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<document id="FD12BA85997058026ED9A80000DD3995" ID-CLB-Dataset="45466" ID-DOI="10.5281/zenodo.215224" ID-GBIF-Dataset="3a53b495-17db-421b-be56-da4dadaea543" ID-ISSN="1175-5326" ID-Zenodo-Dep="215224" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460437889326" checkinUser="plazi" docAuthor="Magnacca, Karl N. &amp; Price, Donald K." docDate="2012" docId="039CB000FFC6FFB6F5EFD122AE88FEC9" docLanguage="en" docName="zt03188p030.pdf" docOrigin="Zootaxa 3188" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Drosophila moli Magnacca, new species" docType="treatment" docVersion="10" lastPageNumber="18" masterDocId="FFA5C878FFD7FFA4F578D04DAF04FFF6" masterDocTitle="New species of Hawaiian picture wing Drosophila (Diptera: Drosophilidae), with a key to species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="18" updateTime="1698332463146" updateUser="ExternalLinkService">
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<mods:title id="50896809FCA1087D337B98E37FE2D42D">New species of Hawaiian picture wing Drosophila (Diptera: Drosophilidae), with a key to species</mods:title>
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<mods:namePart id="1E06CFA258338211E3AB7A223D8994A5">Magnacca, Karl N.</mods:namePart>
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<mods:namePart id="9D6963BA2E1025BF13B8B8D21B0ACBD4">Price, Donald K.</mods:namePart>
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<subSubSection id="C32F529DFFC6FFB5F5EFD122AFDCFE5C" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8B8A0116FFC6FFB5F5EFD122AD85FE7F" blockId="17.[151,641,367,426]" box="[151,641,367,393]" pageId="17" pageNumber="18">
<heading id="D0C2B67AFFC6FFB5F5EFD122AD85FE7F" bold="true" box="[151,641,367,393]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="B941DD04FFC6FFB5F5EFD122AD85FE7F" bold="true" box="[151,641,367,393]" pageId="17" pageNumber="18">
<taxonomicName id="4C357A95FFC6FFB5F5EFD122AEE0FE7F" ID-CoL="8BXY4" authority="Magnacca" authorityName="Magnacca" box="[151,484,367,393]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="moli" status="sp. nov.">
<emphasis id="B941DD04FFC6FFB5F5EFD122AE5FFE7F" bold="true" box="[151,347,367,393]" italics="true" pageId="17" pageNumber="18">Drosophila moli</emphasis>
Magnacca
</taxonomicName>
,
<taxonomicNameLabel id="A272607FFFC6FFB5F48AD122AD85FE7F" box="[498,641,367,393]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A0116FFC6FFB5F5EFD1DFAFDCFE5C" blockId="17.[151,641,367,426]" box="[151,216,402,426]" pageId="17" pageNumber="18">
<figureCitation id="130E1D93FFC6FFB5F5EFD1DFAFDCFE5C" box="[151,216,402,426]" captionStart="FIGURE 4" captionStartId="18.[151,250,1587,1610]" captionTargetBox="[151,1436,377,1565]" captionTargetId="figure@18.[151,1436,375,1565]" captionTargetPageId="18" captionText="FIGURE 4. Drosophila moli. A. Right front leg of male, anterior view. B. Male wing. C. Female wing." httpUri="https://zenodo.org/record/215228/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32F529DFFC6FFB5F5EFD197ABF8FDCC" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8B8A0116FFC6FFB5F5EFD197ABF8FDCC" blockId="17.[151,1437,474,2025]" pageId="17" pageNumber="18">
<emphasis id="B941DD04FFC6FFB5F5EFD197AE15FE05" bold="true" box="[151,273,474,499]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
The wing and body coloration patterns resemble others in this species complex, including
<taxonomicName id="4C357A95FFC6FFB5F07AD191AA91FE05" box="[1282,1429,475,499]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lanaiensis">
<emphasis id="B941DD04FFC6FFB5F07AD191AA91FE05" box="[1282,1429,475,499]" italics="true" pageId="17" pageNumber="18">D. lanaiensis</emphasis>
</taxonomicName>
, with which it was formerly grouped, and females of
<taxonomicName id="4C357A95FFC6FFB5F79FD24DAC7FFDE0" box="[743,891,511,535]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="conspicua">
<emphasis id="B941DD04FFC6FFB5F79FD24DAC7FFDE0" box="[743,891,511,535]" italics="true" pageId="17" pageNumber="18">D. conspicua</emphasis>
</taxonomicName>
. Males can be immediately distinguished by the elongate, narrow wings, and both sexes have the mark over the dm-cu crossvein distinctly L-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C32F529DFFC6FFB5F5B1D20BAB20F9C0" pageId="17" pageNumber="18" type="description">
<paragraph id="8B8A0116FFC6FFB5F5B1D20BABE8FACF" blockId="17.[151,1437,474,2025]" pageId="17" pageNumber="18">
<emphasis id="B941DD04FFC6FFB5F5B1D20BAE5FFDA9" bold="true" box="[201,347,582,607]" pageId="17" pageNumber="18">Description.</emphasis>
Male.
<emphasis id="B941DD04FFC6FFB5F4D6D20AAEF1FDA9" bold="true" box="[430,501,583,607]" pageId="17" pageNumber="18">Head.</emphasis>
Front pale, faintly tinged with brown; ocellar triangle black. Fronto-orbital setae normal, anterior reclinate very small, about half as long as the proclinate and one-third as long as the posterior reclinate. Face and gena pale yellow except for a small black spot below the eye. Antenna with the second segment brown laterally, otherwise yellow; arista with about 910 dorsal and 4 ventral rays in addition to the apical fork, and several rather long median branches. One strong oral vibrissa, followed posteriorly by three progressively smaller setae, with numerous small setulae in between them. Palp largely yellow, slightly darker at the tip, nearly cylindrical; with a long, thin, yellow apical seta.
<emphasis id="B941DD04FFC6FFB5F7C7D352AC1BFCC1" bold="true" box="[703,799,799,823]" pageId="17" pageNumber="18">Thorax.</emphasis>
Mesonotum largely yellow with three strongly contrasting dark brown to black stripes; lateral stripes dark, extending from the anterior margin of the humeri nearly to the scutellum; median stripe fainter, broadening between the dorsocentral setae to fill the space between them, and extending onto the scutellum. Pleura almost entirely yellow except for a large dark mark on the posterior half of the anepisternum. Two pairs of strong dorsocentral setae, the anterior about 2/3 as long as the posterior.
<emphasis id="B941DD04FFC6FFB5F184D3E2AA3FFC31" bold="true" box="[1276,1339,943,967]" pageId="17" pageNumber="18">Legs.</emphasis>
Entirely yellow. Front tibia with regular anterodorsal and posterodorsal rows of 1012 long cilia along its entire length, slightly longer at the base, with a few scattered dorsal cilia in between. Front tarsus with about 20 dorsal cilia arranged in three irregular rows, similar to those of the tibia. Second and third tarsal segments also with sparse cilia.
<emphasis id="B941DD04FFC6FFB5F5ADD473AE2CFBA1" bold="true" box="[213,296,1086,1111]" pageId="17" pageNumber="18">Wings.</emphasis>
Extremely long and narrow, about four times as long as wide; apically pointed and slightly bent posteriorly, posterior margin between M and CuA1 weakly concave. Subbasal wing spot small, scarcely reaching into cell R2. Anterior mark elongate, extending along about 1/3 the length of the wing; connected to the large L-shaped mark on the dm-cu crossvein via a small circular mark over R2+3 (the latter rarely reduced or absent). Apical marks confluent or nearly so. Costal fringe extending about 1/2 the distance between apex of R2+3 and R4+5.
<emphasis id="B941DD04FFC6FFB5F058D49BAA98FB19" bold="true" box="[1312,1436,1238,1263]" pageId="17" pageNumber="18">Abdomen.</emphasis>
Coloration variable, from each segment predominantly yellow with only a narrow posterior band dark brown, to predominantly brown with yellow anterolateral areas. Cerci yellow-brown. Genitalia not dissected.
</paragraph>
<paragraph id="8B8A0116FFC6FFB5F5B1D50BAC6CFA18" blockId="17.[151,1437,474,2025]" pageId="17" pageNumber="18">
Female. Identical to the male with the following exceptions.
<emphasis id="B941DD04FFC6FFB5F615D50BACB0FAA8" bold="true" box="[877,948,1350,1374]" pageId="17" pageNumber="18">Head.</emphasis>
Third antennal segment tinged with brown. Apical seta of palp shorter, stronger, black.
<emphasis id="B941DD04FFC6FFB5F7FBD527ADE7FA74" bold="true" box="[643,739,1386,1410]" pageId="17" pageNumber="18">Thorax.</emphasis>
Median stripe of mesonotum broader on the anterior half and dark, similar to the lateral stripes.
<emphasis id="B941DD04FFC6FFB5F76FD5C3AD52FA50" bold="true" box="[535,598,1422,1446]" pageId="17" pageNumber="18">Legs.</emphasis>
Front legs without elongate cilia or setae.
<emphasis id="B941DD04FFC6FFB5F14DD5C0AB83FA50" bold="true" box="[1077,1159,1421,1446]" pageId="17" pageNumber="18">Wings.</emphasis>
Not as long and narrow, about three times as long as wide; shape normal, posterior margin convex, apex only very weakly pointed.
<emphasis id="B941DD04FFC6FFB5F029D5FCAFD5FA18" bold="true" pageId="17" pageNumber="18">Abdomen.</emphasis>
Ovipositor long, straight, yellow, pointed vertically at rest.
</paragraph>
<paragraph id="8B8A0116FFC6FFB5F5B1D5B7AB20F9C0" blockId="17.[151,1437,474,2025]" pageId="17" pageNumber="18">
<emphasis id="B941DD04FFC6FFB5F5B1D5B7AE6CF9E4" bold="true" box="[201,360,1529,1554]" pageId="17" pageNumber="18">
<typeStatus id="548EBFB4FFC6FFB5F5B1D5B7AE09F9E4" box="[201,269,1530,1554]" pageId="17" pageNumber="18">Types</typeStatus>
. Oahu
</emphasis>
:
<typeStatus id="548EBFB4FFC6FFB5F402D5B4AEE6F9E7" box="[378,482,1529,1553]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
3 and allotype Ƥ, above Nuuanu Pali lookout,
<quantity id="4CCDACF3FFC6FFB5F16ED5B4AB68F9E7" box="[1046,1132,1529,1553]" metricMagnitude="2" metricUnit="m" metricValue="4.2672" pageId="17" pageNumber="18" unit="ft" value="1400.0">1400 ft</quantity>
.,
<geoCoordinate id="EE0167D1FFC6FFB5F106D5B4ABEEF9E7" box="[1150,1258,1529,1553]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="55" value="21.365">21.365°N</geoCoordinate>
<geoCoordinate id="EE0167D1FFC6FFB5F18BD5B4AA77F9E7" box="[1267,1395,1529,1553]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="55" value="-157.761">157.761°W</geoCoordinate>
, on bait sponge,
<date id="FF8B27D6FFC6FFB5F45DD653AEA5F9C0" box="[293,417,1566,1590]" pageId="17" pageNumber="18" value="2009-08-02">2.viii.2009</date>
, K. Magnacca (UHIM);
<typeStatus id="548EBFB4FFC6FFB5F7CDD652AC1CF9C1" box="[693,792,1567,1591]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
, 3, same data (BPBM).
</paragraph>
</subSubSection>
<subSubSection id="C32F529DFFC6FFB5F5B1D60CAE60F988" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8B8A0116FFC6FFB5F5B1D60CAE60F988" blockId="17.[151,1437,474,2025]" pageId="17" pageNumber="18">
<emphasis id="B941DD04FFC6FFB5F5B1D60CAEFCF9AC" bold="true" box="[201,504,1601,1626]" pageId="17" pageNumber="18">Distribution and ecology.</emphasis>
Oahu. Probably breeds in
<taxonomicName id="4C357A95FFC6FFB5F64ED60FACC9F9AC" box="[822,973,1602,1626]" class="Magnoliopsida" family="Amaranthaceae" genus="Charpentiera" kingdom="Plantae" order="Caryophyllales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B941DD04FFC6FFB5F64ED60FACC9F9AC" box="[822,973,1602,1626]" italics="true" pageId="17" pageNumber="18">Charpentiera</emphasis>
</taxonomicName>
; females observed ovipositing in rotting trunk (not reared).
</paragraph>
</subSubSection>
<subSubSection id="C32F529DFFC6FFB5F5B1D6C7AA45F957" box="[201,1345,1673,1698]" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8B8A0116FFC6FFB5F5B1D6C7AA45F957" blockId="17.[151,1437,474,2025]" box="[201,1345,1673,1698]" pageId="17" pageNumber="18">
<emphasis id="B941DD04FFC6FFB5F5B1D6C7AE54F954" bold="true" box="[201,336,1674,1698]" pageId="17" pageNumber="18">Etymology.</emphasis>
From the Hawaiian
<emphasis id="B941DD04FFC6FFB5F74DD6C6AD4DF954" box="[565,585,1675,1698]" italics="true" pageId="17" pageNumber="18">m</emphasis>
ō
<emphasis id="B941DD04FFC6FFB5F720D6C7AD64F954" box="[600,608,1674,1698]" italics="true" pageId="17" pageNumber="18">l</emphasis>
ī, albatross, referring to the similarly long, narrow, pointed wings.
</paragraph>
</subSubSection>
<subSubSection id="C32F529DFFC6FFB6F5B1D6E0AE88FEC9" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8B8A0116FFC6FFB6F5B1D6E0AE88FEC9" blockId="17.[151,1437,474,2025]" lastBlockId="18.[151,1436,151,319]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B941DD04FFC6FFB5F5B1D6E0AE4AF930" bold="true" box="[201,334,1709,1734]" pageId="17" pageNumber="18">Discussion.</emphasis>
<bibRefCitation id="EFA47CE7FFC6FFB5F424D6E3AEFEF930" author="Hardy" box="[348,506,1710,1734]" pageId="17" pageNumber="18" refString="Hardy, D. E. (1965) Insects of Hawaii vol. 12. Diptera: Cyclorrhapha II, Series Schizophora, Section Acalypterae I. Family Drosophilidae. University of Hawaii Press, Honolulu, 814 pp." type="book" year="1965">Hardy (1965)</bibRefCitation>
considered this population from Oahu to be conspecific with
<taxonomicName id="4C357A95FFC6FFB5F182D6E2AA91F930" box="[1274,1429,1710,1734]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lanaiensis">
<emphasis id="B941DD04FFC6FFB5F182D6E2AA91F930" box="[1274,1429,1710,1734]" italics="true" pageId="17" pageNumber="18">D. lanaiensis</emphasis>
</taxonomicName>
, described from only females. However, it is now clear that the
<taxonomicName id="4C357A95FFC6FFB5F62CD69EACE3F91C" box="[852,999,1746,1770]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lanaiensis">
<emphasis id="B941DD04FFC6FFB5F62CD69EACE3F91C" box="[852,999,1746,1770]" italics="true" pageId="17" pageNumber="18">D. lanaiensis</emphasis>
</taxonomicName>
<typeStatus id="548EBFB4FFC6FFB5F696D69FAB2DF91C" box="[1006,1065,1746,1770]" pageId="17" pageNumber="18">types</typeStatus>
are identical to recent specimens from Lānai and those from Maui (previously known as
<taxonomicName id="4C357A95FFC6FFB5F674D6BAAC97F8F8" box="[780,915,1782,1806]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="virgulata">
<emphasis id="B941DD04FFC6FFB5F674D6BAAC97F8F8" box="[780,915,1782,1806]" italics="true" pageId="17" pageNumber="18">D. virgulata</emphasis>
</taxonomicName>
), and different from the Oahu specimens (see Discussion under
<taxonomicName id="4C357A95FFC6FFB5F427D756AEF0F8C4" box="[351,500,1818,1842]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lanaiensis">
<emphasis id="B941DD04FFC6FFB5F427D756AEF0F8C4" box="[351,500,1818,1842]" italics="true" pageId="17" pageNumber="18">D. lanaiensis</emphasis>
</taxonomicName>
above). The drawing of the male wing in
<bibRefCitation id="EFA47CE7FFC6FFB5F6B3D754AB66F8C7" author="Hardy" box="[971,1122,1817,1841]" pageId="17" pageNumber="18" refString="Hardy, D. E. (1965) Insects of Hawaii vol. 12. Diptera: Cyclorrhapha II, Series Schizophora, Section Acalypterae I. Family Drosophilidae. University of Hawaii Press, Honolulu, 814 pp." type="book" year="1965">Hardy (1965)</bibRefCitation>
appears to be of an unusual specimen; all those I have seen possess a small but distinct spot in the middle of vein R4+5, as in
<taxonomicName id="4C357A95FFC6FFB5F1AFD772AA68F8A0" box="[1239,1388,1854,1878]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lanaiensis">
<emphasis id="B941DD04FFC6FFB5F1AFD772AA68F8A0" box="[1239,1388,1854,1878]" italics="true" pageId="17" pageNumber="18">D. lanaiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C357A95FFC6FFB5F5EFD72BAE13F88B" box="[151,279,1893,1917]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="digressa">
<emphasis id="B941DD04FFC6FFB5F5EFD72BAE13F88B" box="[151,279,1893,1917]" italics="true" pageId="17" pageNumber="18">D. digressa</emphasis>
</taxonomicName>
, although it is missing on one wing of the
<taxonomicName id="4C357A95FFC6FFB5F782D72BAC4BF88B" box="[762,847,1893,1917]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="moli">
<emphasis id="B941DD04FFC6FFB5F782D72BAC4BF88B" box="[762,847,1893,1917]" italics="true" pageId="17" pageNumber="18">D. moli</emphasis>
</taxonomicName>
allotype. This variation is occasionally seen in individuals of the other two species as well. None of the specimens examined by Hardy, or any other earlier ones, could be located in the BPBM or UHIM collections. Although it has not been reared from
<taxonomicName id="4C357A95FFC6FFB5F130D7E0ABDBF833" box="[1096,1247,1965,1989]" class="Magnoliopsida" family="Amaranthaceae" genus="Charpentiera" kingdom="Plantae" order="Caryophyllales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B941DD04FFC6FFB5F130D7E0ABDBF833" box="[1096,1247,1965,1989]" italics="true" pageId="17" pageNumber="18">Charpentiera</emphasis>
</taxonomicName>
, it was observed ovipositing in a rotting tree, and
<taxonomicName id="4C357A95FFC6FFB5F77BD79FAD87F81F" box="[515,643,2001,2025]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="digressa">
<emphasis id="B941DD04FFC6FFB5F77BD79FAD87F81F" box="[515,643,2001,2025]" italics="true" pageId="17" pageNumber="18">D. digressa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C357A95FFC6FFB5F7C1D79FAC49F81F" box="[697,845,2001,2025]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lanaiensis">
<emphasis id="B941DD04FFC6FFB5F7C1D79FAC49F81F" box="[697,845,2001,2025]" italics="true" pageId="17" pageNumber="18">D. lanaiensis</emphasis>
</taxonomicName>
have both been reared from it. This species is probably highly vulnerable due to habitat loss; although
<taxonomicName id="4C357A95FFC5FFB6F7A9D0D5AC6CFF46" box="[721,872,152,176]" class="Magnoliopsida" family="Amaranthaceae" genus="Charpentiera" kingdom="Plantae" order="Caryophyllales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis id="B941DD04FFC5FFB6F7A9D0D5AC6CFF46" box="[721,872,152,176]" italics="true" pageId="18" pageNumber="19">Charpentiera</emphasis>
</taxonomicName>
is still relatively common on Oahu, the restricted range suggests that
<taxonomicName id="4C357A95FFC5FFB6F40CD0F0AECDFF22" box="[372,457,188,212]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="moli">
<emphasis id="B941DD04FFC5FFB6F40CD0F0AECDFF22" box="[372,457,188,212]" italics="true" pageId="18" pageNumber="19">D. moli</emphasis>
</taxonomicName>
requires large trees to breed in, and these are increasingly scarce. In recent years it has only been taken at the
<typeStatus id="548EBFB4FFC5FFB6F4ECD0ADAEC0FF0E" box="[404,452,224,248]" pageId="18" pageNumber="19">type</typeStatus>
locality. It has been reported from Hālawa Valley (S.L. Montgomery, pers. comm.), but the specimens could not be located. Other potential sites may exist in the middle and northern Koolau range, but are difficult to access.
</paragraph>
</subSubSection>
</treatment>
</document>