189 lines
22 KiB
XML
189 lines
22 KiB
XML
<document ID-ISSN="1464-5262" approvalRequired="33" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="14" approvalRequired_for_textStreams="17" approvalRequired_for_treatments="1" checkinTime="1630026853140" checkinUser="felipe" docAuthor="Lee, Won Je & Patterson, David J." docDate="2000" docId="313A87D7FF9E6B05AE2B2085FEB74A3C" docLanguage="en" docName="JNatHist.34.4.483-562.pdf" docOrigin="Journal of Natural History 34" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Petalomonas poosilla Larsen and Patterson 1990" docType="treatment" docVersion="5" lastPageNumber="524" masterDocId="CD03FFAFFFB46B2EAF3D2531FFA64F40" masterDocTitle="Heterotrophic ¯ agellates (Protista) from marine sediments of Botany Bay, Australia" masterLastPageNumber="562" masterPageNumber="483" pageNumber="523" updateTime="1674653143356" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Heterotrophic ¯ agellates (Protista) from marine sediments of Botany Bay, Australia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lee, Won Je</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Patterson, David J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2000</mods:date>
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<mods:detail type="volume">
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<mods:number>34</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>483</mods:start>
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<mods:end>562</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ISSN">1464-5262</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5281783" ID-Zenodo-Dep="5281783" LSID="urn:lsid:plazi:treatment:313A87D7FF9E6B05AE2B2085FEB74A3C" httpUri="http://treatment.plazi.org/id/313A87D7FF9E6B05AE2B2085FEB74A3C" lastPageId="43" lastPageNumber="524" pageId="42" pageNumber="523">
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<subSubSection pageId="42" pageNumber="523" type="nomenclature">
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<paragraph blockId="42.[278,836,1460,1516]" box="[278,836,1460,1484]" pageId="42" pageNumber="523">
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<heading box="[278,836,1460,1484]" centered="true" fontSize="10" level="2" pageId="42" pageNumber="523" reason="3">
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<taxonomicName authority="Larsen and Patterson, 1990" authorityName="Larsen and Patterson" authorityYear="1990" box="[278,836,1460,1484]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="42" pageNumber="523" phylum="Cercozoa" rank="species" species="poosilla">
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<emphasis bold="true" box="[278,513,1460,1484]" italics="true" pageId="42" pageNumber="523">Petalomonas poosilla</emphasis>
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<bibRefCitation author="LARSEN, J. & PATTERSON, D. J." box="[521,836,1460,1484]" pageId="42" pageNumber="523" pagination="937" refId="ref41333" refString="LARSEN, J. and PATTERSON, D. J., 1990, Some ¯ agellates (Protista) from tropical marine sediments, Journal of Natural History, 24, 801 ± 937." type="journal article" year="1990">Larsen and Patterson, 1990</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph blockId="42.[278,836,1460,1516]" box="[429,685,1492,1516]" pageId="42" pageNumber="523">
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(®gures
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<quantity box="[522,561,1492,1516]" metricMagnitude="-2" metricUnit="kg" metricValue="1.4" pageId="42" pageNumber="523" unit="g" value="14.0">14g</quantity>
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, h, 16e±g)
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="43" lastPageNumber="524" pageId="42" pageNumber="523" type="description">
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<paragraph blockId="42.[79,1035,1540,1691]" box="[120,691,1540,1564]" pageId="42" pageNumber="523">
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<emphasis box="[120,253,1540,1563]" italics="true" pageId="42" pageNumber="523">Description.</emphasis>
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We describe two populations of cells.
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</paragraph>
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<paragraph blockId="42.[79,1035,1540,1691]" lastBlockId="43.[79,1035,137,1404]" lastPageId="43" lastPageNumber="524" pageId="42" pageNumber="523">Population 1: small cells; ovate, 5±9 Mm long, 3±5 Mm wide, dorso-ventrally ¯attened. This species may or may not have up to three dorsal ridges and the ridges may or may not be well developed. The organism may also have two ®ne ventral ridges which are di cult to observe. The right ventral ridge runs beneath the reservoir along the cell. With one ¯agellum as long as the cell. The reservoir is in the right side of the cell and the nucleus is in the left side. Moves by gliding. Common.</paragraph>
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<paragraph blockId="43.[79,1035,137,1404]" pageId="43" pageNumber="524">Population 2: Larger cells; cell outline oblong-oval, 10±12 Mm long and 5±6 Mm wide, dorso-ventrally ¯attened. It has four ridges extending along the cell: two indistinct dorsal ridges located at the lateral margins of the cell; two ®ne ventral ridges, the right one of which runs beneath the reservoir. One ¯agellum is about as long as the cell or slightly longer. The ovate reservoir is anteriorly situated in the right-hand side of the cell and the nucleus in the left-hand side of the cell. Often co-occurs with the smaller cell.</paragraph>
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<paragraph blockId="43.[79,1035,137,1404]" pageId="43" pageNumber="524">
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<emphasis box="[120,226,423,447]" italics="true" pageId="43" pageNumber="524">Remarks.</emphasis>
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<bibRefCitation author="LARSEN, J. & PATTERSON, D. J." box="[232,557,423,447]" pageId="43" pageNumber="524" pagination="937" refId="ref41333" refString="LARSEN, J. and PATTERSON, D. J., 1990, Some ¯ agellates (Protista) from tropical marine sediments, Journal of Natural History, 24, 801 ± 937." type="journal article" year="1990">Larsen and Patterson (1990)</bibRefCitation>
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provided a new name for
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<taxonomicName authority="Skuja, 1948" authorityName="Skuja" authorityYear="1948" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="pusilla">
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<emphasis box="[853,955,423,447]" italics="true" pageId="43" pageNumber="524">P. pusilla</emphasis>
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<bibRefCitation author="SKUJA, H." pageId="43" pageNumber="524" pagination="399" refId="ref43143" refString="SKUJA, H., 1948, Taxonomie des Phytoplanktons einiger Seen in Uppland, Schweden, Symbolae Botanicae Upsalienses, 9, 5 ± 399." type="journal article" year="1948">Skuja, 1948</bibRefCitation>
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</taxonomicName>
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(
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<emphasis box="[145,274,455,479]" italics="true" pageId="43" pageNumber="524">
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<taxonomicName authorityName="Larsen and Patterson" authorityYear="1990" box="[145,260,455,479]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="poosilla">P. poosilla</taxonomicName>
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)
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</emphasis>
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because under the principle of co-ordination of the ICZN,
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<taxonomicName authority="Skuja, 1948" authorityName="Skuja" authorityYear="1948" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="pusilla">
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<emphasis box="[932,1034,455,479]" italics="true" pageId="43" pageNumber="524">P. pusilla</emphasis>
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<bibRefCitation author="SKUJA, H." box="[79,225,487,511]" pageId="43" pageNumber="524" pagination="399" refId="ref43143" refString="SKUJA, H., 1948, Taxonomie des Phytoplanktons einiger Seen in Uppland, Schweden, Symbolae Botanicae Upsalienses, 9, 5 ± 399." type="journal article" year="1948">Skuja, 1948</bibRefCitation>
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</taxonomicName>
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is a homonym of
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<taxonomicName authority="Klebs, 1893" authorityName="Klebs" authorityYear="1893" box="[499,1034,487,511]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="variety" species="mediocanellata" variety="pusilla">
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<emphasis box="[499,710,487,511]" italics="true" pageId="43" pageNumber="524">P. mediocanellata</emphasis>
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var.
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<emphasis box="[796,867,487,511]" italics="true" pageId="43" pageNumber="524">pusilla</emphasis>
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<bibRefCitation author="KLEBS, G." box="[888,1034,487,511]" pageId="43" pageNumber="524" pagination="445" refId="ref41149" refString="KLEBS, G., 1893, Flagellatenstudien, Zeitschrift fuEr Wissenschaftliche Zoologie, 55, 265 ± 445." type="journal article" year="1893">Klebs, 1893</bibRefCitation>
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</taxonomicName>
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(
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<bibRefCitation author="LEMMERMANN, E." box="[90,324,519,543]" pageId="43" pageNumber="524" pagination="174" refId="ref41582" refString="LEMMERMANN, E., 1913, Chrysomonadinae, Cryptomonadinae, Eugleninae, Chloromonadinae und Gefarbte Flagellaten Unsicherer Stellung, in A. Pascher (ed.) Die SuEsswasserora Deutschlands, OEsterreich und der Schweiz, II (Jena: Fischer Verlag), pp. 115 ± 174." type="journal article" year="1913">Lemmermann, 1913</bibRefCitation>
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)Ðwhich we refer to as
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<taxonomicName authorityName="Hollande" authorityYear="1942" box="[632,747,520,543]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="minuta">
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<emphasis box="[632,747,520,543]" italics="true" pageId="43" pageNumber="524">P. minuta</emphasis>
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</taxonomicName>
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(see
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<taxonomicName authorityName="Hollande" authorityYear="1942" box="[820,935,520,543]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="minuta">
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<emphasis box="[820,935,520,543]" italics="true" pageId="43" pageNumber="524">P. minuta</emphasis>
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</taxonomicName>
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above).
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<taxonomicName authorityName="Larsen and Patterson" authorityYear="1990" box="[79,315,551,575]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="poosilla">
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<emphasis box="[79,315,551,575]" italics="true" pageId="43" pageNumber="524">Petalomonas poosilla</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Hollande" authorityYear="1942" box="[378,488,552,575]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="minuta">
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<emphasis box="[378,488,552,575]" italics="true" pageId="43" pageNumber="524">P. minuta</emphasis>
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</taxonomicName>
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resemble each other in shape and appearance, but can be distinguised by the presence of what we regard as a longitudinal groove in
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<emphasis box="[110,226,616,639]" italics="true" pageId="43" pageNumber="524">
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<taxonomicName authorityName="Hollande" authorityYear="1942" box="[110,222,616,639]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="minuta">P. minuta</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="524" type="reference_group">
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<paragraph blockId="43.[79,1035,137,1404]" pageId="43" pageNumber="524">
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<taxonomicName authorityName="Larsen and Patterson" authorityYear="1990" box="[120,352,647,671]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="poosilla">
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<emphasis box="[120,352,647,671]" italics="true" pageId="43" pageNumber="524">Petalomonas poosilla</emphasis>
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</taxonomicName>
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was originally described under the name
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<taxonomicName authorityName="Skuja" authorityYear="1948" box="[818,920,647,671]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="pusilla">
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<emphasis box="[818,920,647,671]" italics="true" pageId="43" pageNumber="524">P. pusilla</emphasis>
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</taxonomicName>
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(5±12 Mm long) from freshwater sites (
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<bibRefCitation author="SKUJA, H." box="[407,544,679,703]" pageId="43" pageNumber="524" pagination="399" refId="ref43143" refString="SKUJA, H., 1948, Taxonomie des Phytoplanktons einiger Seen in Uppland, Schweden, Symbolae Botanicae Upsalienses, 9, 5 ± 399." type="journal article" year="1948">Skuja, 1948</bibRefCitation>
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) and has since been found in marine sites in
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<collectingCountry box="[113,217,710,734]" name="Australia" pageId="43" pageNumber="524">Australia</collectingCountry>
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,
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<collectingCountry box="[237,304,710,734]" name="Brazil" pageId="43" pageNumber="524">Brazil</collectingCountry>
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, Danish Wadden Sea,
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<collectingCountry box="[581,621,710,734]" name="Fiji" pageId="43" pageNumber="524">Fiji</collectingCountry>
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, Gulf of
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<collectingCountry box="[739,828,710,734]" name="Finland" pageId="43" pageNumber="524">Finland</collectingCountry>
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; Hawaii and NE
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<collectingCountry box="[79,167,742,766]" name="Canada" pageId="43" pageNumber="524">Canada</collectingCountry>
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(
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<bibRefCitation author="LACKEY, J. B. & LACKEY, E. W." box="[186,474,742,766]" pageId="43" pageNumber="524" pagination="795" refId="ref41196" refString="LACKEY, J. B. and LACKEY, E. W., 1970, A late summer checklist of the marine microbiota around Logy Bay, Newfoundland, Canadian Journal of Zoology, 48, 789 ± 795." type="journal article" year="1970">Lackey and Lackey, 1970</bibRefCitation>
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;
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<bibRefCitation author="LARSEN, J." box="[489,633,742,766]" pageId="43" pageNumber="524" pagination="607" refId="ref41306" refString="LARSEN, J., 1987, Algal studies of the interstitial euglenoid ¯ agellates, Nordic Journal of Botany, 7, 589 ± 607." type="journal article" year="1987">Larsen, 1987</bibRefCitation>
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;
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<bibRefCitation author="LARSEN, J. & PATTERSON, D. J." box="[648,958,742,766]" pageId="43" pageNumber="524" pagination="937" refId="ref41333" refString="LARSEN, J. and PATTERSON, D. J., 1990, Some ¯ agellates (Protista) from tropical marine sediments, Journal of Natural History, 24, 801 ± 937." type="journal article" year="1990">Larsen and Patterson, 1990</bibRefCitation>
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; Vùrs, 1992a;
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<bibRefCitation author="PATTERSON, D. J. & SIMPSON, A. G. B." box="[163,500,774,798]" pageId="43" pageNumber="524" pagination="448" refId="ref42397" refString="PATTERSON, D. J. and SIMPSON, A. G. B., 1996, Heterotrophic ¯ agellates from coastal marine and hypersaline sediments in Western Australia, European Journal of Protistology, 32, 423 ± 448." type="journal article" year="1996">Patterson and Simpson, 1996</bibRefCitation>
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;
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<bibRefCitation author="TONG, S. M. & NYGAARD, K. & BERNARD, C. & VORS, N. & PATTERSON, D. J." box="[518,714,774,799]" pageId="43" pageNumber="524" pagination="194" refId="ref43851" refString="TONG, S. M., NYGAARD, K., BERNARD, C., VORS, N. and PATTERSON, D. J., 1998, Heterotrophic ¯ agellates from the water column in Port Jackson, Sydney, Australia, European Journal of Protistology, 34, 162 ± 194." type="journal article" year="1998">
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Tong
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<emphasis box="[586,636,774,798]" italics="true" pageId="43" pageNumber="524">et al</emphasis>
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., 1998
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</bibRefCitation>
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). We observed cells (in the small population) which are similar in size and outline to
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<taxonomicName authorityName="Larsen and Patterson" authorityYear="1990" box="[764,883,806,830]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="poosilla">
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<emphasis box="[764,883,806,830]" italics="true" pageId="43" pageNumber="524">P. poosilla</emphasis>
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</taxonomicName>
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as described by
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<bibRefCitation author="LARSEN, J. & PATTERSON, D. J." box="[116,452,838,862]" pageId="43" pageNumber="524" pagination="937" refId="ref41333" refString="LARSEN, J. and PATTERSON, D. J., 1990, Some ¯ agellates (Protista) from tropical marine sediments, Journal of Natural History, 24, 801 ± 937." type="journal article" year="1990">Larsen and Patterson (1990)</bibRefCitation>
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. This species has been described without ridges or grooves (
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<bibRefCitation author="SKUJA, H." box="[186,324,870,894]" pageId="43" pageNumber="524" pagination="399" refId="ref43143" refString="SKUJA, H., 1948, Taxonomie des Phytoplanktons einiger Seen in Uppland, Schweden, Symbolae Botanicae Upsalienses, 9, 5 ± 399." type="journal article" year="1948">Skuja, 1948</bibRefCitation>
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;
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<bibRefCitation author="LARSEN, J. & PATTERSON, D. J." box="[344,667,870,894]" pageId="43" pageNumber="524" pagination="937" refId="ref41333" refString="LARSEN, J. and PATTERSON, D. J., 1990, Some ¯ agellates (Protista) from tropical marine sediments, Journal of Natural History, 24, 801 ± 937." type="journal article" year="1990">Larsen and Patterson, 1990</bibRefCitation>
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). Cells assigned to this species have been described more recently as having ridges (
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<bibRefCitation author="PATTERSON, D. J. & SIMPSON, A. G. B." box="[691,1026,901,925]" pageId="43" pageNumber="524" pagination="448" refId="ref42397" refString="PATTERSON, D. J. and SIMPSON, A. G. B., 1996, Heterotrophic ¯ agellates from coastal marine and hypersaline sediments in Western Australia, European Journal of Protistology, 32, 423 ± 448." type="journal article" year="1996">Patterson and Simpson, 1996</bibRefCitation>
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; Schroeckh and Patterson, unpubl.). We suspect that this species has previously been under-described and regard the latter descriptions as more accurate than the former. The ventral face of this species is very similar to that of
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<taxonomicName authorityName="Larsen and Patterson" authorityYear="1990" box="[746,845,998,1021]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="minor">
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<emphasis box="[746,845,998,1021]" italics="true" pageId="43" pageNumber="524">P. minor</emphasis>
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</taxonomicName>
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, described here, in having two ®ne ventral ridges and in the position of the ridges, but the two species can be distinguished by cell shape and the dorsal ¯ange in
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<taxonomicName authorityName="Larsen and Patterson" authorityYear="1990" box="[923,1027,1062,1085]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="minor">
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<emphasis box="[923,1027,1062,1085]" italics="true" pageId="43" pageNumber="524">P. minor</emphasis>
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</taxonomicName>
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.
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<taxonomicName authorityName="Larsen and Patterson" authorityYear="1990" box="[79,315,1093,1117]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="poosilla">
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<emphasis box="[79,315,1093,1117]" italics="true" pageId="43" pageNumber="524">Petalomonas poosilla</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Hollande" authorityYear="1942" box="[378,488,1094,1117]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="minuta">
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<emphasis box="[378,488,1094,1117]" italics="true" pageId="43" pageNumber="524">P. minuta</emphasis>
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</taxonomicName>
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resemble each other in shape and appearance, but can be distinguised by the presence of what we regard as a longitudinal groove in
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<emphasis box="[112,230,1158,1181]" italics="true" pageId="43" pageNumber="524">
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<taxonomicName authorityName="Hollande" authorityYear="1942" box="[112,225,1158,1181]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="minuta">P. minuta</taxonomicName>
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.
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</emphasis>
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This species resembles
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<taxonomicName authorityName="Cann & Pennick" authorityYear="1986" box="[509,674,1158,1181]" class="Imbricatea" family="Thaumatomonadidae" genus="Petalomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thaumatomonadida" pageId="43" pageNumber="524" phylum="Cercozoa" rank="species" species="cantuscygni">
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<emphasis box="[509,674,1158,1181]" italics="true" pageId="43" pageNumber="524">P. cantuscygni</emphasis>
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</taxonomicName>
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(see
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<bibRefCitation author="CANN, J. P. & PENNICK, N. C." box="[737,1024,1157,1181]" pageId="43" pageNumber="524" pagination="71" refId="ref39402" refString="CANN, J. P. and PENNICK, N. C., 1986, Observations on Petalomonas cantuscygni, n. sp., a new halo-tolerant strain, Archiv fuEr Protistenkunde, 132, 63 ± 71." type="journal article" year="1986">Cann and Pennick, 1986</bibRefCitation>
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) from which it can be distinguished by never having fewer than six dorsal ridges.
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</paragraph>
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<paragraph blockId="43.[79,1035,137,1404]" pageId="43" pageNumber="524">We have observed larger cells about 10±12 Mm long, but they consistently had four discrete ridges on both sides of the cell. The two populations often co-occur. Although these may represent a separate species, we have been unable to establish morphological discontinuities by which single cells could be easily identi®ed when comparisons are made. Further studies are required to establish the status of the two populations.</paragraph>
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</subSubSection>
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</treatment>
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</document> |