; M, 100 µ; N–O, 300 µ. Photos: Alexander Martynov." pageId="59" pageNumber="60">Figs 12K, L</figureCitation>
). Adradial sides of the jaws distally bear few indistinct straight folds and irregularly-shaped tubercles (
<figureCitation id="ACCFF487FF8D5271FAB4FD2484C1FCB9" captionStart="FIGURE 12" captionStartId="21.[151,255,1049,1073]" captionTargetBox="[158,1428,194,1027]" captionTargetId="figure-465@21.[158,1428,194,1027]" captionTargetPageId="21" captionText="FIGURE 12. Dental plates and jaws of the genera Ophioscolex, Ophiocymbium and Ophiologimus. A–B, Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, dental plates; C–D, Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, abradial and adradial views of the jaws respectively; E. same, dental plate; F. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-803, dental plate; G. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-805, dental plate; H. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, dental plate; I–J, Ophiologimus cf. secundus Koehler, 1914, ZMMU D-815, 4 mm dd, dental plates; K–L, Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-805, abradial and adradial views of the jaws respectively; M. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, jaw, adradial view; N–O, Ophiologimus cf. secundus Koehler, 1914, ZMMU D-815, 4 mm dd, abradial and adradial views of the jaws respectively; Scale bars: A, 200 µ; B, 200 µ; C–D, 300 µ; E, 100 µ; F, 300 µ; G, 100 µ; H, 300 µ; I–J, 100 µ; K–L, 300 µ; M, 100 µ; N–O, 300 µ. Photos: Alexander Martynov." pageId="59" pageNumber="60">Fig. 12L</figureCitation>
). The dental plate is devoid of folds, with few small rounded sockets (
<figureCitation id="ACCFF487FF8D5271FBBEFD0180F1FCB9" box="[1121,1278,749,775]" captionStart="FIGURE 12" captionStartId="21.[151,255,1049,1073]" captionTargetBox="[158,1428,194,1027]" captionTargetId="figure-465@21.[158,1428,194,1027]" captionTargetPageId="21" captionText="FIGURE 12. Dental plates and jaws of the genera Ophioscolex, Ophiocymbium and Ophiologimus. A–B, Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, dental plates; C–D, Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, abradial and adradial views of the jaws respectively; E. same, dental plate; F. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-803, dental plate; G. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-805, dental plate; H. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, dental plate; I–J, Ophiologimus cf. secundus Koehler, 1914, ZMMU D-815, 4 mm dd, dental plates; K–L, Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-805, abradial and adradial views of the jaws respectively; M. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, jaw, adradial view; N–O, Ophiologimus cf. secundus Koehler, 1914, ZMMU D-815, 4 mm dd, abradial and adradial views of the jaws respectively; Scale bars: A, 200 µ; B, 200 µ; C–D, 300 µ; E, 100 µ; F, 300 µ; G, 100 µ; H, 300 µ; I–J, 100 µ; K–L, 300 µ; M, 100 µ; N–O, 300 µ. Photos: Alexander Martynov." pageId="59" pageNumber="60">Figs 12F, G</figureCitation>
). Arm spine articulations are well defined and belong to the double-opening 
<typeStatus id="EB4F56A0FF8D5271FC49FCFE87C6FC92" box="[918,969,786,812]" pageId="59" pageNumber="60">type</typeStatus>
(
<figureCitation id="ACCFF487FF8D5271FC05FCFE8077FC92" box="[986,1144,786,812]" captionStart="FIGURE 55" captionStartId="80.[151,258,1764,1788]" captionTargetBox="[209,1364,194,1740]" captionTargetId="figure-174@80.[209,1364,194,1740]" captionTargetPageId="80" captionText="FIGURE 55. Ophioplexa condita gen. nov., sp. nov. and Ophiocymbium ninae sp. nov., details. A. Ophioplexa condita gen. nov., sp. nov., paratype 19 mm dd, ZMMU D-791, disk ventral view; B. same oral frame showing clusters of the ventral teeth (apical papillae); C. same, one interradius showing apical, oral and adoral shield papillae; D-F, same, most proximal arm segments showing tentacle scales and spines; G-H, arm spine articulations, distal segments; I. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-803, details, part of the oral frame and ventral arm side; J. same, jaw and oral papillae, one interradius; K. ventral arm view showing spines and tentacle scales; L. most proximal segments showing details of tentacle scales and spines; M-N, arm spine articulations, proximal segments; Scale bars: A– B, 1 mm; C–D, 500 µ; E–F, 200 µ; G–H, 100 µ; I, 1 mm; J–N, 300 µ. Photos: Alexander Martynov." pageId="59" pageNumber="60">Figs 55M, N</figureCitation>
). The muscle opening is placed on a distinct round or elongated elevation, some articulations are slightly ribbed distally. Nerve opening placed laterally below the level of the muscle opening. Vertebrae rather short, not keeled and with well developed zygospondylous articulation (
<figureCitation id="ACCFF487FF8D5271FD62FC6E8721FC22" box="[701,814,898,924]" captionStart="FIGURE 41" captionStartId="53.[151,255,1505,1529]" captionTargetBox="[158,1428,194,1484]" captionTargetId="figure-189@53.[158,1428,194,1484]" captionTargetPageId="53" captionText="FIGURE 41. External views and vertebrae of Ophiocymbium ninae sp. nov. A. paratype, ZMMU D-804, 5.5 mm dd, dorsal view; B. holotype 6.8 mm dd, ZMMU D-802, 6.8 mm dd, ventral view; C. paratype 6.5 mm dd, ZMMU D-803, ventral view; D. paratype, ZMMU D-803, 6.1 mm dd, ventral view; E. holotype, dorsal view, ZMMU D-802, 6.8 mm dd, F. vertebra, ventral view; G. vertebra, dorsal view; Scale bars: F,G, 300 µ. Photos: A–F Tatiana Korshunova; F–G Alexander Martynov." pageId="59" pageNumber="60">Figs 41G</figureCitation>
, 
<figureCitation id="ACCFF487FF8D5271FCE4FC6E8786FC22" box="[827,905,898,924]" captionStart="FIGURE 17" captionStartId="27.[151,255,1036,1060]" captionTargetBox="[161,1426,186,1015]" captionTargetId="figure-399@27.[161,1426,186,1015]" captionTargetPageId="27" captionText="FIGURE 17. Vertebrae of various taxa of the family Ophiomyxidae. A. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, proximal segments, dorsal and ventral views; B. same, distal segments, dorsal and ventral views; C. same, proximal segments, proximal view; D. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, 10.5 mm dd, proximal segments, dorsal view; E. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, proximal segments, dorsal view; F. Ophioscolex glacialis, proximal segments, distal view; G. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, proximal segments, proximal view; H. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, ventral view; I. same, proximal view; J. same, distal view; K. Ophiomyxa pentagona, ZMMU D-553, 21 mm dd, distal view; L. Ophioplexa condita gen. et sp. nov., holotype ca. 18 mm dd, ZMMU D-788, proximal segments, dorsal view; M. same, ventral view; N. same, proximal view; O. same, distal view; P. Ophiocymbium tanyae sp. nov., holotype 9.5 mm dd, ZMMU D-799, proximal segments, dorsal view; Q. same, ventral view; R. same, distal view; S. same, proximal view. Scale bars: A, 1000 µ; B, C, 200 µ, D, 200 µ; E, 300 µ; F, 200 µ; G, 200 µ; H, 300 µ; I, 100 µ; J, 100 µ; K, 1000 µ; L–S, 300 µ. Photos: Alexander Martynov." pageId="59" pageNumber="60">17E, J</figureCitation>
). Vertebral dorsal median groove is distinct (
<figureCitation id="ACCFF487FF8D5271FF40FC448503FC7C" box="[159,268,936,962]" captionStart="FIGURE 41" captionStartId="53.[151,255,1505,1529]" captionTargetBox="[158,1428,194,1484]" captionTargetId="figure-189@53.[158,1428,194,1484]" captionTargetPageId="53" captionText="FIGURE 41. External views and vertebrae of Ophiocymbium ninae sp. nov. A. paratype, ZMMU D-804, 5.5 mm dd, dorsal view; B. holotype 6.8 mm dd, ZMMU D-802, 6.8 mm dd, ventral view; C. paratype 6.5 mm dd, ZMMU D-803, ventral view; D. paratype, ZMMU D-803, 6.1 mm dd, ventral view; E. holotype, dorsal view, ZMMU D-802, 6.8 mm dd, F. vertebra, ventral view; G. vertebra, dorsal view; Scale bars: F,G, 300 µ. Photos: A–F Tatiana Korshunova; F–G Alexander Martynov." pageId="59" pageNumber="60">Figs 41G</figureCitation>
, 
<figureCitation id="ACCFF487FF8D5271FEC7FC448544FC7C" box="[280,331,936,962]" captionStart="FIGURE 17" captionStartId="27.[151,255,1036,1060]" captionTargetBox="[161,1426,186,1015]" captionTargetId="figure-399@27.[161,1426,186,1015]" captionTargetPageId="27" captionText="FIGURE 17. Vertebrae of various taxa of the family Ophiomyxidae. A. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, proximal segments, dorsal and ventral views; B. same, distal segments, dorsal and ventral views; C. same, proximal segments, proximal view; D. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, 10.5 mm dd, proximal segments, dorsal view; E. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, proximal segments, dorsal view; F. Ophioscolex glacialis, proximal segments, distal view; G. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, proximal segments, proximal view; H. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, ventral view; I. same, proximal view; J. same, distal view; K. Ophiomyxa pentagona, ZMMU D-553, 21 mm dd, distal view; L. Ophioplexa condita gen. et sp. nov., holotype ca. 18 mm dd, ZMMU D-788, proximal segments, dorsal view; M. same, ventral view; N. same, proximal view; O. same, distal view; P. Ophiocymbium tanyae sp. nov., holotype 9.5 mm dd, ZMMU D-799, proximal segments, dorsal view; Q. same, ventral view; R. same, distal view; S. same, proximal view. Scale bars: A, 1000 µ; B, C, 200 µ, D, 200 µ; E, 300 µ; F, 200 µ; G, 200 µ; H, 300 µ; I, 100 µ; J, 100 µ; K, 1000 µ; L–S, 300 µ. Photos: Alexander Martynov." pageId="59" pageNumber="60">17E</figureCitation>
). Podial basins are large (
<figureCitation id="ACCFF487FF8D5271FDA8FC4486EFFC7C" box="[631,736,936,962]" captionStart="FIGURE 41" captionStartId="53.[151,255,1505,1529]" captionTargetBox="[158,1428,194,1484]" captionTargetId="figure-189@53.[158,1428,194,1484]" captionTargetPageId="53" captionText="FIGURE 41. External views and vertebrae of Ophiocymbium ninae sp. nov. A. paratype, ZMMU D-804, 5.5 mm dd, dorsal view; B. holotype 6.8 mm dd, ZMMU D-802, 6.8 mm dd, ventral view; C. paratype 6.5 mm dd, ZMMU D-803, ventral view; D. paratype, ZMMU D-803, 6.1 mm dd, ventral view; E. holotype, dorsal view, ZMMU D-802, 6.8 mm dd, F. vertebra, ventral view; G. vertebra, dorsal view; Scale bars: F,G, 300 µ. Photos: A–F Tatiana Korshunova; F–G Alexander Martynov." pageId="59" pageNumber="60">Figs 41F</figureCitation>
, 
<figureCitation id="ACCFF487FF8D5271FD33FC44872DFC7C" box="[748,802,936,962]" captionStart="FIGURE 17" captionStartId="27.[151,255,1036,1060]" captionTargetBox="[161,1426,186,1015]" captionTargetId="figure-399@27.[161,1426,186,1015]" captionTargetPageId="27" captionText="FIGURE 17. Vertebrae of various taxa of the family Ophiomyxidae. A. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, proximal segments, dorsal and ventral views; B. same, distal segments, dorsal and ventral views; C. same, proximal segments, proximal view; D. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, 10.5 mm dd, proximal segments, dorsal view; E. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, proximal segments, dorsal view; F. Ophioscolex glacialis, proximal segments, distal view; G. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, proximal segments, proximal view; H. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, ventral view; I. same, proximal view; J. same, distal view; K. Ophiomyxa pentagona, ZMMU D-553, 21 mm dd, distal view; L. Ophioplexa condita gen. et sp. nov., holotype ca. 18 mm dd, ZMMU D-788, proximal segments, dorsal view; M. same, ventral view; N. same, proximal view; O. same, distal view; P. Ophiocymbium tanyae sp. nov., holotype 9.5 mm dd, ZMMU D-799, proximal segments, dorsal view; Q. same, ventral view; R. same, distal view; S. same, proximal view. Scale bars: A, 1000 µ; B, C, 200 µ, D, 200 µ; E, 300 µ; F, 200 µ; G, 200 µ; H, 300 µ; I, 100 µ; J, 100 µ; K, 1000 µ; L–S, 300 µ. Photos: Alexander Martynov." pageId="59" pageNumber="60">17H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7CEEBB89FF8D5271FF18FC218122FBE9" pageId="59" pageNumber="60" type="discussion">
<paragraph id="344BE802FF8D5271FF18FC218122FBE9" blockId="59.[151,1436,152,1111]" pageId="59" pageNumber="60">
<emphasis id="06803410FF8D5271FF18FC21854FFC59" bold="true" box="[199,320,973,999]" pageId="59" pageNumber="60">Remarks.</emphasis>
<taxonomicName id="F3F49381FF8D5271FE96FC218645FC59" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[329,586,973,999]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiocymbium" kingdom="Animalia" order="Ophioscolecida" pageId="59" pageNumber="141" phylum="Echinodermata" rank="species" species="ninae" status="sp. nov.">
<emphasis id="06803410FF8D5271FE96FC218645FC59" box="[329,586,973,999]" italics="true" pageId="59" pageNumber="60">
Ophiocymbium 
<emphasis id="06803410FF8D5271FDDAFC218645FC59" bold="true" box="[517,586,973,999]" pageId="59" pageNumber="60">ninae</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1DB3896BFF8D5271FD8DFC2186BEFC59" box="[594,689,973,999]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
is clearly distinguished from both 
<taxonomicName id="F3F49381FF8D5271FB8EFC218193FC58" authorityName="Lyman" authorityYear="1880" box="[1105,1436,973,998]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiocymbium" kingdom="Animalia" order="Ophioscolecida" pageId="59" pageNumber="60" phylum="Echinodermata" rank="species" species="cavernosum">
<emphasis id="06803410FF8D5271FB8EFC218193FC58" box="[1105,1436,973,998]" italics="true" pageId="59" pageNumber="60">Ophiocymbium cavernosum</emphasis>
</taxonomicName>
and 
<taxonomicName id="F3F49381FF8D5271FF15FC1E85D5FBB2" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[202,474,1010,1036]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiocymbium" kingdom="Animalia" order="Ophioscolecida" pageId="59" pageNumber="141" phylum="Echinodermata" rank="species" species="tanyae" status="sp. nov.">
<emphasis id="06803410FF8D5271FF15FC1E85D5FBB2" box="[202,474,1010,1036]" italics="true" pageId="59" pageNumber="60">
Ophiocymbium 
<emphasis id="06803410FF8D5271FE56FC1E85D5FBB2" bold="true" box="[393,474,1010,1036]" pageId="59" pageNumber="60">tanyae</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1DB3896BFF8D5271FE3CFC1E864CFBB2" box="[483,579,1010,1036]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
by the presence of conspicuous short disk spinelets, less evident distal block-shaped oral papillae, and the shape of the arm spine articulations and vertebrae. 
<taxonomicName id="F3F49381FF8D5271FB4DFBF48194FB8F" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[1170,1435,1047,1073]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiocymbium" kingdom="Animalia" order="Ophioscolecida" pageId="59" pageNumber="141" phylum="Echinodermata" rank="species" species="tanyae" status="sp. nov.">
<emphasis id="06803410FF8D5271FB4DFBF48194FB8F" box="[1170,1435,1047,1073]" italics="true" pageId="59" pageNumber="60">
Ophiocymbium 
<emphasis id="06803410FF8D5271FA93FBFB8194FB8F" bold="true" box="[1356,1435,1047,1073]" pageId="59" pageNumber="60">tanyae</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1DB3896BFF8D5271FF48FBD184FBFBE9" box="[151,244,1085,1111]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
also differs from 
<taxonomicName id="F3F49381FF8D5271FE19FBD186CBFBE9" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[454,708,1085,1111]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiocymbium" kingdom="Animalia" order="Ophioscolecida" pageId="59" pageNumber="141" phylum="Echinodermata" rank="species" species="ninae" status="sp. nov.">
<emphasis id="06803410FF8D5271FE19FBD186CBFBE9" box="[454,708,1085,1111]" italics="true" pageId="59" pageNumber="60">
Ophiocymbium 
<emphasis id="06803410FF8D5271FD5FFBD186CBFBE9" bold="true" box="[640,708,1085,1111]" pageId="59" pageNumber="60">ninae</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1DB3896BFF8D5271FD13FBD18726FBE9" box="[716,809,1085,1111]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
by the absence of the adradial genital plate.
</paragraph>
</subSubSection>
</treatment>
</document>