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<mods:title id="ED80CF3EAC446298AF7966D46F7B86B8">Zwicknia gen. n., a new genus for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera)</mods:title>
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<mods:namePart id="269F30416895DEE7CA018CB514A41F13">Murányi, Dávid</mods:namePart>
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<taxonomicName id="4C5059A3FF25FFC21BE45D79D7E6F988" ID-CoL="7GGKF" authority="Muranyi, Orci &amp; Gamboa" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[151,735,1594,1621]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rupprechti" status="sp. nov.">
<emphasis id="B924FE32FF25FFC21BE45D79D4B4F989" bold="true" box="[151,397,1594,1620]" italics="true" pageId="45" pageNumber="46">Zwicknia rupprechti</emphasis>
Murányi, Orci &amp; Gamboa
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<paragraph id="8BEF2220FF25FFC21BE45D1ED047F9AB" blockId="45.[151,1406,1594,1654]" box="[151,1406,1629,1654]" pageId="45" pageNumber="46">
(
<figureCitation id="136B3EA5FF25FFC21BEC5D1ED433F9AB" box="[159,266,1629,1654]" captionStart="FIGURES 7–10" captionStartId="9.[151,264,1084,1107]" captionTargetBox="[151,1436,179,1048]" captionTargetId="figure-317@9.[151,1436,179,1048]" captionTargetPageId="9" captionText="FIGURES 7–10. Male genitalia of Zwicknia Murányi, gen. n.—7, 10: Z. bifrons (Newman); 8–9: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 7: Fp disconnected to the epiproct, latero-caudal view; 8: epiproct tip with contorted Ec, dorsal view; 9: epiproct with partly everted Ec and Fp connected to the Ep-scl, dorsal view; 10: epiproct with everted Ec, dorso-lateral view (B-scl: basal sclerite; Ec: eversible crest; Ep-scl: main epiproct sclerite; Fp: fusion plate; Lb-scl: laterobasal sclerite; Pp: paraproct; Sg: subgenital plate; Tg 9 p: process of tergite 9; Tg 10: tergite 10)—scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919083" httpUri="https://zenodo.org/record/4919083/files/figure.png" pageId="45" pageNumber="46">Figs. 8–9</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC21A655D1ED40DF9AB" box="[278,308,1629,1654]" captionStart="FIGURES 58–61" captionStartId="31.[151,264,1086,1109]" captionTargetBox="[151,1436,193,1065]" captionTargetId="figure-492@31.[151,1436,193,1065]" captionTargetPageId="31" captionText="FIGURES 58–61. Micropterous wings of Zwicknia Murányi, gen. n. males—58: Z. bifrons (Newman, 1838); 59: Z. acuta Murányi &amp; Orci, sp. n.; 60: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 61: Z. sevanica (Zhiltzova, 1964)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919108" httpUri="https://zenodo.org/record/4919108/files/figure.png" pageId="45" pageNumber="46">60</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC21A335D1ED467F9AB" box="[320,350,1629,1654]" captionStart="FIGURES 62–65" captionStartId="33.[151,264,1508,1531]" captionTargetBox="[151,1436,231,1487]" captionTargetId="figure-310@33.[151,1436,193,1487]" captionTargetPageId="33" captionText="FIGURES 62–65. Female genitalia of Zwicknia Murányi, gen. n.—62–63: Z. bifrons (Newman, 1838); 64: Z. acuta Murányi &amp; Orci, sp. n.; 65: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 62: terminalia, ventral view; 63: inner genital organs, dorsal view; 64–65: subgenital plates, ventral view (gc: genital cavity; go: genital opening; Sg: subgenital plate)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919112" httpUri="https://zenodo.org/record/4919112/files/figure.png" pageId="45" pageNumber="46">65</figureCitation>
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<figureCitation id="136B3EA5FF25FFC21A195D1ED4B0F9AB" box="[362,393,1629,1654]" captionStart="FIGURES 66–71" captionStartId="34.[151,264,1663,1686]" captionTargetBox="[151,1436,326,1642]" captionTargetId="figure-254@34.[151,1436,326,1642]" captionTargetPageId="34" captionText="FIGURES 66–71. Larva of Zwicknia Murányi, gen. n.—66: Z. bifrons (Newman, 1838); 67-69, 71: Z. acuta Murányi &amp; Orci, sp. n.; 70: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 66: head, dorso-lateral view; 67: mouthparts, ventral view; 68: pro- and mesothorax, ventral view; 69: meso- and metathorax, ventral view; 70: Tergite 5, dorsal view; 71: female terminalia, ventral view (Es: ecdysial suture; Etn: eutrochantin; Gl: glossa; Lb: labrum; Lp: labial palp; Me: mentum; MeFs: mesothoracic furcasternum; MeFsa: mesothoracic furcasternal arm; MeFsp: mesothoracic furcasternal pit; MeTn: mesothoracic trochantin; MFs: metathoracic furcasternum; Mp: maxillary palp; MTn: metathoracic trochantin; Oc: frontal ocellus; Pc: pedicell; PFs: prothoracic furcasternum; Pg: paraglossa; Pp: paraproct; St 8: Sternite 8)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919114" httpUri="https://zenodo.org/record/4919114/files/figure.png" pageId="45" pageNumber="46">70</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC21AE75D1ED4E4F9AB" box="[404,477,1629,1654]" captionStart="FIGURES 72–77" captionStartId="35.[151,264,1828,1851]" captionTargetBox="[151,1436,483,1807]" captionTargetId="figure-202@35.[151,1436,483,1807]" captionTargetPageId="35" captionText="FIGURES 72–77. Larva of Zwicknia Murányi, gen. n.—72, 74, 76: Z. bifrons (Newman, 1838); 73, 75: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 77: Z. acuta Murányi &amp; Orci, sp. n.; 72-73: hind femur, lateral view; 74-75: male terminalia, lateral view; 76-77: 11-13 cercal segments, lateral view (Pp: paraproct; Tg 10: tergite 10)— scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919116" httpUri="https://zenodo.org/record/4919116/files/figure.png" pageId="45" pageNumber="46">73, 75</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC21A9B5D1ED708F9AB" box="[488,561,1629,1654]" captionStart="FIGURES 90–93" captionStartId="47.[151,264,1938,1961]" captionTargetBox="[257,1337,206,1906]" captionTargetId="figure-60@47.[235,1353,193,1922]" captionTargetPageId="47" captionText="FIGURES 90–93. Male terminalia of Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n.; paratype, Croatia, Krapina- Zagorje County, Ivanščica Mts., Stari Golubovec—90: dorsal view; 91: ventral view; 92: lateral view; 93: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919124" httpUri="https://zenodo.org/record/4919124/files/figure.png" pageId="45" pageNumber="46">90–93</figureCitation>
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<figureCitation id="136B3EA5FF25FFC2194F5D1ED750F9AB" box="[572,617,1629,1654]" captionStart="FIGURES 106–109" captionStartId="54.[151,264,1087,1110]" captionTargetBox="[151,1436,193,1067]" captionTargetId="figure-252@54.[151,1436,193,1067]" captionTargetPageId="54" captionText="FIGURES 106–109. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—106: Z. bifrons (Newman, 1838); 107: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 108–109: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919136" httpUri="https://zenodo.org/record/4919136/files/figure.png" pageId="45" pageNumber="46">107</figureCitation>
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<figureCitation id="136B3EA5FF25FFC219075D1ED7EFF9AB" box="[628,726,1629,1654]" captionStart="FIGURES 114–119" captionStartId="56.[151,265,1540,1563]" captionTargetBox="[151,1436,193,1520]" captionTargetId="figure-203@56.[151,1436,193,1520]" captionTargetPageId="56" captionText="FIGURES 114–119. Dorsal view of male epiproct of Zwicknia Murányi, gen. n.—114: Z. bifrons (Newman, 1838); 115, 117: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 116: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 118: Z. sevanica (Zhiltzova, 1964); 119: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919140" httpUri="https://zenodo.org/record/4919140/files/figure.png" pageId="45" pageNumber="46">115, 117</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC219915D1ED637F9AB" box="[738,782,1629,1654]" captionStart="FIGURES 120–123" captionStartId="57.[151,264,1761,1784]" captionTargetBox="[165,1401,468,1695]" captionTargetId="figure-157@57.[151,1436,457,1720]" captionTargetPageId="57" captionText="FIGURES 120–123. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—120: Z. bifrons (Newman, 1838); 121: Z. acuta Murányi &amp; Orci, sp. n.; 122: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 123: Z. kovacsi Murányi &amp; Gamboa, sp. n.—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919142" httpUri="https://zenodo.org/record/4919142/files/figure.png" pageId="45" pageNumber="46">122</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC218695D1ED646F9AB" box="[794,895,1629,1654]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="45" pageNumber="46">133–138</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC218F85D1ED6C9F9AB" box="[907,1008,1629,1654]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="45" pageNumber="46">157–159</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC218885D1ED11EF9AB" box="[1019,1063,1629,1654]" captionStart="FIGURES 165–168" captionStartId="61.[151,264,1330,1353]" captionTargetBox="[162,1428,857,1282]" captionTargetId="figure-628@61.[151,1436,850,1309]" captionTargetPageId="61" captionText="FIGURES 165–168. Slow oscillograms of male calls of four Zwicknia Murányi, gen. n. species, showing call sequences versus sporadic call production—165: two call sequence of Z. bifrons (Newman, 1838), &quot;//&quot; indicates where 2 minutes 23 seconds of silence was cut out from the recording; 166: three call sequences of Z. acuta Murányi &amp; Orci, sp. n.; 167: sporadic calls of Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 168: sporadic call production in Z. kovacsi Murányi &amp; Gamboa, sp. n.—time scale is the same for the four oscillograms; for detailed data on the animals and recording circumstances, see Table 9." figureDoi="http://doi.org/10.5281/zenodo.4919154" httpUri="https://zenodo.org/record/4919154/files/figure.png" pageId="45" pageNumber="46">167</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC21F405D1ED1AEF9AB" box="[1075,1175,1629,1654]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="45" pageNumber="46">179–182</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC21FD05D1ED03EF9AB" box="[1187,1287,1629,1654]" captionStart-0="FIGURES 185–189" captionStart-1="FIGURE 190" captionStart-2="FIGURE 191" captionStart-3="FIGURE 192" captionStartId-0="63.[151,264,1012,1035]" captionStartId-1="64.[151,250,1125,1147]" captionStartId-2="65.[151,250,1357,1379]" captionStartId-3="66.[151,249,1180,1202]" captionTargetBox-0="[166,1430,178,988]" captionTargetBox-1="[163,1430,185,1044]" captionTargetBox-2="[164,1416,208,1328]" captionTargetBox-3="[156,1424,193,1155]" captionTargetId-0="figure-458@63.[151,1436,178,1004]" captionTargetId-1="figure-392@64.[151,1436,177,1058]" captionTargetId-2="figure-231@65.[151,1436,193,1336]" captionTargetId-3="figure-240@66.[151,1433,185,1171]" captionTargetPageId-0="63" captionTargetPageId-1="64" captionTargetPageId-2="65" captionTargetPageId-3="66" captionText-0="FIGURES 185–189. Male-female drumming duets of three Zwicknia Murányi, gen. n. species.—185: Z. bifrons (Newman, 1838) from Hungary, Pilis Mts., Bükkös Stream, recorded at 16.9 °C; 186–187: Z. acuta Murányi &amp; Orci, sp. n. from Slovakia, Krupinská Planina, Stará Rieka, recorded at 18.2 °C; 188: call-answer duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 18.2 °C; 189: call-answer-reply duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 17.9 °C—arrows with a gender sign point to the beginning of a section of the oscillogram containing the beats of that gender." captionText-1="FIGURE 190. Inter-beat interval variation during male calls in the four examined Zwicknia Murányi, gen. n. species.—Box plot colors: white—Z. bifrons (Newman, 1838), red—Z. acuta Murányi &amp; Orci, sp. n., blue—Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. In Z. kovacsi Murányi &amp; Gamboa, sp. n. we have drumming samples only from two specimens therefore in the case of that species two simple mark series are used to show inter-beat intervals in their calls (circles for specimen 1, and triangles for specimen 2). i1, i2, i3 and so on are labels containing the serial number of inter-beat intervals counting them from the beginning of each call. Box plots show the duration values measured in 7 specimens in Z. bifrons and Z. acuta, and 5 specimens in Z. rupprechti. Vertical axis has a logarithmic scale. Only calls recorded at an ambient air temperature of 17–19 oC were used. (see the measurement data in Appendix Tables 1–4)." captionText-2="FIGURE 191. A scatter plot showing the clear separation of male calls of Zwicknia Murányi, gen. n. along two drumming signal characters—Z. bifrons (Newman, 1838): triangles; Z. acuta Murányi &amp; Orci, sp. n.: circles; Z. kovacsi Murányi &amp; Gamboa, sp. n.: +; Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.: X—Each mark represents the mean values of the characters for a given specimen. Note that both axes have logarithmic scale. Air temperature varied between 15–21 oC in every species (for details see Tables 6–8)." captionText-3="FIGURE 192. Phylogenetic tree of cox1 including Zwicknia Murányi, gen. n. with our accession codes (300941-301003), and GenBank closest species (with alphanumeric accession codes) as outgroup. MP and ML had same topology. Numbers on braches are ML/MP bootstrap values&gt;50%. Species names right to the tree refer to morphological species. P-Distance table is showed." figureDoi-0="http://doi.org/10.5281/zenodo.4919158" figureDoi-1="http://doi.org/10.5281/zenodo.4919160" figureDoi-2="http://doi.org/10.5281/zenodo.4919162" figureDoi-3="http://doi.org/10.5281/zenodo.4919164" httpUri-0="https://zenodo.org/record/4919158/files/figure.png" httpUri-1="https://zenodo.org/record/4919160/files/figure.png" httpUri-2="https://zenodo.org/record/4919162/files/figure.png" httpUri-3="https://zenodo.org/record/4919164/files/figure.png" pageId="45" pageNumber="46">188–192</figureCitation>
, 
<figureCitation id="136B3EA5FF25FFC21E605D1ED04FF9AB" box="[1299,1398,1629,1654]" captionStart-0="FIGURES 193–195" captionStart-1="FIGURES 196–197" captionStartId-0="67.[151,264,1829,1852]" captionStartId-1="68.[151,264,1908,1931]" captionTargetBox-0="[199,1367,231,1794]" captionTargetBox-1="[188,1393,187,1898]" captionTargetId-0="figure-90@67.[199,1367,231,1810]" captionTargetId-1="figure-93@68.[188,1393,187,1898]" captionTargetPageId-0="67" captionTargetPageId-1="68" captionText-0="FIGURES 193–195. Type localities of the new species described in Zwicknia Murányi, gen. n.—193: Zwicknia acuta Murányi &amp; Orci, sp. n.: Stará rieka, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor; 194: Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.: Cimpoies Stream, Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa; 195: Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n.: Völgységi Stream, Hungary, Baranya County, Mecsek Mts., Komló-Zobákpuszta." captionText-1="FIGURES 196–197. Distribution of Zwicknia Murányi, gen. n.—196: global distribution; 197: studied populations of the Carpathian Basin and the northern Balkans—in Fig. 196, white dot on Z. bifrons symbols indicate populations studied by Rupprecht (1997), dotted lines indicate limits of s.l. Z. bifrons literature reports; in Fig. 197, white dot on symbols indicate populations where drumming signals were studied by us, white star indicate type localities." figureDoi-0="http://doi.org/10.5281/zenodo.4919166" figureDoi-1="http://doi.org/10.5281/zenodo.4919168" httpUri-0="https://zenodo.org/record/4919166/files/figure.png" httpUri-1="https://zenodo.org/record/4919168/files/figure.png" pageId="45" pageNumber="46">195–197</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF25FFC21BE45DE6D152F925" pageId="45" pageNumber="46" type="reference_group">
<paragraph id="8BEF2220FF25FFC21BE45DE6D025F966" blockId="45.[151,1436,1701,1785]" box="[151,1308,1701,1723]" pageId="45" pageNumber="46">
<taxonomicName id="4C5059A3FF25FFC21BE45DE6D4E4F966" ID-CoL="QQPJ" authority="(Newman, 1838)" authorityName="Capnor" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[151,477,1701,1723]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF25FFC21BE45DE6D416F966" box="[151,303,1701,1723]" italics="true" pageId="45" pageNumber="46">Capnia bifrons</emphasis>
(
<bibRefCitation id="EFC15FD1FF25FFC21A4F5DE6D4EFF966" author="Newman, E." box="[316,470,1701,1723]" pageId="45" pageNumber="46" pagination="372 - 402" refId="ref44180" refString="Newman, E. (1838) Entomological notes. The Entomological Magazine, 5 (4), 372 - 402." type="journal article" year="1838">Newman, 1838</bibRefCitation>
)
</taxonomicName>
—drumming signals in 
<bibRefCitation id="EFC15FD1FF25FFC219A35DE6D64AF966" author="Rupprecht, R." box="[720,883,1701,1723]" pageId="45" pageNumber="46" pagination="93 - 103" refId="ref45195" refString="Rupprecht, R. (1982) Drumming signals of Danish Plecoptera. Aquatic Insects, 4 (2), 93 - 103. http: // dx. doi. org / 10.1080 / 01650428209361089" type="journal article" year="1982">Rupprecht 1982</bibRefCitation>
(
<figureCitation id="136B3EA5FF25FFC218F35DE6D6F1F966" box="[896,968,1701,1723]" captionStart="FIGURES 1–6" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[193,1399,221,1867]" captionTargetId="figure-86@8.[151,1436,195,1900]" captionTargetPageId="8" captionText="FIGURES 1–6. Male terminalia of West Palaearctic Capniidae—1–2: Zwicknia bifrons (Newman); 3–4: Capnia s.s. nigra (Pictet); 5–6: Capnia s.l. vidua Klapálek; 1, 3, 5: dorso-caudal view; 2, 4, 6: ventro-caudal view (B-scl: basal sclerite; Cs: caudal setae; Ec: eversible crest; Ep-scl: main epiproct sclerite (shaded in grey); I-scl: inner sclerite; Lb-scl: laterobasal sclerite; Ll: lower limb; Tg 10: tergite 10)—not to scale." figureDoi="http://doi.org/10.5281/zenodo.4919081" httpUri="https://zenodo.org/record/4919081/files/figure.png" pageId="45" pageNumber="46">Fig. 2c</figureCitation>
) probably refers to 
<emphasis id="B924FE32FF25FFC21FE35DE5D025F966" box="[1168,1308,1701,1723]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="4C5059A3FF25FFC21FE35DE5D021F966" ID-CoL="7GGKF" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[1168,1304,1701,1723]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rupprechti">Z. rupprechti</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BEF2220FF25FFC21BE45D87D152F925" blockId="45.[151,1436,1701,1785]" pageId="45" pageNumber="46">
<treatmentCitationGroup id="AB40050EFF25FFC21BE45D87D152F925" pageId="45" pageNumber="46">
<taxonomicName id="4C5059A3FF25FFC21BE45D87D714F907" ID-CoL="QQPJ" authority="(Newman, 1838) Capbif" authorityName="Capbif" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[151,557,1732,1754]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF25FFC21BE45D87D416F907" box="[151,303,1732,1754]" italics="true" pageId="45" pageNumber="46">Capnia bifrons</emphasis>
(
<bibRefCitation id="EFC15FD1FF25FFC21A4F5D87D4E3F907" author="Newman, E." box="[316,474,1732,1754]" pageId="45" pageNumber="46" pagination="372 - 402" refId="ref44180" refString="Newman, E. (1838) Entomological notes. The Entomological Magazine, 5 (4), 372 - 402." type="journal article" year="1838">Newman, 1838</bibRefCitation>
) Capbif
</taxonomicName>
<taxonomicName id="4C5059A3FF25FFC219405D86D67AF907" authority="Rupprecht 1997" authorityName="Rupprecht" authorityYear="1997" box="[563,835,1732,1754]" pageId="45" pageNumber="46" rank="subSpecies" subSpecies="sensu">
race sensu 
<bibRefCitation id="EFC15FD1FF25FFC219D05D87D67AF907" author="Rupprecht, R." box="[675,835,1732,1754]" pageId="45" pageNumber="46" pagination="93 - 98" refId="ref45232" refString="Rupprecht, R. (1997) An attempt to explain different drumming signals within Capnia bifrons. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera, Biology-Ecology-Systematics. MTL-Mauron + Tinguely &amp; Lachat SA., Fribourg, pp. 93 - 98." type="book chapter" year="1997">Rupprecht 1997</bibRefCitation>
</taxonomicName>
— 
<treatmentCitation id="0AF10431FF25FFC2182E5D87D115F907" author="Rupprecht, R." box="[861,1068,1732,1754]" page="94" pageId="45" pageNumber="46" year="1997">
<bibRefCitation id="EFC15FD1FF25FFC2182E5D87D115F907" author="Rupprecht, R." box="[861,1068,1732,1754]" pageId="45" pageNumber="46" pagination="93 - 98" refId="ref45232" refString="Rupprecht, R. (1997) An attempt to explain different drumming signals within Capnia bifrons. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera, Biology-Ecology-Systematics. MTL-Mauron + Tinguely &amp; Lachat SA., Fribourg, pp. 93 - 98." type="book chapter" year="1997">Rupprecht 1997: 94</bibRefCitation>
</treatmentCitation>
. (drumming signals, probably refers to 
<taxonomicName id="4C5059A3FF25FFC21B925DA7D45CF924" ID-CoL="7GGKF" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[225,357,1763,1785]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF25FFC21B925DA7D45CF924" box="[225,357,1763,1785]" italics="true" pageId="45" pageNumber="46">Z. rupprechti</emphasis>
</taxonomicName>
); 
<treatmentCitation id="0AF10431FF25FFC21A085DA0D790F924" author="Enting, K. &amp; Rupprecht, R." box="[379,681,1763,1785]" page="71" pageId="45" pageNumber="46" year="2001">
<bibRefCitation id="EFC15FD1FF25FFC21A085DA0D790F924" author="Enting, K. &amp; Rupprecht, R." box="[379,681,1763,1785]" pageId="45" pageNumber="46" pagination="63 - 77" refId="ref41636" refString="Enting, K. &amp; Rupprecht, R. (2001) Zur Kenntnis der Steinfliegenfauna (Insecta: Plecoptera) im Taunus. Lauterbornia, 41, 63 - 77." type="journal article" year="2001">Enting &amp; Rupprecht 2001: 71</bibRefCitation>
</treatmentCitation>
(clarification as not the nominal 
<taxonomicName id="4C5059A3FF25FFC2188B5DA0D165F924" ID-CoL="QQPJ" authorityName="Capbif" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[1016,1116,1763,1785]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF25FFC2188B5DA0D165F924" box="[1016,1116,1763,1785]" italics="true" pageId="45" pageNumber="46">C. bifrons</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF25FFC11BE45C66D443FF25" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="8BEF2220FF25FFC11BE45C66D443FF25" blockId="45.[151,1437,1829,2034]" lastBlockId="46.[151,1437,151,2013]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<emphasis id="B924FE32FF25FFC21BE45C66D428F8E3" bold="true" box="[151,273,1829,1854]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
Male epiproct: Ep-scl wide and blunt in dorsal view, tip upcurved in lateral view; ventral membranous section nearly reach the base in lateral view, apical spines distributed only on the membranous part of the apex. Process of male Tg 9: very low, caudally projecting, &lt;1½× wider than Ep-scl, rounded and with continuously narrow sides caudally. Male drumming calls are short beat groups with 60–80 ms duration and including 4–6 percussive beats produced with a mean inter beat interval of 17–23 ms. Inter-beat intervals gradually decrease, while the amplitude of beats noticeably increase during a call. Sometimes two calls follow each other with an inter- call interval of 1.5–5.5 seconds, but generally single calls are produced sporadically. The male-female drumming duet consists of a single male call—female answer or male call—female answer—male response signal exchanges as in 
<figureCitation id="136B3EA5FF26FFC11BA15B9CD44FFF25" box="[210,374,223,248]" captionStart="FIGURES 185–189" captionStartId="63.[151,264,1012,1035]" captionTargetBox="[166,1430,178,988]" captionTargetId="figure-458@63.[151,1436,178,1004]" captionTargetPageId="63" captionText="FIGURES 185–189. Male-female drumming duets of three Zwicknia Murányi, gen. n. species.—185: Z. bifrons (Newman, 1838) from Hungary, Pilis Mts., Bükkös Stream, recorded at 16.9 °C; 186–187: Z. acuta Murányi &amp; Orci, sp. n. from Slovakia, Krupinská Planina, Stará Rieka, recorded at 18.2 °C; 188: call-answer duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 18.2 °C; 189: call-answer-reply duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 17.9 °C—arrows with a gender sign point to the beginning of a section of the oscillogram containing the beats of that gender." figureDoi="http://doi.org/10.5281/zenodo.4919158" httpUri="https://zenodo.org/record/4919158/files/figure.png" pageId="46" pageNumber="47">Figs. 188–189</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF26FFC11BB45A47D781FB85" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BEF2220FF26FFC11BB45A47D4AFFC5D" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
<emphasis id="B924FE32FF26FFC11BB45A47D44FFEC0" bold="true" box="[199,374,260,285]" pageId="46" pageNumber="47">Type material.</emphasis>
<materialsCitation id="3B38287DFF26FFC11A0D5A47D766FEB8" ID-GBIF-Occurrence="3325453325" collectionCode="HNHM" collectorName="K. M. Orci" country="Hungary" county="Mecsek Mts." elevation="305" latitude="46.194035" location="Volgysegi Stream" longLatPrecision="1" longitude="18.31785" municipality="Komlo-Zobakpuszta" pageId="46" pageNumber="47" specimenCode="PLH1277" specimenCount="1" stateProvince="Baranya" typeStatus="holotype">
<typeStatus id="54EB9C82FF26FFC11A0D5A47D4DFFEC0" box="[382,486,260,285]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
male: 
<collectingCountry id="F34762B0FF26FFC119465A47D7FBFEC0" box="[565,706,260,285]" name="Hungary" pageId="46" pageNumber="47">HUNGARY</collectingCountry>
: 
<collectingRegion id="4994ECC2FF26FFC119BE5A47D6BFFEC0" box="[717,902,260,285]" country="Hungary" name="Baranya" pageId="46" pageNumber="47">Baranya County</collectingRegion>
, 
<collectingCounty id="628E5AACFF26FFC118E15A47D11EFEC6" box="[914,1063,260,285]" pageId="46" pageNumber="47">Mecsek Mts.</collectingCounty>
, 
<collectingMunicipality id="6B8BB85AFF26FFC11F405A47D025FEC0" box="[1075,1308,260,285]" pageId="46" pageNumber="47">Komló-Zobákpuszta</collectingMunicipality>
, 
<location id="8E8F74FBFF26FFC11E5A5A47D5D0FE9D" LSID="urn:lsid:plazi:treatment:03F99336FF25FFDF1BE45D79D688F990:8E8F74FBFF26FFC11E5A5A47D5D0FE9D" country="Hungary" county="Mecsek Mts." latitude="46.194035" longLatPrecision="1" longitude="18.31785" municipality="Komlo-Zobakpuszta" name="Volgysegi Stream" pageId="46" pageNumber="47" stateProvince="Baranya">Völgységi Stream</location>
, 
<geoCoordinate id="EE6444E7FF26FFC11B855A6BD4A9FE9D" box="[246,400,295,320]" degrees="46" direction="north" minutes="11.642" orientation="latitude" pageId="46" pageNumber="47" precision="1" value="46.194035">N 46°11.642’</geoCoordinate>
<geoCoordinate id="EE6444E7FF26FFC11AE35A64D715FE9D" box="[400,556,295,320]" degrees="18" direction="east" minutes="19.071" orientation="longitude" pageId="46" pageNumber="47" precision="1" value="18.31785">E 18°19.071’</geoCoordinate>
, 
<elevation id="007DC513FF26FFC119485A64D7FDFE9D" box="[571,708,295,320]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="46" pageNumber="47" unit="m" value="305.0">
<quantity id="4CA88FC5FF26FFC119485A64D7BCFEE2" box="[571,645,295,320]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="46" pageNumber="47" unit="m" value="305.0">305 m</quantity>
a.s.l.
</elevation>
, 09.03.201 0, leg. D. 
<taxonomicName id="4C5059A3FF26FFC118C85A64D127FE9D" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[955,1054,295,320]" genus="Muranyi" pageId="46" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
, 
<collectorName id="26A547F6FF26FFC11F5F5A6BD194FE9D" box="[1068,1197,295,320]" pageId="46" pageNumber="47">K. M. Orci</collectorName>
(
<collectionCode id="ED41BAE5FF26FFC11FCC5A64D023FE9D" box="[1215,1306,295,320]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC11E5B5A64D0A1FE9D" box="[1320,1432,295,320]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1277</specimenCode>
; drumming recorded as duett 2010/No.6)
</materialsCitation>
. 
<materialsCitation id="3B38287DFF26FFC119195A0FD6CDFDE0" ID-GBIF-Occurrence="3325453372" collectionCode="HNHM, BYUC" collectorName="K. M. Orci" country="Hungary" county="Mecsek Mts." elevation="0" latitude="46.194035" location="Volgysegi Stream" longLatPrecision="1" longitude="18.31785" municipality="Komlo-Zobakpuszta" pageId="46" pageNumber="47" specimenCode="PLH1184, PLH1274, PLH1275, PLH1276, PLH1278, PLH1279, PLH1280" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Baranya" typeStatus="paratype">
<typeStatus id="54EB9C82FF26FFC119195A0FD7E2FEB9" box="[618,731,332,356]" pageId="46" pageNumber="47" type="paratype">Paratypes</typeStatus>
: same locality and date: 
<elevation id="007DC513FF26FFC118825A0FD171FEB8" box="[1009,1096,332,357]" isEnumeration="true" pageId="46" pageNumber="47">
<quantity id="4CA88FC5FF26FFC118825A0FD12EFEB8" box="[1009,1047,332,357]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="46" pageNumber="47" unit="m" value="6.0">6m</quantity>
, 
<quantity id="4CA88FC5FF26FFC11F575A0FD171FEB8" box="[1060,1096,332,357]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="m" value="2.0">2m</quantity>
</elevation>
2f larvae (
<collectionCode id="ED41BAE5FF26FFC11FB15A0FD027FEB8" box="[1218,1310,332,357]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC11E5A5A0FD0A1FEB8" box="[1321,1432,332,357]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1184</specimenCode>
; 
<specimenCount id="9D56E9A9FF26FFC11BE45A2CD435FE55" box="[151,268,367,392]" pageId="46" pageNumber="47" type="male">three male</specimenCount>
terminalias, 
<specimenCount id="9D56E9A9FF26FFC11AED5A33D723FE55" box="[414,538,367,392]" pageId="46" pageNumber="47" type="female">one female</specimenCount>
and 
<specimenCount id="9D56E9A9FF26FFC119225A33D78FFE55" box="[593,694,367,392]" pageId="46" pageNumber="47" type="male">one male</specimenCount>
larva prepared for SEM, specimens used for drawings and photos 
<figureCitation id="136B3EA5FF26FFC11BE45AD7D5DCFE70" box="[151,229,404,429]" captionStart="FIGURES 7–10" captionStartId="9.[151,264,1084,1107]" captionTargetBox="[151,1436,179,1048]" captionTargetId="figure-317@9.[151,1436,179,1048]" captionTargetPageId="9" captionText="FIGURES 7–10. Male genitalia of Zwicknia Murányi, gen. n.—7, 10: Z. bifrons (Newman); 8–9: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 7: Fp disconnected to the epiproct, latero-caudal view; 8: epiproct tip with contorted Ec, dorsal view; 9: epiproct with partly everted Ec and Fp connected to the Ep-scl, dorsal view; 10: epiproct with everted Ec, dorso-lateral view (B-scl: basal sclerite; Ec: eversible crest; Ep-scl: main epiproct sclerite; Fp: fusion plate; Lb-scl: laterobasal sclerite; Pp: paraproct; Sg: subgenital plate; Tg 9 p: process of tergite 9; Tg 10: tergite 10)—scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919083" httpUri="https://zenodo.org/record/4919083/files/figure.png" pageId="46" pageNumber="47">Figs. 8</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11B825AD7D436FE70" box="[241,271,404,429]" captionStart="FIGURES 66–71" captionStartId="34.[151,264,1663,1686]" captionTargetBox="[151,1436,326,1642]" captionTargetId="figure-254@34.[151,1436,326,1642]" captionTargetPageId="34" captionText="FIGURES 66–71. Larva of Zwicknia Murányi, gen. n.—66: Z. bifrons (Newman, 1838); 67-69, 71: Z. acuta Murányi &amp; Orci, sp. n.; 70: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 66: head, dorso-lateral view; 67: mouthparts, ventral view; 68: pro- and mesothorax, ventral view; 69: meso- and metathorax, ventral view; 70: Tergite 5, dorsal view; 71: female terminalia, ventral view (Es: ecdysial suture; Etn: eutrochantin; Gl: glossa; Lb: labrum; Lp: labial palp; Me: mentum; MeFs: mesothoracic furcasternum; MeFsa: mesothoracic furcasternal arm; MeFsp: mesothoracic furcasternal pit; MeTn: mesothoracic trochantin; MFs: metathoracic furcasternum; Mp: maxillary palp; MTn: metathoracic trochantin; Oc: frontal ocellus; Pc: pedicell; PFs: prothoracic furcasternum; Pg: paraglossa; Pp: paraproct; St 8: Sternite 8)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919114" httpUri="https://zenodo.org/record/4919114/files/figure.png" pageId="46" pageNumber="47">70</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11A695AD7D45BFE70" box="[282,354,404,429]" captionStart="FIGURES 72–77" captionStartId="35.[151,264,1828,1851]" captionTargetBox="[151,1436,483,1807]" captionTargetId="figure-202@35.[151,1436,483,1807]" captionTargetPageId="35" captionText="FIGURES 72–77. Larva of Zwicknia Murányi, gen. n.—72, 74, 76: Z. bifrons (Newman, 1838); 73, 75: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 77: Z. acuta Murányi &amp; Orci, sp. n.; 72-73: hind femur, lateral view; 74-75: male terminalia, lateral view; 76-77: 11-13 cercal segments, lateral view (Pp: paraproct; Tg 10: tergite 10)— scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919116" httpUri="https://zenodo.org/record/4919116/files/figure.png" pageId="46" pageNumber="47">73, 75</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11A1D5AD7D4A0FE70" box="[366,409,404,429]" captionStart="FIGURES 114–119" captionStartId="56.[151,265,1540,1563]" captionTargetBox="[151,1436,193,1520]" captionTargetId="figure-203@56.[151,1436,193,1520]" captionTargetPageId="56" captionText="FIGURES 114–119. Dorsal view of male epiproct of Zwicknia Murányi, gen. n.—114: Z. bifrons (Newman, 1838); 115, 117: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 116: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 118: Z. sevanica (Zhiltzova, 1964); 119: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919140" httpUri="https://zenodo.org/record/4919140/files/figure.png" pageId="46" pageNumber="47">115</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11AD65AD7D730FE70" box="[421,521,404,429]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="46" pageNumber="47">135–136</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC119665AD7D77BFE70" box="[533,578,404,429]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="46" pageNumber="47">158</figureCitation>
), 
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</quantity>
1f (
<collectionCode id="ED41BAE5FF26FFC119D55AD7D63BFE70" box="[678,770,404,429]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC1187F5AD7D645FE70" box="[780,892,404,429]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1274</specimenCode>
; drumming recorded as 2010/No.8), 1f (
<collectionCode id="ED41BAE5FF26FFC11E4E5AD7D0A0FE70" box="[1341,1433,404,429]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC11BE45AF4D431FE0D" box="[151,264,439,464]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1275</specimenCode>
; drumming recorded as 2010/No.2, male escaped), 
<quantity id="4CA88FC5FF26FFC118365AF4D650FE0D" box="[837,873,439,464]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">
<elevation id="007DC513FF26FFC118365AF4D650FE0D" box="[837,873,439,464]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">1m</elevation>
</quantity>
1f (
<collectionCode id="ED41BAE5FF26FFC118E55AF4D6C8FE0D" box="[918,1009,439,464]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC118885AF4D155FE0D" box="[1019,1132,439,464]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1276</specimenCode>
; used for molecular studies as 300970, 300971, drummings recorded as 2010/No.1), 1f (
<collectionCode id="ED41BAE5FF26FFC118495A9FD6ACFE28" box="[826,917,476,501]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC118EC5A9FD129FE28" box="[927,1040,476,501]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1278</specimenCode>
; drumming recorded as duett 2010/ No.6, pair of the 
<typeStatus id="54EB9C82FF26FFC11A1D5ABCD4EDFDC5" box="[366,468,511,536]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
), 
<quantity id="4CA88FC5FF26FFC11A9E5ABCD728FDC5" box="[493,529,511,536]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">
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</quantity>
1f (
<collectionCode id="ED41BAE5FF26FFC119395ABCD79FFDC5" box="[586,678,511,536]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC119C55ABCD61FFDC5" box="[694,806,511,536]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1279</specimenCode>
; drumming recorded as 2010/No.5), 
<quantity id="4CA88FC5FF26FFC11F935ABCD03DFDC5" box="[1248,1284,511,536]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">
<elevation id="007DC513FF26FFC11F935ABCD03DFDC5" box="[1248,1284,511,536]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">1m</elevation>
</quantity>
1f (
<collectionCode id="ED41BAE5FF26FFC11E4C5943D0A0FDCA" box="[1343,1433,512,535]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC11BE45967D43EFDE0" box="[151,263,548,573]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1280</specimenCode>
; drumming recorded as 2010/No.7, 
<figureCitation id="136B3EA5FF26FFC119D45967D674FDE0" box="[679,845,548,573]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="46" pageNumber="47">Figs. 179–180</figureCitation>
), 
<quantity id="4CA88FC5FF26FFC118175967D6B1FDE0" box="[868,904,548,573]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="46" pageNumber="47" unit="m" value="3.0">
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</quantity>
(
<collectionCode id="ED41BAE5FF26FFC118EA5967D6D5FDE0" box="[921,1004,548,573]" pageId="46" pageNumber="47">BYUC</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B38287DFF26FFC11F715967D42BFC5D" ID-GBIF-Occurrence="3325453370" collectionCode="HNHM, GVC" collectorName="L. Czigany &amp; Czigany, D." country="Croatia" county="Ivanscica Mts." elevation="1" latitude="46.175667" location="Reka Stream" longLatPrecision="11" longitude="16.049833" municipality="Stari Golubovec" pageId="46" pageNumber="47" specimenCode="PLH1297, PLH1298, PLH1299, PLP3401, PLP3878, PLP3879, PLP3880" specimenCount="1" specimenCount-male="1" stateProvince="Krapina-Zagorje County" typeStatus="paratype">
<quantity id="4CA88FC5FF26FFC11F715967D11FFDE0" box="[1026,1062,548,573]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">
<elevation id="007DC513FF26FFC11F715967D11FFDE0" box="[1026,1062,548,573]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">1m</elevation>
</quantity>
1f (
<collectionCode id="ED41BAE5FF26FFC11F2B5967D18DFDE0" box="[1112,1204,548,573]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC11FB35967D008FDE0" box="[1216,1329,548,573]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1297</specimenCode>
; used for molecular studies as 300948, 300949, drummings recorded as 2010/No.3), 
<quantity id="4CA88FC5FF26FFC1188D5904D11BFDBD" box="[1022,1058,583,608]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">
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</quantity>
1f (
<collectionCode id="ED41BAE5FF26FFC11F255904D18BFDBD" box="[1110,1202,583,608]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC11FCC5904D009FDBD" box="[1215,1328,583,608]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1298</specimenCode>
; used for molecular studies as 300963, 300964, drummings recorded as 2010/No.4), 1f (
<collectionCode id="ED41BAE5FF26FFC11F38592FD19EFD58" box="[1099,1191,620,645]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC11FCB592FD011FD58" box="[1208,1320,620,645]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLH1299</specimenCode>
; used for molecular studies as 300967, drumming recorded as 2010/No.9); 
<collectingCountry id="F34762B0FF26FFC118F759CCD13BFD75" box="[900,1026,655,680]" name="Croatia" pageId="46" pageNumber="47">CROATIA</collectingCountry>
: 
<collectingRegion id="4994ECC2FF26FFC11F7D59CCD01FFD75" box="[1038,1318,655,680]" inferredBySuffix="true" pageId="46" pageNumber="47">Krapina-Zagorje County</collectingRegion>
, 
<collectingCounty id="628E5AACFF26FFC11E4159CCD5F5FD16" pageId="46" pageNumber="47">Ivanščica Mts.</collectingCounty>
, 
<collectingMunicipality id="6B8BB85AFF26FFC11BAF59F7D4A3FD10" box="[220,410,692,717]" pageId="46" pageNumber="47">Stari Golubovec</collectingMunicipality>
, 
<location id="8E8F74FBFF26FFC11AD959F7D707FD10" LSID="urn:lsid:plazi:treatment:03F99336FF25FFDF1BE45D79D688F990:8E8F74FBFF26FFC11AD959F7D707FD10" box="[426,574,692,717]" country="Croatia" county="Ivanscica Mts." latitude="46.175667" longLatPrecision="11" longitude="16.049833" municipality="Stari Golubovec" name="Reka Stream" pageId="46" pageNumber="47" stateProvince="Krapina-Zagorje County">Reka Stream</location>
below the village, 
<geoCoordinate id="EE6444E7FF26FFC1185559F7D68CFD10" box="[806,949,692,717]" degrees="46" direction="north" minutes="10.54" orientation="latitude" pageId="46" pageNumber="47" precision="9" value="46.175667">N 46°10.54’</geoCoordinate>
<geoCoordinate id="EE6444E7FF26FFC118B159F7D168FD10" box="[962,1105,692,717]" degrees="16" direction="east" minutes="02.99" orientation="longitude" pageId="46" pageNumber="47" precision="9" value="16.049833">E 16°02.99’</geoCoordinate>
, 
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<elevation id="007DC513FF26FFC11F1359F7D197FD16" box="[1120,1198,692,717]" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="46" pageNumber="47" unit="m" value="345.0">345 m</elevation>
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, 06.04.201 0, leg. 
<collectorName id="26A547F6FF26FFC11EF759F7D5CCFD2D" pageId="46" pageNumber="47">L. Czigány</collectorName>
, D. 
<taxonomicName id="4C5059A3FF26FFC11A585994D4A9FD2D" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[299,400,727,752]" genus="Muranyi" pageId="46" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
: 
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<elevation id="007DC513FF26FFC11AEC599BD4FAFD32" box="[415,451,728,751]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="46" pageNumber="47" unit="m" value="7.0">7m</elevation>
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(
<collectionCode id="ED41BAE5FF26FFC11AA55994D70BFD2D" box="[470,562,727,752]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC119335994D795FD2D" box="[576,684,727,752]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLP3401</specimenCode>
; 
<specimenCount id="9D56E9A9FF26FFC119CF599BD61CFD2D" box="[700,805,727,752]" pageId="46" pageNumber="47" type="male">one male</specimenCount>
prepared for SEM, specimens used for drawings and photos 
<figureCitation id="136B3EA5FF26FFC11B9959BFD47EFCC8" box="[234,327,764,789]" captionStart="FIGURES 58–61" captionStartId="31.[151,264,1086,1109]" captionTargetBox="[151,1436,193,1065]" captionTargetId="figure-492@31.[151,1436,193,1065]" captionTargetPageId="31" captionText="FIGURES 58–61. Micropterous wings of Zwicknia Murányi, gen. n. males—58: Z. bifrons (Newman, 1838); 59: Z. acuta Murányi &amp; Orci, sp. n.; 60: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 61: Z. sevanica (Zhiltzova, 1964)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919108" httpUri="https://zenodo.org/record/4919108/files/figure.png" pageId="46" pageNumber="47">Figs. 60</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11A2759BFD4A4FCC8" box="[340,413,764,789]" captionStart="FIGURES 90–93" captionStartId="47.[151,264,1938,1961]" captionTargetBox="[257,1337,206,1906]" captionTargetId="figure-60@47.[235,1353,193,1922]" captionTargetPageId="47" captionText="FIGURES 90–93. Male terminalia of Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n.; paratype, Croatia, Krapina- Zagorje County, Ivanščica Mts., Stari Golubovec—90: dorsal view; 91: ventral view; 92: lateral view; 93: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919124" httpUri="https://zenodo.org/record/4919124/files/figure.png" pageId="46" pageNumber="47">90–93</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11ADA59BFD4ECFCC8" box="[425,469,764,789]" captionStart="FIGURES 120–123" captionStartId="57.[151,264,1761,1784]" captionTargetBox="[165,1401,468,1695]" captionTargetId="figure-157@57.[151,1436,457,1720]" captionTargetPageId="57" captionText="FIGURES 120–123. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—120: Z. bifrons (Newman, 1838); 121: Z. acuta Murányi &amp; Orci, sp. n.; 122: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 123: Z. kovacsi Murányi &amp; Gamboa, sp. n.—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919142" httpUri="https://zenodo.org/record/4919142/files/figure.png" pageId="46" pageNumber="47">122</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11A9159BFD781FCC8" box="[482,696,764,789]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="46" pageNumber="47">133–134, 137–138</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC119B759BFD612FCC8" box="[708,811,764,789]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="46" pageNumber="47">157, 159</figureCitation>
), 
<quantity id="4CA88FC5FF26FFC1183359BFD65DFCC8" box="[832,868,764,789]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">
<elevation id="007DC513FF26FFC1183359BFD65DFCC8" box="[832,868,764,789]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">1m</elevation>
</quantity>
1f (
<collectionCode id="ED41BAE5FF26FFC118E059BFD6D7FCC8" box="[915,1006,764,789]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC1188959BFD15FFCC8" box="[1018,1126,764,789]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLP3878</specimenCode>
; used for molecular studies as 300973, 300974, drummings recorded as 2010/No.2, 
<figureCitation id="136B3EA5FF26FFC11874585CD692FCE5" box="[775,939,799,824]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="46" pageNumber="47">Figs. 181–182</figureCitation>
), 
<quantity id="4CA88FC5FF26FFC118CD585CD6DBFCE5" box="[958,994,799,824]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">
<elevation id="007DC513FF26FFC118CD585CD6DBFCE5" box="[958,994,799,824]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">1m</elevation>
</quantity>
(
<collectionCode id="ED41BAE5FF26FFC11883585CD172FCE5" box="[1008,1099,799,824]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC11F26585CD1F8FCE5" box="[1109,1217,799,824]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLP3879</specimenCode>
; used for molecular studies as 300968, drumming recorded as 2010/No.1), 
<quantity id="4CA88FC5FF26FFC118735807D61DFC80" box="[768,804,836,861]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">
<elevation id="007DC513FF26FFC118735807D61DFC80" box="[768,804,836,861]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">1m</elevation>
</quantity>
(
<collectionCode id="ED41BAE5FF26FFC118405807D6B6FC80" box="[819,911,836,861]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
: 
<specimenCode id="DBF68A5BFF26FFC118EA5807D13CFC80" box="[921,1029,836,861]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">PLP3880</specimenCode>
; drumming recorded as 2010/No.3), 
<quantity id="4CA88FC5FF26FFC11BE45824D582FC5D" box="[151,187,871,896]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="m" value="2.0">
<elevation id="007DC513FF26FFC11BE45824D582FC5D" box="[151,187,871,896]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="m" value="2.0">2m</elevation>
</quantity>
(
<collectionCode id="ED41BAE5FF26FFC11BB95824D433FC5D" box="[202,266,871,896]" pageId="46" pageNumber="47">GVC</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B38287DFF26FFC11A6E5824D4ABFC5D" ID-GBIF-Occurrence="3325453377" box="[285,402,871,896]" collectionCode="PZC" collectorName="L. Czigany &amp; Czigany, D." country="Croatia" county="Ivanscica Mts." elevation="2" isTailEnumeration="true" latitude="46.175667" location="Reka Stream" longLatPrecision="11" longitude="16.049833" municipality="Stari Golubovec" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Krapina-Zagorje County" typeStatus="paratype">
<quantity id="4CA88FC5FF26FFC11A6E5824D478FC5D" box="[285,321,871,896]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="m" value="2.0">
<elevation id="007DC513FF26FFC11A6E5824D478FC5D" box="[285,321,871,896]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="m" value="2.0">2m</elevation>
</quantity>
(
<collectionCode id="ED41BAE5FF26FFC11A235824D4B0FC5D" box="[336,393,871,896]" pageId="46" pageNumber="47">PZC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF2220FF26FFC11BB458CFD781FB85" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
<emphasis id="B924FE32FF26FFC11BB458CFD61AFC78" bold="true" box="[199,803,908,933]" pageId="46" pageNumber="47">Other material—Records based on morphology:</emphasis>
<materialsCitation id="3B38287DFF26FFC1184358CFD1ACFC30" ID-GBIF-Occurrence="3325453320" collectionCode="HNHM" collectorName="J. Kontschan" country="Croatia" county="Ivanscica Mts." elevation="330" latitude="46.15" location="Reka Stream" longLatPrecision="1125" longitude="16.083334" municipality="Lobor" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Krapina-Zagorje County">
<collectingCountry id="F34762B0FF26FFC1184358CFD697FC78" box="[816,942,908,933]" name="Croatia" pageId="46" pageNumber="47">CROATIA</collectingCountry>
: 
<collectingRegion id="4994ECC2FF26FFC118CD58CFD1E3FC78" box="[958,1242,908,933]" inferredBySuffix="true" pageId="46" pageNumber="47">Krapina-Zagorje County</collectingRegion>
, 
<collectingCounty id="628E5AACFF26FFC11F9858CFD0A1FC7E" box="[1259,1432,908,933]" pageId="46" pageNumber="47">Ivanščica Mts.</collectingCounty>
, 
<collectingMunicipality id="6B8BB85AFF26FFC11BE458ECD5E4FC15" box="[151,221,943,968]" pageId="46" pageNumber="47">Lobor</collectingMunicipality>
, 
<location id="8E8F74FBFF26FFC11B9A58ECD440FC15" LSID="urn:lsid:plazi:treatment:03F99336FF25FFDF1BE45D79D688F990:8E8F74FBFF26FFC11B9A58ECD440FC15" box="[233,377,943,968]" country="Croatia" county="Ivanscica Mts." latitude="46.15" longLatPrecision="1125" longitude="16.083334" municipality="Lobor" name="Reka Stream" pageId="46" pageNumber="47" stateProvince="Krapina-Zagorje County">Reka Stream</location>
above the village, 
<geoCoordinate id="EE6444E7FF26FFC1193C58F3D781FC15" box="[591,696,943,968]" degrees="46" direction="north" minutes="09" orientation="latitude" pageId="46" pageNumber="47" precision="925" value="46.15">N 46°09’</geoCoordinate>
<geoCoordinate id="EE6444E7FF26FFC119CC58F3D61FFC15" box="[703,806,943,968]" degrees="16" direction="east" minutes="05" orientation="longitude" pageId="46" pageNumber="47" precision="925" value="16.083334">E 16°05’</geoCoordinate>
, 
<quantity id="4CA88FC5FF26FFC1184058ECD645FC1A" box="[819,892,943,968]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="46" pageNumber="47" unit="m" value="330.0">
<elevation id="007DC513FF26FFC1184058ECD645FC1A" box="[819,892,943,968]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="46" pageNumber="47" unit="m" value="330.0">330 m</elevation>
</quantity>
, 01.04.200 6, leg. 
<collectorName id="26A547F6FF26FFC11F3558F3D1E2FC15" box="[1094,1243,943,968]" pageId="46" pageNumber="47">J. Kontschán</collectorName>
, D. 
<taxonomicName id="4C5059A3FF26FFC11E7A58ECD057FC15" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[1289,1390,943,968]" genus="Muranyi" pageId="46" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
: 
<quantity id="4CA88FC5FF26FFC11E0B58ECD0A5FC15" box="[1400,1436,943,968]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="m" value="5.0">
<elevation id="007DC513FF26FFC11E0B58ECD0A5FC15" box="[1400,1436,943,968]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="m" value="5.0">5m</elevation>
</quantity>
1f (
<collectionCode id="ED41BAE5FF26FFC11BB25897D425FC30" box="[193,284,980,1005]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
; 
<specimenCount id="9D56E9A9FF26FFC11A585897D4ADFC30" box="[299,404,980,1005]" pageId="46" pageNumber="47" type="male">one male</specimenCount>
terminalia prepared for SEM, used for photos 
<figureCitation id="136B3EA5FF26FFC118B35897D12AFC30" box="[960,1043,980,1005]" captionStart="FIGURES 7–10" captionStartId="9.[151,264,1084,1107]" captionTargetBox="[151,1436,179,1048]" captionTargetId="figure-317@9.[151,1436,179,1048]" captionTargetPageId="9" captionText="FIGURES 7–10. Male genitalia of Zwicknia Murányi, gen. n.—7, 10: Z. bifrons (Newman); 8–9: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 7: Fp disconnected to the epiproct, latero-caudal view; 8: epiproct tip with contorted Ec, dorsal view; 9: epiproct with partly everted Ec and Fp connected to the Ep-scl, dorsal view; 10: epiproct with everted Ec, dorso-lateral view (B-scl: basal sclerite; Ec: eversible crest; Ep-scl: main epiproct sclerite; Fp: fusion plate; Lb-scl: laterobasal sclerite; Pp: paraproct; Sg: subgenital plate; Tg 9 p: process of tergite 9; Tg 10: tergite 10)—scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919083" httpUri="https://zenodo.org/record/4919083/files/figure.png" pageId="46" pageNumber="47">Figs. 9</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11F505897D176FC30" box="[1059,1103,980,1005]" captionStart="FIGURES 106–109" captionStartId="54.[151,264,1087,1110]" captionTargetBox="[151,1436,193,1067]" captionTargetId="figure-252@54.[151,1436,193,1067]" captionTargetPageId="54" captionText="FIGURES 106–109. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—106: Z. bifrons (Newman, 1838); 107: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 108–109: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919136" httpUri="https://zenodo.org/record/4919136/files/figure.png" pageId="46" pageNumber="47">107</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11F2C5897D1B5FC30" box="[1119,1164,980,1005]" captionStart="FIGURES 114–119" captionStartId="56.[151,265,1540,1563]" captionTargetBox="[151,1436,193,1520]" captionTargetId="figure-203@56.[151,1436,193,1520]" captionTargetPageId="56" captionText="FIGURES 114–119. Dorsal view of male epiproct of Zwicknia Murányi, gen. n.—114: Z. bifrons (Newman, 1838); 115, 117: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 116: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 118: Z. sevanica (Zhiltzova, 1964); 119: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919140" httpUri="https://zenodo.org/record/4919140/files/figure.png" pageId="46" pageNumber="47">117</figureCitation>
)
</materialsCitation>
; 
<materialsCitation id="3B38287DFF26FFC11FD75897D00AFBCD" ID-GBIF-Occurrence="3325453386" collectionCode="HNHM" country="Hungary" county="Mecsek Mts." elevation="360" location="Hidas Valley" municipality="Komlo" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Baranya">
<collectingCountry id="F34762B0FF26FFC11FD75897D008FC30" box="[1188,1329,980,1005]" name="Hungary" pageId="46" pageNumber="47">HUNGARY</collectingCountry>
: 
<collectingRegion id="4994ECC2FF26FFC11E4C5897D5D2FBCD" country="Hungary" name="Baranya" pageId="46" pageNumber="47">Baranya County</collectingRegion>
, 
<collectingCounty id="628E5AACFF26FFC11B8558B4D4B3FBD2" box="[246,394,1015,1040]" pageId="46" pageNumber="47">Mecsek Mts.</collectingCounty>
, 
<collectingMunicipality id="6B8BB85AFF26FFC11AE558B4D4DEFBCD" box="[406,487,1015,1040]" pageId="46" pageNumber="47">Komló</collectingMunicipality>
, 
<location id="8E8F74FBFF26FFC11A8058B4D7BCFBCD" LSID="urn:lsid:plazi:treatment:03F99336FF25FFDF1BE45D79D688F990:8E8F74FBFF26FFC11A8058B4D7BCFBCD" box="[499,645,1015,1040]" country="Hungary" county="Mecsek Mts." municipality="Komlo" name="Hidas Valley" pageId="46" pageNumber="47" stateProvince="Baranya">Hidas Valley</location>
, 
<quantity id="4CA88FC5FF26FFC119E358B4D7E0FBD2" box="[656,729,1015,1040]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="46" pageNumber="47" unit="m" value="360.0">
<elevation id="007DC513FF26FFC119E358B4D7E0FBD2" box="[656,729,1015,1040]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="46" pageNumber="47" unit="m" value="360.0">360 m</elevation>
</quantity>
, 28.02.200 4, leg. D. 
<taxonomicName id="4C5059A3FF26FFC118B558B4D112FBCD" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[966,1067,1015,1040]" genus="Muranyi" pageId="46" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
: 
<quantity id="4CA88FC5FF26FFC11F4558B4D165FBCD" box="[1078,1116,1015,1040]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="46" pageNumber="47" unit="m" value="3.0">
<elevation id="007DC513FF26FFC11F4558B4D165FBCD" box="[1078,1116,1015,1040]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="46" pageNumber="47" unit="m" value="3.0">3m</elevation>
</quantity>
, 1f larva (
<collectionCode id="ED41BAE5FF26FFC11FBC58B4D012FBCD" box="[1231,1323,1015,1040]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B38287DFF26FFC11E4C58BBD4B0FBEE" ID-GBIF-Occurrence="3325453337" collectorName="Mecsek Mts" country="Hungary" location="Baranya County" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Baranya">
<collectingRegion id="4994ECC2FF26FFC11E4C58BBD5D2FBE8" country="Hungary" name="Baranya" pageId="46" pageNumber="47">Baranya County</collectingRegion>
, 
<collectorName id="26A547F6FF26FFC11B8B5F5FD4B0FBEE" box="[248,393,1052,1077]" pageId="46" pageNumber="47">Mecsek Mts</collectorName>
</materialsCitation>
., 
<materialsCitation id="3B38287DFF26FFC11AE85F5FD78DFB85" ID-GBIF-Occurrence="3325453383" collectionCode="HNHM" country="Hungary" elevation="250" location="Valley" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Pecs">
<collectingRegion id="4994ECC2FF26FFC11AE85F5FD4E8FBE8" box="[411,465,1052,1077]" country="Hungary" name="Pecs" pageId="46" pageNumber="47">Pécs</collectingRegion>
, 
<collectingRegion id="4994ECC2FF26FFC11AAD5F5FD72AFBE8" box="[478,531,1052,1077]" country="Hungary" name="Eger" pageId="46" pageNumber="47">Éger</collectingRegion>
<location id="8E8F74FBFF26FFC1196F5F5FD75CFBE8" LSID="urn:lsid:plazi:treatment:03F99336FF25FFDF1BE45D79D688F990:8E8F74FBFF26FFC1196F5F5FD75CFBE8" box="[540,613,1052,1077]" country="Hungary" name="Valley" pageId="46" pageNumber="47" stateProvince="Pecs">Valley</location>
, 
<quantity id="4CA88FC5FF26FFC119015F5FD784FBEE" box="[626,701,1052,1077]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="47" unit="m" value="250.0">
<elevation id="007DC513FF26FFC119015F5FD784FBEE" box="[626,701,1052,1077]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="47" unit="m" value="250.0">250 m</elevation>
</quantity>
, 29.02.200 4, leg. D. 
<taxonomicName id="4C5059A3FF26FFC118C35F5FD12CFBE8" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[944,1045,1052,1077]" genus="Muranyi" pageId="46" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
: 
<quantity id="4CA88FC5FF26FFC11F525F5FD17CFBE8" box="[1057,1093,1052,1077]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">
<elevation id="007DC513FF26FFC11F525F5FD17CFBE8" box="[1057,1093,1052,1077]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="m" value="1.0">1m</elevation>
</quantity>
2f, 
<quantity id="4CA88FC5FF26FFC11F055F5FD1A0FBE8" box="[1142,1177,1052,1077]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="m" value="5.0">
<elevation id="007DC513FF26FFC11F055F5FD1A0FBE8" box="[1142,1177,1052,1077]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="m" value="5.0">5m</elevation>
</quantity>
6f larvae, 
<quantity id="4CA88FC5FF26FFC11E6B5F5FD002FBE8" box="[1304,1339,1052,1077]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="m" value="2.0">
<elevation id="007DC513FF26FFC11E6B5F5FD002FBE8" box="[1304,1339,1052,1077]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="m" value="2.0">2m</elevation>
</quantity>
exuviae (
<collectionCode id="ED41BAE5FF26FFC11BED5F7CD5C0FB85" box="[158,249,1087,1112]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
; 
<specimenCount id="9D56E9A9FF26FFC11A765F03D4B9FB85" box="[261,384,1087,1112]" pageId="46" pageNumber="47" type="female">one female</specimenCount>
prepared for SEM 
<figureCitation id="136B3EA5FF26FFC1192B5F7CD795FB85" box="[600,684,1087,1112]" captionStart="FIGURES 62–65" captionStartId="33.[151,264,1508,1531]" captionTargetBox="[151,1436,231,1487]" captionTargetId="figure-310@33.[151,1436,193,1487]" captionTargetPageId="33" captionText="FIGURES 62–65. Female genitalia of Zwicknia Murányi, gen. n.—62–63: Z. bifrons (Newman, 1838); 64: Z. acuta Murányi &amp; Orci, sp. n.; 65: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 62: terminalia, ventral view; 63: inner genital organs, dorsal view; 64–65: subgenital plates, ventral view (gc: genital cavity; go: genital opening; Sg: subgenital plate)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919112" httpUri="https://zenodo.org/record/4919112/files/figure.png" pageId="46" pageNumber="47">Fig. 65</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF26FFDF1BB45F27D076FAF0" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
<paragraph id="8BEF2220FF26FFC11BB45F27D4D2FB3A" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
<emphasis id="B924FE32FF26FFC11BB45F27D461FBA0" bold="true" box="[199,344,1124,1149]" pageId="46" pageNumber="47">Description.</emphasis>
Head, thorax, appendages and basal segments of the abdomen generotypic. Occipital rugosities present on teneral specimens. Males micropterous, females macropterous. Body length: 
<typeStatus id="54EB9C82FF26FFC11FE35FC4D1CBFB7D" box="[1168,1266,1159,1184]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
6.5 mm, male 
<typeStatus id="54EB9C82FF26FFC11BE45FEFD43BFB19" box="[151,258,1196,1220]" pageId="46" pageNumber="47" type="paratype">paratypes</typeStatus>
6.0–6.5, female 
<typeStatus id="54EB9C82FF26FFC11AB25FEFD715FB19" box="[449,556,1196,1220]" pageId="46" pageNumber="47" type="paratype">paratypes</typeStatus>
8.0–8.5 mm; forewing length: 
<typeStatus id="54EB9C82FF26FFC118FE5FEFD6D6FB18" box="[909,1007,1196,1221]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
1.6 mm, male 
<typeStatus id="54EB9C82FF26FFC11FE85FEFD03FFB19" box="[1179,1286,1196,1220]" pageId="46" pageNumber="47" type="paratype">paratypes</typeStatus>
1.3–2.0 mm, female 
<typeStatus id="54EB9C82FF26FFC11B995F93D46CFB35" box="[234,341,1232,1256]" pageId="46" pageNumber="47" type="paratype">paratypes</typeStatus>
8.0–8.5 mm.
</paragraph>
<paragraph id="8BEF2220FF26FFC11BB45FB7D077F9F0" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
Male terminalia (
<figureCitation id="136B3EA5FF26FFC11AFB5FB7D729FAD0" box="[392,528,1268,1293]" captionStart="FIGURES 90–93" captionStartId="47.[151,264,1938,1961]" captionTargetBox="[257,1337,206,1906]" captionTargetId="figure-60@47.[235,1353,193,1922]" captionTargetPageId="47" captionText="FIGURES 90–93. Male terminalia of Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n.; paratype, Croatia, Krapina- Zagorje County, Ivanščica Mts., Stari Golubovec—90: dorsal view; 91: ventral view; 92: lateral view; 93: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919124" httpUri="https://zenodo.org/record/4919124/files/figure.png" pageId="46" pageNumber="47">Figs. 90–93</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC119685FB7D770FAD0" box="[539,585,1268,1293]" captionStart="FIGURES 120–123" captionStartId="57.[151,264,1761,1784]" captionTargetBox="[165,1401,468,1695]" captionTargetId="figure-157@57.[151,1436,457,1720]" captionTargetPageId="57" captionText="FIGURES 120–123. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—120: Z. bifrons (Newman, 1838); 121: Z. acuta Murányi &amp; Orci, sp. n.; 122: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 123: Z. kovacsi Murányi &amp; Gamboa, sp. n.—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919142" httpUri="https://zenodo.org/record/4919142/files/figure.png" pageId="46" pageNumber="47">122</figureCitation>
): Process of Tg 9 very low, caudally projecting, its apex is,1½ × wider than the medial section of Ep-scl; its shape is flattened elliptical, the apex smooth, rounded; sides continuously narrow in caudal view, the membranous portion narrowest apically (
<figureCitation id="136B3EA5FF26FFC118305E7FD6D0FA88" box="[835,1001,1340,1365]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="46" pageNumber="47">Figs. 157–159</figureCitation>
). Tg 10, B-scl and Lb-scl generotypic. Ep-scl wide and blunt in dorsal view, medially not swollen, its medial width ⅔ basal width; tip upcurved in lateral view, divided section short. Ventral membranous part between the division of Ep-scl nearly reach the base in lateral view; apical spines short, distributed only on the membranous part (
<figureCitation id="136B3EA5FF26FFC118915EE4D16FFA1D" box="[994,1110,1447,1472]" captionStart="FIGURES 106–109" captionStartId="54.[151,264,1087,1110]" captionTargetBox="[151,1436,193,1067]" captionTargetId="figure-252@54.[151,1436,193,1067]" captionTargetPageId="54" captionText="FIGURES 106–109. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—106: Z. bifrons (Newman, 1838); 107: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 108–109: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919136" httpUri="https://zenodo.org/record/4919136/files/figure.png" pageId="46" pageNumber="47">Figs. 107</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11F195EE4D1ECFA1D" box="[1130,1237,1447,1472]" captionStart="FIGURES 114–119" captionStartId="56.[151,265,1540,1563]" captionTargetBox="[151,1436,193,1520]" captionTargetId="figure-203@56.[151,1436,193,1520]" captionTargetPageId="56" captionText="FIGURES 114–119. Dorsal view of male epiproct of Zwicknia Murányi, gen. n.—114: Z. bifrons (Newman, 1838); 115, 117: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 116: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 118: Z. sevanica (Zhiltzova, 1964); 119: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919140" httpUri="https://zenodo.org/record/4919140/files/figure.png" pageId="46" pageNumber="47">115, 117</figureCitation>
, 
<figureCitation id="136B3EA5FF26FFC11F9A5EE4D076FA1D" box="[1257,1359,1447,1472]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="46" pageNumber="47">133–138</figureCitation>
). I-scl generotypic, Ec short and often partly or fully everted on the non in-copula specimens, even on tenerals (
<figureCitation id="136B3EA5FF26FFC11E175E8FD5C4F9D5" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="46" pageNumber="47">Figs. 133–138</figureCitation>
). St 9 slightly projecting medially, vesicle small to medium-sized, 
<figureCitation id="136B3EA5FF26FFC11F7A5EACD165F9D5" box="[1033,1116,1519,1544]" captionStart="FIGURES 90–93" captionStartId="47.[151,264,1938,1961]" captionTargetBox="[257,1337,206,1906]" captionTargetId="figure-60@47.[235,1353,193,1922]" captionTargetPageId="47" captionText="FIGURES 90–93. Male terminalia of Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n.; paratype, Croatia, Krapina- Zagorje County, Ivanščica Mts., Stari Golubovec—90: dorsal view; 91: ventral view; 92: lateral view; 93: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919124" httpUri="https://zenodo.org/record/4919124/files/figure.png" pageId="46" pageNumber="47">Fig. 91</figureCitation>
illustrates the smallest size range. Sg rounded with pronounced triangular shape, tip usually rounded. Pp, Fp, Rp and cerci generotypic.
</paragraph>
<paragraph id="8BEF2220FF26FFC11BB45D74D490F945" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
Female subgenital plate (
<figureCitation id="136B3EA5FF26FFC11A915D74D70EF98D" box="[482,567,1591,1616]" captionStart="FIGURES 62–65" captionStartId="33.[151,264,1508,1531]" captionTargetBox="[151,1436,231,1487]" captionTargetId="figure-310@33.[151,1436,193,1487]" captionTargetPageId="33" captionText="FIGURES 62–65. Female genitalia of Zwicknia Murányi, gen. n.—62–63: Z. bifrons (Newman, 1838); 64: Z. acuta Murányi &amp; Orci, sp. n.; 65: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 62: terminalia, ventral view; 63: inner genital organs, dorsal view; 64–65: subgenital plates, ventral view (gc: genital cavity; go: genital opening; Sg: subgenital plate)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919112" httpUri="https://zenodo.org/record/4919112/files/figure.png" pageId="46" pageNumber="47">Fig. 65</figureCitation>
): Rectangular, posterior margin slightly rounded, usually slightly incised and equal to the segment`s posterior margin. Antero-lateral recess usually distinct, the plate is entirely brown; lateral sclerites relatively large.
</paragraph>
<paragraph id="8BEF2220FF26FFC11BB45DE7D467F801" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
Drumming: Males produce generally single, monophasic calls in a sporadic pattern. Within each call inter-beat intervals decrease gradually (
<figureCitation id="136B3EA5FF26FFC11A965D84D778F93D" box="[485,577,1735,1760]" captionStart="FIGURE 190" captionStartId="64.[151,250,1125,1147]" captionTargetBox="[163,1430,185,1044]" captionTargetId="figure-392@64.[151,1436,177,1058]" captionTargetPageId="64" captionText="FIGURE 190. Inter-beat interval variation during male calls in the four examined Zwicknia Murányi, gen. n. species.—Box plot colors: white—Z. bifrons (Newman, 1838), red—Z. acuta Murányi &amp; Orci, sp. n., blue—Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. In Z. kovacsi Murányi &amp; Gamboa, sp. n. we have drumming samples only from two specimens therefore in the case of that species two simple mark series are used to show inter-beat intervals in their calls (circles for specimen 1, and triangles for specimen 2). i1, i2, i3 and so on are labels containing the serial number of inter-beat intervals counting them from the beginning of each call. Box plots show the duration values measured in 7 specimens in Z. bifrons and Z. acuta, and 5 specimens in Z. rupprechti. Vertical axis has a logarithmic scale. Only calls recorded at an ambient air temperature of 17–19 oC were used. (see the measurement data in Appendix Tables 1–4)." figureDoi="http://doi.org/10.5281/zenodo.4919160" httpUri="https://zenodo.org/record/4919160/files/figure.png" pageId="46" pageNumber="47">Fig 190</figureCitation>
, Appendix 
<tableCitation id="C6D2179BFF26FFC119B45D84D627F93D" box="[711,798,1735,1760]" captionStart="TABLE 3" captionStartId="10.[147,237,543,566]" captionTargetBox="[194,1631,634,1733]" captionText="TABLE 3. Male epiproct of valid extant Capniidae genera. — in case of two character states the first one refers to the type species; for abbreviations of parts see Figs. 1-22." httpUri="http://table.plazi.org/id/DF2F72A8FF02FFE51BE0595CD720FD88" pageId="46" pageNumber="47" tableUuid="DF2F72A8FF02FFE51BE0595CD720FD88">Table 3</tableCitation>
). Peak amplitude of beats generally increases along the beat sequence of a call. See 
<figureCitation id="136B3EA5FF26FFC11ABC5DAFD756F8D8" box="[463,623,1772,1797]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="46" pageNumber="47">Figs. 179–182</figureCitation>
for the oscillographic pattern of the male drumming calls of this species, and also 
<tableCitation id="C6D2179BFF26FFC11B8A5C4CD472F8F5" box="[249,331,1807,1832]" captionStart="TABLE 8" captionStartId="65.[151,239,1605,1628]" captionTargetPageId="65" captionText="TABLE 8. Descriptive statistics for the examined rhythmic characters of the male drumming calls of Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n., and also for the ambient air temperature during the sound recordings.—&quot;n&quot; is the number of male specimens examined. In each character data points where mean values obtained for each examined male on the basis of 1-10 signal measurements per specimen. Number of examined specimens per population: Hungary: Mecsek Mts—7; Croatia: Ivanščica Mts—3." pageId="46" pageNumber="47">Table 8</tableCitation>
for descriptive statistics of the examined three sonometric parameters. The male-female drumming duet consists of a single male call—female answer signal exchanges, but also male call—female answer—male response intersexual signalling can be observed (
<figureCitation id="136B3EA5FF26FFC119CD5C14D618F8AD" box="[702,801,1879,1904]" captionStart="FIGURES 185–189" captionStartId="63.[151,264,1012,1035]" captionTargetBox="[166,1430,178,988]" captionTargetId="figure-458@63.[151,1436,178,1004]" captionTargetPageId="63" captionText="FIGURES 185–189. Male-female drumming duets of three Zwicknia Murányi, gen. n. species.—185: Z. bifrons (Newman, 1838) from Hungary, Pilis Mts., Bükkös Stream, recorded at 16.9 °C; 186–187: Z. acuta Murányi &amp; Orci, sp. n. from Slovakia, Krupinská Planina, Stará Rieka, recorded at 18.2 °C; 188: call-answer duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 18.2 °C; 189: call-answer-reply duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 17.9 °C—arrows with a gender sign point to the beginning of a section of the oscillogram containing the beats of that gender." figureDoi="http://doi.org/10.5281/zenodo.4919158" httpUri="https://zenodo.org/record/4919158/files/figure.png" pageId="46" pageNumber="47">Fig. 189</figureCitation>
.) Female answer signals are monophasic beat sequences of a similar beat repetition rate as observable in the male call, but are longer and contain more beats than the male signals. For a more detailed quantitative analysis of the male-female duet pattern, recordings from more females will be necessary.
</paragraph>
<caption id="DF2F72A8FF27FFC01BE45CD1D5FCF83B" ID-DOI="http://doi.org/10.5281/zenodo.4919124" ID-Zenodo-Dep="4919124" httpUri="https://zenodo.org/record/4919124/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,264,1938,1961]" targetBox="[257,1337,206,1906]" targetPageId="47">
<paragraph id="8BEF2220FF27FFC01BE45CD1D5FCF83B" blockId="47.[151,1436,1938,2022]" pageId="47" pageNumber="48">
<emphasis id="B924FE32FF27FFC01BE45CD1D461F875" bold="true" box="[151,344,1938,1961]" pageId="47" pageNumber="48">FIGURES 90–93.</emphasis>
Male terminalia of 
<taxonomicName id="4C5059A3FF27FFC019425CD0D11BF874" authority="Muranyi, Orci &amp; Gamboa" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[561,1058,1939,1961]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="rupprechti" status="sp. nov.">
<emphasis id="B924FE32FF27FFC019425CD0D63BF874" box="[561,770,1939,1961]" italics="true" pageId="47" pageNumber="48">Zwicknia rupprechti</emphasis>
Murányi, Orci &amp; Gamboa
</taxonomicName>
, 
<emphasis id="B924FE32FF27FFC01F435CD0D154F875" bold="true" box="[1072,1133,1939,1960]" pageId="47" pageNumber="48">
<taxonomicNameLabel id="A2174349FF27FFC01F435CD0D154F875" box="[1072,1133,1939,1960]" pageId="47" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; paratype, Croatia, Krapina- Zagorje County, Ivanščica Mts., Stari Golubovec 
<emphasis id="B924FE32FF27FFC019F75CF0D7A3F815" box="[644,666,1971,1992]" italics="true" pageId="47" pageNumber="48">—</emphasis>
90: dorsal view; 91: ventral view; 92: lateral view; 93: caudal view—scale 1 mm.
</paragraph>
</caption>
<caption id="DF2F72A8FF38FFDF1BE45BDBD68EFEF2" pageId="48" pageNumber="49" startId="48.[151,241,152,175]" targetBox="[155,1432,336,622]" targetIsTable="true" targetPageId="48">
<paragraph id="8BEF2220FF38FFDF1BE45BDBD68EFEF2" blockId="48.[151,1436,151,303]" pageId="48" pageNumber="49">
<emphasis id="B924FE32FF38FFDF1BE45BDBD432FF72" bold="true" box="[151,267,152,175]" pageId="48" pageNumber="49">TABLE 7.</emphasis>
Descriptive statistics for the examined rhythmic characters of the male drumming calls of 
<taxonomicName id="4C5059A3FF38FFDF1F945BD4D468FF12" authority="(Newman, 1838)" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Newman" baseAuthorityYear="1838" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF38FFDF1F945BD4D0A5FF72" box="[1255,1436,151,175]" italics="true" pageId="48" pageNumber="49">Zwicknia bifrons</emphasis>
(
<bibRefCitation id="EFC15FD1FF38FFDF1BED5BFBD470FF12" author="Newman, E." box="[158,329,184,207]" pageId="48" pageNumber="49" pagination="372 - 402" refId="ref44180" refString="Newman, E. (1838) Entomological notes. The Entomological Magazine, 5 (4), 372 - 402." type="journal article" year="1838">Newman, 1838</bibRefCitation>
)
</taxonomicName>
, and also for the ambient air temperature during the sound recordings.—&quot;n&quot; is the number of male specimens examined. In each character data points where mean values obtained for each examined male on the basis of 1-10 signal measurements per specimen. Number of examined specimens per population: Hungary: Pilis Mts— 7, Bakony Mts—2, Börzsöny Mts—4, Mátra Mts—5;Serbia: Maljen Mts—7.
</paragraph>
</caption>
<paragraph id="8BEF2220FF38FFDF18E95A12D0AEFDB3" pageId="48" pageNumber="49">
<table id="F950D080FF3800101BE85A13D0A1FDB3" box="[155,1432,336,622]" colsContinueFrom="43.[159,1386,918,1191]" colsContinueIn="65.[159,1380,1796,1986]" gridcols="6" gridrows="7" pageId="48" pageNumber="49">
<tr id="35602062FF3800101BE85A13D0A1FEBB" box="[155,1432,336,358]" gridrow="0" pageId="48" pageNumber="49" rowspan-0="1">
<th id="76B1491EFF38001018E75A13D697FEBB" box="[916,942,336,358]" gridcol="1" gridrow="0" pageId="48" pageNumber="49">n</th>
<th id="76B1491EFF38001018805A13D178FEBB" box="[1011,1089,336,358]" gridcol="2" gridrow="0" pageId="48" pageNumber="49">mean</th>
<th id="76B1491EFF3800101F005A13D186FEBB" box="[1139,1215,336,358]" gridcol="3" gridrow="0" pageId="48" pageNumber="49">SD</th>
<th id="76B1491EFF3800101F965A13D015FEBB" box="[1253,1324,336,358]" gridcol="4" gridrow="0" pageId="48" pageNumber="49">min</th>
<th id="76B1491EFF3800101E3C5A13D0A1FEBB" box="[1359,1432,336,358]" gridcol="5" gridrow="0" pageId="48" pageNumber="49">max</th>
</tr>
<tr id="35602062FF3800101BE85A3FD0A1FE4F" box="[155,1432,380,402]" gridrow="1" pageId="48" pageNumber="49">
<th id="76B1491EFF3800101BE85A3FD75FFE4F" box="[155,614,380,402]" gridcol="0" gridrow="1" pageId="48" pageNumber="49">Call duration (ms)</th>
<td id="76B1491EFF38001018E75A3FD697FE4F" box="[916,942,380,402]" gridcol="1" gridrow="1" pageId="48" pageNumber="49">25</td>
<td id="76B1491EFF38001018805A3FD178FE4F" box="[1011,1089,380,402]" gridcol="2" gridrow="1" pageId="48" pageNumber="49">1811</td>
<td id="76B1491EFF3800101F005A3FD186FE4F" box="[1139,1215,380,402]" gridcol="3" gridrow="1" pageId="48" pageNumber="49">264.7</td>
<td id="76B1491EFF3800101F965A3FD015FE4F" box="[1253,1324,380,402]" gridcol="4" gridrow="1" pageId="48" pageNumber="49">1443</td>
<td id="76B1491EFF3800101E3C5A3FD0A1FE4F" box="[1359,1432,380,402]" gridcol="5" gridrow="1" pageId="48" pageNumber="49">2389</td>
</tr>
<tr id="35602062FF3800101BE85AEBD0A1FE63" box="[155,1432,424,446]" gridrow="2" pageId="48" pageNumber="49">
<th id="76B1491EFF3800101BE85AEBD75FFE63" box="[155,614,424,446]" gridcol="0" gridrow="2" pageId="48" pageNumber="49">Number of beats per call</th>
<td id="76B1491EFF38001018E75AEBD697FE63" box="[916,942,424,446]" gridcol="1" gridrow="2" pageId="48" pageNumber="49">25</td>
<td id="76B1491EFF38001018805AEBD178FE63" box="[1011,1089,424,446]" gridcol="2" gridrow="2" pageId="48" pageNumber="49">11.26</td>
<td id="76B1491EFF3800101F005AEBD186FE63" box="[1139,1215,424,446]" gridcol="3" gridrow="2" pageId="48" pageNumber="49">1.684</td>
<td id="76B1491EFF3800101F965AEBD015FE63" box="[1253,1324,424,446]" gridcol="4" gridrow="2" pageId="48" pageNumber="49">7.33</td>
<td id="76B1491EFF3800101E3C5AEBD0A1FE63" box="[1359,1432,424,446]" gridcol="5" gridrow="2" pageId="48" pageNumber="49">15.5</td>
</tr>
<tr id="35602062FF3800101BE85A97D0A1FE37" box="[155,1432,468,490]" gridrow="3" pageId="48" pageNumber="49">
<th id="76B1491EFF3800101BE85A97D75FFE37" box="[155,614,468,490]" gridcol="0" gridrow="3" pageId="48" pageNumber="49">Mean beat inteval (ms)</th>
<td id="76B1491EFF38001018E75A97D697FE37" box="[916,942,468,490]" gridcol="1" gridrow="3" pageId="48" pageNumber="49">25</td>
<td id="76B1491EFF38001018805A97D178FE37" box="[1011,1089,468,490]" gridcol="2" gridrow="3" pageId="48" pageNumber="49">178.1</td>
<td id="76B1491EFF3800101F005A97D186FE37" box="[1139,1215,468,490]" gridcol="3" gridrow="3" pageId="48" pageNumber="49">19.2</td>
<td id="76B1491EFF3800101F965A97D015FE37" box="[1253,1324,468,490]" gridcol="4" gridrow="3" pageId="48" pageNumber="49">147.6</td>
<td id="76B1491EFF3800101E3C5A97D0A1FE37" box="[1359,1432,468,490]" gridcol="5" gridrow="3" pageId="48" pageNumber="49">229</td>
</tr>
<tr id="35602062FF3800101BE85943D0A1FDCB" box="[155,1432,512,534]" gridrow="4" pageId="48" pageNumber="49">
<th id="76B1491EFF3800101BE85943D75FFDCB" box="[155,614,512,534]" gridcol="0" gridrow="4" pageId="48" pageNumber="49">Inter-call interval within a call sequence (ms)</th>
<td id="76B1491EFF38001018E75943D697FDCB" box="[916,942,512,534]" gridcol="1" gridrow="4" pageId="48" pageNumber="49">21</td>
<td id="76B1491EFF38001018805943D178FDCB" box="[1011,1089,512,534]" gridcol="2" gridrow="4" pageId="48" pageNumber="49">3061</td>
<td id="76B1491EFF3800101F005943D186FDCB" box="[1139,1215,512,534]" gridcol="3" gridrow="4" pageId="48" pageNumber="49">1411.1</td>
<td id="76B1491EFF3800101F965943D015FDCB" box="[1253,1324,512,534]" gridcol="4" gridrow="4" pageId="48" pageNumber="49">1447</td>
<td id="76B1491EFF3800101E3C5943D0A1FDCB" box="[1359,1432,512,534]" gridcol="5" gridrow="4" pageId="48" pageNumber="49">6295</td>
</tr>
<tr id="35602062FF3800101BE8596FD0A1FD9F" box="[155,1432,556,578]" gridrow="5" pageId="48" pageNumber="49">
<th id="76B1491EFF3800101BE8596FD75FFD9F" box="[155,614,556,578]" gridcol="0" gridrow="5" pageId="48" pageNumber="49">Number of calls in a call sequence</th>
<td id="76B1491EFF38001018E7596FD697FD9F" box="[916,942,556,578]" gridcol="1" gridrow="5" pageId="48" pageNumber="49">24</td>
<td id="76B1491EFF3800101880596FD178FD9F" box="[1011,1089,556,578]" gridcol="2" gridrow="5" pageId="48" pageNumber="49">1.992</td>
<td id="76B1491EFF3800101F00596FD186FD9F" box="[1139,1215,556,578]" gridcol="3" gridrow="5" pageId="48" pageNumber="49">0.575</td>
<td id="76B1491EFF3800101F96596FD015FD9F" box="[1253,1324,556,578]" gridcol="4" gridrow="5" pageId="48" pageNumber="49">1</td>
<td id="76B1491EFF3800101E3C596FD0A1FD9F" box="[1359,1432,556,578]" gridcol="5" gridrow="5" pageId="48" pageNumber="49">3</td>
</tr>
<tr id="35602062FF3800101BE85915D0A1FDB3" box="[155,1432,598,622]" gridrow="6" pageId="48" pageNumber="49">
<th id="76B1491EFF3800101BE85915D75FFDB3" box="[155,614,598,622]" gridcol="0" gridrow="6" pageId="48" pageNumber="49">Air temperature (o C)</th>
<td id="76B1491EFF38001018E75915D697FDB3" box="[916,942,598,622]" gridcol="1" gridrow="6" pageId="48" pageNumber="49">25</td>
<td id="76B1491EFF38001018805915D178FDB3" box="[1011,1089,598,622]" gridcol="2" gridrow="6" pageId="48" pageNumber="49">18.5</td>
<td id="76B1491EFF3800101F005915D186FDB3" box="[1139,1215,598,622]" gridcol="3" gridrow="6" pageId="48" pageNumber="49">1.407</td>
<td id="76B1491EFF3800101F965915D015FDB3" box="[1253,1324,598,622]" gridcol="4" gridrow="6" pageId="48" pageNumber="49">15.8</td>
<td id="76B1491EFF3800101E3C5915D0A1FDB3" box="[1359,1432,598,622]" gridcol="5" gridrow="6" pageId="48" pageNumber="49">20.8</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEF2220FF38FFDF1BB459F3D487FCCC" blockId="48.[151,1436,688,1613]" pageId="48" pageNumber="49">
Genetics: Two different haplotypes from southwestern 
<collectingCountry id="F34762B0FF38FFDF184C59F2D698FD14" box="[831,929,689,713]" name="Hungary" pageId="48" pageNumber="49">Hungary</collectingCountry>
and northern 
<collectingCountry id="F34762B0FF38FFDF1F3559F3D1A0FD14" box="[1094,1177,688,713]" name="Croatia" pageId="48" pageNumber="49">Croatia</collectingCountry>
clustered on the same node with 1% divergence with 5 informative characters separating them. All individuals collected in 
<collectingCountry id="F34762B0FF38FFDF1E795997D055FD31" box="[1290,1388,724,748]" name="Hungary" pageId="48" pageNumber="49">Hungary</collectingCountry>
and 
<collectingCountry id="F34762B0FF38FFDF1BE459BBD5D0FCCC" box="[151,233,760,785]" name="Croatia" pageId="48" pageNumber="49">Croatia</collectingCountry>
clustered together.
</paragraph>
<paragraph id="8BEF2220FF38FFDF1BB4585FD076FAF0" blockId="48.[151,1436,688,1613]" pageId="48" pageNumber="49">
<emphasis id="B924FE32FF38FFDF1BB4585FD405FCEE" bold="true" box="[199,316,796,821]" pageId="48" pageNumber="49">Affinities.</emphasis>
The relationship of 
<taxonomicName id="4C5059A3FF38FFDF1957585ED781FCE8" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[548,696,796,821]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF38FFDF1957585ED781FCE8" box="[548,696,796,821]" italics="true" pageId="48" pageNumber="49">Z. rupprechti</emphasis>
</taxonomicName>
to 
<taxonomicName id="4C5059A3FF38FFDF19AD585ED672FCE8" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[734,843,796,821]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF38FFDF19AD585ED672FCE8" box="[734,843,796,821]" italics="true" pageId="48" pageNumber="49">Z. bifrons</emphasis>
</taxonomicName>
is discussed above. In addition, 
<taxonomicName id="4C5059A3FF38FFDF1FCD585ED06BFCE8" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[1214,1362,796,821]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF38FFDF1FCD585ED06BFCE8" box="[1214,1362,796,821]" italics="true" pageId="48" pageNumber="49">Z. rupprechti</emphasis>
</taxonomicName>
males are easily distinguishable from other 
<taxonomicName id="4C5059A3FF38FFDF19365803D795FC84" authorityName="Muranyi" authorityYear="2014" box="[581,684,832,857]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B924FE32FF38FFDF19365803D795FC84" box="[581,684,832,857]" italics="true" pageId="48" pageNumber="49">Zwicknia</emphasis>
</taxonomicName>
on the basis of upcurved Ep-scl tip and low, caudally projecting process of Tg 9. Females cannot be distinguished from other species of the genus as are larvae morphologically indistinguishable. The male drumming calls of 
<taxonomicName id="4C5059A3FF38FFDF19C258CAD67CFC7C" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[689,837,904,929]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF38FFDF19C258CAD67CFC7C" box="[689,837,904,929]" italics="true" pageId="48" pageNumber="49">Z. rupprechti</emphasis>
</taxonomicName>
are of short duration and have a fast beat repetition, differing from those of 
<taxonomicName id="4C5059A3FF38FFDF1AD158EED736FC18" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[418,527,940,965]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF38FFDF1AD158EED736FC18" box="[418,527,940,965]" italics="true" pageId="48" pageNumber="49">Z. bifrons</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5059A3FF38FFDF193B58EED79CFC19" authorityName="Muranyi &amp; Orci" authorityYear="2014" box="[584,677,941,964]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="B924FE32FF38FFDF193B58EED79CFC19" box="[584,677,941,964]" italics="true" pageId="48" pageNumber="49">Z. acuta</emphasis>
</taxonomicName>
, but similar to the calls of 
<taxonomicName id="4C5059A3FF38FFDF18A858EED177FC18" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[987,1102,940,965]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF38FFDF18A858EED177FC18" box="[987,1102,940,965]" italics="true" pageId="48" pageNumber="49">Z. kovacsi</emphasis>
</taxonomicName>
. Distinctive characters of the male calls of 
<taxonomicName id="4C5059A3FF38FFDF1A4F5892D4EAFC34" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[316,467,976,1001]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF38FFDF1A4F5892D4EAFC34" box="[316,467,976,1001]" italics="true" pageId="48" pageNumber="49">Z. rupprechti</emphasis>
</taxonomicName>
that differ from those of 
<taxonomicName id="4C5059A3FF38FFDF18785892D6BBFC34" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[779,898,976,1001]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF38FFDF18785892D6BBFC34" box="[779,898,976,1001]" italics="true" pageId="48" pageNumber="49">Z. kovacsi</emphasis>
</taxonomicName>
are shorter call durations, inter-beat intervals exhibiting a gradual decrease, and the beat peak amplitude with a characteristic increase during the beat call sequence. The calls of 
<taxonomicName id="4C5059A3FF38FFDF1AEA5F5AD732FBEC" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[409,523,1048,1073]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF38FFDF1AEA5F5AD732FBEC" box="[409,523,1048,1073]" italics="true" pageId="48" pageNumber="49">Z. kovacsi</emphasis>
</taxonomicName>
exhibit less of a change in those signal parameters during a call. With their short mean inter-beat intervals the calls of 
<taxonomicName id="4C5059A3FF38FFDF19485F7FD7F7FB88" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[571,718,1084,1109]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rupprechti">Z. rupprechti</taxonomicName>
are closer to Rupprecht's &quot;Capbif&quot; call variant than the slightly slower calls of 
<taxonomicName id="4C5059A3FF38FFDF1A375F22D48FFBA4" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[324,438,1120,1145]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF38FFDF1A375F22D48FFBA4" box="[324,438,1120,1145]" italics="true" pageId="48" pageNumber="49">Z. kovacsi</emphasis>
</taxonomicName>
, which stand a little farther form &quot;Capbif&quot;. However, regarding the number of beats per call, it is 
<taxonomicName id="4C5059A3FF38FFDF1A775FC6D441FB40" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[260,376,1156,1181]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF38FFDF1A775FC6D441FB40" box="[260,376,1156,1181]" italics="true" pageId="48" pageNumber="49">Z. kovacsi</emphasis>
</taxonomicName>
that is closer to &quot;Capbif&quot; (compare our results to the &quot;Capbif&quot; drumming data in 
<bibRefCitation id="EFC15FD1FF38FFDF1E545FC7D5EAFB1C" author="Rupprecht, R." pageId="48" pageNumber="49" pagination="93 - 98" refId="ref45232" refString="Rupprecht, R. (1997) An attempt to explain different drumming signals within Capnia bifrons. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera, Biology-Ecology-Systematics. MTL-Mauron + Tinguely &amp; Lachat SA., Fribourg, pp. 93 - 98." type="book chapter" year="1997">Rupprecht 1997</bibRefCitation>
). Male call differences are minor and to examine their significance in female mate choice will require additional studies including play-back experiments. However, the male terminalia of 
<taxonomicName id="4C5059A3FF38FFDF1F025F8ED03EFB38" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[1137,1287,1228,1253]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF38FFDF1F025F8ED03EFB38" box="[1137,1287,1228,1253]" italics="true" pageId="48" pageNumber="49">Z. rupprechti</emphasis>
</taxonomicName>
is similar to populations producing &quot;Capbif&quot; 
<typeStatus id="54EB9C82FF38FFDF19775FB2D70DFAD4" box="[516,564,1265,1289]" pageId="48" pageNumber="49">type</typeStatus>
signals and 
<taxonomicName id="4C5059A3FF38FFDF19CC5FB2D608FAD4" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[703,817,1264,1289]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF38FFDF19CC5FB2D608FAD4" box="[703,817,1264,1289]" italics="true" pageId="48" pageNumber="49">Z. kovacsi</emphasis>
</taxonomicName>
exhibits clear distinctive features from them. 
<taxonomicName id="4C5059A3FF38FFDF1E465FB3D435FAF0" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF38FFDF1E465FB3D435FAF0" italics="true" pageId="48" pageNumber="49">Zwicknia rupprechti</emphasis>
</taxonomicName>
differs from other species of the genus by 8% genetic divergence with 17 informative characters.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF38FFDF1BB45E7BD1EAFA3C" pageId="48" pageNumber="49" type="distribution">
<paragraph id="8BEF2220FF38FFDF1BB45E7BD1EAFA3C" blockId="48.[151,1436,688,1613]" pageId="48" pageNumber="49">
<emphasis id="B924FE32FF38FFDF1BB45E7BD4CCFA8C" bold="true" box="[199,501,1336,1361]" pageId="48" pageNumber="49">Distribution and ecology.</emphasis>
<taxonomicName id="4C5059A3FF38FFDF19755E7BD7D3FA8C" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[518,746,1336,1361]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF38FFDF19755E7BD7D3FA8C" box="[518,746,1336,1361]" italics="true" pageId="48" pageNumber="49">Zwicknia rupprechti</emphasis>
</taxonomicName>
occurs in southwestern 
<collectingCountry id="F34762B0FF38FFDF188D5E7AD159FA8C" box="[1022,1120,1337,1361]" name="Hungary" pageId="48" pageNumber="49">Hungary</collectingCountry>
and northern 
<collectingCountry id="F34762B0FF38FFDF1E725E7BD06AFA8C" box="[1281,1363,1336,1361]" name="Croatia" pageId="48" pageNumber="49">Croatia</collectingCountry>
(
<figureCitation id="136B3EA5FF38FFDF1E175E7BD5C4FAA8" captionStart="FIGURES 196–197" captionStartId="68.[151,264,1908,1931]" captionTargetBox="[188,1393,187,1898]" captionTargetId="figure-93@68.[188,1393,187,1898]" captionTargetPageId="68" captionText="FIGURES 196–197. Distribution of Zwicknia Murányi, gen. n.—196: global distribution; 197: studied populations of the Carpathian Basin and the northern Balkans—in Fig. 196, white dot on Z. bifrons symbols indicate populations studied by Rupprecht (1997), dotted lines indicate limits of s.l. Z. bifrons literature reports; in Fig. 197, white dot on symbols indicate populations where drumming signals were studied by us, white star indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.4919168" httpUri="https://zenodo.org/record/4919168/files/figure.png" pageId="48" pageNumber="49">Figs. 196–197</figureCitation>
). Populations from the southern Alps have similar drumming signals (
<bibRefCitation id="EFC15FD1FF38FFDF1F315E1FD038FAA8" author="Rupprecht, R." box="[1090,1281,1372,1397]" pageId="48" pageNumber="49" pagination="93 - 98" refId="ref45232" refString="Rupprecht, R. (1997) An attempt to explain different drumming signals within Capnia bifrons. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera, Biology-Ecology-Systematics. MTL-Mauron + Tinguely &amp; Lachat SA., Fribourg, pp. 93 - 98." type="book chapter" year="1997">Rupprecht 1997</bibRefCitation>
), but are not considered to be this species because of the close similarities of the drumming calls with those of 
<taxonomicName id="4C5059A3FF38FFDF1FA05EC2D07DFA44" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[1235,1348,1408,1433]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF38FFDF1FA05EC2D07DFA44" box="[1235,1348,1408,1433]" italics="true" pageId="48" pageNumber="49">Z. kovacsi</emphasis>
</taxonomicName>
. Adults of 
<taxonomicName id="4C5059A3FF38FFDF1BC45EE6D475FA60" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[183,332,1444,1469]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF38FFDF1BC45EE6D475FA60" box="[183,332,1444,1469]" italics="true" pageId="48" pageNumber="49">Z. rupprechti</emphasis>
</taxonomicName>
are active from March to early April and are associated with slow or moderately fast, mediumsized streams flowing through alder (
<taxonomicName id="4C5059A3FF38FFDF194B5E8BD7DEFA3C" box="[568,743,1480,1505]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="glutinosa">
<emphasis id="B924FE32FF38FFDF194B5E8BD7DEFA3C" box="[568,743,1480,1505]" italics="true" pageId="48" pageNumber="49">Alnus glutinosa</emphasis>
</taxonomicName>
) forests between 250 and 
<quantity id="4CA88FC5FF38FFDF1F7D5E8BD16CFA3D" box="[1038,1109,1480,1505]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="48" pageNumber="49" unit="m" value="350.0">350 m</quantity>
(
<figureCitation id="136B3EA5FF38FFDF1F175E8BD1FFFA3C" box="[1124,1222,1480,1505]" captionStart="FIGURES 193–195" captionStartId="67.[151,264,1829,1852]" captionTargetBox="[199,1367,231,1794]" captionTargetId="figure-90@67.[199,1367,231,1810]" captionTargetPageId="67" captionText="FIGURES 193–195. Type localities of the new species described in Zwicknia Murányi, gen. n.—193: Zwicknia acuta Murányi &amp; Orci, sp. n.: Stará rieka, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor; 194: Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.: Cimpoies Stream, Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa; 195: Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n.: Völgységi Stream, Hungary, Baranya County, Mecsek Mts., Komló-Zobákpuszta." figureDoi="http://doi.org/10.5281/zenodo.4919166" httpUri="https://zenodo.org/record/4919166/files/figure.png" pageId="48" pageNumber="49">Fig. 195</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF38FFDF1BB45EAFD688F990" pageId="48" pageNumber="49" type="etymology">
<paragraph id="8BEF2220FF38FFDF1BB45EAFD688F990" blockId="48.[151,1436,688,1613]" pageId="48" pageNumber="49">
<emphasis id="B924FE32FF38FFDF1BB45EAFD477F9D8" bold="true" box="[199,334,1516,1541]" pageId="48" pageNumber="49">Etymology.</emphasis>
The species is dedicated to Dr. Rainer Rupprecht, Mainz, 
<collectingCountry id="F34762B0FF38FFDF18915EAFD172F9D8" box="[994,1099,1516,1541]" name="Germany" pageId="48" pageNumber="49">Germany</collectingCountry>
, in recognition of his primary contribution to our knowledge on 
<taxonomicName id="4C5059A3FF38FFDF19665D53D7B2F9F4" box="[533,651,1552,1577]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
drumming calls; furthermore, he recognized the distinctiveness of the present drumming 
<typeStatus id="54EB9C82FF38FFDF1A195D77D4A7F991" box="[362,414,1588,1612]" pageId="48" pageNumber="49">type</typeStatus>
. Used as the genitive of a noun of male gender.
</paragraph>
</subSubSection>
</treatment>
</document>