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<mods:title id="D0CA65E4DFB28D45C38C708BBD0B1012">Zwicknia gen. n., a new genus for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera)</mods:title>
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<mods:namePart id="9520EE6938A408F8EA646FF5679CC884">Gamboa, Maribet</mods:namePart>
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<taxonomicName id="4C5059A3FF1BFFFC1BE45BD4D4FCFF6F" ID-CoL="8MQDB" authority="Morton, 1896" authorityName="Morton" authorityYear="1896" box="[151,453,151,178]" class="Insecta" family="Capniidae" genus="Capnopsis" kingdom="Animalia" order="Plecoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B924FE32FF1BFFFC1BE45BD4D4FCFF6F" bold="true" box="[151,453,151,178]" pageId="19" pageNumber="20">
<emphasis id="B924FE32FF1BFFFC1BE45BD4D42DFF6C" bold="true" box="[151,276,151,177]" italics="true" pageId="19" pageNumber="20">Capnopsis</emphasis>
<bibRefCitation id="EFC15FD1FF1BFFFC1A6F5BDBD4FCFF6F" author="Morton, K. J." box="[284,453,152,178]" pageId="19" pageNumber="20" pagination="55 - 63" refId="ref43808" refString="Morton, K. J. (1896) New and little-known Palaearctic Perlidae. The Transactions of the Entomological Society of London, 1896 (1), 55 - 63." type="journal article" year="1896">Morton, 1896</bibRefCitation>
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<paragraph id="8BEF2220FF1BFFFC1BE45BF9D5C0FF0E" blockId="19.[151,453,151,211]" box="[151,249,186,211]" pageId="19" pageNumber="20">
(
<figureCitation id="136B3EA5FF1BFFFC1BEC5BF9D5C8FF0E" box="[159,241,186,211]" captionStart="FIGURES 3248" captionStartId="22.[151,264,1817,1840]" captionTargetBox="[184,1414,205,1789]" captionTargetId="figure-138@22.[151,1436,163,1822]" captionTargetPageId="22" captionText="FIGURES 3248. Male terminalia of Capniidae, ventral view—32: Zwicknia bifrons (Newman); 33: Bolshecapnia gregsoni (Ricker); 34: Eucapnopsis stigmatica transversa Aubert; 35: Isocapnia grandis (Banks); 36: Capnia s.l. arensi Zhiltzova; 37: C. s.l. prolongata; 38: C. s.l. pedestris Kimmins; 39: C. s.s. nigra (Pictet); 40: Arsapnia decepta Banks; 41: Allocapnia granulata (Claassen); 42: Mesocapnia silvatica Raušer; 43: Capnura venosa Banks; 44: Utacapnia lemoniana (Nebeker &amp; Gaufin); 45: Paracapnia opis (Newman); 46: Nemocapnia carolina Banks; 47: Capnopsis schilleri schilleri (Rostock); 48: Capnioneura nemuroides Ris (C: cercus; Ep-scl: main epiproct sclerite; Fp: fusion plate; Pp: paraproct; Sg: subgenital plate; St 9: sternite 9; Tg 9: tergite 9; Tg 10: tergite 10; Vv: ventral vesicle)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919093" httpUri="https://zenodo.org/record/4919093/files/figure.png" pageId="19" pageNumber="20">Fig. 47</figureCitation>
)
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<taxonomicName id="4C5059A3FF1BFFFC1BE45A41D4AAFEC5" ID-CoL="8N6GJ" authority="Rostock, 1892" authorityName="Rostock" authorityYear="1892" box="[151,403,258,280]" class="Insecta" family="Capniidae" genus="Capnodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B924FE32FF1BFFFC1BE45A41D5C4FEC5" box="[151,253,258,280]" italics="true" pageId="19" pageNumber="20">Capnodes</emphasis>
<bibRefCitation id="EFC15FD1FF1BFFFC1A715A41D4AAFEC5" author="Rostock, M." box="[258,403,258,280]" pageId="19" pageNumber="20" pagination="1 - 5" refId="ref45022" refString="Rostock, M. (1892) Capnodes schilleri, eine neue deutsche Perlide. Berliner Entomologische Zeitschrift, 37 (1), 1 - 5. http: // dx. doi. org / 10.1002 / mmnd. 18920370103" type="journal article" year="1892">Rostock, 1892</bibRefCitation>
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<bibRefCitation id="EFC15FD1FF1BFFFC1ADE5A41D749FEC5" author="Rostock, M." box="[429,624,258,280]" pageId="19" pageNumber="20" pagination="1 - 5" refId="ref45022" refString="Rostock, M. (1892) Capnodes schilleri, eine neue deutsche Perlide. Berliner Entomologische Zeitschrift, 37 (1), 1 - 5. http: // dx. doi. org / 10.1002 / mmnd. 18920370103" type="journal article" year="1892">Rostock 1892: 398</bibRefCitation>
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. (original description,
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<taxonomicName id="4C5059A3FF1BFFFC18A55A41D008FEC5" authority="Rostock, 1892" authorityName="Rostock" authorityYear="1892" box="[982,1329,258,280]" class="Insecta" family="Capniidae" genus="Capnodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="schilleri">
<emphasis id="B924FE32FF1BFFFC18A55A41D1ADFEC5" box="[982,1172,258,280]" italics="true" pageId="19" pageNumber="20">Capnodes schilleri</emphasis>
<bibRefCitation id="EFC15FD1FF1BFFFC1FE85A41D008FEC5" author="Rostock, M." box="[1179,1329,258,280]" pageId="19" pageNumber="20" pagination="1 - 5" refId="ref45022" refString="Rostock, M. (1892) Capnodes schilleri, eine neue deutsche Perlide. Berliner Entomologische Zeitschrift, 37 (1), 1 - 5. http: // dx. doi. org / 10.1002 / mmnd. 18920370103" type="journal article" year="1892">Rostock, 1892</bibRefCitation>
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).
<taxonomicName id="4C5059A3FF1BFFFC1BE45A62D4ACFEEA" ID-CoL="8MQDB" authority="Morton, 1896" authorityName="Morton" authorityYear="1896" box="[151,405,289,311]" class="Insecta" family="Capniidae" genus="Capnopsis" kingdom="Animalia" order="Plecoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B924FE32FF1BFFFC1BE45A62D43BFEEA" box="[151,258,289,311]" italics="true" pageId="19" pageNumber="20">Capnopsis</emphasis>
<bibRefCitation id="EFC15FD1FF1BFFFC1A795A61D4ACFEEA" author="Morton, K. J." box="[266,405,289,311]" pageId="19" pageNumber="20" pagination="55 - 63" refId="ref43808" refString="Morton, K. J. (1896) New and little-known Palaearctic Perlidae. The Transactions of the Entomological Society of London, 1896 (1), 55 - 63." type="journal article" year="1896">Morton, 1896</bibRefCitation>
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<treatmentCitation id="0AF10431FF1BFFFC1ADC5A62D75BFEEA" author="Morton, K. J." box="[431,610,289,311]" page="61" pageId="19" pageNumber="20" year="1896">
<bibRefCitation id="EFC15FD1FF1BFFFC1ADC5A62D75BFEEA" author="Morton, K. J." box="[431,610,289,311]" pageId="19" pageNumber="20" pagination="55 - 63" refId="ref43808" refString="Morton, K. J. (1896) New and little-known Palaearctic Perlidae. The Transactions of the Entomological Society of London, 1896 (1), 55 - 63." type="journal article" year="1896">Morton 1896: 61</bibRefCitation>
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. (replace name for
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<emphasis id="B924FE32FF1BFFFC185F5A62D6A8FEEA" box="[812,913,289,311]" italics="true" pageId="19" pageNumber="20">Capnodes</emphasis>
<bibRefCitation id="EFC15FD1FF1BFFFC18EA5A62D117FEEA" author="Rostock, M." box="[921,1070,289,311]" pageId="19" pageNumber="20" pagination="1 - 5" refId="ref45022" refString="Rostock, M. (1892) Capnodes schilleri, eine neue deutsche Perlide. Berliner Entomologische Zeitschrift, 37 (1), 1 - 5. http: // dx. doi. org / 10.1002 / mmnd. 18920370103" type="journal article" year="1892">Rostock, 1892</bibRefCitation>
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preoccupied by
<taxonomicName id="4C5059A3FF1BFFFC1FA85A62D0A5FEEA" authority="Guenee, 1852" authorityName="Guenee" authorityYear="1852" box="[1243,1436,289,311]" class="Insecta" family="Capniidae" genus="Capnodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B924FE32FF1BFFFC1FA85A62D079FEEA" box="[1243,1344,289,311]" italics="true" pageId="19" pageNumber="20">Capnodes</emphasis>
<bibRefCitation id="EFC15FD1FF1BFFFC1E3B5A62D0A5FEEA" author="Guenee" box="[1352,1436,289,311]" pageId="19" pageNumber="20" refId="ref42210" refString="Guenee (1852) Histoire Naturelle des Insectes. Species General des Lepidopteres V. 7 (Noctuelites, V. 3). Librairie Encyclopedique de Roret, Paris, 442 pp." type="book" year="1852">Guénée,</bibRefCitation>
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1852; inherited
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species
<taxonomicName id="4C5059A3FF1BFFFC1A9B5A03D67DFE8B" authority="Rostock, 1892" authorityName="Rostock" authorityYear="1892" box="[488,836,320,342]" class="Insecta" family="Capniidae" genus="Capnodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="schilleri">
<emphasis id="B924FE32FF1BFFFC1A9B5A03D79EFE8B" box="[488,679,320,342]" italics="true" pageId="19" pageNumber="20">Capnodes schilleri</emphasis>
<bibRefCitation id="EFC15FD1FF1BFFFC19DE5A03D67DFE8B" author="Rostock, M." box="[685,836,320,342]" pageId="19" pageNumber="20" pagination="1 - 5" refId="ref45022" refString="Rostock, M. (1892) Capnodes schilleri, eine neue deutsche Perlide. Berliner Entomologische Zeitschrift, 37 (1), 1 - 5. http: // dx. doi. org / 10.1002 / mmnd. 18920370103" type="journal article" year="1892">Rostock, 1892</bibRefCitation>
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);
<bibRefCitation id="EFC15FD1FF1BFFFC18265A03D6D2FE8B" author="Zwick, P." box="[853,1003,320,342]" pageId="19" pageNumber="20" pagination="1 - 7" refId="ref46795" refString="Zwick, P. (1984) Geographische Rassen und Verbreitungsgeschichte von Capnopsis schilleri (Plecoptera, Capniidae). Deutsche Entomologische Zeitschrift, 31 (1 - 3), 1 - 7." type="journal article" year="1984">Zwick 1984: 2</bibRefCitation>
. (revision).
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF1BFFFC1BE45AC1D768FD66" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BEF2220FF1BFFFC1BE45AC1D70CFDAE" blockId="19.[151,1437,386,699]" pageId="19" pageNumber="20">
<emphasis id="B924FE32FF1BFFFC1BE45AC1D428FE46" bold="true" box="[151,273,386,411]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Male epiproct: B-scl small, divided from Ep-scl; Lb-scl small, fused with Ep-scl; Ep-scl ventrally connected in the apical part, laterally entire, caudal setae absent; I-scl long, erect stick, Ec absent. Male Pp: apical part short and wide; Fp medium long and wide, divided from Rp. Male Sg: fused with St 9 and Tg 9, vesicle lacking. Female Sg: rectangular, entire; inner and lateral sclerites absent. Male tergites: process lacking. Cercus: with less than ten segments. Ventral thoracic sclerites: MPrs elliptical, MeFs triangular, MeFsp separated from MePfs. Macropterous wings: forewing without crossvein between M and Cu; single A vein beyond a and R1 before r straight; hindwing without A veins.
</paragraph>
<paragraph id="8BEF2220FF1BFFFC1BB4593DD768FD66" blockId="19.[151,1437,386,699]" pageId="19" pageNumber="20">
<emphasis id="B924FE32FF1BFFFC1BB4593DD4ADFD4A" bold="true" box="[199,404,638,663]" pageId="19" pageNumber="20">Species included.</emphasis>
1 valid species with 3 subspecies from the West Palaearctic (
<bibRefCitation id="EFC15FD1FF1BFFFC1F22593DD016FD4A" author="DeWalt, R. E. &amp; Neu-Becker, U. &amp; Steuber, G." box="[1105,1327,638,663]" pageId="19" pageNumber="20" refId="ref41558" refString="DeWalt, R. E., Neu-Becker, U. &amp; Steuber, G. (2014) Plecoptera Species File Online. Version 5.0 / 5.0. Available from: http: // plecoptera. speciesfile. org (accessed 12 May 2014)" type="url" year="2014">
DeWalt
<emphasis id="B924FE32FF1BFFFC1FC3593CD1D3FD4A" box="[1200,1258,638,663]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2014
</bibRefCitation>
); 2 of the subspecies examined (see Appendix 1).
</paragraph>
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