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<document id="3D9BB32A872712CAD532955B648745DB" ID-DOI="10.5281/zenodo.191105" ID-GBIF-Dataset="03517f19-f5a0-490d-b967-46594ab736b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="191105" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461124192379" checkinUser="plazi" docAuthor="Britz, Ralf" docDate="2009" docId="03B3E452FFFCFFC3BF922F82A465BB66" docLanguage="en" docName="zt02277p060.pdf" docOrigin="Zootaxa 2277" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Danionella priapus Britz, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="FF8A9C2AFFFCFFC7BF052905A129BC5A" masterDocTitle="Danionella priapus, a new species of miniature cyprinid fish from West Bengal, India (Teleostei: Cypriniformes: Cyprinidae)" masterLastPageNumber="60" masterPageNumber="53" pageNumber="53" updateTime="1698596414738" updateUser="plazi">
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<mods:title id="D5DEA2F0675DBBCB195D7C8229CC5E43">Danionella priapus, a new species of miniature cyprinid fish from West Bengal, India (Teleostei: Cypriniformes: Cyprinidae)</mods:title>
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<mods:name id="8B08920B4EABBB7B0410F38F4CC7AB99" type="personal">
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<mods:namePart id="3D8CB0A9765979CC10B392737648B4D1">Britz, Ralf</mods:namePart>
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<mods:typeOfResource id="21A8AEB774AFDD882E1F337BF3BEB2E8">text</mods:typeOfResource>
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<mods:title id="6490B3FEAFBC638FDB2022EFB5AC6224">Zootaxa</mods:title>
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<mods:date id="44DF451386E78877ED001355BBFC7341">2009</mods:date>
<mods:detail id="B24EE45B6A425ACF666B80EA88ED86A2" type="volume">
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<mods:classification id="B8E890CEAD61D2C080AB70D9F73F54CA">journal article</mods:classification>
<mods:identifier id="92AEDA9069E2A81B1CD34F453C4C8E3B" type="DOI">10.5281/zenodo.191105</mods:identifier>
<mods:identifier id="90A73900E269D4836EB04ABAB2E2AA1F" type="GBIF-Dataset">03517f19-f5a0-490d-b967-46594ab736b8</mods:identifier>
<mods:identifier id="7E24C6BE3696BF12FE9B7710D36F9C92" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3A40410AB36531A1A99F97EABCC4FA78" type="Zenodo-Dep">191105</mods:identifier>
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<treatment id="03B3E452FFFCFFC3BF922F82A465BB66" ID-DOI="http://doi.org/10.5281/zenodo.6213021" ID-GBIF-Taxon="119626042" ID-Zenodo-Dep="6213021" LSID="urn:lsid:plazi:treatment:03B3E452FFFCFFC3BF922F82A465BB66" httpUri="http://treatment.plazi.org/id/03B3E452FFFCFFC3BF922F82A465BB66" lastPageId="4" lastPageNumber="57" pageId="0" pageNumber="53">
<subSubSection id="C30006CFFFFCFFC7BF922F82A03EBA9E" pageId="0" pageNumber="53" type="nomenclature">
<paragraph id="8BA55544FFFCFFC7BF922F82A0C5BAFB" blockId="0.[151,492,1671,1732]" box="[151,492,1671,1697]" pageId="0" pageNumber="53">
<heading id="D0EDE228FFFCFFC7BF922F82A0C5BAFB" bold="true" box="[151,492,1671,1697]" fontSize="11" level="1" pageId="0" pageNumber="53" reason="1">
<emphasis id="B96E8956FFFCFFC7BF922F82A0C5BAFB" bold="true" box="[151,492,1671,1697]" pageId="0" pageNumber="53">
<taxonomicName id="4C1A2EC7FFFCFFC7BF922F82A0A9BAFB" ID-CoL="34447" box="[151,384,1671,1697]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="0" pageNumber="53" phylum="Chordata" rank="species" species="priapus" status="sp. nov.">
<emphasis id="B96E8956FFFCFFC7BF922F82A0A9BAFB" bold="true" box="[151,384,1671,1697]" italics="true" pageId="0" pageNumber="53">Danionella priapus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25D342DFFFCFFC7BE8A2F82A0C5BAFB" box="[399,492,1671,1697]" pageId="0" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA55544FFFCFFC7BF922FAFA03EBA9E" blockId="0.[151,492,1671,1732]" box="[151,279,1706,1732]" pageId="0" pageNumber="53">
(
<figureCitation id="132149C1FFFCFFC7BF9A2FAFA026BA9E" box="[159,271,1706,1732]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,260,727,751]" captionStartId-1="1.[151,255,1831,1855]" captionStartId-2="2.[151,258,1913,1937]" captionStartId-3="5.[151,255,710,734]" captionTargetBox-0="[327,1258,197,702]" captionTargetBox-1="[440,1147,874,1806]" captionTargetBox-2="[226,1360,197,1887]" captionTargetBox-3="[437,1150,197,685]" captionTargetId-0="figure@1.[327,1259,197,703]" captionTargetId-1="figure@1.[440,1147,874,1807]" captionTargetId-2="figure@2.[226,1360,197,1888]" captionTargetId-3="figure@5.[437,1150,197,687]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionTargetPageId-2="2" captionTargetPageId-3="5" captionText-0="FIGURE 1. Lateral view of Danionella priapus: a. BMNH 2009.9. 9.1, holotype, male, 14.4 mm SL; b. BMNH 2009.9.9.2 - 37, paratype, female, 15.0 mm SL. Note presence of eggs and larval preanal fin fold in female, and difference in size and shape of pelvic fins and in position of anus and genital papilla between male and female." captionText-1="FIGURE 2. Difference in pelvic-fin structure in Danionella priapus: a holotype, BMNH 2009.9. 9.1, male, 14.4 mm SL, close-up of modified pelvic fins and conically projecting genital papilla; b, BMNH 2009.9.9.2 - 37, paratype, female, 15.0 mm SL, close-up of pelvic fins and preanal fin fold." captionText-2="FIGURE 3. Dorsal and ventral view of colour pattern of head and anterior body in Danionella priapus, holotype, BMNH 2009.9. 9.1, male, 14.4 mm SL." captionText-3="FIGURE 4. Live colouration of male (above) and female (below) Danionella priapus. Specimens not preserved. Note eggs in body cavity of female." httpUri-0="https://zenodo.org/record/191106/files/figure.png" httpUri-1="https://zenodo.org/record/191107/files/figure.png" httpUri-2="https://zenodo.org/record/191108/files/figure.png" httpUri-3="https://zenodo.org/record/191109/files/figure.png" pageId="0" pageNumber="53">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30006CFFFFCFFC7BF922FFFA354BB4A" box="[151,637,1784,1809]" pageId="0" pageNumber="53" type="description">
<paragraph id="8BA55544FFFCFFC7BF922FFFA354BB4A" blockId="0.[151,637,1784,1809]" box="[151,637,1784,1809]" pageId="0" pageNumber="53">
<taxonomicName id="4C1A2EC7FFFCFFC7BF922FFFA026BB4B" box="[151,271,1786,1809]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="0" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B96E8956FFFCFFC7BF922FFFA026BB4B" box="[151,271,1786,1809]" italics="true" pageId="0" pageNumber="53">Danionella</emphasis>
</taxonomicName>
sp.: Jorai River,
<bibRefCitation id="EF8B28B5FFFCFFC7BEC12FFCA354BB4A" author="Britz" box="[452,637,1784,1809]" pageId="0" pageNumber="60" refString="Britz, R., Conway, K. W. &amp; Ruber, L. (2009) Spectacular morphological novelty in a miniature cyprinid fish, Danionella dracula n. sp. Proceedings of the Royal Society B, 276, 2179 - 2186." type="journal article" year="2009">
Britz
<emphasis id="B96E8956FFFCFFC7BD052FFDA313BB4B" box="[512,570,1784,1809]" italics="true" pageId="0" pageNumber="53">et al.</emphasis>
, 2009
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30006CFFFFCFFC7BF922E45A2C8BBAC" pageId="0" pageNumber="53" type="materials_examined">
<paragraph id="8BA55544FFFCFFC7BF922E45A3E7BBF2" blockId="0.[151,1436,1856,2038]" pageId="0" pageNumber="53">
<emphasis id="B96E8956FFFCFFC7BF922E45A07ABB00" bold="true" box="[151,339,1856,1882]" pageId="0" pageNumber="53">
<typeStatus id="54A1EBE6FFFCFFC7BF922E45A1FFBB00" box="[151,214,1856,1882]" pageId="0" pageNumber="53">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54A1EBE6FFFCFFC7BE5E2E44A0F9BB01" box="[347,464,1857,1883]" pageId="0" pageNumber="53" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED0BCD81FFFCFFC7BED92E44A31FBB01" box="[476,566,1857,1883]" pageId="0" pageNumber="53">BMNH</collectionCode>
2009.9.9.1, male,
<quantity id="4CE2F8A1FFFCFFC7BC1F2E44A2ACBB01" box="[794,901,1857,1883]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="0" pageNumber="53" unit="mm" value="14.4">14.4 mm</quantity>
SL;
<collectingCountry id="F30D15D4FFFCFFC7BCC72E44A529BB01" box="[962,1024,1857,1883]" name="India" pageId="0" pageNumber="53">India</collectingCountry>
, West Bengal, Jalpaiguri District, Brahmaputra drainage, Jorai River, a tributary of the Sankosh at Laskarpara, outskirts of Barobisha town.
<geoCoordinate id="EE2E3383FFFCFFC7BA762E6DA03EBBF2" direction="north" orientation="latitude" pageId="0" pageNumber="53" precision="1" value="26.481195">26° 28' 52.3''N</geoCoordinate>
,
<geoCoordinate id="EE2E3383FFFCFFC7BE272E8BA0E6BBF2" box="[290,463,1934,1960]" direction="east" orientation="longitude" pageId="0" pageNumber="53" precision="1" value="89.82494">89° 49' 29.8''E</geoCoordinate>
; M. Das,
<date id="FFA47384FFFCFFC7BD442E8BA3E3BBF2" box="[577,714,1934,1960]" pageId="0" pageNumber="53" value="2008-04-02">2 Apr 2008</date>
.
</paragraph>
<paragraph id="8BA55544FFFCFFC7BFC02EB0A2C8BBAC" blockId="0.[151,1436,1856,2038]" pageId="0" pageNumber="53">
<typeStatus id="54A1EBE6FFFCFFC7BFC02EB0A017BB95" box="[197,318,1973,1999]" pageId="0" pageNumber="53" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="ED0BCD81FFFCFFC7BE4C2EB0A08BBB95" box="[329,418,1973,1999]" pageId="0" pageNumber="53">BMNH</collectionCode>
2009.9.9.2-37 (36), 11.8–16.0 mm SL; data as for
<typeStatus id="54A1EBE6FFFCFFC7BB0D2EB0A55DBB95" box="[1032,1140,1973,1999]" pageId="0" pageNumber="53" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED0BCD81FFFCFFC7BB842EB0A5F3BB95" box="[1153,1242,1973,1999]" pageId="0" pageNumber="53">BMNH</collectionCode>
2009.
<date id="FFA47384FFFCFFC7BA202EB0A41DBB95" box="[1317,1332,1973,1999]" pageId="0" pageNumber="53" value="1938-09-09">9</date>
.9.38–43 (6), cleared and double stained,
<quantity id="4CE2F8A1FFFCFFC7BD082ED9A391BBAC" box="[525,696,2012,2038]" metricMagnitude="-2" metricUnit="m" metricValue="1.475" metricValueMax="1.59" metricValueMin="1.3599999999999999" pageId="0" pageNumber="53" unit="mm" value="14.75" valueMax="15.9" valueMin="13.6">13.6–15.9 mm</quantity>
SL; data as for
<typeStatus id="54A1EBE6FFFCFFC7BC772ED9A2F4BBAC" box="[882,989,2012,2038]" pageId="0" pageNumber="53" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="DF6505CCFFFDFFC6BF922BD2A5EABF6E" httpUri="https://zenodo.org/record/191106/files/figure.png" pageId="1" pageNumber="54" targetBox="[327,1258,197,702]" targetPageId="1">
<paragraph id="8BA55544FFFDFFC6BF922BD2A5EABF6E" blockId="1.[151,1437,727,820]" pageId="1" pageNumber="54">
<emphasis id="B96E8956FFFDFFC6BF922BD2A00BBEB5" bold="true" box="[151,290,727,751]" pageId="1" pageNumber="54">FIGURE 1.</emphasis>
Lateral view of
<taxonomicName id="4C1A2EC7FFFDFFC6BEE72BDDA3E8BEB5" box="[482,705,728,751]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="priapus">
<emphasis id="B96E8956FFFDFFC6BEE72BDDA3E8BEB5" box="[482,705,728,751]" italics="true" pageId="1" pageNumber="54">Danionella priapus</emphasis>
</taxonomicName>
: a. BMNH 2009.9.9.1, holotype, male, 14.4 mm SL; b. BMNH 2009.9.9.2-37, paratype, female, 15.0 mm SL. Note presence of eggs and larval preanal fin fold in female, and difference in size and shape of pelvic fins and in position of anus and genital papilla between male and female.
</paragraph>
</caption>
<caption id="DF6505CCFFFDFFC6BF922E22A3E9BBDE" httpUri="https://zenodo.org/record/191107/files/figure.png" pageId="1" pageNumber="54" targetBox="[440,1147,874,1806]" targetPageId="1">
<paragraph id="8BA55544FFFDFFC6BF922E22A3E9BBDE" blockId="1.[151,1436,1831,1924]" pageId="1" pageNumber="54">
<emphasis id="B96E8956FFFDFFC6BF922E22A030BB65" bold="true" box="[151,281,1831,1855]" pageId="1" pageNumber="54">FIGURE 2.</emphasis>
Difference in pelvic-fin structure in
<taxonomicName id="4C1A2EC7FFFDFFC6BDA52E2DA259BB65" box="[672,880,1832,1855]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="priapus">
<emphasis id="B96E8956FFFDFFC6BDA52E2DA259BB65" box="[672,880,1832,1855]" italics="true" pageId="1" pageNumber="54">Danionella priapus</emphasis>
</taxonomicName>
: a holotype, BMNH 2009.9.9.1, male, 14.4 mm SL, close-up of modified pelvic fins and conically projecting genital papilla; b, BMNH 2009.9.9.2-37, paratype, female, 15.0 mm SL, close-up of pelvic fins and preanal fin fold.
</paragraph>
</caption>
<caption id="DF6505CCFFFEFFC5BF922E7CA310BBE9" httpUri="https://zenodo.org/record/191108/files/figure.png" pageId="2" pageNumber="55" targetBox="[226,1360,197,1887]" targetPageId="2">
<paragraph id="8BA55544FFFEFFC5BF922E7CA310BBE9" blockId="2.[151,1436,1913,1971]" pageId="2" pageNumber="55">
<emphasis id="B96E8956FFFEFFC5BF922E7CA036BBCB" bold="true" box="[151,287,1913,1937]" pageId="2" pageNumber="55">FIGURE 3.</emphasis>
Dorsal and ventral view of colour pattern of head and anterior body in
<taxonomicName id="4C1A2EC7FFFEFFC5BB4C2E7FA40DBBCB" box="[1097,1316,1914,1937]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="priapus">
<emphasis id="B96E8956FFFEFFC5BB4C2E7FA40DBBCB" box="[1097,1316,1914,1937]" italics="true" pageId="2" pageNumber="55">Danionella priapus</emphasis>
</taxonomicName>
, holotype, BMNH 2009.9.9.1, male, 14.4 mm SL.
</paragraph>
</caption>
<subSubSection id="C30006CFFFFFFFC4BFC02992A08EBF46" pageId="3" pageNumber="56" type="diagnosis">
<paragraph id="8BA55544FFFFFFC4BFC02992A08EBF46" blockId="3.[151,1437,151,2034]" pageId="3" pageNumber="56">
<emphasis id="B96E8956FFFFFFC4BFC02992A061BCEB" bold="true" box="[197,328,151,177]" pageId="3" pageNumber="56">Diagnosis.</emphasis>
<taxonomicName id="4C1A2EC7FFFFFFC4BE54299DA314BCEB" box="[337,573,152,177]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="priapus">
<emphasis id="B96E8956FFFFFFC4BE54299DA314BCEB" box="[337,573,152,177]" italics="true" pageId="3" pageNumber="56">Danionella priapus</emphasis>
</taxonomicName>
is distinguished from its congeners,
<emphasis id="B96E8956FFFFFFC4BCFA299DA46BBCEB" box="[1023,1346,152,177]" italics="true" pageId="3" pageNumber="56">
<taxonomicName id="4C1A2EC7FFFFFFC4BCFA299DA5AFBCEB" box="[1023,1158,152,177]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="dracula">D. dracula</taxonomicName>
,
<taxonomicName id="4C1A2EC7FFFFFFC4BB96299DA46BBCEB" box="[1171,1346,152,177]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="translucida">D. translucida</taxonomicName>
</emphasis>
and
<taxonomicName id="4C1A2EC7FFFFFFC4BA7A299DA1DEBC82" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="mirifica">
<emphasis id="B96E8956FFFFFFC4BA7A299DA1DEBC82" italics="true" pageId="3" pageNumber="56">D. mirifica</emphasis>
</taxonomicName>
, by the presence in adult males of a conical projection of the genital papilla situated between the pelvic fins, which form a funnel-like structure (vs. genital papilla not developed as a conical projection and pelvic fins not funnel-shaped), by possessing 8 pectoral-fin rays (vs. 6–7) and 20–21 anal–fin rays (vs.
<quantity id="4CE2F8A1FFFFFFC4BA572809A187BD16" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.556" metricValueMin="3.048" pageId="3" pageNumber="56" unit="in" value="13.0" valueMax="14.0" valueMin="12.0">12–14 in</quantity>
<taxonomicName id="4C1A2EC7FFFFFFC4BFB32837A01EBD11" box="[182,311,306,331]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="dracula">
<emphasis id="B96E8956FFFFFFC4BFB32837A01EBD11" box="[182,311,306,331]" italics="true" pageId="3" pageNumber="56">D. dracula</emphasis>
</taxonomicName>
,
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<taxonomicName id="4C1A2EC7FFFFFFC4BEBD2837A34DBD11" box="[440,612,306,331]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="translucida">
<emphasis id="B96E8956FFFFFFC4BEBD2837A34DBD11" box="[440,612,306,331]" italics="true" pageId="3" pageNumber="56">D. translucida</emphasis>
</taxonomicName>
, and 17–19, rarely
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<emphasis id="B96E8956FFFFFFC4BC912837A53EBD11" box="[916,1047,306,331]" italics="true" pageId="3" pageNumber="56">D. mirifica</emphasis>
</taxonomicName>
), and by having the last anal-fin pterygiophore inserted in front of haemal spine of vertebra 27 or 28 (vs. 21 or
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<taxonomicName id="4C1A2EC7FFFFFFC4BB7E285CA40DBD28" box="[1147,1316,345,370]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="translucida">
<emphasis id="B96E8956FFFFFFC4BB7E285CA40DBD28" box="[1147,1316,345,370]" italics="true" pageId="3" pageNumber="56">D. translucida</emphasis>
</taxonomicName>
,
<quantity id="4CE2F8A1FFFFFFC4BA31285CA4B5BD29" box="[1332,1436,345,371]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" metricValueMax="6.35" metricValueMin="5.842" pageId="3" pageNumber="56" unit="in" value="24.0" valueMax="25.0" valueMin="23.0">23–25 in</quantity>
<taxonomicName id="4C1A2EC7FFFFFFC4BF922885A031BDC3" box="[151,280,384,409]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="mirifica">
<emphasis id="B96E8956FFFFFFC4BF922885A031BDC3" box="[151,280,384,409]" italics="true" pageId="3" pageNumber="56">D. mirifica</emphasis>
</taxonomicName>
, and
<quantity id="4CE2F8A1FFFFFFC4BE5D2885A096BDC0" box="[344,447,384,410]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.096" metricValueMin="5.588" pageId="3" pageNumber="56" unit="in" value="23.0" valueMax="24.0" valueMin="22.0">22–24 in</quantity>
<taxonomicName id="4C1A2EC7FFFFFFC4BEC32885A36CBDC3" box="[454,581,384,409]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="dracula">
<emphasis id="B96E8956FFFFFFC4BEC32885A36CBDC3" box="[454,581,384,409]" italics="true" pageId="3" pageNumber="56">D. dracula</emphasis>
</taxonomicName>
). It differs further from
<taxonomicName id="4C1A2EC7FFFFFFC4BC672885A522BDC3" box="[866,1035,384,409]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="translucida">
<emphasis id="B96E8956FFFFFFC4BC672885A522BDC3" box="[866,1035,384,409]" italics="true" pageId="3" pageNumber="56">D. translucida</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1A2EC7FFFFFFC4BB412885A5ECBDC3" box="[1092,1221,384,409]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="mirifica">
<emphasis id="B96E8956FFFFFFC4BB412885A5ECBDC3" box="[1092,1221,384,409]" italics="true" pageId="3" pageNumber="56">D. mirifica</emphasis>
</taxonomicName>
in the presence of a median and two paramedian rows of pigment cells on the dorsal side of the body (vs. pigment rows absent). It is further distinguished from
<taxonomicName id="4C1A2EC7FFFFFFC4BD0D28C8A3AEBDBC" box="[520,647,461,486]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="dracula">
<emphasis id="B96E8956FFFFFFC4BD0D28C8A3AEBDBC" box="[520,647,461,486]" italics="true" pageId="3" pageNumber="56">D. dracula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1A2EC7FFFFFFC4BDC428C8A26ABDBC" box="[705,835,461,486]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="mirifica">
<emphasis id="B96E8956FFFFFFC4BDC428C8A26ABDBC" box="[705,835,461,486]" italics="true" pageId="3" pageNumber="56">D. mirifica</emphasis>
</taxonomicName>
by 7–8 dorsal caudal procurrent rays (vs. 5–6) and 6–8 ventral procurrent rays (vs. 4–5). In addition it differs from
<taxonomicName id="4C1A2EC7FFFFFFC4BC7A28F1A2D4BE57" box="[895,1021,500,525]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="dracula">
<emphasis id="B96E8956FFFFFFC4BC7A28F1A2D4BE57" box="[895,1021,500,525]" italics="true" pageId="3" pageNumber="56">D. dracula</emphasis>
</taxonomicName>
in having two upper jaw bones (vs. one); edges of jaw bones entire, without processes (vs. dorsal face of dentary and ventral face of upper jaw in males each with a single row of 6–13 odontoid processes, anterior most large and canine-like); presence of maxillary-mandibular cartilage (vs. absence); absence of membrane bone flanges on basipterygium (vs. presence); and 9+9 principal caudal fin rays (vs. 7–8 + 7–8). It differs also from
<taxonomicName id="4C1A2EC7FFFFFFC4BB4C2B8AA5DABEF2" box="[1097,1267,655,680]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="translucida">
<emphasis id="B96E8956FFFFFFC4BB4C2B8AA5DABEF2" box="[1097,1267,655,680]" italics="true" pageId="3" pageNumber="56">D. translucida</emphasis>
</taxonomicName>
in having 15– 16 abdominal (vs. 13–14) and 22–23 caudal (vs. 19–20) vertebrae and a well developed lateral stripe extending from ear capsule to caudal peduncle (vs. a row of few melanophores midlaterally, restricted to posterior part of body).
</paragraph>
</subSubSection>
<subSubSection id="C30006CFFFFFFFC3BFC32A2DA281BED8" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56" type="description">
<paragraph id="8BA55544FFFFFFC4BFC32A2DA51EB8C5" blockId="3.[151,1437,151,2034]" pageId="3" pageNumber="56">
<emphasis id="B96E8956FFFFFFC4BFC32A2DA077BF18" bold="true" box="[198,350,808,834]" pageId="3" pageNumber="56">Description.</emphasis>
Morphometric information, based on
<specimenCount id="9D1C9ECDFFFFFFC4BC4D2A2CA2DBBF19" box="[840,1010,809,835]" pageId="3" pageNumber="56" type="generic">15 specimens</specimenCount>
, is presented in
<tableCitation id="C69860FFFFFFFFC4BBD52A2CA419BF19" box="[1232,1328,809,835]" captionStart="TABLE 1" captionStartId="4.[151,240,847,871]" captionText="TABLE 1. Selected morphometric information on 15 specimens (7 males, 8 females) of Danionella priapus, including the holotype." pageId="3" pageNumber="56">Table 1</tableCitation>
. Known maximum size 16.0 mm SL. Head and eye large, mouth supraterminal (
<figureCitation id="132149C1FFFFFFC4BCF12A55A513BF30" box="[1012,1082,848,874]" captionStart="FIGURE 1" captionStartId="1.[151,260,727,751]" captionTargetBox="[327,1258,197,702]" captionTargetId="figure@1.[327,1259,197,703]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of Danionella priapus: a. BMNH 2009.9. 9.1, holotype, male, 14.4 mm SL; b. BMNH 2009.9.9.2 - 37, paratype, female, 15.0 mm SL. Note presence of eggs and larval preanal fin fold in female, and difference in size and shape of pelvic fins and in position of anus and genital papilla between male and female." httpUri="https://zenodo.org/record/191106/files/figure.png" pageId="3" pageNumber="56">Fig. 1</figureCitation>
). Body laterally compressed, elongate; dorsal fin short, situated opposite posterior half of long anal fin. Caudal fin furcate, with remnants of larval fin fold in front of its dorsal and ventral margins. Remnant of pre-anal larval fin fold present in females only (
<figureCitation id="132149C1FFFFFFC4BFDA2AC1A00CBF84" box="[223,293,964,990]" captionStart="FIGURE 1" captionStartId="1.[151,260,727,751]" captionTargetBox="[327,1258,197,702]" captionTargetId="figure@1.[327,1259,197,703]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of Danionella priapus: a. BMNH 2009.9. 9.1, holotype, male, 14.4 mm SL; b. BMNH 2009.9.9.2 - 37, paratype, female, 15.0 mm SL. Note presence of eggs and larval preanal fin fold in female, and difference in size and shape of pelvic fins and in position of anus and genital papilla between male and female." httpUri="https://zenodo.org/record/191106/files/figure.png" pageId="3" pageNumber="56">Fig. 1</figureCitation>
b). Anus and genital papilla of adult males located between pelvic fins, in females at normal position in front of anal fin. Genital opening of adult males located at tip of short, conical projection at proximal end of funnel-like structure formed by pelvic fins being fused along their midline (
<figureCitation id="132149C1FFFFFFC4BA0D2D14A472B871" box="[1288,1371,1041,1067]" captionStart="FIGURE 1" captionStartId="1.[151,260,727,751]" captionTargetBox="[327,1258,197,702]" captionTargetId="figure@1.[327,1259,197,703]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of Danionella priapus: a. BMNH 2009.9. 9.1, holotype, male, 14.4 mm SL; b. BMNH 2009.9.9.2 - 37, paratype, female, 15.0 mm SL. Note presence of eggs and larval preanal fin fold in female, and difference in size and shape of pelvic fins and in position of anus and genital papilla between male and female." httpUri="https://zenodo.org/record/191106/files/figure.png" pageId="3" pageNumber="56">Figs. 1</figureCitation>
a, 2). Pseudotypanum present in body musculature at lateral side of anterior gas-bladder chamber, rendering its pigmented surface visible (
<figureCitation id="132149C1FFFFFFC4BEDE2D5BA30BB822" box="[475,546,1118,1144]" captionStart="FIGURE 1" captionStartId="1.[151,260,727,751]" captionTargetBox="[327,1258,197,702]" captionTargetId="figure@1.[327,1259,197,703]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of Danionella priapus: a. BMNH 2009.9. 9.1, holotype, male, 14.4 mm SL; b. BMNH 2009.9.9.2 - 37, paratype, female, 15.0 mm SL. Note presence of eggs and larval preanal fin fold in female, and difference in size and shape of pelvic fins and in position of anus and genital papilla between male and female." httpUri="https://zenodo.org/record/191106/files/figure.png" pageId="3" pageNumber="56">Fig. 1</figureCitation>
). Body muscles greatly thinned out at lateral side of posterior gas bladder chamber. Scales absent. Lateral-line canals and pores on head and body absent.
</paragraph>
<paragraph id="8BA55544FFFFFFC4BFC02DA9A2EEBA35" blockId="3.[151,1437,151,2034]" pageId="3" pageNumber="56">Skull, hyopalatine arch, gill arches, and endoskeletal shoulder girdle mostly cartilaginous with perichondral ossifications giving skeleton overall larval appearance. Numerous bones and cartilages absent: kinethmoid (and kinethmoid cartilage), preethmoid, vomer, dermethmoid, nasal, parietal, intercalar, extrascapular, infraorbitals 2–5, ectopterygoid, metapterygoid, urohyal, hypobranchials 1–3, sclerotic bones, posttemporal, postcleithrum, mesocoracoid, pectoral radials 3–4 (and pectoral radial cartilages 3–4), pelvic radials 1–3 (and pelvic radial cartilages 2–3), supraneurals 2 and 5–9 (and supraneural cartilages 5–9), middle and distal radials in dorsal and anal fins, epineurals, epipleurals, uroneural 2, and scales. Ceratobranchial 5 heavily ossified, bearing 4–5 tri- to quadricuspid teeth plus the tips of one or two replacement teeth. Outer arm of os suspensorium sexually dimorphic, being enlarged and hyperossified in males and in confluence with lateral process of second vertebra via anterior process and with inner arm of os suspensorium below tripus via broad median flange. Males with a relatively large conical nodule of cartilage between fifth rib and outer arm of suspensorium, its proximal surface approaching swimbladder wall.</paragraph>
<paragraph id="8BA55544FFFFFFC4BFC02F79A3EBBB24" blockId="3.[151,1437,151,2034]" pageId="3" pageNumber="56">Vertebrae totalling 37 (2) or 38 (4), abdominal vertebrae 15 (3) or 16 (3); caudal vertebrae 22 (5) or 23 (1). Ribs present on vertebrae 5 to 14. Dorsal-fin rays ii,6,i (5) or ii,7,i (1). First dorsal-fin ray pterygiophore inserted behind neural spine of vertebra 21 (3) or 22 (3), and last in front of neural spine of vertebra 25 (2) or 26 (4). Anal-fin rays ii,17,i (2) or ii,18,i (4). Number of anal-fin pterygiophores in front of first hemal spine: 0 (2), 1 (2) or 2 (2). Last anal-fin pterygiphore inserted in front of neural spine of vertebra 27 (3) or 28 (3). Principal caudal-fin rays 9+9 (6) plus 7 (5) or 8 (1) dorsal and 6 (2), 7 (1) or 8 (3) ventral procurrent rays. Pectoral-fin rays 8 (6) and pelvic-fin rays 5 (6).</paragraph>
<paragraph id="8BA55544FFFFFFC3BFC02E8FA599BE54" blockId="3.[151,1437,151,2034]" lastBlockId="4.[151,1437,152,796]" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
<emphasis id="B96E8956FFFFFFC4BFC02E8FA04DBBFE" bold="true" box="[197,356,1930,1956]" pageId="3" pageNumber="56">Colouration.</emphasis>
Pigmentation in alcohol specimens restricted to several rows of melanophores (
<figureCitation id="132149C1FFFFFFC4BA1F2E8FA445BBFE" box="[1306,1388,1930,1956]" captionStart="FIGURE 1" captionStartId="1.[151,260,727,751]" captionTargetBox="[327,1258,197,702]" captionTargetId="figure@1.[327,1259,197,703]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of Danionella priapus: a. BMNH 2009.9. 9.1, holotype, male, 14.4 mm SL; b. BMNH 2009.9.9.2 - 37, paratype, female, 15.0 mm SL. Note presence of eggs and larval preanal fin fold in female, and difference in size and shape of pelvic fins and in position of anus and genital papilla between male and female." httpUri="https://zenodo.org/record/191106/files/figure.png" pageId="3" pageNumber="56">Figs. 1</figureCitation>
,
<figureCitation id="132149C1FFFFFFC4BA7E2E8FA4A5BBFE" box="[1403,1420,1930,1956]" captionStart="FIGURE 3" captionStartId="2.[151,258,1913,1937]" captionTargetBox="[226,1360,197,1887]" captionTargetId="figure@2.[226,1360,197,1888]" captionTargetPageId="2" captionText="FIGURE 3. Dorsal and ventral view of colour pattern of head and anterior body in Danionella priapus, holotype, BMNH 2009.9. 9.1, male, 14.4 mm SL." httpUri="https://zenodo.org/record/191108/files/figure.png" pageId="3" pageNumber="56">3</figureCitation>
): mid-lateral row along horizontal septum from shoulder girdle to hypural plate; ventral row from in front of and slightly above anal-fin base along ventral larval fin fold to end of hypural plate; row along anal-fin base; mid-dorsal row starting behind head with several enlarged melanophores forming small blotch and extending usually halfway between head and anterior base of dorsal fin, rarely to anterior dorsal-fin base; paired dorsal paramedian rows from head to base of caudal fin; dorsal-fin base row; abdominal mid-ventral row from ventral tip of cleithrum to anus; several melanophores on otic capsule; melanophores capping dorsal and dorso-lateral face of gasbladder chambers and their connecting duct and lining peritoneum covering gonads; streaks of melanophores lining anterior edge of lower cleithrum, ventral edge of opercle and preopercle, branchiostegal rays and fin rays in dorsal, anal and caudal fins and anterior two or three rays of pectoral fin; and several larger melanophores near symphysis of lower jaw. Head with supraoccipital blotch formed by several large contiguous melanophores, with numerous large scattered melanophores between supraoccipital blotch and epiphyseal bar, with a preepiphyseal blotch, and a few prenasal melanophores.
</paragraph>
<paragraph id="8BA55544FFF8FFC3BFC02B1FA281BED8" blockId="4.[151,1437,152,796]" pageId="4" pageNumber="57">
In life, body colourless and largely translucent (
<figureCitation id="132149C1FFF8FFC3BDF32B1FA217BE6E" box="[758,830,538,564]" captionStart="FIGURE 4" captionStartId="5.[151,255,710,734]" captionTargetBox="[437,1150,197,685]" captionTargetId="figure@5.[437,1150,197,687]" captionTargetPageId="5" captionText="FIGURE 4. Live colouration of male (above) and female (below) Danionella priapus. Specimens not preserved. Note eggs in body cavity of female." httpUri="https://zenodo.org/record/191109/files/figure.png" pageId="4" pageNumber="57">Fig. 4</figureCitation>
), except for melanophore patterns described above, a thin yellowish-greenish line running along body at level of neural tube; golden chromatophores forming ring around pupil, capping swimbladder chambers and connecting duct.
</paragraph>
</subSubSection>
<subSubSection id="C30006CFFFF8FFC3BFC32B8BA4B5BEF2" box="[198,1436,654,680]" pageId="4" pageNumber="57" type="distribution">
<paragraph id="8BA55544FFF8FFC3BFC32B8BA4B5BEF2" blockId="4.[151,1437,152,796]" box="[198,1436,654,680]" pageId="4" pageNumber="57">
<emphasis id="B96E8956FFF8FFC3BFC32B8BA04EBEF2" bold="true" box="[198,359,654,680]" pageId="4" pageNumber="57">Distribution.</emphasis>
The species is known only from the
<typeStatus id="54A1EBE6FFF8FFC3BC112B8BA26EBEF2" box="[788,839,654,680]" pageId="4" pageNumber="57">type</typeStatus>
locality, Jorai River in West Bengal,
<collectingCountry id="F30D15D4FFF8FFC3BBFF2B8BA41FBEF2" box="[1274,1334,654,680]" name="India" pageId="4" pageNumber="57">India</collectingCountry>
(
<figureCitation id="132149C1FFF8FFC3BA432B8BA4A4BEF2" box="[1350,1421,654,680]" captionStart="FIGURE 5" captionStartId="5.[151,255,1775,1799]" captionTargetBox="[424,1163,822,1751]" captionTargetId="figure@5.[424,1163,822,1751]" captionTargetPageId="5" captionText="FIGURE 5. Distribution of the four species of Danionella in India and Myanmar." httpUri="https://zenodo.org/record/191110/files/figure.png" pageId="4" pageNumber="57">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30006CFFFF8FFC3BFC32BB0A465BB66" pageId="4" pageNumber="57" type="etymology">
<paragraph id="8BA55544FFF8FFC3BFC32BB0A2EEBF46" blockId="4.[151,1437,152,796]" pageId="4" pageNumber="57">
<emphasis id="B96E8956FFF8FFC3BFC32BB0A07ABE95" bold="true" box="[198,339,693,719]" pageId="4" pageNumber="57">Etymology.</emphasis>
The species name
<taxonomicName id="4C1A2EC7FFF8FFC3BD3E2BB0A3BCBE94" box="[571,661,693,718]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="priapus">
<emphasis id="B96E8956FFF8FFC3BD3E2BB0A3BCBE94" box="[571,661,693,718]" italics="true" pageId="4" pageNumber="57">priapus</emphasis>
</taxonomicName>
is derived from the greek ρίαπος, the god of fertility of Greek mythology. It was inspired by the conical projection of the genital papilla in males, which—at a superficial level—is reminiscent of the penis of mammals. A noun in apposition.
</paragraph>
<caption id="DF6505CCFFF8FFC3BF922A4AA00ABFD3" pageId="4" pageNumber="57">
<paragraph id="8BA55544FFF8FFC3BF922A4AA00ABFD3" blockId="4.[151,1436,847,1041]" pageId="4" pageNumber="57">
<emphasis id="B96E8956FFF8FFC3BF922A4AA022BF3D" bold="true" box="[151,267,847,871]" pageId="4" pageNumber="57">TABLE 1.</emphasis>
Selected morphometric information on 15 specimens (7 males, 8 females) of
<taxonomicName id="4C1A2EC7FFF8FFC3BB532A55A401BF3D" box="[1110,1320,848,871]" class="Actinopterygii" family="Cyprinidae" genus="Danionella" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="priapus">
<emphasis id="B96E8956FFF8FFC3BB532A55A401BF3D" box="[1110,1320,848,871]" italics="true" pageId="4" pageNumber="57">Danionella priapus</emphasis>
</taxonomicName>
, including the holotype.
</paragraph>
</caption>
<paragraph id="8BA55544FFF8FFC3BD172AA0A2D2BFE6" blockId="4.[151,1436,847,1041]" box="[530,1019,933,956]" pageId="4" pageNumber="57">
<typeStatus id="54A1EBE6FFF8FFC3BD172AA0A35CBFE6" box="[530,629,933,956]" pageId="4" pageNumber="57" type="holotype">Holotype</typeStatus>
<typeStatus id="54A1EBE6FFF8FFC3BC912AA0A2D2BFE6" box="[916,1019,933,956]" pageId="4" pageNumber="57" type="paratype">Paratypes</typeStatus>
</paragraph>
<paragraph id="8BA55544FFF8FFC3BD172AD5A5D5BFBD" blockId="4.[151,1436,847,1041]" box="[530,1276,976,999]" pageId="4" pageNumber="57">Male Males Females</paragraph>
<paragraph id="8BA55544FFF8FFC3BD962AFFA450B84B" blockId="4.[151,1436,847,1041]" box="[659,1401,1018,1041]" pageId="4" pageNumber="57">Range Mean St. Dev Range Mean St. Dev</paragraph>
<paragraph id="8BA55544FFF8FFC3BF9A2D55A551B83D" blockId="4.[159,1144,1104,1127]" box="[159,1144,1104,1127]" pageId="4" pageNumber="57">Standard length (SL) 14.4 12.5–14.1 13.7–15.0</paragraph>
<paragraph id="8BA55544FFF8FFC3BF9A2DA0A03DB8E6" blockId="4.[159,1356,1189,1681]" box="[159,276,1189,1212]" pageId="4" pageNumber="57">In % of SL</paragraph>
<paragraph id="8BA55544FFF8FFC3BF9A2DD5A465BACB" blockId="4.[159,1356,1189,1681]" pageId="4" pageNumber="57">Head length 17.4 17.4–20.0 18.3 0.9 19.0–20.7 19.8 0.7 Predorsal-fin length 70.1 68.7–70.4 69.4 0.7 68.7–71.0 70.1 0.8 Prepelvic-fin length 30.6 29.8–32.8 30.8 1.1 34.0–35.6 34.8 0.5 Preanal-fin length 52.1 51.1–52.1 51.5 0.4 53.3–54.8 54.2 0.6 Snout to anus 29.9 29.1–32.0 30.1 1.0 51.4–52.7 51.9 0.4 Body depth at dorsal-fin origin 13.2 13.1–15.2 14.1 0.7 14.4–16.0 15.4 0.5 Caudal-peduncle depth 7.6 7.5–8.5 8.0 0.3 7.5–8.9 8.0 0.5 Caudal-peduncle length 18.8 17.9–21.6 19.6 1.4 18.1–21.3 19.6 1.1 Length of dorsal-fin base 9.7 9.7–11.3 10.7 0.5 10.3–11.7 10.9 0.5 Length of anal-fin base 30.6 28.4–30.7 29.5 0.9 26.0–28.3 27.4 0.7 Length of pelvic fin 8.7 7.2–8.7 7.4 0.3 6.3–8.3 7.0 0.6</paragraph>
<paragraph id="8BA55544FFF8FFC3BF9A2FD5A030BABD" blockId="4.[159,1356,1744,1852]" box="[159,281,1744,1767]" pageId="4" pageNumber="57">In % of HL</paragraph>
<paragraph id="8BA55544FFF8FFC3BF9A2FFFA465BB66" blockId="4.[159,1356,1744,1852]" pageId="4" pageNumber="57">Eye diameter 44.0 40.0–44.0 40.8 1.5 35.0–39.3 37.3 1.3 Snout length 16.0 16.0–16.7 16.1 0.3 17.9–21.4 20.0 1.2</paragraph>
</subSubSection>
</treatment>
</document>
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