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<mods:title id="27D385F560F2C998B523DBDF2F2E6898">Taxonomy of the clam shrimp Halysestheriidae (Diplostraca: Spinicaudata) from the Upper Cretaceous Nenjiang Formation in central Dayangshu Basin of eastern Inner Mongolia, northeastern China</mods:title>
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<treatment id="03B3BB0AFFA2AA551E9CF8AAFF05FEDE" ID-DOI="http://doi.org/10.5281/zenodo.5559979" ID-GBIF-Taxon="188606833" ID-Zenodo-Dep="5559979" LSID="urn:lsid:plazi:treatment:03B3BB0AFFA2AA551E9CF8AAFF05FEDE" httpUri="http://treatment.plazi.org/id/03B3BB0AFFA2AA551E9CF8AAFF05FEDE" lastPageId="7" lastPageNumber="159" pageId="2" pageNumber="158">
<subSubSection id="C3005997FFA2AA501E9CF8AAFEB7F839" pageId="2" pageNumber="158" type="nomenclature">
<paragraph id="8BA50A1CFFA2AA501E9CF8AAFEB7F839" blockId="2.[136,777,1883,1982]" pageId="2" pageNumber="158">
<heading id="D0EDBD70FFA2AA501E9CF8AAFEB7F839" bold="true" centered="true" fontSize="10" level="2" pageId="2" pageNumber="158" reason="3">
<emphasis id="B96ED60EFFA2AA501E9CF8AAFEB7F839" bold="true" pageId="2" pageNumber="158">
<taxonomicName id="4C1A719FFFA2AA501E9CF8AAFDA9F8D6" authority="Zhang &amp; Chen" authorityName="Zhang &amp; Chen" box="[136,589,1883,1910]" class="Branchiopoda" family="Halysestheriidae" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="2" pageNumber="158" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B96ED60EFFA2AA501E9CF8AAFE6BF8D5" bold="true" box="[136,399,1883,1910]" italics="true" pageId="2" pageNumber="158">Halysestheria biformis</emphasis>
Zhang &amp; Chen
</taxonomicName>
<emphasis id="B96ED60EFFA2AA501C40F8AAFD8FF8D6" bold="true" box="[596,619,1883,1909]" italics="true" pageId="2" pageNumber="158">in</emphasis>
<bibRefCitation id="EF8B77EDFFA2AA501C60F8ADFF11F83A" author="Zhang, W. T. &amp; Chen, P. J. &amp; Shen, Y. B." pageId="2" pageNumber="158" refId="ref5922" refString="Zhang, W. T., Chen, P. J. &amp; Shen, Y. B. (1976) Fossil Conchostraca of China. Science Press, Beijing, 325 pp., 138 pls. [In Chinese]." type="book" year="1976">
Zhang
<emphasis id="B96ED60EFFA2AA501CDEF8ADFD18F8D5" bold="true" box="[714,764,1883,1910]" italics="true" pageId="2" pageNumber="158">et al</emphasis>
., 1976
</bibRefCitation>
emend.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3005997FFA2AA501E9CF852FB28FF7B" pageId="2" pageNumber="158" type="description">
<paragraph id="8BA50A1CFFA2AA501E9CF852FF18F81D" blockId="2.[136,777,1883,1982]" box="[136,252,1955,1982]" pageId="2" pageNumber="158">
(
<figureCitation id="13211699FFA2AA501E84F852FF10F81D" box="[144,244,1955,1982]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="3.[136,229,1498,1520]" captionStartId-1="4.[136,229,1681,1703]" captionStartId-2="5.[136,229,1681,1703]" captionStartId-3="6.[136,229,1689,1711]" captionTargetBox-0="[151,1436,190,1473]" captionTargetBox-1="[248,1340,190,1657]" captionTargetBox-2="[248,1340,190,1657]" captionTargetBox-3="[244,1342,190,1665]" captionTargetId-0="figure-146@3.[151,1436,190,1473]" captionTargetId-1="figure-15@4.[248,1340,190,1657]" captionTargetId-2="figure-20@5.[248,1340,190,1657]" captionTargetId-3="figure-15@6.[244,1342,190,1665]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 2. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend.All are SEM images except A, which is a light micrograph. The specimen was collected from the First Member of the Upper Cretaceous Nenjiang Formation in Jilin, northeastern China. A, An external mould of a right valve, holotype NIGP27106. B, The furrows intercalated nodular platform on a growth band near the umbo. C, A reverse image of B, widely spaced radial lirae with intercalated reticulation and puncta, lirae simple or branched. D, Widely spaced furrow with intercalated nodular platform on growth bands near the umbo. E, Radially aligned, discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould, and serration along the lower margins of growth lines. F, A reverse image of E, showing the dense radial lirae with intercalated chain-like ornamentation on growth bands in the ventral part of the carapace, growth lines with serrated lower margins. Scale bars are 1 mm in A; 20 µm in B, C; 100 µm in DF." captionText-1="FIGURE 3. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are SEM images. The specimen is from the First Member of the Upper Cretaceous Nenjiang Formation at Ganhe, Oroqen Autonomous Banner of Inner Mongolia, northeastern China. A, An external mould of a right valve, NIGP174748. B, A reverse image showing widely spaced radial lirae with intercalated reticulation and puncta on growth bands near the umbo, lirae simple or branched. C, Widely spaced furrow with intercalated nodules on growth bands in the anterodorsal part of the carapace external mould. D, A reverse image of C, showing widely spaced radial lirae with intercalated reticulation and puncta. E, Sparse nodules along the lower part of each growth band in the anteroventral part of the carapace, growth lines are serrated in the lower margins. F, A reverse image of E, showing almost smooth growth bands with pitted ornament along the lower part of each growth band in the anteroventral part of the carapace. G, Radially aligned discontinuous short ridges on a growth band in the ventral part of the carapace external mould, growth lines with serrated lower margins. H, A reverse image of G, showing thickened radial lirae with intercalated chain-like ornament on growth band in the ventral part of the carapace. Scale bars are 1 mm in A; 10 µm in C, D; 20 µm in B, EH." captionText-2="FIGURE 4. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are SEM images of the specimen of Fig. 3A. A, Widely spaced furrows with intercalated nodules on a growth band in the posterodoral part of the carapace external mould. B, A reverse image of A, showing widely spaced radial lirae with intercalated reticulation and puncta on a growth band in the posterodoral part of the carapace. C, Radially aligned nodules on a growth band in the posteroventral part of the carapace external mould. D, A reverse image of C, showing thickened radial lirae with intercalated chain-like ornament on a growth band in the ventral part of the carapace. E, Radially aligned nodules on growth bands in the posteroventral part of the carapace external mould. F, A reverse image of E, showing dense radial lirae with intercalated chain-like ornament on growth bands in the posteroventral part of the carapace. G, Radially aligned discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould. H, A reverse image of G, showing dense and thickened radial lirae with intercalated chain-like ornament on growth bands in the ventral part of the carapace. Growth lines with serrated lower margins. Scale bars are 10 µm in A, B; 20 µm in C, D; 100 µm in EH." captionText-3="FIGURE 5. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are light micrographs of specimens form the First Member of the Upper Cretaceous Nenjiang Formation of Ganhe, Oroqen Autonomous Banner, Inner Mongolia, northeastern China. A, An external mould of a left valve, NIGP174749. B, A left valve, NIGP174750. C, The left one: a left valve, NIGP174751; the right one: an external mould of a right valve, NIGP174752. D, An external mould of a left valve, NIGP174753. E, An external mould of a right valve, NIGP174754. F, External moulds of articulated two valves, NIGP174755. G, Displace valves, NIGP174756. H, Articulated valves, NIGP174757. Scale bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5507754" figureDoi-1="http://doi.org/10.5281/zenodo.5507756" figureDoi-2="http://doi.org/10.5281/zenodo.5507758" figureDoi-3="http://doi.org/10.5281/zenodo.5507762" httpUri-0="https://zenodo.org/record/5507754/files/figure.png" httpUri-1="https://zenodo.org/record/5507756/files/figure.png" httpUri-2="https://zenodo.org/record/5507758/files/figure.png" httpUri-3="https://zenodo.org/record/5507762/files/figure.png" pageId="2" pageNumber="158">Figs 25</figureCitation>
)
</paragraph>
<paragraph id="8BA50A1CFFA2AA501E9CF81EFD97F78B" blockId="2.[136,778,2031,2089]" pageId="2" pageNumber="158">
1976
<taxonomicName id="4C1A719FFFA2AA501EAAF81EFDC9F7A6" authority="Zhang &amp; Chen" authorityName="Zhang &amp; Chen" box="[190,557,2031,2053]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="2" pageNumber="158" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B96ED60EFFA2AA501EAAF81EFE7DF7A7" box="[190,409,2031,2052]" italics="true" pageId="2" pageNumber="158">Halysestheria biformis</emphasis>
Zhang &amp; Chen
</taxonomicName>
<emphasis id="B96ED60EFFA2AA501C26F801FDA1F7A6" box="[562,581,2032,2053]" italics="true" pageId="2" pageNumber="158">in</emphasis>
Zhang
<emphasis id="B96ED60EFFA2AA501C9AF801FD5CF7A7" box="[654,696,2031,2053]" italics="true" pageId="2" pageNumber="158">et al</emphasis>
., p. 211, pl. 91, figs 12; pl. 92, figs 14; pl. 93, fig. 8.
</paragraph>
<paragraph id="8BA50A1CFFA2AA501D3EFF6EFB28FF7B" blockId="2.[810,1451,159,217]" pageId="2" pageNumber="158">
1985
<taxonomicName id="4C1A719FFFA2AA501D4BFF6EFB28FF16" authority="Zhang &amp; Chen" authorityName="Zhang &amp; Chen" box="[863,1228,159,181]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="2" pageNumber="158" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B96ED60EFFA2AA501D4BFF6EFBDDFF17" box="[863,1081,159,180]" italics="true" pageId="2" pageNumber="158">Halysestheria biformis</emphasis>
Zhang &amp; Chen
</taxonomicName>
(
<emphasis id="B96ED60EFFA2AA501ACDFF51FB08FF16" box="[1241,1260,160,181]" italics="true" pageId="2" pageNumber="158">in</emphasis>
<bibRefCitation id="EF8B77EDFFA2AA501AE5FF6EFA44FF16" author="Zhang, W. T. &amp; Chen, P. J. &amp; Shen, Y. B." box="[1265,1440,159,181]" pageId="2" pageNumber="158" refId="ref5922" refString="Zhang, W. T., Chen, P. J. &amp; Shen, Y. B. (1976) Fossil Conchostraca of China. Science Press, Beijing, 325 pp., 138 pls. [In Chinese]." type="book" year="1976">
Zhang
<emphasis id="B96ED60EFFA2AA501B27FF51FAB9FF17" box="[1331,1373,159,181]" italics="true" pageId="2" pageNumber="158">et al</emphasis>
., 1976
</bibRefCitation>
), Chen &amp; Shen, p. 133, pl. 13, figs 12.
</paragraph>
</subSubSection>
<subSubSection id="C3005997FFA2AA501D3EFEF9FC42FD72" pageId="2" pageNumber="158" type="diagnosis">
<paragraph id="8BA50A1CFFA2AA501D3EFEF9FA8FFDE2" blockId="2.[810,1452,263,2090]" pageId="2" pageNumber="158">
<emphasis id="B96ED60EFFA2AA501D3EFEF9FBCFFE82" bold="true" box="[810,1067,263,290]" pageId="2" pageNumber="158">Emended diagnosis.</emphasis>
Carapace moderate in size; elliptical, long oval or subquadrate in outline; umbo located in the anterior part of the dorsal margin. Growth lines are serrated in the lower margins. Growth bands in the umbonal area ornamented with widely spaced radial lirae, intercalated with fine reticulation and puncta; growth bands in the ventral part of the carapace ornamented with chain-like reticulation, which appears as radially aligned, discontinuous, short, fine ridges on external mould.
</paragraph>
<paragraph id="8BA50A1CFFA2AA501D4BFDBAFC42FD72" blockId="2.[810,1452,263,2090]" pageId="2" pageNumber="158">
<emphasis id="B96ED60EFFA2AA501D4BFDBAFB8DFDC6" bold="true" box="[863,1129,587,614]" pageId="2" pageNumber="158">Locality and horizon.</emphasis>
<collectingRegion id="49DEC4FEFFA2AA501A60FDBAFAF3FDC6" box="[1140,1303,587,613]" country="China" name="Jilin" pageId="2" pageNumber="158">Jilin Province</collectingRegion>
, and Ganhe, OroqenAutonomous Banner,
<materialsCitation id="3B720041FFA2AA501A78FD81FC46FD72" ID-GBIF-Occurrence="3382545301" country="China" location="Upper Cretaceous Nenjiang Formation" pageId="2" pageNumber="158" specimenCount="1" stateProvince="Nei Mongol">
<collectingRegion id="49DEC4FEFFA2AA501A78FD81FAFEFD2A" box="[1132,1306,623,650]" country="China" name="Nei Mongol" pageId="2" pageNumber="158">Inner Mongolia</collectingRegion>
,northeastern
<collectingCountry id="F30D4A8CFFA2AA501D3EFD62FC94FD0E" box="[810,880,659,685]" name="China" pageId="2" pageNumber="158">China</collectingCountry>
; First Member of the
<location id="8EC55CC7FFA2AA501A7BFD65FC46FD72" LSID="urn:lsid:plazi:treatment:03B3BB0AFFA2AA551E9CF8AAFF05FEDE:8EC55CC7FFA2AA501A7BFD65FC46FD72" country="China" name="Upper Cretaceous Nenjiang Formation" pageId="2" pageNumber="158" stateProvince="Nei Mongol">Upper Cretaceous Nenjiang Formation</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3005997FFA2AA501D4BFD2AFB76F946" pageId="2" pageNumber="158" type="description">
<paragraph id="8BA50A1CFFA2AA501D4BFD2AFB29FA01" blockId="2.[810,1452,263,2090]" pageId="2" pageNumber="158">
<emphasis id="B96ED60EFFA2AA501D4BFD2AFC15FD56" bold="true" box="[863,1009,731,757]" pageId="2" pageNumber="158">Description.</emphasis>
Carapace moderate in size, elliptical, long oval, or subquadrate in outline. Anterior margin widely rounded, posterior margin pointed. Umbo located in the anterior part of the dorsal margin. Growth lines prominent, 2030 or more in number, serrated along their lower margins (
<figureCitation id="13211699FFA2AA501DF5FC7EFBA4FC09" box="[993,1088,911,938]" captionStart="FIGURE 2" captionStartId="3.[136,229,1498,1520]" captionTargetBox="[151,1436,190,1473]" captionTargetId="figure-146@3.[151,1436,190,1473]" captionTargetPageId="3" captionText="FIGURE 2. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend.All are SEM images except A, which is a light micrograph. The specimen was collected from the First Member of the Upper Cretaceous Nenjiang Formation in Jilin, northeastern China. A, An external mould of a right valve, holotype NIGP27106. B, The furrows intercalated nodular platform on a growth band near the umbo. C, A reverse image of B, widely spaced radial lirae with intercalated reticulation and puncta, lirae simple or branched. D, Widely spaced furrow with intercalated nodular platform on growth bands near the umbo. E, Radially aligned, discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould, and serration along the lower margins of growth lines. F, A reverse image of E, showing the dense radial lirae with intercalated chain-like ornamentation on growth bands in the ventral part of the carapace, growth lines with serrated lower margins. Scale bars are 1 mm in A; 20 µm in B, C; 100 µm in DF." figureDoi="http://doi.org/10.5281/zenodo.5507754" httpUri="https://zenodo.org/record/5507754/files/figure.png" pageId="2" pageNumber="158">Figs 2E</figureCitation>
,
<figureCitation id="13211699FFA2AA501A5BFC61FB58FC09" box="[1103,1212,912,938]" captionStart="FIGURE 3" captionStartId="4.[136,229,1681,1703]" captionTargetBox="[248,1340,190,1657]" captionTargetId="figure-15@4.[248,1340,190,1657]" captionTargetPageId="4" captionText="FIGURE 3. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are SEM images. The specimen is from the First Member of the Upper Cretaceous Nenjiang Formation at Ganhe, Oroqen Autonomous Banner of Inner Mongolia, northeastern China. A, An external mould of a right valve, NIGP174748. B, A reverse image showing widely spaced radial lirae with intercalated reticulation and puncta on growth bands near the umbo, lirae simple or branched. C, Widely spaced furrow with intercalated nodules on growth bands in the anterodorsal part of the carapace external mould. D, A reverse image of C, showing widely spaced radial lirae with intercalated reticulation and puncta. E, Sparse nodules along the lower part of each growth band in the anteroventral part of the carapace, growth lines are serrated in the lower margins. F, A reverse image of E, showing almost smooth growth bands with pitted ornament along the lower part of each growth band in the anteroventral part of the carapace. G, Radially aligned discontinuous short ridges on a growth band in the ventral part of the carapace external mould, growth lines with serrated lower margins. H, A reverse image of G, showing thickened radial lirae with intercalated chain-like ornament on growth band in the ventral part of the carapace. Scale bars are 1 mm in A; 10 µm in C, D; 20 µm in B, EH." figureDoi="http://doi.org/10.5281/zenodo.5507756" httpUri="https://zenodo.org/record/5507756/files/figure.png" pageId="2" pageNumber="158">3B, E, G</figureCitation>
,
<figureCitation id="13211699FFA2AA501AD8FC61FB16FC09" box="[1228,1266,912,938]" captionStart="FIGURE 4" captionStartId="5.[136,229,1681,1703]" captionTargetBox="[248,1340,190,1657]" captionTargetId="figure-20@5.[248,1340,190,1657]" captionTargetPageId="5" captionText="FIGURE 4. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are SEM images of the specimen of Fig. 3A. A, Widely spaced furrows with intercalated nodules on a growth band in the posterodoral part of the carapace external mould. B, A reverse image of A, showing widely spaced radial lirae with intercalated reticulation and puncta on a growth band in the posterodoral part of the carapace. C, Radially aligned nodules on a growth band in the posteroventral part of the carapace external mould. D, A reverse image of C, showing thickened radial lirae with intercalated chain-like ornament on a growth band in the ventral part of the carapace. E, Radially aligned nodules on growth bands in the posteroventral part of the carapace external mould. F, A reverse image of E, showing dense radial lirae with intercalated chain-like ornament on growth bands in the posteroventral part of the carapace. G, Radially aligned discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould. H, A reverse image of G, showing dense and thickened radial lirae with intercalated chain-like ornament on growth bands in the ventral part of the carapace. Growth lines with serrated lower margins. Scale bars are 10 µm in A, B; 20 µm in C, D; 100 µm in EH." figureDoi="http://doi.org/10.5281/zenodo.5507758" httpUri="https://zenodo.org/record/5507758/files/figure.png" pageId="2" pageNumber="158">4A</figureCitation>
). Growth bands narrow in the umbonal area, become wider in the middle part of the carapace (
<figureCitation id="13211699FFA2AA501A00FC26FB95FC51" box="[1044,1137,983,1010]" captionStart="FIGURE 2" captionStartId="3.[136,229,1498,1520]" captionTargetBox="[151,1436,190,1473]" captionTargetId="figure-146@3.[151,1436,190,1473]" captionTargetPageId="3" captionText="FIGURE 2. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend.All are SEM images except A, which is a light micrograph. The specimen was collected from the First Member of the Upper Cretaceous Nenjiang Formation in Jilin, northeastern China. A, An external mould of a right valve, holotype NIGP27106. B, The furrows intercalated nodular platform on a growth band near the umbo. C, A reverse image of B, widely spaced radial lirae with intercalated reticulation and puncta, lirae simple or branched. D, Widely spaced furrow with intercalated nodular platform on growth bands near the umbo. E, Radially aligned, discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould, and serration along the lower margins of growth lines. F, A reverse image of E, showing the dense radial lirae with intercalated chain-like ornamentation on growth bands in the ventral part of the carapace, growth lines with serrated lower margins. Scale bars are 1 mm in A; 20 µm in B, C; 100 µm in DF." figureDoi="http://doi.org/10.5281/zenodo.5507754" httpUri="https://zenodo.org/record/5507754/files/figure.png" pageId="2" pageNumber="158">Figs 2A</figureCitation>
,
<figureCitation id="13211699FFA2AA501A69FC26FB26FC51" box="[1149,1218,983,1010]" captionStart="FIGURE 5" captionStartId="6.[136,229,1689,1711]" captionTargetBox="[244,1342,190,1665]" captionTargetId="figure-15@6.[244,1342,190,1665]" captionTargetPageId="6" captionText="FIGURE 5. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are light micrographs of specimens form the First Member of the Upper Cretaceous Nenjiang Formation of Ganhe, Oroqen Autonomous Banner, Inner Mongolia, northeastern China. A, An external mould of a left valve, NIGP174749. B, A left valve, NIGP174750. C, The left one: a left valve, NIGP174751; the right one: an external mould of a right valve, NIGP174752. D, An external mould of a left valve, NIGP174753. E, An external mould of a right valve, NIGP174754. F, External moulds of articulated two valves, NIGP174755. G, Displace valves, NIGP174756. H, Articulated valves, NIGP174757. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5507762" httpUri="https://zenodo.org/record/5507762/files/figure.png" pageId="2" pageNumber="158">5EH</figureCitation>
); those in the ventral part of the carapace become narrower again. Growth bands near the umbo ornamented with widely spaced long radial lirae, which are simple or branched, intercalated with fine reticulation, puncta occur within the fine mesh (
<figureCitation id="13211699FFA2AA501D26FB7AFC75FB05" box="[818,913,1163,1190]" captionStart="FIGURE 2" captionStartId="3.[136,229,1498,1520]" captionTargetBox="[151,1436,190,1473]" captionTargetId="figure-146@3.[151,1436,190,1473]" captionTargetPageId="3" captionText="FIGURE 2. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend.All are SEM images except A, which is a light micrograph. The specimen was collected from the First Member of the Upper Cretaceous Nenjiang Formation in Jilin, northeastern China. A, An external mould of a right valve, holotype NIGP27106. B, The furrows intercalated nodular platform on a growth band near the umbo. C, A reverse image of B, widely spaced radial lirae with intercalated reticulation and puncta, lirae simple or branched. D, Widely spaced furrow with intercalated nodular platform on growth bands near the umbo. E, Radially aligned, discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould, and serration along the lower margins of growth lines. F, A reverse image of E, showing the dense radial lirae with intercalated chain-like ornamentation on growth bands in the ventral part of the carapace, growth lines with serrated lower margins. Scale bars are 1 mm in A; 20 µm in B, C; 100 µm in DF." figureDoi="http://doi.org/10.5281/zenodo.5507754" httpUri="https://zenodo.org/record/5507754/files/figure.png" pageId="2" pageNumber="158">Figs 2B</figureCitation>
,
<figureCitation id="13211699FFA2AA501DB5FB7DFC21FB05" box="[929,965,1164,1190]" captionStart="FIGURE 3" captionStartId="4.[136,229,1681,1703]" captionTargetBox="[248,1340,190,1657]" captionTargetId="figure-15@4.[248,1340,190,1657]" captionTargetPageId="4" captionText="FIGURE 3. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are SEM images. The specimen is from the First Member of the Upper Cretaceous Nenjiang Formation at Ganhe, Oroqen Autonomous Banner of Inner Mongolia, northeastern China. A, An external mould of a right valve, NIGP174748. B, A reverse image showing widely spaced radial lirae with intercalated reticulation and puncta on growth bands near the umbo, lirae simple or branched. C, Widely spaced furrow with intercalated nodules on growth bands in the anterodorsal part of the carapace external mould. D, A reverse image of C, showing widely spaced radial lirae with intercalated reticulation and puncta. E, Sparse nodules along the lower part of each growth band in the anteroventral part of the carapace, growth lines are serrated in the lower margins. F, A reverse image of E, showing almost smooth growth bands with pitted ornament along the lower part of each growth band in the anteroventral part of the carapace. G, Radially aligned discontinuous short ridges on a growth band in the ventral part of the carapace external mould, growth lines with serrated lower margins. H, A reverse image of G, showing thickened radial lirae with intercalated chain-like ornament on growth band in the ventral part of the carapace. Scale bars are 1 mm in A; 10 µm in C, D; 20 µm in B, EH." figureDoi="http://doi.org/10.5281/zenodo.5507756" httpUri="https://zenodo.org/record/5507756/files/figure.png" pageId="2" pageNumber="158">3C</figureCitation>
). Growth bands in the ventral part of the carapace, radial lirae are thickened, fine mesh between the lirae are radially aligned to form chain-like ornamentation (
<figureCitation id="13211699FFA2AA501D26FB06FC63FAB1" box="[818,903,1271,1298]" captionStart="FIGURE 2" captionStartId="3.[136,229,1498,1520]" captionTargetBox="[151,1436,190,1473]" captionTargetId="figure-146@3.[151,1436,190,1473]" captionTargetPageId="3" captionText="FIGURE 2. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend.All are SEM images except A, which is a light micrograph. The specimen was collected from the First Member of the Upper Cretaceous Nenjiang Formation in Jilin, northeastern China. A, An external mould of a right valve, holotype NIGP27106. B, The furrows intercalated nodular platform on a growth band near the umbo. C, A reverse image of B, widely spaced radial lirae with intercalated reticulation and puncta, lirae simple or branched. D, Widely spaced furrow with intercalated nodular platform on growth bands near the umbo. E, Radially aligned, discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould, and serration along the lower margins of growth lines. F, A reverse image of E, showing the dense radial lirae with intercalated chain-like ornamentation on growth bands in the ventral part of the carapace, growth lines with serrated lower margins. Scale bars are 1 mm in A; 20 µm in B, C; 100 µm in DF." figureDoi="http://doi.org/10.5281/zenodo.5507754" httpUri="https://zenodo.org/record/5507754/files/figure.png" pageId="2" pageNumber="158">Figs 2F</figureCitation>
,
<figureCitation id="13211699FFA2AA501D85FB09FC51FAB1" box="[913,949,1272,1298]" captionStart="FIGURE 3" captionStartId="4.[136,229,1681,1703]" captionTargetBox="[248,1340,190,1657]" captionTargetId="figure-15@4.[248,1340,190,1657]" captionTargetPageId="4" captionText="FIGURE 3. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are SEM images. The specimen is from the First Member of the Upper Cretaceous Nenjiang Formation at Ganhe, Oroqen Autonomous Banner of Inner Mongolia, northeastern China. A, An external mould of a right valve, NIGP174748. B, A reverse image showing widely spaced radial lirae with intercalated reticulation and puncta on growth bands near the umbo, lirae simple or branched. C, Widely spaced furrow with intercalated nodules on growth bands in the anterodorsal part of the carapace external mould. D, A reverse image of C, showing widely spaced radial lirae with intercalated reticulation and puncta. E, Sparse nodules along the lower part of each growth band in the anteroventral part of the carapace, growth lines are serrated in the lower margins. F, A reverse image of E, showing almost smooth growth bands with pitted ornament along the lower part of each growth band in the anteroventral part of the carapace. G, Radially aligned discontinuous short ridges on a growth band in the ventral part of the carapace external mould, growth lines with serrated lower margins. H, A reverse image of G, showing thickened radial lirae with intercalated chain-like ornament on growth band in the ventral part of the carapace. Scale bars are 1 mm in A; 10 µm in C, D; 20 µm in B, EH." figureDoi="http://doi.org/10.5281/zenodo.5507756" httpUri="https://zenodo.org/record/5507756/files/figure.png" pageId="2" pageNumber="158">3H</figureCitation>
,
<figureCitation id="13211699FFA2AA501DD4FB09FBC4FAB1" box="[960,1056,1272,1298]" captionStart="FIGURE 4" captionStartId="5.[136,229,1681,1703]" captionTargetBox="[248,1340,190,1657]" captionTargetId="figure-20@5.[248,1340,190,1657]" captionTargetPageId="5" captionText="FIGURE 4. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are SEM images of the specimen of Fig. 3A. A, Widely spaced furrows with intercalated nodules on a growth band in the posterodoral part of the carapace external mould. B, A reverse image of A, showing widely spaced radial lirae with intercalated reticulation and puncta on a growth band in the posterodoral part of the carapace. C, Radially aligned nodules on a growth band in the posteroventral part of the carapace external mould. D, A reverse image of C, showing thickened radial lirae with intercalated chain-like ornament on a growth band in the ventral part of the carapace. E, Radially aligned nodules on growth bands in the posteroventral part of the carapace external mould. F, A reverse image of E, showing dense radial lirae with intercalated chain-like ornament on growth bands in the posteroventral part of the carapace. G, Radially aligned discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould. H, A reverse image of G, showing dense and thickened radial lirae with intercalated chain-like ornament on growth bands in the ventral part of the carapace. Growth lines with serrated lower margins. Scale bars are 10 µm in A, B; 20 µm in C, D; 100 µm in EH." figureDoi="http://doi.org/10.5281/zenodo.5507758" httpUri="https://zenodo.org/record/5507758/files/figure.png" pageId="2" pageNumber="158">4D, F, H</figureCitation>
), which appears as radially aligned, discontinuous, short, fine ridges on external mould (
<figureCitation id="13211699FFA2AA501B6EFAEAFCAFFAF9" captionStart="FIGURE 2" captionStartId="3.[136,229,1498,1520]" captionTargetBox="[151,1436,190,1473]" captionTargetId="figure-146@3.[151,1436,190,1473]" captionTargetPageId="3" captionText="FIGURE 2. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend.All are SEM images except A, which is a light micrograph. The specimen was collected from the First Member of the Upper Cretaceous Nenjiang Formation in Jilin, northeastern China. A, An external mould of a right valve, holotype NIGP27106. B, The furrows intercalated nodular platform on a growth band near the umbo. C, A reverse image of B, widely spaced radial lirae with intercalated reticulation and puncta, lirae simple or branched. D, Widely spaced furrow with intercalated nodular platform on growth bands near the umbo. E, Radially aligned, discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould, and serration along the lower margins of growth lines. F, A reverse image of E, showing the dense radial lirae with intercalated chain-like ornamentation on growth bands in the ventral part of the carapace, growth lines with serrated lower margins. Scale bars are 1 mm in A; 20 µm in B, C; 100 µm in DF." figureDoi="http://doi.org/10.5281/zenodo.5507754" httpUri="https://zenodo.org/record/5507754/files/figure.png" pageId="2" pageNumber="158">Figs 2E</figureCitation>
,
<figureCitation id="13211699FFA2AA501D4FFAB1FC9BFAF9" box="[859,895,1344,1370]" captionStart="FIGURE 3" captionStartId="4.[136,229,1681,1703]" captionTargetBox="[248,1340,190,1657]" captionTargetId="figure-15@4.[248,1340,190,1657]" captionTargetPageId="4" captionText="FIGURE 3. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are SEM images. The specimen is from the First Member of the Upper Cretaceous Nenjiang Formation at Ganhe, Oroqen Autonomous Banner of Inner Mongolia, northeastern China. A, An external mould of a right valve, NIGP174748. B, A reverse image showing widely spaced radial lirae with intercalated reticulation and puncta on growth bands near the umbo, lirae simple or branched. C, Widely spaced furrow with intercalated nodules on growth bands in the anterodorsal part of the carapace external mould. D, A reverse image of C, showing widely spaced radial lirae with intercalated reticulation and puncta. E, Sparse nodules along the lower part of each growth band in the anteroventral part of the carapace, growth lines are serrated in the lower margins. F, A reverse image of E, showing almost smooth growth bands with pitted ornament along the lower part of each growth band in the anteroventral part of the carapace. G, Radially aligned discontinuous short ridges on a growth band in the ventral part of the carapace external mould, growth lines with serrated lower margins. H, A reverse image of G, showing thickened radial lirae with intercalated chain-like ornament on growth band in the ventral part of the carapace. Scale bars are 1 mm in A; 10 µm in C, D; 20 µm in B, EH." figureDoi="http://doi.org/10.5281/zenodo.5507756" httpUri="https://zenodo.org/record/5507756/files/figure.png" pageId="2" pageNumber="158">3G</figureCitation>
,
<figureCitation id="13211699FFA2AA501D9BFAB1FC18FAF9" box="[911,1020,1344,1370]" captionStart="FIGURE 4" captionStartId="5.[136,229,1681,1703]" captionTargetBox="[248,1340,190,1657]" captionTargetId="figure-20@5.[248,1340,190,1657]" captionTargetPageId="5" captionText="FIGURE 4. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are SEM images of the specimen of Fig. 3A. A, Widely spaced furrows with intercalated nodules on a growth band in the posterodoral part of the carapace external mould. B, A reverse image of A, showing widely spaced radial lirae with intercalated reticulation and puncta on a growth band in the posterodoral part of the carapace. C, Radially aligned nodules on a growth band in the posteroventral part of the carapace external mould. D, A reverse image of C, showing thickened radial lirae with intercalated chain-like ornament on a growth band in the ventral part of the carapace. E, Radially aligned nodules on growth bands in the posteroventral part of the carapace external mould. F, A reverse image of E, showing dense radial lirae with intercalated chain-like ornament on growth bands in the posteroventral part of the carapace. G, Radially aligned discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould. H, A reverse image of G, showing dense and thickened radial lirae with intercalated chain-like ornament on growth bands in the ventral part of the carapace. Growth lines with serrated lower margins. Scale bars are 10 µm in A, B; 20 µm in C, D; 100 µm in EH." figureDoi="http://doi.org/10.5281/zenodo.5507758" httpUri="https://zenodo.org/record/5507758/files/figure.png" pageId="2" pageNumber="158">4C, E, G</figureCitation>
). Growth bands in the antero-ventral part of the carapace are almost smooth with some pits in the lower part of each band (
<figureCitation id="13211699FFA2AA501A7FFA76FB24FA01" box="[1131,1216,1415,1442]" captionStart="FIGURE 3" captionStartId="4.[136,229,1681,1703]" captionTargetBox="[248,1340,190,1657]" captionTargetId="figure-15@4.[248,1340,190,1657]" captionTargetPageId="4" captionText="FIGURE 3. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend. All are SEM images. The specimen is from the First Member of the Upper Cretaceous Nenjiang Formation at Ganhe, Oroqen Autonomous Banner of Inner Mongolia, northeastern China. A, An external mould of a right valve, NIGP174748. B, A reverse image showing widely spaced radial lirae with intercalated reticulation and puncta on growth bands near the umbo, lirae simple or branched. C, Widely spaced furrow with intercalated nodules on growth bands in the anterodorsal part of the carapace external mould. D, A reverse image of C, showing widely spaced radial lirae with intercalated reticulation and puncta. E, Sparse nodules along the lower part of each growth band in the anteroventral part of the carapace, growth lines are serrated in the lower margins. F, A reverse image of E, showing almost smooth growth bands with pitted ornament along the lower part of each growth band in the anteroventral part of the carapace. G, Radially aligned discontinuous short ridges on a growth band in the ventral part of the carapace external mould, growth lines with serrated lower margins. H, A reverse image of G, showing thickened radial lirae with intercalated chain-like ornament on growth band in the ventral part of the carapace. Scale bars are 1 mm in A; 10 µm in C, D; 20 µm in B, EH." figureDoi="http://doi.org/10.5281/zenodo.5507756" httpUri="https://zenodo.org/record/5507756/files/figure.png" pageId="2" pageNumber="158">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="8BA50A1CFFA2AA501D4BFA5AFB76F946" blockId="2.[810,1452,263,2090]" pageId="2" pageNumber="158">
<emphasis id="B96ED60EFFA2AA501D4BFA5AFC15FA66" bold="true" box="[863,1009,1451,1477]" pageId="2" pageNumber="158">Dimensions.</emphasis>
In order: specimen no., number of growth bands, length (mm), height (mm), height/length ratio: NIGP27106, 26, 6.8, 4.6, 0.67; NIGP174748, 31,8.4, 5.9, 0.71; NIGP174749, 33, 7.7, 5.3, 0.7; NIGP174750, 25, 8.5, 5.9, 0.69; NIGP 174751, 21, 7.8, 5.2, 0.67; NIGP 174752,&gt;14, 9.2, 6.8, 0.74; NIGP174753, 34, 7.5, 4.7, 0.63; NIGP174754,&gt;21, 7.3, 4.6, 0.64; NIGP174755, 26, 7.0, 4.3, 0.61; NIGP174756, 25, 7.6, 5.1, 0.66; NIGP174757, 22, 7.7, 4.6, 0.60.
</paragraph>
</subSubSection>
<subSubSection id="C3005997FFA2AA551D4BF901FF05FEDE" lastPageId="7" lastPageNumber="163" pageId="2" pageNumber="158" type="discussion">
<paragraph id="8BA50A1CFFA2AA511D4BF901FEDDF8C0" blockId="2.[810,1452,263,2090]" lastBlockId="3.[136,778,1829,2071]" lastPageId="3" lastPageNumber="159" pageId="2" pageNumber="158">
<emphasis id="B96ED60EFFA2AA501D4BF901FC37F8A9" bold="true" box="[863,979,1776,1802]" pageId="2" pageNumber="158">Remarks.</emphasis>
In order to make a comparison, the
<typeStatus id="54A1B4BEFFA2AA501D3EF8E2FC68F88E" box="[810,908,1811,1837]" pageId="2" pageNumber="158" type="holotype">holotype</typeStatus>
specimen of the type species (an external mould of a right valve) was also examined under the SEM (
<figureCitation id="13211699FFA2AA501B6AF8C6FC88F8D5" captionStart="FIGURE 2" captionStartId="3.[136,229,1498,1520]" captionTargetBox="[151,1436,190,1473]" captionTargetId="figure-146@3.[151,1436,190,1473]" captionTargetPageId="3" captionText="FIGURE 2. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend.All are SEM images except A, which is a light micrograph. The specimen was collected from the First Member of the Upper Cretaceous Nenjiang Formation in Jilin, northeastern China. A, An external mould of a right valve, holotype NIGP27106. B, The furrows intercalated nodular platform on a growth band near the umbo. C, A reverse image of B, widely spaced radial lirae with intercalated reticulation and puncta, lirae simple or branched. D, Widely spaced furrow with intercalated nodular platform on growth bands near the umbo. E, Radially aligned, discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould, and serration along the lower margins of growth lines. F, A reverse image of E, showing the dense radial lirae with intercalated chain-like ornamentation on growth bands in the ventral part of the carapace, growth lines with serrated lower margins. Scale bars are 1 mm in A; 20 µm in B, C; 100 µm in DF." figureDoi="http://doi.org/10.5281/zenodo.5507754" httpUri="https://zenodo.org/record/5507754/files/figure.png" pageId="2" pageNumber="158">Fig. 2BF</figureCitation>
). Although the specimen was varnished in order to protect it, clear micrographs have been taken from growth bands near the umbo (
<figureCitation id="13211699FFA2AA501A32F852FB44F81D" box="[1062,1184,1955,1982]" captionStart="FIGURE 2" captionStartId="3.[136,229,1498,1520]" captionTargetBox="[151,1436,190,1473]" captionTargetId="figure-146@3.[151,1436,190,1473]" captionTargetPageId="3" captionText="FIGURE 2. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend.All are SEM images except A, which is a light micrograph. The specimen was collected from the First Member of the Upper Cretaceous Nenjiang Formation in Jilin, northeastern China. A, An external mould of a right valve, holotype NIGP27106. B, The furrows intercalated nodular platform on a growth band near the umbo. C, A reverse image of B, widely spaced radial lirae with intercalated reticulation and puncta, lirae simple or branched. D, Widely spaced furrow with intercalated nodular platform on growth bands near the umbo. E, Radially aligned, discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould, and serration along the lower margins of growth lines. F, A reverse image of E, showing the dense radial lirae with intercalated chain-like ornamentation on growth bands in the ventral part of the carapace, growth lines with serrated lower margins. Scale bars are 1 mm in A; 20 µm in B, C; 100 µm in DF." figureDoi="http://doi.org/10.5281/zenodo.5507754" httpUri="https://zenodo.org/record/5507754/files/figure.png" pageId="2" pageNumber="158">Fig. 2B, D</figureCitation>
) and in the ventral part of the carapace external mould (
<figureCitation id="13211699FFA2AA501AB1F836FAE5F841" box="[1189,1281,1991,2018]" captionStart="FIGURE 2" captionStartId="3.[136,229,1498,1520]" captionTargetBox="[151,1436,190,1473]" captionTargetId="figure-146@3.[151,1436,190,1473]" captionTargetPageId="3" captionText="FIGURE 2. Halysestheria biformis Zhang &amp; Chen in Zhang et al., 1976 emend.All are SEM images except A, which is a light micrograph. The specimen was collected from the First Member of the Upper Cretaceous Nenjiang Formation in Jilin, northeastern China. A, An external mould of a right valve, holotype NIGP27106. B, The furrows intercalated nodular platform on a growth band near the umbo. C, A reverse image of B, widely spaced radial lirae with intercalated reticulation and puncta, lirae simple or branched. D, Widely spaced furrow with intercalated nodular platform on growth bands near the umbo. E, Radially aligned, discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould, and serration along the lower margins of growth lines. F, A reverse image of E, showing the dense radial lirae with intercalated chain-like ornamentation on growth bands in the ventral part of the carapace, growth lines with serrated lower margins. Scale bars are 1 mm in A; 20 µm in B, C; 100 µm in DF." figureDoi="http://doi.org/10.5281/zenodo.5507754" httpUri="https://zenodo.org/record/5507754/files/figure.png" pageId="2" pageNumber="158">Fig. 2E</figureCitation>
). Clear widely spaced radial lirae with intercalated reticulation and puncta in the dorsal part of the carapace, and a chain-like ornamentation occurs on growth bands in the ventral part of the carapace.
</paragraph>
<caption id="DF655A94FFA3AA511E9CFA2BFC36F94F" ID-DOI="http://doi.org/10.5281/zenodo.5507754" ID-Zenodo-Dep="5507754" httpUri="https://zenodo.org/record/5507754/files/figure.png" pageId="3" pageNumber="159" startId="3.[136,229,1498,1520]" targetBox="[151,1436,190,1473]" targetPageId="3">
<paragraph id="8BA50A1CFFA3AA511E9CFA2BFC36F94F" blockId="3.[136,1453,1498,1772]" pageId="3" pageNumber="159">
<emphasis id="B96ED60EFFA3AA511E9CFA2BFF18FA4C" bold="true" box="[136,252,1498,1520]" pageId="3" pageNumber="159">FIGURE 2.</emphasis>
<taxonomicName id="4C1A719FFFA3AA511F15FA2BFD8FFA53" authority="Zhang &amp; Chen" authorityName="Zhang &amp; Chen" box="[257,619,1498,1520]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B96ED60EFFA3AA511F15FA2BFE3FFA4C" box="[257,475,1498,1519]" italics="true" pageId="3" pageNumber="159">Halysestheria biformis</emphasis>
Zhang &amp; Chen
</taxonomicName>
<emphasis id="B96ED60EFFA3AA511C64FA2AFD67FA53" box="[624,643,1499,1520]" italics="true" pageId="3" pageNumber="159">in</emphasis>
<bibRefCitation id="EF8B77EDFFA3AA511C93FA2BFCD6FA53" author="Zhang, W. T. &amp; Chen, P. J. &amp; Shen, Y. B." box="[647,818,1498,1520]" pageId="3" pageNumber="159" refId="ref5922" refString="Zhang, W. T., Chen, P. J. &amp; Shen, Y. B. (1976) Fossil Conchostraca of China. Science Press, Beijing, 325 pp., 138 pls. [In Chinese]." type="book" year="1976">
Zhang
<emphasis id="B96ED60EFFA3AA511CDDFA2AFD16FA4C" box="[713,754,1498,1520]" italics="true" pageId="3" pageNumber="159">et al</emphasis>
., 1976
</bibRefCitation>
emend.All are SEM images except
<emphasis id="B96ED60EFFA3AA511A93FA2BFB7CFA4C" bold="true" box="[1159,1176,1498,1519]" pageId="3" pageNumber="159">A</emphasis>
, which is a light micrograph. The specimen was collected from the First Member of the Upper Cretaceous Nenjiang Formation in Jilin, northeastern China.
<emphasis id="B96ED60EFFA3AA511B64FA0FFA65F9B0" bold="true" box="[1392,1409,1534,1555]" pageId="3" pageNumber="159">A</emphasis>
, An external mould of a right valve, holotype NIGP27106.
<emphasis id="B96ED60EFFA3AA511C85F9D3FD45F994" bold="true" box="[657,673,1570,1591]" pageId="3" pageNumber="159">B</emphasis>
, The furrows intercalated nodular platform on a growth band near the umbo.
<emphasis id="B96ED60EFFA3AA511B94F9D3FA75F99B" bold="true" box="[1408,1425,1570,1592]" pageId="3" pageNumber="159">C</emphasis>
, A reverse image of
<emphasis id="B96ED60EFFA3AA511F38F9B7FED8F9F8" bold="true" box="[300,316,1606,1627]" pageId="3" pageNumber="159">B</emphasis>
, widely spaced radial lirae with intercalated reticulation and puncta, lirae simple or branched.
<emphasis id="B96ED60EFFA3AA511AA8F9B7FB29F9F8" bold="true" box="[1212,1229,1606,1627]" pageId="3" pageNumber="159">D</emphasis>
, Widely spaced furrow with intercalated nodular platform on growth bands near the umbo.
<emphasis id="B96ED60EFFA3AA511D00F99BFCC0F9DC" bold="true" box="[788,804,1642,1663]" pageId="3" pageNumber="159">E</emphasis>
, Radially aligned, discontinuous short ridges or nodules on growth bands in the ventral part of the carapace external mould, and serration along the lower margins of growth lines.
<emphasis id="B96ED60EFFA3AA511AAAF97FFB29F900" bold="true" box="[1214,1229,1678,1699]" pageId="3" pageNumber="159">F</emphasis>
, A reverse image of
<emphasis id="B96ED60EFFA3AA511B81F97FFA41F900" bold="true" box="[1429,1445,1678,1699]" pageId="3" pageNumber="159">E</emphasis>
, showing the dense radial lirae with intercalated chain-like ornamentation on growth bands in the ventral part of the carapace, growth lines with serrated lower margins. Scale bars are 1 mm in
<emphasis id="B96ED60EFFA3AA511C95F927FD76F948" bold="true" box="[641,658,1750,1771]" pageId="3" pageNumber="159">A</emphasis>
; 20 µm in
<emphasis id="B96ED60EFFA3AA511CE9F927FCE9F948" bold="true" box="[765,781,1750,1771]" pageId="3" pageNumber="159">B</emphasis>
,
<emphasis id="B96ED60EFFA3AA511D0DF927FCCEF94F" bold="true" box="[793,810,1750,1772]" pageId="3" pageNumber="159">C</emphasis>
; 100 µm in
<emphasis id="B96ED60EFFA3AA511DB4F927FC28F948" bold="true" box="[928,972,1750,1772]" pageId="3" pageNumber="159">DF</emphasis>
.
</paragraph>
</caption>
<paragraph id="8BA50A1CFFA3AA511EA9F89CFDE4F7B4" blockId="3.[136,778,1829,2071]" pageId="3" pageNumber="159">
The specimens from
<collectingRegion id="49DEC4FEFFA3AA511FB8F89CFDB8F824" box="[428,604,1901,1927]" country="China" name="Nei Mongol" pageId="3" pageNumber="159">Inner Mongolia</collectingRegion>
have elliptical, long oval or subquadrate outline, the height/length ratio between 0.61 and 0.74. In consideration of the sex dimorphism, the long oval form may be male, and subquadrate form may be female.
</paragraph>
<paragraph id="8BA50A1CFFA3AA551D4BF8D4FF05FEDE" blockId="3.[810,1452,1829,2071]" lastBlockId="7.[136,778,173,381]" lastPageId="7" lastPageNumber="163" pageId="3" pageNumber="159">
Except for the
<typeStatus id="54A1B4BEFFA3AA511DEAF8D4FBCAF89C" box="[1022,1070,1829,1855]" pageId="3" pageNumber="159">type</typeStatus>
species, several other species have also been described from the First Member of the Nenjiang Formation, such as
<taxonomicName id="4C1A719FFFA3AA511A14F89CFA62F82B" authority="(Chang, 1957)" baseAuthorityName="Chang" baseAuthorityYear="1957" box="[1024,1414,1901,1928]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B96ED60EFFA3AA511A14F89CFB3AF824" box="[1024,1246,1901,1927]" italics="true" pageId="3" pageNumber="159">Halysestheria ovata</emphasis>
(
<bibRefCitation id="EF8B77EDFFA3AA511AF8F89CFA9BF82B" author="Chang, W. T." box="[1260,1407,1901,1928]" pageId="3" pageNumber="159" pagination="479 - 502" refId="ref4060" refString="Chang, W. T. (1957) Fossil Conchostraca from the Nenjiang Shale, NW Heilongjiang. Acta Palaeontologica Sinica, 5, 479 - 502. 6 pls." type="book chapter" year="1957">Chang, 1957</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C1A719FFFA3AA511B84F89FFBF4F808" authority="Chen" authorityName="Chen" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="qinggangensis">
<emphasis id="B96ED60EFFA3AA511B84F89FFA40F824" box="[1424,1444,1902,1927]" italics="true" pageId="3" pageNumber="159">H</emphasis>
.
<emphasis id="B96ED60EFFA3AA511D3EF863FC28F808" box="[810,972,1938,1963]" italics="true" pageId="3" pageNumber="159">qinggangensis</emphasis>
Chen
</taxonomicName>
<emphasis id="B96ED60EFFA3AA511A0DF863FBCBF808" box="[1049,1071,1938,1963]" italics="true" pageId="3" pageNumber="159">in</emphasis>
<bibRefCitation id="EF8B77EDFFA3AA511A2CF860FAEBF808" author="Zhang, W. T. &amp; Chen, P. J. &amp; Shen, Y. B." box="[1080,1295,1937,1963]" pageId="3" pageNumber="159" refId="ref5922" refString="Zhang, W. T., Chen, P. J. &amp; Shen, Y. B. (1976) Fossil Conchostraca of China. Science Press, Beijing, 325 pp., 138 pls. [In Chinese]." type="book" year="1976">
Zhang
<emphasis id="B96ED60EFFA3AA511A9DF863FB59F808" box="[1161,1213,1937,1963]" italics="true" pageId="3" pageNumber="159">et al</emphasis>
., 1976
</bibRefCitation>
,
<taxonomicName id="4C1A719FFFA3AA511B08F863FC28F86C" authority="Zhang &amp; Chen" authorityName="Zhang &amp; Chen" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="rhombica">
<emphasis id="B96ED60EFFA3AA511B08F863FAD4F808" box="[1308,1328,1938,1963]" italics="true" pageId="3" pageNumber="159">H</emphasis>
.
<emphasis id="B96ED60EFFA3AA511B55F860FA48F808" box="[1345,1452,1937,1963]" italics="true" pageId="3" pageNumber="159">rhombica</emphasis>
Zhang &amp; Chen
</taxonomicName>
<emphasis id="B96ED60EFFA3AA511DC5F847FC03F86C" box="[977,999,1974,1999]" italics="true" pageId="3" pageNumber="159">in</emphasis>
<bibRefCitation id="EF8B77EDFFA3AA511DF8F844FB51F86C" author="Zhang, W. T. &amp; Chen, P. J. &amp; Shen, Y. B." box="[1004,1205,1973,1999]" pageId="3" pageNumber="159" refId="ref5922" refString="Zhang, W. T., Chen, P. J. &amp; Shen, Y. B. (1976) Fossil Conchostraca of China. Science Press, Beijing, 325 pp., 138 pls. [In Chinese]." type="book" year="1976">
Zhang
<emphasis id="B96ED60EFFA3AA511A2CF847FB83F86C" box="[1080,1127,1973,1999]" italics="true" pageId="3" pageNumber="159">et al</emphasis>
., 1976
</bibRefCitation>
,
<emphasis id="B96ED60EFFA3AA511AAAF847FB36F86C" box="[1214,1234,1974,1999]" italics="true" pageId="3" pageNumber="159">H</emphasis>
.?
<emphasis id="B96ED60EFFA3AA511AFEF844FABBF86C" box="[1258,1375,1973,1999]" italics="true" pageId="3" pageNumber="159">sublineata</emphasis>
Zhang &amp; Chen
<emphasis id="B96ED60EFFA3AA511D87F82BFC4DF850" box="[915,937,2010,2035]" italics="true" pageId="3" pageNumber="159">in</emphasis>
<bibRefCitation id="EF8B77EDFFA3AA511DA0F828FB77F850" author="Zhang, W. T. &amp; Chen, P. J. &amp; Shen, Y. B." box="[948,1171,2009,2035]" pageId="3" pageNumber="159" refId="ref5922" refString="Zhang, W. T., Chen, P. J. &amp; Shen, Y. B. (1976) Fossil Conchostraca of China. Science Press, Beijing, 325 pp., 138 pls. [In Chinese]." type="book" year="1976">
Zhang
<emphasis id="B96ED60EFFA3AA511A1CF82BFBDAF850" box="[1032,1086,2009,2035]" italics="true" pageId="3" pageNumber="159">et al</emphasis>
., 1976
</bibRefCitation>
,
<taxonomicName id="4C1A719FFFA3AA511AB7F82BFC62F7B4" authority="Zhang &amp; Chen" authorityName="Zhang &amp; Chen" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="subquadrata">
<emphasis id="B96ED60EFFA3AA511AB7F82BFB53F850" box="[1187,1207,2010,2035]" italics="true" pageId="3" pageNumber="159">H</emphasis>
.
<emphasis id="B96ED60EFFA3AA511ADEF828FABCF850" box="[1226,1368,2009,2035]" italics="true" pageId="3" pageNumber="159">subquadrata</emphasis>
Zhang &amp; Chen
</taxonomicName>
<emphasis id="B96ED60EFFA3AA511D86F80FFC4CF7B4" box="[914,936,2046,2071]" italics="true" pageId="3" pageNumber="159">in</emphasis>
<bibRefCitation id="EF8B77EDFFA3AA511DA0F80CFB75F7B4" author="Zhang, W. T. &amp; Chen, P. J. &amp; Shen, Y. B." box="[948,1169,2045,2071]" pageId="3" pageNumber="159" refId="ref5922" refString="Zhang, W. T., Chen, P. J. &amp; Shen, Y. B. (1976) Fossil Conchostraca of China. Science Press, Beijing, 325 pp., 138 pls. [In Chinese]." type="book" year="1976">
Zhang
<emphasis id="B96ED60EFFA3AA511A13F80FFBD9F7B4" box="[1031,1085,2045,2071]" italics="true" pageId="3" pageNumber="159">et al</emphasis>
., 1976
</bibRefCitation>
,
<taxonomicName id="4C1A719FFFA3AA541AB5F80FFF28F863" authority="(Chang, 1957)" baseAuthorityName="Chang" baseAuthorityYear="1957" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="6" lastPageNumber="162" order="Diplostraca" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="subtermina">
<emphasis id="B96ED60EFFA3AA511AB5F80FFB51F7B4" box="[1185,1205,2046,2071]" italics="true" pageId="3" pageNumber="159">H</emphasis>
.
<emphasis id="B96ED60EFFA3AA511ADCF80CFAA2F7B4" box="[1224,1350,2045,2071]" italics="true" pageId="3" pageNumber="159">subtermina</emphasis>
(
<bibRefCitation id="EF8B77EDFFA3AA541B4EF80CFF20F863" author="Chang, W. T." lastPageId="6" lastPageNumber="162" pageId="3" pageNumber="159" pagination="479 - 502" refId="ref4060" refString="Chang, W. T. (1957) Fossil Conchostraca from the Nenjiang Shale, NW Heilongjiang. Acta Palaeontologica Sinica, 5, 479 - 502. 6 pls." type="book chapter" year="1957">Chang, 1957</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C1A719FFFA6AA541EC3F857FE20F863" authority="(Chang, 1957)" baseAuthorityName="Chang" baseAuthorityYear="1957" box="[215,452,1957,1984]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="yui">
<emphasis id="B96ED60EFFA6AA541EC3F857FF0FF81C" box="[215,235,1958,1983]" italics="true" pageId="6" pageNumber="162">H</emphasis>
.
<emphasis id="B96ED60EFFA6AA541EECF857FEFEF81C" box="[248,282,1958,1983]" italics="true" pageId="6" pageNumber="162">yui</emphasis>
(
<bibRefCitation id="EF8B77EDFFA6AA541F3CF854FE58F863" author="Chang, W. T." box="[296,444,1957,1984]" pageId="6" pageNumber="162" pagination="479 - 502" refId="ref4060" refString="Chang, W. T. (1957) Fossil Conchostraca from the Nenjiang Shale, NW Heilongjiang. Acta Palaeontologica Sinica, 5, 479 - 502. 6 pls." type="book chapter" year="1957">Chang, 1957</bibRefCitation>
)
</taxonomicName>
. Li (2005) considered that
<taxonomicName id="4C1A719FFFA6AA541CFAF857FECEF840" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="qinggangensis">
<emphasis id="B96ED60EFFA6AA541CFAF857FCE6F81C" box="[750,770,1958,1983]" italics="true" pageId="6" pageNumber="162">H</emphasis>
.
<emphasis id="B96ED60EFFA6AA541E9CF83BFECEF840" box="[136,298,1994,2019]" italics="true" pageId="6" pageNumber="162">qinggangensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1A719FFFA6AA541F74F83BFDE4F840" baseAuthorityName="Chang" baseAuthorityYear="1957" box="[352,512,1993,2020]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="subtermina">
<emphasis id="B96ED60EFFA6AA541F74F83BFE90F840" box="[352,372,1994,2019]" italics="true" pageId="6" pageNumber="162">H</emphasis>
.
<emphasis id="B96ED60EFFA6AA541F96F838FDE4F840" box="[386,512,1993,2019]" italics="true" pageId="6" pageNumber="162">subtermina</emphasis>
</taxonomicName>
are junior synonyms of
<taxonomicName id="4C1A719FFFA6AA541E9CF81FFF28F7A4" baseAuthorityName="Chang" baseAuthorityYear="1957" box="[136,204,2030,2056]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="yui">
<emphasis id="B96ED60EFFA6AA541E9CF81FFF78F7A4" box="[136,156,2030,2055]" italics="true" pageId="6" pageNumber="162">H</emphasis>
.
<emphasis id="B96ED60EFFA6AA541EBEF81FFF28F7A4" box="[170,204,2030,2055]" italics="true" pageId="6" pageNumber="162">yui</emphasis>
</taxonomicName>
, the former two species have an elliptical carapace, and
<taxonomicName id="4C1A719FFFA6AA541EA3F7E3FF1FF788" baseAuthorityName="Chang" baseAuthorityYear="1957" box="[183,251,2066,2092]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="yui">
<emphasis id="B96ED60EFFA6AA541EA3F7E3FF2FF788" box="[183,203,2066,2091]" italics="true" pageId="6" pageNumber="162">H</emphasis>
.
<emphasis id="B96ED60EFFA6AA541ECDF7E3FF1FF788" box="[217,251,2066,2091]" italics="true" pageId="6" pageNumber="162">yui</emphasis>
</taxonomicName>
has a rounded carapace. The rounded carapace may be female, and the elliptical carapace may represent the male. While the
<emphasis id="B96ED60EFFA6AA541A0DF838FBC9F841" box="[1049,1069,1993,2018]" italics="true" pageId="6" pageNumber="162">H</emphasis>
.?
<emphasis id="B96ED60EFFA6AA541A5FF839FB24F841" box="[1099,1216,1992,2018]" italics="true" pageId="6" pageNumber="162">sublineata</emphasis>
, having a long oval carapace, was named basing on only
<specimenCount id="9D1CC195FFA6AA541AA3F81FFAB4F7A4" box="[1207,1360,2029,2056]" pageId="6" pageNumber="162" type="generic">one specimen</specimenCount>
.It could be a junior synonym of the
<typeStatus id="54A1B4BEFFA6AA541A77F7E2FB77F78E" box="[1123,1171,2067,2093]" pageId="6" pageNumber="162">type</typeStatus>
species.
<taxonomicName id="4C1A719FFFA6AA541AEFF7E2FA48F78F" box="[1275,1452,2066,2093]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="subquadrata">
<emphasis id="B96ED60EFFA6AA541AEFF7E2FAEBF78F" box="[1275,1295,2067,2092]" italics="true" pageId="6" pageNumber="162">H</emphasis>
.
<emphasis id="B96ED60EFFA6AA541B0AF7E3FA48F78F" box="[1310,1452,2066,2092]" italics="true" pageId="6" pageNumber="162">subquadrata</emphasis>
</taxonomicName>
has a subquadrate carapace, which associates with
<taxonomicName id="4C1A719FFFA7AA551CABFF5FFCE6FF64" baseAuthorityName="Chang" baseAuthorityYear="1957" box="[703,770,174,200]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="7" pageNumber="163" phylum="Arthropoda" rank="species" species="yui">
<emphasis id="B96ED60EFFA7AA551CABFF5FFD37FF64" box="[703,723,174,199]" italics="true" pageId="7" pageNumber="163">H</emphasis>
.
<emphasis id="B96ED60EFFA7AA551CF4FF5FFCE6FF64" box="[736,770,174,199]" italics="true" pageId="7" pageNumber="163">yui</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1A719FFFA7AA551E9CFF22FF08FF4F" baseAuthorityName="Chang" baseAuthorityYear="1957" box="[136,236,210,236]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="7" pageNumber="163" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B96ED60EFFA7AA551E9CFF22FF78FF4F" box="[136,156,211,236]" italics="true" pageId="7" pageNumber="163">H</emphasis>
.
<emphasis id="B96ED60EFFA7AA551EBAFF22FF08FF4F" box="[174,236,211,236]" italics="true" pageId="7" pageNumber="163">ovata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1A719FFFA7AA551F3EFF22FD21FF4E" authority="(Zhang et al., 1976)" baseAuthorityName="Zhang" baseAuthorityYear="1976" box="[298,709,210,237]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="7" pageNumber="163" phylum="Arthropoda" rank="species" species="subtermina">
<emphasis id="B96ED60EFFA7AA551F3EFF22FEDAFF4F" box="[298,318,211,236]" italics="true" pageId="7" pageNumber="163">H</emphasis>
.
<emphasis id="B96ED60EFFA7AA551F44FF23FE2AFF4F" box="[336,462,210,236]" italics="true" pageId="7" pageNumber="163">subtermina</emphasis>
(
<bibRefCitation id="EF8B77EDFFA7AA551FF4FF23FD59FF4E" author="Zhang, W. T. &amp; Chen, P. J. &amp; Shen, Y. B." box="[480,701,210,237]" pageId="7" pageNumber="163" refId="ref5922" refString="Zhang, W. T., Chen, P. J. &amp; Shen, Y. B. (1976) Fossil Conchostraca of China. Science Press, Beijing, 325 pp., 138 pls. [In Chinese]." type="book" year="1976">
Zhang
<emphasis id="B96ED60EFFA7AA551C20FF22FD8DFF4F" box="[564,617,210,236]" italics="true" pageId="7" pageNumber="163">et al</emphasis>
., 1976
</bibRefCitation>
)
</taxonomicName>
, thus, may be they are the same species.
<taxonomicName id="4C1A719FFFA7AA551C19FF06FD7FFEB3" box="[525,667,246,273]" class="Branchiopoda" genus="Halysestheria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplostraca" pageId="7" pageNumber="163" phylum="Arthropoda" rank="species" species="rhombica">
<emphasis id="B96ED60EFFA7AA551C19FF06FDC5FEB3" box="[525,545,247,272]" italics="true" pageId="7" pageNumber="163">H</emphasis>
.
<emphasis id="B96ED60EFFA7AA551C24FF07FD7FFEB3" box="[560,667,246,272]" italics="true" pageId="7" pageNumber="163">rhombica</emphasis>
</taxonomicName>
is similar with the
<typeStatus id="54A1B4BEFFA7AA551EFEFEEAFEFEFE96" box="[234,282,283,309]" pageId="7" pageNumber="163">type</typeStatus>
species, and was named basing on only one broken specimen. It could be a junior synonym of the
<typeStatus id="54A1B4BEFFA7AA551CCEFEB1FCEEFEF9" box="[730,778,320,346]" pageId="7" pageNumber="163">type</typeStatus>
species.
</paragraph>
</subSubSection>
</treatment>
</document>