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<document id="E5A3A8CB89343BC97EAD1F8FB1AAF1A3" ID-CLB-Dataset="55464" ID-DOI="10.5852/ejt.2022.819.1787" ID-GBIF-Dataset="0f6a0aa7-3aaa-4d95-a674-2327ec91945a" ID-ISSN="2118-9773" ID-Zenodo-Dep="6564531" ID-ZooBank="80DE5F71-8A1C-472F-9205-F956639D0605" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652984580463" checkinUser="felipe" docAuthor="Johanson, Kjell Arne, Malm, Tobias &amp; Sjöberg, Tin" docDate="2022" docId="A703A9650D32A406FD55FAC93C16024D" docLanguage="en" docName="ejt-819_158-165.pdf" docOrigin="European Journal of Taxonomy 819" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Oecetis apelqvisti Johanson &amp; Malm &amp; Sjöberg 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="163" masterDocId="5B3AD11D0D31A403FFD5FFC13F7B0614" masterDocTitle="Description of two new species of Oecetis (Trichoptera, Leptoceridae) from Borneo" masterLastPageNumber="165" masterPageNumber="158" pageNumber="161" updateTime="1699337803888" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6CF4FEA51CBD8E2752079E19134E70DF">Description of two new species of Oecetis (Trichoptera, Leptoceridae) from Borneo</mods:title>
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<mods:namePart id="A8BF422B435BA0EB33B2D7E796287E41">Johanson, Kjell Arne</mods:namePart>
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<mods:affiliation id="00CA485C96FA77D105C22ABF427F77C2">Department of Zoology, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden.</mods:affiliation>
<mods:nameIdentifier id="F8D265841FD763C693B03BB7765AB779" type="email">kjell.arne.johanson@nrm.se</mods:nameIdentifier>
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<mods:namePart id="FCE6840A91CC4C931EEFF7F8C3C870E5">Malm, Tobias</mods:namePart>
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<mods:affiliation id="020469C6E303BA135F37CDBE3A17E290">Department of Zoology, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden.</mods:affiliation>
<mods:nameIdentifier id="FEECD3B1E67C1D6B6CC828EE04814BC3" type="email">tobias.malm@nrm.se</mods:nameIdentifier>
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<mods:namePart id="658502E7B091EDD1BA04757B32400063">Sjöberg, Tin</mods:namePart>
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<mods:affiliation id="A0D2E09220A11A54042C7D2B6340AA23">Department of Zoology, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden.</mods:affiliation>
<mods:nameIdentifier id="F96A6E57D6EA73D3E48589B637AA6083" type="email">tin.sjoberg@nrm.se</mods:nameIdentifier>
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<mods:title id="0A849F012F0001DABEC03C106CB03570">European Journal of Taxonomy</mods:title>
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<mods:part id="2EC3CDE050A619E1E53546A06D29F204">
<mods:date id="DD0B53995481B3631EFAA5B45916DDC4">2022</mods:date>
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<treatment id="A703A9650D32A406FD55FAC93C16024D" ID-DOI="http://doi.org/10.5281/zenodo.6564542" ID-GBIF-Taxon="195537921" ID-Zenodo-Dep="6564542" LSID="urn:lsid:plazi:treatment:A703A9650D32A406FD55FAC93C16024D" httpUri="http://treatment.plazi.org/id/A703A9650D32A406FD55FAC93C16024D" lastPageId="5" lastPageNumber="163" pageId="3" pageNumber="161">
<subSubSection id="67B04BF80D32A400FD55FAC93CCF0337" box="[640,948,1288,1315]" pageId="3" pageNumber="161" type="nomenclature">
<paragraph id="2F1518730D32A400FD55FAC93CCF0337" blockId="3.[357,1230,1288,1384]" box="[640,948,1288,1315]" pageId="3" pageNumber="161">
<heading id="745DAF1F0D32A400FD55FAC93CCF0337" box="[640,948,1288,1315]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="161" reason="2">
<taxonomicName id="E8AA63F00D32A400FD55FAC93C290337" authority="Johanson &amp; Malm &amp; Sjöberg, 2022" authorityName="Johanson &amp; Malm &amp; Sjöberg" authorityYear="2022" box="[640,850,1288,1315]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="161" phylum="Arthropoda" rank="species" species="apelqvisti" status="sp. nov.">
<emphasis id="1DDEC4610D32A400FD55FAC93C290337" bold="true" box="[640,850,1288,1315]" italics="true" pageId="3" pageNumber="161">Oecetis apelqvisti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06ED791A0D32A400FC8CFAC83CCF0337" box="[857,948,1289,1315]" pageId="3" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="67B04BF80D32A400FEB0FAEA3C28037C" pageId="3" pageNumber="161" type="description">
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<uri id="5B3B14710D32A400FEB0FAEA3BB50352" box="[357,1230,1323,1350]" pageId="3" pageNumber="161">
urn:lsid:zoobank.org:act:
<uuid id="5B0C22A60D32A400FD54FAEA3BB50352" box="[641,1230,1323,1350]" pageId="3" pageNumber="161">0463DEDE-CF3B-43BA-9B3D-AF23699C165C</uuid>
</uri>
</paragraph>
<paragraph id="2F1518730D32A400FD35FA8F3C28037C" blockId="3.[357,1230,1288,1384]" box="[736,851,1358,1384]" pageId="3" pageNumber="161">
<figureCitation id="B79104F60D32A400FD35FA8F3C28037C" box="[736,851,1358,1384]" captionStart-0="Figs 610" captionStart-1="Fig" captionStartId-0="4.[189,242,1925,1951]" captionStartId-1="5.[189,232,1954,1980]" captionTargetBox-0="[248,1332,280,1880]" captionTargetBox-1="[189,1398,1220,1916]" captionTargetId-0="figure-12@4.[222,1366,265,1890]" captionTargetId-1="figure-295@5.[189,1398,1220,1916]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figs 610. Oecetis apelqvisti sp. nov., holotype, ♂ (NHRS). 6. Genitalia, lateral view. 7. Genitalia, dorsal view. 8. Genitalia, ventral view. 9. Phallus, lateral view. 10. Phallus, ventral view. Scale bar = 0.1 mm." captionText-1="Fig. 11. Map with the position of the two sampling sites in Maliau Basin shown as orange dots. The position of the Maliau Basin area on Borneo is indicated with a blue rectangular in the upper-left map." figureDoi-0="http://doi.org/10.5281/zenodo.6564537" figureDoi-1="http://doi.org/10.5281/zenodo.6564540" httpUri-0="https://zenodo.org/record/6564537/files/figure.png" httpUri-1="https://zenodo.org/record/6564540/files/figure.png" pageId="3" pageNumber="161">Figs 611</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="67B04BF80D32A400FF68FA513C41009D" pageId="3" pageNumber="161" type="diagnosis">
<paragraph id="2F1518730D32A400FF68FA513E4D03BE" blockId="3.[189,310,1424,1450]" box="[189,310,1424,1450]" pageId="3" pageNumber="161">
<heading id="745DAF1F0D32A400FF68FA513E4D03BE" bold="true" box="[189,310,1424,1450]" fontSize="11" level="3" pageId="3" pageNumber="161" reason="3">
<emphasis id="1DDEC4610D32A400FF68FA513E4D03BE" bold="true" box="[189,310,1424,1450]" pageId="3" pageNumber="161">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="2F1518730D32A400FF68FA7F3C41009D" blockId="3.[188,1400,1470,1673]" pageId="3" pageNumber="161">
The species is distinguished from many of the Oriental and Palearctic species by the absence of modified tergites on segment VIVIII. It resembles
<taxonomicName id="E8AA63F00D32A400FD60FA233CC403E8" authority="Malicky, 2005" authorityName="Malicky" authorityYear="2005" box="[693,959,1505,1532]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="161" phylum="Arthropoda" rank="species" species="idas">
<emphasis id="1DDEC4610D32A400FD60FA233C7703E8" box="[693,780,1506,1532]" italics="true" pageId="3" pageNumber="161">O. idas</emphasis>
<bibRefCitation id="4B3B65820D32A400FCC1FA233CC403E8" author="Malicky H." box="[788,959,1505,1532]" pageId="3" pageNumber="161" pagination="605 - 669" refId="ref2697" refString="Malicky H. 2005. Beitrage zur Kenntnis asiatischer Oecetis (Trichoptera, Leptoceridae). Linzer biologische Beitrage 27: 605 - 669." type="journal article" year="2005">
Malicky,
<quantity id="E852B5960D32A400FC51FA203CC403E8" box="[900,959,1505,1532]" metricMagnitude="1" metricUnit="m" metricValue="5.0927" pageId="3" pageNumber="161" unit="in" value="2005.0">2005</quantity>
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in genitalic features, particularly the long and slightly curving superior appendages, long and strongly ventrally curving tergum X, with right branch longer than left branch, and long pair of dorsal processes.
<taxonomicName id="E8AA63F00D32A400FC35F9E93BCE0056" authority="Johanson &amp; Malm &amp; Sjöberg, 2022" authorityName="Johanson &amp; Malm &amp; Sjöberg" authorityYear="2022" box="[992,1205,1576,1602]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="161" phylum="Arthropoda" rank="species" species="apelqvisti" status="sp. nov.">
<emphasis id="1DDEC4610D32A400FC35F9E93BCE0056" box="[992,1205,1576,1602]" italics="true" pageId="3" pageNumber="161">Oecetis apelqvisti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06ED791A0D32A400FB15F9E83A660057" box="[1216,1309,1577,1603]" pageId="3" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="E8AA63F00D32A400FE75F98D3E8F0071" authorityName="Malicky" authorityYear="2005" box="[416,500,1611,1638]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="161" phylum="Arthropoda" rank="species" species="idas">
<emphasis id="1DDEC4610D32A400FE75F98D3E8F0071" box="[416,500,1611,1638]" italics="true" pageId="3" pageNumber="161">O. idas</emphasis>
</taxonomicName>
in lateral view by the clearly bifid inferior appendages, and the apical part of phallus with a single hook-shaped sclerotized process.
</paragraph>
</subSubSection>
<subSubSection id="67B04BF80D32A400FF68F9713BAC00ED" box="[189,1239,1712,1785]" pageId="3" pageNumber="161" type="etymology">
<paragraph id="2F1518730D32A400FF68F9713E3F00DE" blockId="3.[189,324,1712,1738]" box="[189,324,1712,1738]" pageId="3" pageNumber="161">
<heading id="745DAF1F0D32A400FF68F9713E3F00DE" bold="true" box="[189,324,1712,1738]" fontSize="11" level="3" pageId="3" pageNumber="161" reason="3">
<emphasis id="1DDEC4610D32A400FF68F9713E3F00DE" bold="true" box="[189,324,1712,1738]" pageId="3" pageNumber="161">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="2F1518730D32A400FF68F91F3BAC00ED" blockId="3.[189,1239,1758,1785]" box="[189,1239,1758,1785]" pageId="3" pageNumber="161">
<emphasis id="1DDEC4610D32A400FF68F91F3E4900EC" box="[189,306,1758,1784]" italics="true" pageId="3" pageNumber="161">Apelqvisti</emphasis>
, named after one of the collectors, Niklas Apelqvist (formerly Niklas Jönsson).
</paragraph>
</subSubSection>
<subSubSection id="67B04BF80D32A400FF68F8E13C4E01C6" pageId="3" pageNumber="161" type="materials_examined">
<paragraph id="2F1518730D32A400FF68F8E13E17012E" blockId="3.[189,364,1824,1850]" box="[189,364,1824,1850]" pageId="3" pageNumber="161">
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<emphasis id="1DDEC4610D32A400FF68F8E13E17012E" bold="true" box="[189,364,1824,1850]" pageId="3" pageNumber="161">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="2F1518730D32A400FF3FF88E3C4E01C6" blockId="3.[189,1399,1871,2003]" pageId="3" pageNumber="161">
<materialsCitation id="9FC2122E0D32A400FF3FF88E3C4E01C6" ID-GBIF-Occurrence="3776288301" collectingDate="2007-12-13" collectingMethod="light trap" collectionCode="NHRS" collectorName="N. Jonsson &amp; T. Malm &amp; B. Viklund" country="Malaysia" county="Tawau" elevation="1031" latitude="4.73225" location="Nepenthes Camp" longLatPrecision="1" longitude="116.88111" municipality="Maliau Basin" pageId="3" pageNumber="161" specimenCode="VKBS-2007-47" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="1DDEC4610D32A400FF3FF88E3E27017D" bold="true" box="[234,348,1871,1897]" pageId="3" pageNumber="161">
<typeStatus id="F011A6D10D32A400FF3FF88E3E27017D" box="[234,348,1871,1897]" pageId="3" pageNumber="161" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="57BD58E30D32A400FF68F8B33E2F0198" box="[189,340,1906,1932]" name="Malaysia" pageId="3" pageNumber="161">MALAYSIA</collectingCountry>
<specimenCount id="39ACD3FA0D32A400FEBBF8B33EFF0198" box="[366,388,1906,1932]" pageId="3" pageNumber="161" type="male"></specimenCount>
(in alcohol);
<collectingRegion id="ED6ED6910D32A400FDF0F8B33D0B0198" box="[549,624,1906,1932]" country="Malaysia" name="Sabah" pageId="3" pageNumber="161">Sabah</collectingRegion>
,
<collectingCounty id="C67460FF0D32A400FDAEF8B33DB70198" box="[635,716,1906,1932]" pageId="3" pageNumber="161">Tawau</collectingCounty>
,
<collectingMunicipality id="CF7182090D32A400FD0DF8B33C020198" box="[728,889,1906,1932]" pageId="3" pageNumber="161">Maliau Basin</collectingMunicipality>
,
<location id="2A754EA80D32A400FC53F8B33B290198" LSID="urn:lsid:plazi:treatment:A703A9650D32A406FD55FAC93C16024D:2A754EA80D32A400FC53F8B33B290198" box="[902,1106,1906,1932]" country="Malaysia" county="Tawau" latitude="4.73225" longLatPrecision="1" longitude="116.88111" municipality="Maliau Basin" name="Nepenthes Camp" pageId="3" pageNumber="161" stateProvince="Sabah">Nepenthes Camp</location>
,
<locationDeviation id="3EA701510D32A400FB8AF8B03A090198" box="[1119,1394,1905,1932]" pageId="3" pageNumber="161">
<quantity id="E852B5960D32A400FB8AF8B03BD10198" box="[1119,1194,1905,1932]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="161" unit="m" value="150.0">150 m</quantity>
from Jalan Babi
</locationDeviation>
;
<geoCoordinate id="4A9E7EB40D32A400FF68F8523E2001BB" box="[189,347,1939,1967]" degrees="4" direction="north" minutes="43" orientation="latitude" pageId="3" pageNumber="161" precision="1" seconds="56.1" value="4.73225">4°4356.1″ N</geoCoordinate>
,
<geoCoordinate id="4A9E7EB40D32A400FEB2F8523D6501BB" box="[359,542,1939,1967]" degrees="116" direction="east" minutes="52" orientation="longitude" pageId="3" pageNumber="161" precision="1" seconds="52.0" value="116.88111">116°5252.0″ E</geoCoordinate>
;
<elevation id="A487FF400D32A400FDFEF8543DBE01BB" box="[555,709,1941,1967]" metricMagnitude="3" metricUnit="m" metricValue="1.031" pageId="3" pageNumber="161" unit="m" value="1031.0">
<quantity id="E852B5960D32A400FDFEF8543DFF01BB" box="[555,644,1941,1967]" metricMagnitude="3" metricUnit="m" metricValue="1.031" pageId="3" pageNumber="161" unit="m" value="1031.0">1031 m</quantity>
a.s.l.
</elevation>
;
<date id="5B143EB30D32A400FD06F8543C0F01BB" box="[723,884,1941,1967]" pageId="3" pageNumber="161" value="2007-12-13">
<collectingDate id="4B50C75B0D32A400FD06F8543C0F01BB" box="[723,884,1941,1967]" pageId="3" pageNumber="161" value="2007-12-13">13 Dec. 2007</collectingDate>
</date>
;
<collectorName id="825F7DA50D32A400FC54F8543B7E01BB" box="[897,1029,1941,1967]" pageId="3" pageNumber="161">N. Jönsson</collectorName>
,
<collectorName id="825F7DA50D32A400FBC4F8543B0D01BB" box="[1041,1142,1941,1967]" pageId="3" pageNumber="161">T. Malm</collectorName>
and
<collectorName id="825F7DA50D32A400FB65F8543A4F01BB" box="[1200,1332,1941,1967]" pageId="3" pageNumber="161">B. Viklund</collectorName>
;
<collectingMethod id="F6EB60640D32A400FA94F8543F9501C7" pageId="3" pageNumber="161">light trap</collectingMethod>
, loc#
<specimenCode id="7F0CB0080D32A400FEF8F8793E9101C6" box="[301,490,1976,2002]" pageId="3" pageNumber="161">VKBS-2007-47</specimenCode>
; DNA voucher JQ1;
<collectionCode id="49BB80B60D32A400FD0AF8793C4A01C6" box="[735,817,1976,2002]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="3" pageNumber="161">NHRS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="7BD548FB0D35A407FF68F8443E5A01F2" ID-DOI="http://doi.org/10.5281/zenodo.6564537" ID-Zenodo-Dep="6564537" httpUri="https://zenodo.org/record/6564537/files/figure.png" pageId="4" pageNumber="162" startId="4.[189,242,1925,1951]" targetBox="[248,1332,280,1880]" targetPageId="4">
<paragraph id="2F1518730D35A407FF68F8443E5A01F2" blockId="4.[189,1399,1925,2022]" pageId="4" pageNumber="162">
<emphasis id="1DDEC4610D35A407FF68F8443E44018B" bold="true" box="[189,319,1925,1951]" pageId="4" pageNumber="162">Figs 610.</emphasis>
<taxonomicName id="E8AA63F00D35A407FE9FF8473D5A018B" authority="Johanson &amp; Malm &amp; Sjöberg, 2022" authorityName="Johanson &amp; Malm &amp; Sjöberg" authorityYear="2022" box="[330,545,1925,1952]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="162" phylum="Arthropoda" rank="species" species="apelqvisti" status="sp. nov.">
<emphasis id="1DDEC4610D35A407FE9FF8473D5A018B" box="[330,545,1925,1952]" italics="true" pageId="4" pageNumber="162">Oecetis apelqvisti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06ED791A0D35A407FDF9F8473DF701B4" box="[556,652,1926,1952]" pageId="4" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHRS).
<emphasis id="1DDEC4610D35A407FC7DF8443CCC018B" bold="true" box="[936,951,1925,1951]" pageId="4" pageNumber="162">6</emphasis>
. Genitalia, lateral view.
<emphasis id="1DDEC4610D35A407FB36F8443B89018B" bold="true" box="[1251,1266,1925,1951]" pageId="4" pageNumber="162">7</emphasis>
. Genitalia, dorsal view.
<emphasis id="1DDEC4610D35A407FE8DF8693E1E01D6" bold="true" box="[344,357,1960,1986]" pageId="4" pageNumber="162">8</emphasis>
. Genitalia, ventral view.
<emphasis id="1DDEC4610D35A407FD42F8693DDF01D6" bold="true" box="[663,676,1960,1986]" pageId="4" pageNumber="162">9</emphasis>
. Phallus, lateral view.
<emphasis id="1DDEC4610D35A407FC6DF8693CA801D6" bold="true" box="[952,979,1960,1986]" pageId="4" pageNumber="162">10</emphasis>
. Phallus, ventral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="67B04BF80D34A406FF68FEC93B7E05FD" pageId="5" pageNumber="163" type="description">
<paragraph id="2F1518730D34A406FF68FEC93ECA0737" blockId="5.[189,433,264,291]" box="[189,433,264,291]" pageId="5" pageNumber="163">
<heading id="745DAF1F0D34A406FF68FEC93ECA0737" bold="true" box="[189,433,264,291]" fontSize="11" level="3" pageId="5" pageNumber="163" reason="3">
<emphasis id="1DDEC4610D34A406FF68FEC93ECA0737" bold="true" box="[189,433,264,291]" pageId="5" pageNumber="163">Description of male</emphasis>
</heading>
</paragraph>
<paragraph id="2F1518730D34A406FF68FEF93CD00745" blockId="5.[189,939,311,338]" box="[189,939,311,338]" pageId="5" pageNumber="163">
BODY. Forewing:
<quantity id="E852B5960D34A406FE59FEF63E9D0746" box="[396,486,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="5" pageNumber="163" unit="mm" value="6.7">6.7 mm</quantity>
(N = 1). Tergites VIVIII unmodified.
</paragraph>
<paragraph id="2F1518730D34A406FF68FEB83B7E05FD" blockId="5.[188,1401,376,1001]" pageId="5" pageNumber="163">
GENITALIA (
<figureCitation id="B79104F60D34A406FE97FEB93EC70787" box="[322,444,376,403]" captionStart="Figs 610" captionStartId="4.[189,242,1925,1951]" captionTargetBox="[248,1332,280,1880]" captionTargetId="figure-12@4.[222,1366,265,1890]" captionTargetPageId="4" captionText="Figs 610. Oecetis apelqvisti sp. nov., holotype, ♂ (NHRS). 6. Genitalia, lateral view. 7. Genitalia, dorsal view. 8. Genitalia, ventral view. 9. Phallus, lateral view. 10. Phallus, ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6564537" httpUri="https://zenodo.org/record/6564537/files/figure.png" pageId="5" pageNumber="163">Figs 610</figureCitation>
). Longer than high in lateral view. Segment IX oriented almost vertically, anterior margin uniformly convex along its height; ventral part almost circularly expanded posteriorly; dorsal part reduced. Acrotergite of segment IX well developed, forming single oval aggregation of setal bases about width of dorsal processes combined. Pair of dorsal processes of segment IX elongate, tube-like; segment lacking membranous area. Lower posterolateral plate of segment IX short, high, located from basis of inferior appendages to mid-height of segment; upper posterolateral plate of segment IX lacking. Longitudinal apodeme absent. Tergum X in lateral view twice length of superior appendages, curving ventrally along its length to pointed apex; in dorsal view bilobed from half length, both lobes tapering. Superior appendages very long, tubular, originating at anterior margin of segment IX at junction between dorsal and ventral parts; oriented dorsally before curving posteriorly over segment IX. Each inferior appendage high and wide, about as long as superior appendages, all setae shorter than length of inferior appendages; in lateral view 2-branched, basal two-thirds widening from base, sharply triangular dorsal branch, ventral margin expanded around mid-length; apical branch digitate, narrow, oriented posterodorsally; in ventral view, each inferior appendage with basimesal expansions, sharply narrowing after basal one-third, apical one-third narrow, digitate. Phallus about as long as tergum X in lateral view, uniformly slender, oriented posteriorly before curving ventrally at mid-length, apex with single hook-shaped sclerotized process; in ventral view about length of inferior appendages, tubular, slightly widening apically, hook-shaped sclerotized process located on left side.
</paragraph>
</subSubSection>
<subSubSection id="67B04BF80D34A406FF68FBD13C16024D" box="[189,877,1040,1113]" pageId="5" pageNumber="163" type="distribution">
<paragraph id="2F1518730D34A406FF68FBD13E2D023E" blockId="5.[189,342,1040,1066]" box="[189,342,1040,1066]" pageId="5" pageNumber="163">
<heading id="745DAF1F0D34A406FF68FBD13E2D023E" bold="true" box="[189,342,1040,1066]" fontSize="11" level="3" pageId="5" pageNumber="163" reason="3">
<emphasis id="1DDEC4610D34A406FF68FBD13E2D023E" bold="true" box="[189,342,1040,1066]" pageId="5" pageNumber="163">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="2F1518730D34A406FF68FBFE3C16024D" blockId="5.[189,877,1087,1113]" box="[189,877,1087,1113]" pageId="5" pageNumber="163">
The species is only known from the
<typeStatus id="F011A6D10D34A406FDB2FBFE3DE1024D" box="[615,666,1087,1113]" pageId="5" pageNumber="163">type</typeStatus>
locality (
<figureCitation id="B79104F60D34A406FCDCFBFE3C24024D" box="[777,863,1087,1113]" captionStart="Fig" captionStartId="5.[189,232,1954,1980]" captionTargetBox="[189,1398,1220,1916]" captionTargetId="figure-295@5.[189,1398,1220,1916]" captionTargetPageId="5" captionText="Fig. 11. Map with the position of the two sampling sites in Maliau Basin shown as orange dots. The position of the Maliau Basin area on Borneo is indicated with a blue rectangular in the upper-left map." figureDoi="http://doi.org/10.5281/zenodo.6564540" httpUri="https://zenodo.org/record/6564540/files/figure.png" pageId="5" pageNumber="163">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>