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<document id="CFA111E3304FD9AB6694A65D408F7AB8" ID-CLB-Dataset="25305" ID-DOI="10.5852/ejt.2019.562" ID-GBIF-Dataset="f6af87ae-7f5c-482a-8437-cba6bdcf88f5" ID-Zenodo-Dep="3474305" ID-ZooBank="6567F621-7A92-4D1A-8902-A1E76325AF94" IM.bibliography_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1570394176492" checkinUser="plazi" docAuthor="Galea, Horia R. &amp; Schuchert, Peter" docDate="2019" docId="03C0D256AD7A6116FDEEA2E05A97F923" docLanguage="en" docName="ejt-562_galea_schuchert.pdf.imf" docOrigin="European Journal of Taxonomy 562" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tuberocaulus Galea &amp; Schuchert 2019, gen. nov." docType="treatment" docUuid="317807A0-E2BB-4EBF-9305-9917A818E1E9" docUuidSource="ZooBank" docVersion="15" lastPageNumber="54" masterDocId="FFF9AA2EAD4E6123FFD5A44F5B5CFFB2" masterDocTitle="Some thecate hydroids (Cnidaria: Hydrozoa) from off New Caledonia collected during KANACONO and KANADEEP expeditions of the French Tropical Deep-Sea Benthos Program" masterLastPageNumber="70" masterPageNumber="1" pageNumber="53" updateTime="1737040101154" updateUser="ExternalLinkService">
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<mods:title id="A5005C589914E1F837874C5C9DA1BE04">Some thecate hydroids (Cnidaria: Hydrozoa) from off New Caledonia collected during KANACONO and KANADEEP expeditions of the French Tropical Deep-Sea Benthos Program</mods:title>
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<treatment id="03C0D256AD7A6116FDEEA2E05A97F923" ID-DOI="http://doi.org/10.5281/zenodo.3475335" ID-GBIF-Taxon="159646945" ID-Zenodo-Dep="3475335" LSID="urn:lsid:zoobank.org:act:317807A0-E2BB-4EBF-9305-9917A818E1E9" httpUri="http://treatment.plazi.org/id/03C0D256AD7A6116FDEEA2E05A97F923" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" scope_family="Thyroscyphidae" scope_order="Leptothecata" scope_phylum="Cnidaria" scope_subClass="Hydroidolina">
<subSubSection id="C37330CBAD7A6117FDEEA2E058A5F97B" box="[571,1017,1710,1737]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="8BD66340AD7A6117FDEEA2E058A5F97B" blockId="52.[372,1216,1710,1773]" box="[571,1017,1710,1737]" pageId="52" pageNumber="53">
<heading id="D09ED42CAD7A6117FDEEA2E058A5F97B" box="[571,1017,1710,1737]" centered="true" fontSize="11" level="2" pageId="52" pageNumber="53" reason="2">
Genus
<taxonomicName id="4C6918C3AD7A6117FD58A2E158DEF97B" ID-CoL="845V6" authority="Galea" authorityName="Galea &amp; Schuchert" authorityYear="2019" box="[653,898,1710,1737]" class="Hydrozoa" family="Thyroscyphidae" genus="Tuberocaulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Leptothecata" pageId="52" pageNumber="53" phylum="Cnidaria" rank="genus" status="gen. nov.">
<emphasis id="B91DBF52AD7A6117FD58A2E15868F97B" bold="true" box="[653,820,1710,1737]" italics="true" pageId="52" pageNumber="53">Tuberocaulus</emphasis>
Galea
</taxonomicName>
,
<taxonomicNameLabel id="A22E0229AD7A6117FC5BA2E058A5F97B" box="[910,1017,1711,1737]" pageId="52" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37330CBAD7A6117FEA1A29D5F9CF95F" box="[372,1216,1746,1773]" pageId="52" pageNumber="53" type="description">
<paragraph id="8BD66340AD7A6117FEA1A29D5F9CF95F" blockId="52.[372,1216,1710,1773]" box="[372,1216,1746,1773]" pageId="52" pageNumber="53">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C37330CBAD7A6116FF68A35C5E7EFE26" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="8BD66340AD7A6117FF68A35C5A6AF89F" blockId="52.[189,310,1811,1837]" box="[189,310,1811,1837]" pageId="52" pageNumber="53">
<heading id="D09ED42CAD7A6117FF68A35C5A6AF89F" bold="true" box="[189,310,1811,1837]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="B91DBF52AD7A6117FF68A35C5A6AF89F" bold="true" box="[189,310,1811,1837]" pageId="52" pageNumber="53">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD66340AD7A6116FF68A30D5E7EFE26" blockId="52.[189,1399,1858,2025]" lastBlockId="53.[189,1398,267,404]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">Stems unbranched, strongly fascicled basally, arched downwards distally, and there giving rise to a row of cladia arranged unilaterally; division into internodes indistinct; each equivalent of internode with short, lateral apophysis supporting a cladium, an axillar nematotheca and second nematotheca on opposite side and slightly above; cladia divided into moderately-long internodes, each bearing a hydrotheca distally; the two rows of hydrothecae forming a sharp angle; hydrothecae long, tubular, tapering in lower halves, adnate for a minute portion of their adaxial wall to the internode, aperture with three pointed cusps (one abaxial, two latero-adaxial) separated by deep, rounded embayments, rim thickened. Gonothecae given off laterally from below the hydrothecae; short, club-shaped, aperture subterminal, semi-circular. Perisarc filmy and easily collapsible on both hydro- and gonothecae.</paragraph>
</subSubSection>
<subSubSection id="C37330CBAD7B6116FF68A5F2598BFD9E" pageId="53" pageNumber="54" type="type_taxon">
<paragraph id="8BD66340AD7B6116FF68A5F25A04FE65" blockId="53.[189,344,445,471]" box="[189,344,445,471]" pageId="53" pageNumber="54">
<heading id="D09ED42CAD7B6116FF68A5F25A04FE65" bold="true" box="[189,344,445,471]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="B91DBF52AD7B6116FF68A5F25A04FE65" bold="true" box="[189,344,445,471]" pageId="53" pageNumber="54">
<typeStatus id="54D2DDE2AD7B6116FF68A5F25BA6FE65" box="[189,250,445,471]" pageId="53" pageNumber="54">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD66340AD7B6116FF68A5A2598BFD9E" blockId="53.[189,1398,493,556]" pageId="53" pageNumber="54">
<taxonomicName id="4C6918C3AD7B6116FF68A5A2599CFDB5" authority="Vervoort, 1993" authorityName="Vervoort" authorityYear="1993" box="[189,704,493,520]" class="Hydrozoa" family="Sertulariidae" genus="Thyroscyphus" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="species" species="scorpioides">
<emphasis id="B91DBF52AD7B6116FF68A5A25AAAFDB5" box="[189,502,493,519]" italics="true" pageId="53" pageNumber="54">Thyroscyphus scorpioides</emphasis>
<bibRefCitation id="EFF81EB1AD7B6116FDD0A5A1599CFDB5" author="Vervoort" box="[517,704,493,520]" firstAuthor="Vervoort" pageId="53" pageNumber="54" pagination="89 - 298" refId="ref39987" refString="Vervoort W. 1993. Cnidaria, Hydrozoa, Hydroida: Hydroids from the western Pacific (Philippines, Indonesia and New Caledonia) I. Sertulariidae (Part 1). In: Crosnier A. (ed.) Resultats des Campagnes MUSORSTOM, Vol. 11. Memoires du Museum national d'histoire naturelle 158: 89 - 298." type="journal article" year="1993">Vervoort, 1993</bibRefCitation>
</taxonomicName>
, as
<taxonomicName id="4C6918C3AD7B6116FD2BA5A25E53FDB5" authority="(Vervoort, 1993)" authorityName="Galea &amp; Schuchert" authorityYear="2019" baseAuthorityName="Vervoort" baseAuthorityYear="1993" box="[766,1295,493,519]" class="Hydrozoa" family="Symplectoscyphidae" genus="Tuberocaulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="species" species="scorpioides" status="gen. et comb. nov.">
<emphasis id="B91DBF52AD7B6116FD2BA5A25F6FFDB5" box="[766,1075,493,519]" italics="true" pageId="53" pageNumber="54">Tuberocaulus scorpioides</emphasis>
(
<bibRefCitation id="EFF81EB1AD7B6116FB9EA5A25E5BFDB5" author="Vervoort" box="[1099,1287,493,519]" firstAuthor="Vervoort" pageId="53" pageNumber="54" pagination="89 - 298" refId="ref39987" refString="Vervoort W. 1993. Cnidaria, Hydrozoa, Hydroida: Hydroids from the western Pacific (Philippines, Indonesia and New Caledonia) I. Sertulariidae (Part 1). In: Crosnier A. (ed.) Resultats des Campagnes MUSORSTOM, Vol. 11. Memoires du Museum national d'histoire naturelle 158: 89 - 298." type="journal article" year="1993">Vervoort, 1993</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22E0229AD7B6116FAF5A5A15A63FD9E" pageId="53" pageNumber="54" rank="species">gen. et comb. nov.</taxonomicNameLabel>
The genus is currently monotypic.
</paragraph>
</subSubSection>
<subSubSection id="C37330CBAD7B6116FF68A61A5906FD77" pageId="53" pageNumber="54" type="etymology">
<paragraph id="8BD66340AD7B6116FF68A61A5A18FDDD" blockId="53.[189,324,597,623]" box="[189,324,597,623]" pageId="53" pageNumber="54">
<heading id="D09ED42CAD7B6116FF68A61A5A18FDDD" bold="true" box="[189,324,597,623]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="B91DBF52AD7B6116FF68A61A5A18FDDD" bold="true" box="[189,324,597,623]" pageId="53" pageNumber="54">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD66340AD7B6116FF68A6C95906FD77" blockId="53.[189,1399,645,709]" pageId="53" pageNumber="54">
From the Latin
<emphasis id="B91DBF52AD7B6116FEA8A6CA5AE7FD2D" box="[381,443,645,671]" italics="true" pageId="53" pageNumber="54">tūbĕr</emphasis>
, meaning protuberance, and
<emphasis id="B91DBF52AD7B6116FCE3A6CA5820FD2D" box="[822,892,645,671]" italics="true" pageId="53" pageNumber="54">caulis</emphasis>
, meaning stem, to illustrate the presence of putative nematothecae on stems.
</paragraph>
</subSubSection>
<subSubSection id="C37330CBAD7B6116FF68A6A25A97F923" pageId="53" pageNumber="54" type="discussion">
<paragraph id="8BD66340AD7B6116FF68A6A25A73FCB5" blockId="53.[189,303,749,775]" box="[189,303,749,775]" pageId="53" pageNumber="54">
<heading id="D09ED42CAD7B6116FF68A6A25A73FCB5" bold="true" box="[189,303,749,775]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="B91DBF52AD7B6116FF68A6A25A73FCB5" bold="true" box="[189,303,749,775]" pageId="53" pageNumber="54">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD66340AD7B6116FF68A7525F8BFC42" blockId="53.[189,1398,797,1008]" pageId="53" pageNumber="54">
The genus
<taxonomicName id="4C6918C3AD7B6116FEEDA7525A8BFC85" authority="Galea &amp; Schuchert, 2019" authorityName="Galea &amp; Schuchert" authorityYear="2019" box="[312,471,797,823]" class="Hydrozoa" family="Symplectoscyphidae" genus="Tuberocaulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="genus" status="gen. nov.">
<emphasis id="B91DBF52AD7B6116FEEDA7525A8BFC85" box="[312,471,797,823]" italics="true" pageId="53" pageNumber="54">Tuberocaulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E0229AD7B6116FE0EA751591FFC8A" box="[475,579,798,824]" pageId="53" pageNumber="54" rank="genus">gen. nov.</taxonomicNameLabel>
is monotypic so far and comprises
<taxonomicName id="4C6918C3AD7B6116FC05A7525A5DFCEE" authority="(Vervoort, 1993)" authorityName="Galea &amp; Schuchert" authorityYear="2019" baseAuthorityName="Vervoort" baseAuthorityYear="1993" class="Hydrozoa" family="Symplectoscyphidae" genus="Tuberocaulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="species" species="scorpioides" status="gen. et comb. nov.">
<emphasis id="B91DBF52AD7B6116FC05A7525FA6FC85" box="[976,1274,797,823]" italics="true" pageId="53" pageNumber="54">Tuberocaulus scorpioides</emphasis>
(
<bibRefCitation id="EFF81EB1AD7B6116FAD3A7525BA5FCEE" author="Vervoort" firstAuthor="Vervoort" pageId="53" pageNumber="54" pagination="89 - 298" refId="ref39987" refString="Vervoort W. 1993. Cnidaria, Hydrozoa, Hydroida: Hydroids from the western Pacific (Philippines, Indonesia and New Caledonia) I. Sertulariidae (Part 1). In: Crosnier A. (ed.) Resultats des Campagnes MUSORSTOM, Vol. 11. Memoires du Museum national d'histoire naturelle 158: 89 - 298." type="journal article" year="1993">Vervoort, 1993</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22E0229AD7B6116FEDCA70C5ABEFCEF" box="[265,482,834,861]" pageId="53" pageNumber="54" rank="species">gen. et comb. nov.</taxonomicNameLabel>
This species was originally placed in
<taxonomicName id="4C6918C3AD7B6116FC71A70D5FB2FCEE" authority="Allman, 1877" authorityName="Allman" authorityYear="1877" box="[932,1262,834,860]" class="Hydrozoa" family="Sertulariidae" genus="Thyroscyphus" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="genus">
<emphasis id="B91DBF52AD7B6116FC71A70D5F1AFCEE" box="[932,1094,834,860]" italics="true" pageId="53" pageNumber="54">Thyroscyphus</emphasis>
<bibRefCitation id="EFF81EB1AD7B6116FB99A70D5FB2FCEE" author="Allman" box="[1100,1262,834,860]" firstAuthor="Allman" pageId="53" pageNumber="54" pagination="1 - 66" refId="ref35871" refString="Allman G. J. 1877. Report on the Hydroida collected during the exploration of the Gulf Stream by L. F. de Pourtales, Assistant United States Coast Survey. Memoirs of the Museum of Comparative Zoology at Harvard College 5 (2): 1 - 66. https: // doi. org / 10.5962 / bhl. title. 10420" type="journal article" year="1877">Allman, 1877</bibRefCitation>
</taxonomicName>
on account of the attachment of the hydranth to the inner hydrothecal wall (whose surface is lined by “a sheath of tissue” = mantle) “by means of a circular fold”. However, unlike
<taxonomicName id="4C6918C3AD7B6116FC11A7C35F3AFC14" authorityName="Allman" authorityYear="1877" box="[964,1126,908,934]" class="Hydrozoa" family="Sertulariidae" genus="Thyroscyphus" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="genus">
<emphasis id="B91DBF52AD7B6116FC11A7C35F3AFC14" box="[964,1126,908,934]" italics="true" pageId="53" pageNumber="54">Thyroscyphus</emphasis>
</taxonomicName>
, the hydrotheca of this species is given off from the internode, and it is not borne on a distinct apophysis; additionally, it has a well-defined (though short) adnate adaxial wall, as well as a base (see description above).
</paragraph>
<paragraph id="8BD66340AD7B6116FF68A05759DCFBCE" blockId="53.[189,1399,1048,1149]" pageId="53" pageNumber="54">
The fascicled condition of the stem and its peculiar appearance, the presence of what appears to be stem nematothecae, as well as the unilateral arrangement of cladia and their hydrothecae, are not features shared by the genus of
<bibRefCitation id="EFF81EB1AD7B6116FE1EA02D5920FBCE" author="Allman" box="[459,636,1122,1148]" firstAuthor="Allman" pageId="53" pageNumber="54" pagination="1 - 66" refId="ref35871" refString="Allman G. J. 1877. Report on the Hydroida collected during the exploration of the Gulf Stream by L. F. de Pourtales, Assistant United States Coast Survey. Memoirs of the Museum of Comparative Zoology at Harvard College 5 (2): 1 - 66. https: // doi. org / 10.5962 / bhl. title. 10420" type="journal article" year="1877">Allman (1877)</bibRefCitation>
.
</paragraph>
<paragraph id="8BD66340AD7B6116FF68A0EA5F42FAE1" blockId="53.[189,1399,1189,1363]" pageId="53" pageNumber="54">
The systematic position of
<taxonomicName id="4C6918C3AD7B6116FDD2A0E959F2FB0D" authorityName="Galea &amp; Schuchert" authorityYear="2019" baseAuthorityName="Vervoort" baseAuthorityYear="1993" box="[519,686,1189,1215]" class="Hydrozoa" family="Symplectoscyphidae" genus="Tuberocaulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="species" species="scorpioides">
<emphasis id="B91DBF52AD7B6116FDD2A0E959F2FB0D" box="[519,686,1189,1215]" italics="true" pageId="53" pageNumber="54">T. scorpioides</emphasis>
</taxonomicName>
could not be ascertained on morphological grounds alone. Indeed, its long, tubular, tricuspidate hydrothecae superficially recall those of
<taxonomicName id="4C6918C3AD7B6116FBAFA0855AF9FAB8" authority="(Lamouroux, 1816)" baseAuthorityName="Lamouroux" baseAuthorityYear="1816" class="Hydrozoa" family="Thyroscyphidae" genus="Parascyphus" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="species" species="simplex">
<emphasis id="B91DBF52AD7B6116FBAFA0855E2BFB56" box="[1146,1399,1226,1252]" italics="true" pageId="53" pageNumber="54">Parascyphus simplex</emphasis>
(Lamouroux, 1816)
</taxonomicName>
, but in that species, the thecae are borne on distinct stem apophyses, their bases are represented by an internal annular thickening of the perisarc, and the marginal cusps adopt a different position (one adaxial and two latero-abaxial cusps, see
<bibRefCitation id="EFF81EB1AD7B6116FC90A1765F4EFAE1" author="Galea" box="[837,1042,1337,1363]" etAl="et al." firstAuthor="Galea" pageId="53" pageNumber="54" pagination="1 - 50" refId="ref37340" refString="Galea H. R., Schories D., Forsterra G. &amp; Haussermann V. 2014. New species and new records of hydroids (Cnidaria: Hydrozoa) from Chile. Zootaxa 3852 (1): 1 - 50. https: // doi. org / 10.11646 / zootaxa. 3852.1.1" type="journal article" year="2014">
Galea
<emphasis id="B91DBF52AD7B6116FC45A1765898FAE1" box="[912,964,1337,1363]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. 2014
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD66340AD7B6116FF68A1335AB1F9B6" blockId="53.[189,1399,1404,1541]" pageId="53" pageNumber="54">
Being fused to the corresponding internodes, the hydrothecae of
<taxonomicName id="4C6918C3AD7B6116FC36A1325FD0FA24" authorityName="Galea &amp; Schuchert" authorityYear="2019" baseAuthorityName="Vervoort" baseAuthorityYear="1993" box="[995,1164,1404,1430]" class="Hydrozoa" family="Symplectoscyphidae" genus="Tuberocaulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="species" species="scorpioides">
<emphasis id="B91DBF52AD7B6116FC36A1325FD0FA24" box="[995,1164,1404,1430]" italics="true" pageId="53" pageNumber="54">T. scorpioides</emphasis>
</taxonomicName>
are reminiscent of those met with in members of the Sertulariida
<bibRefCitation id="EFF81EB1AD7B6116FD08A1EE588FFA08" author="Maronna" box="[733,979,1440,1467]" etAl="et al." firstAuthor="Maronna" pageId="53" pageNumber="54" pagination="18075" refId="ref38063" refString="Maronna M. M., Miranda T. P., Pena Cantero A. L., Barbeitos M. S. &amp; Marques A. C. 2016. Towards a phylogenetic classification of Leptothecata (Cnidaria, Hydrozoa). Nature, Scientific Reports 6: 18075. https: // doi. org / 10.1038 / srep 18075" type="journal article" year="2016">
Maronna
<emphasis id="B91DBF52AD7B6116FC9BA1EE58DEFA09" box="[846,898,1441,1467]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
., 2016
</bibRefCitation>
(with only
<taxonomicName id="4C6918C3AD7B6116FB8EA1EE5E2BFA09" authority="Vervoort, 1993" authorityName="Vervoort" authorityYear="1993" box="[1115,1399,1441,1467]" class="Hydrozoa" family="Sertulariidae" genus="Gonaxia" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="genus">
<emphasis id="B91DBF52AD7B6116FB8EA1EE5F9CFA09" box="[1115,1216,1441,1467]" italics="true" pageId="53" pageNumber="54">Gonaxia</emphasis>
<bibRefCitation id="EFF81EB1AD7B6116FB13A1EE5E2BFA09" author="Vervoort" box="[1222,1399,1441,1467]" firstAuthor="Vervoort" pageId="53" pageNumber="54" pagination="89 - 298" refId="ref39987" refString="Vervoort W. 1993. Cnidaria, Hydrozoa, Hydroida: Hydroids from the western Pacific (Philippines, Indonesia and New Caledonia) I. Sertulariidae (Part 1). In: Crosnier A. (ed.) Resultats des Campagnes MUSORSTOM, Vol. 11. Memoires du Museum national d'histoire naturelle 158: 89 - 298." type="journal article" year="1993">Vervoort, 1993</bibRefCitation>
</taxonomicName>
showing the same position of the marginal cusps), although nematothecae have never been reported in any genus belonging to it.
</paragraph>
<paragraph id="8BD66340AD7B6116FF68A2625A97F923" blockId="53.[189,1399,1581,1681]" pageId="53" pageNumber="54">
The relationship of
<taxonomicName id="4C6918C3AD7B6116FE4BA2625911F9F5" authority="Galea &amp; Schuchert, 2019" authorityName="Galea &amp; Schuchert" authorityYear="2019" box="[414,589,1581,1607]" class="Hydrozoa" family="Thyroscyphidae" genus="Tuberoscyphus" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="genus" status="gen. nov.">
<emphasis id="B91DBF52AD7B6116FE4BA2625911F9F5" box="[414,589,1581,1607]" italics="true" pageId="53" pageNumber="54">Tuberoscyphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E0229AD7B6116FD87A26159E7F9FA" box="[594,699,1582,1608]" pageId="53" pageNumber="54" rank="genus">gen. nov.</taxonomicNameLabel>
to the other
<taxonomicName id="4C6918C3AD7B6116FC9DA26258BCF9F5" authorityName="Cornelius" authorityYear="1992" box="[840,992,1581,1607]" class="Hydrozoa" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="order">Leptothecata</taxonomicName>
families remains to be investigated by molecular phylogenies. In the absence of such studies for the time being, the genus is left provisionally in the
<taxonomicName id="4C6918C3AD7B6116FED3A2385A9BF923" authorityName="Stechow" authorityYear="1920" box="[262,455,1655,1681]" class="Hydrozoa" family="Thyroscyphidae" kingdom="Animalia" order="Leptothecata" pageId="53" pageNumber="54" phylum="Cnidaria" rank="family">Thyroscyphidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>