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<document id="F6EE6AE6D2C94597CE2E90A985B81D23" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFE11F4FFF50FAA7FE568F25" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax baracoensis Borhidi &amp; Muniz" docType="treatment" docVersion="1" lastPageNumber="33" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="31" updateTime="1696265644834" updateUser="diego">
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<mods:title id="7ED8995DE5A543A44337E0EF98B4C4F3">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<mods:namePart id="A1B2018B025BA3259F676DCFAEA34AA2">Henderson, Andrew</mods:namePart>
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<mods:title id="5864842A33B0586340D813E03EF45666">Phytotaxa</mods:title>
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<mods:date id="86C0556E3404B2BD60007EBA841776F3">2023</mods:date>
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<mods:number id="DBB991C88519349E526D9A4F504F8137">614</mods:number>
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<mods:classification id="23FA1F76B7B102C8122968D23D076636">journal article</mods:classification>
<mods:identifier id="13CF0936FB6DC75B556A67EBCFEC88CB" type="DOI">10.11646/phytotaxa.614.1.1</mods:identifier>
<mods:identifier id="1F56B6F751103E2EF6489514FDDFE440" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="9605732F749C3CE2BA3C1C9FC6B9C0EA" type="Zenodo-Dep">8389307</mods:identifier>
</mods:mods>
<treatment id="03B387DAFFE11F4FFF50FAA7FE568F25" LSID="urn:lsid:plazi:treatment:03B387DAFFE11F4FFF50FAA7FE568F25" httpUri="http://treatment.plazi.org/id/03B387DAFFE11F4FFF50FAA7FE568F25" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<subSubSection id="C3006547FFE11F71FF50FAA7FB498D23" box="[136,1200,1310,1337]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BA536CCFFE11F71FF50FAA7FB498D23" blockId="30.[136,1452,1310,1408]" box="[136,1200,1310,1337]" pageId="30" pageNumber="31">
<emphasis id="B96EEADEFFE11F71FF50FAA7FB498D23" bold="true" box="[136,1200,1310,1337]" pageId="30" pageNumber="31">
1.4.
<taxonomicName id="4C1A4D4FFFE11F71FF6FFAA7FD4A8D23" ID-CoL="WLXV" authority="Borhidi &amp; Muniz" authorityName="Borhidi &amp; Muniz" box="[183,691,1310,1337]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="baracoensis">
<emphasis id="B96EEADEFFE11F71FF6FFAA7FE1A8D22" bold="true" box="[183,483,1310,1337]" italics="true" pageId="30" pageNumber="31">Coccothrinax baracoensis</emphasis>
Borhidi &amp; Muñiz
</taxonomicName>
in
<bibRefCitation id="EF8B4B3DFFE11F71FD0DFAA7FC1E8D23" author="Muniz, O. &amp; Borhidi, A." box="[725,999,1310,1336]" pageId="30" pageNumber="31" pagination="439 - 454" refId="ref63543" refString="Muniz, O. &amp; Borhidi, A. (1981 publ. 1982) Palmas nuevas del genero Coccothrinax Sarg. en Cuba. Acta Botanica Academiae Scientiarum Hungaricae 27: 439 - 454." type="journal article" year="1981">Muñiz &amp; Borhidi (1981</bibRefCitation>
publ. 1982: 440).
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFE11F71FB6DFAA6FE5A8D9B" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BA536CCFFE11F71FB6DFAA6FE5A8D9B" blockId="30.[136,1452,1310,1408]" pageId="30" pageNumber="31">
<materialsCitation id="3B723C91FFE11F71FB6DFAA6FE5A8D9B" collectingDate="1978-02-19" collectionCode="B, R, HAC" country="Cuba" location="Baracoa" municipality="Arroyo Blanco" pageId="30" pageNumber="31" specimenCount="2" stateProvince="Guantanamo" typeStatus="lectotype">
<typeStatus id="54A1886EFFE11F71FB6DFAA6FADC8D22" box="[1205,1317,1311,1337]" pageId="30" pageNumber="31">Lectotype</typeStatus>
(designated by
<bibRefCitation id="EF8B4B3DFFE11F71FF77FAFAFECD8D47" author="Moya, C." box="[175,308,1346,1373]" pageId="30" pageNumber="31" pagination="1 - 27" refId="ref63288" refString="Moya, C. (2021) An annotated checklist of Cuban palms 6. Coccothrinax, Pt. 2. 1972 - 1995. Nomenclature, typification, and distribution. PalmArbor 3: 1 - 27." type="journal article" year="2021">Moya 2021</bibRefCitation>
):—
<collectingCountry id="F30D765CFFE11F71FE87FAFBFE4B8D47" box="[351,434,1346,1372]" name="Cuba" pageId="30" pageNumber="31">CUBA</collectingCountry>
.
<collectingRegion id="49DEF82EFFE11F71FE18FAFBFDAB8D47" box="[448,594,1346,1372]" country="Cuba" name="Guantanamo" pageId="30" pageNumber="31">Guantánamo</collectingRegion>
,
<collectingMunicipality id="6BC1ACB6FFE11F71FDB8FAFBFCF78D47" box="[608,782,1346,1372]" pageId="30" pageNumber="31">Arroyo Blanco</collectingMunicipality>
,
<location id="8EC56017FFE11F71FCC5FAFAFC858D46" LSID="urn:lsid:plazi:treatment:03B387DAFFE11F4FFF50FAA7FE568F25:8EC56017FFE11F71FCC5FAFAFC858D46" box="[797,892,1347,1373]" country="Cuba" municipality="Arroyo Blanco" name="Baracoa" pageId="30" pageNumber="31" stateProvince="Guantanamo">Baracoa</location>
,
<date id="FFA4100CFFE11F71FC53FAFBFBA48D47" box="[907,1117,1346,1372]" pageId="30" pageNumber="31" value="1978-02-19">
<collectingDate id="EFE0E9E4FFE11F71FC53FAFBFBA48D47" box="[907,1117,1346,1372]" pageId="30" pageNumber="31" value="1978-02-19">19 February 1978</collectingDate>
</date>
,
<emphasis id="B96EEADEFFE11F71FBB4FAFAFA528D47" box="[1132,1451,1346,1373]" italics="true" pageId="30" pageNumber="31">
<collectionCode id="ED0BAE09FFE11F71FBB4FAFAFB798D47" box="[1132,1152,1347,1372]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="30" pageNumber="31" type="Herbarium">B</collectionCode>
. Muñoz &amp;
<collectionCode id="ED0BAE09FFE11F71FADFFAFAFAE28D47" box="[1287,1307,1347,1372]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="30" pageNumber="31">R</collectionCode>
. Fleites 105
</emphasis>
(
<typeStatus id="54A1886EFFE11F71FF48FADFFF0D8D9B" box="[144,244,1382,1408]" pageId="30" pageNumber="31">holotype</typeStatus>
<collectionCode id="ED0BAE09FFE11F71FF22FADFFEC68D9B" box="[250,319,1382,1408]" country="Cuba" httpUri="http://grbio.org/cool/cenz-te88" name="Instituto de Ecología y Sistemática" pageId="30" pageNumber="31" type="Herbarium">HAC</collectionCode>
!).
<figureCitation id="13212A49FFE11F71FE8EFADFFE5A8D9B" box="[342,419,1382,1408]" captionStart="PLATE 4" captionStartId="31.[136,214,1917,1938]" captionTargetBox="[151,1433,194,1891]" captionTargetId="figure-15@31.[150,1434,193,1893]" captionTargetPageId="31" captionText="PLATE 4. Coccothrinax baracoensis, Cuba, with thick stems and wide leaf segments. (Image by Raúl Verdecia Pérez)." figureDoi="http://doi.org/10.5281/zenodo.8389353" httpUri="https://zenodo.org/record/8389353/files/figure.png" pageId="30" pageNumber="31">Plate 4</figureCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFE11F71FF50FA17FD838FDB" pageId="30" pageNumber="31" type="description">
<paragraph id="8BA536CCFFE11F71FF50FA17FD838FDB" blockId="30.[136,1452,1454,2057]" pageId="30" pageNumber="31">
<emphasis id="B96EEADEFFE11F71FF50FA17FF298DD3" bold="true" box="[136,208,1454,1480]" pageId="30" pageNumber="31">Stems</emphasis>
length not recorded, 20.0 cm diameter, solitary.
<emphasis id="B96EEADEFFE11F71FD27FA17FCA88DD3" bold="true" box="[767,849,1454,1480]" pageId="30" pageNumber="31">Leaves</emphasis>
more or less deciduous or only leaf bases persisting on stem; leaf sheath fibers
<quantity id="4CE29B29FFE11F71FE66FA6BFDEC8DF6" box="[446,533,1490,1517]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="30" pageNumber="31" unit="mm" value="1.3">1.3 mm</quantity>
diameter, stout, woody, closely woven, ± joined at the apices; petioles 18.0 mm diameter just below the apex; palmans 14.0 cm long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades not wedge-shaped; segments 30 per leaf, the middle ones 50.0 cm long and
<quantity id="4CE29B29FFE11F71FECEF987FE988E42" box="[278,353,1598,1625]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="30" pageNumber="31" unit="cm" value="6.5">6.5 cm</quantity>
wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments relatively short and broad, abruptly narrowed (shoulder) toward the apex, otherwise parallel-sided, often strongly folded, stiff and leathery, the apices briefly splitting; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially, densely indumentose abaxially, with irregularly shaped, persistent, interlocking, fimbriate hairs, each one with a rounded, raised, light green to greenish-brown or reddish-brown center, without transverse veinlets.
<emphasis id="B96EEADEFFE11F71FD84F94BFCFC8F17" bold="true" box="[604,773,1778,1804]" pageId="30" pageNumber="31">Inflorescences</emphasis>
erect, at least initially, amongst or above the leaves, with few partial inflorescences; rachis bracts narrow, closely sheathing, sparsely tomentose, usually without hairs at the apex; partial inflorescences 4; proximalmost rachillae straight,
<quantity id="4CE29B29FFE11F71FCD3F883FCA18F4E" box="[779,856,1850,1877]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="30" pageNumber="31" unit="cm" value="4.8">4.8 cm</quantity>
long and
<quantity id="4CE29B29FFE11F71FC12F883FBD88F4E" box="[970,1057,1850,1877]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="30" pageNumber="31" unit="mm" value="1.4">1.4 mm</quantity>
diameter; rachillae not recorded at or near anthesis; flowers spirally arranged; stamens 7; fruit pedicels
<quantity id="4CE29B29FFE11F71FC4AF8E7FC138F62" box="[914,1002,1886,1913]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="31" unit="mm" value="0.8">0.8 mm</quantity>
long;
<emphasis id="B96EEADEFFE11F71FBECF8E7FB8C8F63" bold="true" box="[1076,1141,1886,1912]" pageId="30" pageNumber="31">fruits</emphasis>
<quantity id="4CE29B29FFE11F71FBA5F8E7FB2C8F62" box="[1149,1237,1886,1913]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="30" pageNumber="31" unit="mm" value="9.8">9.8 mm</quantity>
long and 11.0 mm diameter, deep purple; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces deeply lobed, the lobes running from base of seeds almost to apices.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFE11F71FF65F873FDF88013" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BA536CCFFE11F71FF65F873FDF88013" blockId="30.[136,1452,1454,2057]" pageId="30" pageNumber="31">
<emphasis id="B96EEADEFFE11F71FF65F873FDFA8FFF" bold="true" box="[189,515,1994,2020]" pageId="30" pageNumber="31">Distribution and habitat:—</emphasis>
<collectingCountry id="F30D765CFFE11F71FDDBF873FDC78FFF" box="[515,574,1994,2020]" name="Cuba" pageId="30" pageNumber="31">Cuba</collectingCountry>
(
<collectingRegion id="49DEF82EFFE11F71FD93F873FD278FFF" box="[587,734,1994,2020]" country="Cuba" name="Guantanamo" pageId="30" pageNumber="31">Guantánamo</collectingRegion>
) (
<figureCitation id="13212A49FFE11F71FD2BF873FCBD8FFF" box="[755,836,1994,2020]" captionStart="FIGURE 10" captionStartId="32.[136,229,1508,1530]" captionTargetBox="[155,1429,198,1479]" captionTargetId="figure-27@32.[150,1434,193,1484]" captionTargetPageId="32" captionText="FIGURE 10. Distribution of Coccothrinax baracoensis, C. bonettiana, C. borhidiana, and C. boschiana." figureDoi="http://doi.org/10.5281/zenodo.8389355" httpUri="https://zenodo.org/record/8389355/files/figure.png" pageId="30" pageNumber="31">Fig. 10</figureCitation>
), habitat and elevation not recorded.
<bibRefCitation id="EF8B4B3DFFE11F71FB04F873FA9F8FFF" author="Craft, P." box="[1244,1382,1994,2020]" pageId="30" pageNumber="31" refId="ref61560" refString="Craft, P. (2017) The Palms of Cuba. Palm Nut Pages, Florida, 232 pp." type="book" year="2017">Craft (2017)</bibRefCitation>
wrote that it occurs on serpentine rocks.
</paragraph>
</subSubSection>
<caption id="DF656644FFE01F70FF50F8C4FAFC8F89" ID-DOI="http://doi.org/10.5281/zenodo.8389353" ID-Zenodo-Dep="8389353" box="[136,1285,1917,1939]" httpUri="https://zenodo.org/record/8389353/files/figure.png" pageId="31" pageNumber="32" startId="31.[136,214,1917,1938]" targetBox="[151,1433,194,1891]" targetPageId="31" targetType="plate">
<paragraph id="8BA536CCFFE01F70FF50F8C4FAFC8F89" blockId="31.[136,1285,1917,1939]" box="[136,1285,1917,1939]" pageId="31" pageNumber="32">
<emphasis id="B96EEADEFFE01F70FF50F8C4FF118F89" bold="true" box="[136,232,1917,1938]" pageId="31" pageNumber="32">PLATE 4</emphasis>
.
<taxonomicName id="4C1A4D4FFFE01F70FF2CF8C4FE088F89" box="[244,497,1917,1938]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="baracoensis">
<emphasis id="B96EEADEFFE01F70FF2CF8C4FE088F89" box="[244,497,1917,1938]" italics="true" pageId="31" pageNumber="32">Coccothrinax baracoensis</emphasis>
</taxonomicName>
, Cuba, with thick stems and wide leaf segments. (Image by Raúl Verdecia Pérez).
</paragraph>
</caption>
<caption id="DF656644FFDF1F4FFF50FA5DFB838DE2" ID-DOI="http://doi.org/10.5281/zenodo.8389355" ID-Zenodo-Dep="8389355" box="[136,1146,1508,1530]" httpUri="https://zenodo.org/record/8389355/files/figure.png" pageId="32" pageNumber="33" startId="32.[136,229,1508,1530]" targetBox="[155,1429,198,1479]" targetPageId="32" targetType="figure">
<paragraph id="8BA536CCFFDF1F4FFF50FA5DFB838DE2" blockId="32.[136,1146,1508,1530]" box="[136,1146,1508,1530]" pageId="32" pageNumber="33">
<emphasis id="B96EEADEFFDF1F4FFF50FA5DFEFA8DE2" bold="true" box="[136,259,1508,1530]" pageId="32" pageNumber="33">FIGURE 10</emphasis>
. Distribution of
<emphasis id="B96EEADEFFDF1F4FFE7BFA5DFC3E8DE2" box="[419,967,1508,1529]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="4C1A4D4FFFDF1F4FFE7BFA5DFD5B8DE2" box="[419,674,1508,1529]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="baracoensis">Coccothrinax baracoensis</taxonomicName>
,
<taxonomicName id="4C1A4D4FFFDF1F4FFD74FA5DFCC88DE2" box="[684,817,1508,1529]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="bonettiana">C. bonettiana</taxonomicName>
,
<taxonomicName id="4C1A4D4FFFDF1F4FFCE3FA5DFC3D8DE2" box="[827,964,1508,1529]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="borhidiana">C. borhidiana</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C1A4D4FFFDF1F4FFC2EFA5DFB8C8DE2" box="[1014,1141,1508,1529]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="boschiana">
<emphasis id="B96EEADEFFDF1F4FFC2EFA5DFB8C8DE2" box="[1014,1141,1508,1529]" italics="true" pageId="32" pageNumber="33">C. boschiana</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3006547FFDF1F4FFF65F991FE568F25" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BA536CCFFDF1F4FFF65F991FE568F25" blockId="32.[136,1452,1576,1855]" pageId="32" pageNumber="33">
<emphasis id="B96EEADEFFDF1F4FFF65F991FE528E59" bold="true" box="[189,427,1576,1602]" pageId="32" pageNumber="33">Taxonomic notes:—</emphasis>
As a preliminary species,
<taxonomicName id="4C1A4D4FFFDF1F4FFD11F991FC168E59" box="[713,1007,1576,1602]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="baracoensis">
<emphasis id="B96EEADEFFDF1F4FFD11F991FC168E59" box="[713,1007,1576,1602]" italics="true" pageId="32" pageNumber="33">Coccothrinax baracoensis</emphasis>
</taxonomicName>
has a unique combination of qualitative character states and is recognized as a phylogenetic species. The
<typeStatus id="54A1886EFFDF1F4FFC84F9F4FC758E7C" box="[860,908,1613,1639]" pageId="32" pageNumber="33">type</typeStatus>
collection label gives the collectors as
<emphasis id="B96EEADEFFDF1F4FFA98F9F4FEB78E91" italics="true" pageId="32" pageNumber="33">B. Muñoz &amp; R. Fleites 105</emphasis>
, although the protologue gives
<emphasis id="B96EEADEFFDF1F4FFD1EF9C8FC7A8E91" box="[710,899,1648,1675]" italics="true" pageId="32" pageNumber="33">O. Muñiz 15105</emphasis>
.
<taxonomicName id="4C1A4D4FFFDF1F4FFC4EF9C9FB398E91" box="[918,1216,1648,1674]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="baracoensis">
<emphasis id="B96EEADEFFDF1F4FFC4EF9C9FB398E91" box="[918,1216,1648,1674]" italics="true" pageId="32" pageNumber="33">Coccothrinax baracoensis</emphasis>
</taxonomicName>
is known from
<specimenCount id="9D1CFD45FFDF1F4FFA5BF9C8FF0B8EB5" pageId="32" pageNumber="33" type="generic">one specimen</specimenCount>
and remains poorly known, and appears somewhat unusual among Cuban species. The leaf sheath fibers were described in the protologue as being densely woven and forming a ligule at the apex. They are here scored as stout, woody, closely woven, and ± joined at the apices. Inflorescences were described as erect and to
<quantity id="4CE29B29FFDF1F4FFAFAF965FA908EEC" box="[1314,1385,1756,1783]" metricMagnitude="-1" metricUnit="m" metricValue="8.5" pageId="32" pageNumber="33" unit="cm" value="85.0">85 cm</quantity>
long. The embryo was described as basal but it is clearly apical.
<taxonomicName id="4C1A4D4FFFDF1F4FFCC2F8B9FBB88F01" box="[794,1089,1792,1818]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="baracoensis">
<emphasis id="B96EEADEFFDF1F4FFCC2F8B9FBB88F01" box="[794,1089,1792,1818]" italics="true" pageId="32" pageNumber="33">Coccothrinax baracoensis</emphasis>
</taxonomicName>
has unusually thick stems, wide segments, and large fruits.
</paragraph>
</subSubSection>
</treatment>
</document>
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