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<document id="0F91188960524D2C384849FA91D2F39E" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFBE1F2CFF50F987FC1B8A5F" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax munizii Borhidi" docType="treatment" docVersion="5" lastPageNumber="68" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="66" updateTime="1696265644834" updateUser="diego">
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<mods:title id="200003764E665C6B9FAC7EFF78850E8B">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<mods:title id="77A649CAB6F6A9B952FA97AC80CB7776">Phytotaxa</mods:title>
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<mods:date id="48A5256E73B6FFA391485FE834A5C1DF">2023</mods:date>
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<treatment id="03B387DAFFBE1F2CFF50F987FC1B8A5F" LSID="urn:lsid:plazi:treatment:03B387DAFFBE1F2CFF50F987FC1B8A5F" httpUri="http://treatment.plazi.org/id/03B387DAFFBE1F2CFF50F987FC1B8A5F" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66">
<subSubSection id="C3006547FFBE1F2EFF50F987FBE48E43" box="[136,1053,1598,1625]" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="8BA536CCFFBE1F2EFF50F987FBE48E43" blockId="65.[136,1452,1598,1697]" box="[136,1053,1598,1625]" pageId="65" pageNumber="66">
<emphasis id="B96EEADEFFBE1F2EFF50F987FBE48E43" bold="true" box="[136,1053,1598,1625]" pageId="65" pageNumber="66">
1.26.
<taxonomicName id="4C1A4D4FFFBE1F2EFF1FF987FDDD8E43" ID-CoL="WLZ4" authority="Borhidi" authorityName="Borhidi" box="[199,548,1598,1625]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="65" pageNumber="66" phylum="Tracheophyta" rank="species" species="munizii">
<emphasis id="B96EEADEFFBE1F2EFF1FF987FE388E43" bold="true" box="[199,449,1598,1625]" italics="true" pageId="65" pageNumber="66">Coccothrinax munizii</emphasis>
Borhidi
</taxonomicName>
in
<bibRefCitation id="EF8B4B3DFFBE1F2EFD90F987FC978E43" author="Borhidi, A. &amp; Muniz, O." box="[584,878,1598,1624]" pageId="65" pageNumber="66" pagination="1 - 36" refId="ref61092" refString="Borhidi, A. &amp; Muniz, O. (1971 b publ. 1972) New plants in Cuba I. Acta Botanica Academiae Scientiarum Hungaricae 17: 1 - 36." type="journal article" year="1971">Borhidi &amp; Muñiz (1971b</bibRefCitation>
publ. 1972: 2).
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFBE1F2EFBFCF987FE048E67" pageId="65" pageNumber="66" type="reference_group">
<paragraph id="8BA536CCFFBE1F2EFBFCF987FE048E67" blockId="65.[136,1452,1598,1697]" pageId="65" pageNumber="66">
<taxonomicName id="4C1A4D4FFFBE1F2EFBFCF987FA558E43" authority="(Borhidi) Borhidi" authorityName="Borhidi" baseAuthorityName="Borhidi" box="[1060,1452,1598,1624]" class="Liliopsida" family="Arecaceae" genus="Haitiella" kingdom="Plantae" order="Arecales" pageId="65" pageNumber="66" phylum="Tracheophyta" rank="species" species="munizii">
<emphasis id="B96EEADEFFBE1F2EFBFCF987FB198E43" box="[1060,1248,1598,1624]" italics="true" pageId="65" pageNumber="66">Haitiella munizii</emphasis>
(Borhidi)
<materialsCitation id="3B723C91FFBE1F2EFA8EF987FA558E43" box="[1366,1452,1598,1624]" collectingDate="1970-03-16" collectionCode="A, O, S, HAC, BP" country="Cuba" county="Imias" location="Type" municipality="San Antonio del Sur" pageId="65" pageNumber="66" specimenCount="1" typeStatus="holotype">Borhidi</materialsCitation>
</taxonomicName>
in
<bibRefCitation id="EF8B4B3DFFBE1F2EFF7EF9DBFE008E67" author="Borhidi, A. &amp; Kereszty, Z." box="[166,505,1634,1661]" pageId="65" pageNumber="66" pagination="1 - 37" refId="ref61155" refString="Borhidi, A. &amp; Kereszty, Z. (1979) New names and new species in the flora of Cuba resp. Antilles. Acta Botanica Academiae Scientiarum Hungaricae 25: 1 - 37." type="journal article" year="1979">Borhidi &amp; Kereszty (1979: 2)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFBE1F2EFDDCF9DBFB108EBB" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="8BA536CCFFBE1F2EFDDCF9DBFB108EBB" blockId="65.[136,1452,1598,1697]" pageId="65" pageNumber="66">
<location id="8EC56017FFBE1F2EFDDCF9DBFDC68E67" LSID="urn:lsid:plazi:treatment:03B387DAFFBE1F2CFF50F987FC1B8A5F:8EC56017FFBE1F2EFDDCF9DBFDC68E67" box="[516,575,1634,1660]" country="Cuba" county="Imias" municipality="San Antonio del Sur" name="Type" pageId="65" pageNumber="66">Type</location>
:—
<collectingCountry id="F30D765CFFBE1F2EFDB9F9DBFD488E67" box="[609,689,1634,1660]" name="Cuba" pageId="65" pageNumber="66">CUBA</collectingCountry>
. Prov. Oriente, ad
<location id="8EC56017FFBE1F2EFC5CF9DBFC0D8E67" LSID="urn:lsid:plazi:treatment:03B387DAFFBE1F2CFF50F987FC1B8A5F:8EC56017FFBE1F2EFC5CF9DBFC0D8E67" box="[900,1012,1634,1660]" country="Cuba" county="Imias" municipality="San Antonio del Sur" name="Macambo" pageId="65" pageNumber="66">Macambo</location>
inter
<collectingMunicipality id="6BC1ACB6FFBE1F2EFBE0F9DBFAE78E67" box="[1080,1310,1634,1660]" pageId="65" pageNumber="66">San Antonio del Sur</collectingMunicipality>
et
<collectingCounty id="62C44E40FFBE1F2EFA9AF9DBFA7A8E67" box="[1346,1411,1634,1660]" pageId="65" pageNumber="66">Imias</collectingCounty>
,
<date id="FFA4100CFFBE1F2EFA57F9DBFEEA8EBB" pageId="65" pageNumber="66" value="1970-03-16">
<collectingDate id="EFE0E9E4FFBE1F2EFA57F9DBFEEA8EBB" pageId="65" pageNumber="66" value="1970-03-16">16 March 1970</collectingDate>
</date>
,
<emphasis id="B96EEADEFFBE1F2EFEC6F93EFD138EBB" box="[286,746,1670,1697]" italics="true" pageId="65" pageNumber="66">A. Borhidi, O Muñiz &amp; S. Vázquez 27117</emphasis>
(
<typeStatus id="54A1886EFFBE1F2EFD20F93FFCA58EBB" box="[760,860,1670,1696]" pageId="65" pageNumber="66">holotype</typeStatus>
HAC!, isotype BP
<emphasis id="B96EEADEFFBE1F2EFBEDF93EFBA58EBB" box="[1077,1116,1671,1696]" italics="true" pageId="65" pageNumber="66">n.v.</emphasis>
, BP image!)
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFBE1F2CFF50F977FC1E8976" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66" type="description">
<paragraph id="8BA536CCFFBE1F2CFF50F977FC1E8976" blockId="65.[136,1451,1742,2093]" lastBlockId="67.[136,1452,158,581]" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66">
<emphasis id="B96EEADEFFBE1F2EFF50F977FF298EF3" bold="true" box="[136,208,1742,1768]" pageId="65" pageNumber="66">Stems</emphasis>
4.3(3.55.0) m long and
<quantity id="4CE29B29FFBE1F2EFE3CF977FDD78EF2" box="[484,558,1742,1769]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="65" pageNumber="66" unit="cm" value="5.3">5.3 cm</quantity>
diameter, solitary.
<emphasis id="B96EEADEFFBE1F2EFD27F977FCA88EF3" bold="true" box="[767,849,1742,1768]" pageId="65" pageNumber="66">Leaves</emphasis>
more or less deciduous or only leaf bases persisting on stem; leaf sheath fibers 2.3(1.33.1) mm diameter, stout, woody, loosely woven, ± joined or briefly free at the apices; petioles 7.3(6.78.0) mm diameter just below the apex; palmans 7.5(6.38.7) cm long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades wedge-shaped; segments 19(1621) per leaf, the middle ones 28.1(26.531.5) cm long and 2.7(1.83.5) cm wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments tapering from base to apex, often folded, stiff and leathery, with or without scarcely developed shoulders, the apices sharply pointed and briefly splitting; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially; leaf segments densely indumentose abaxially, with irregularly shaped, persistent, interlocking, fimbriate hairs, each one with a rounded, raised, light green to greenish-brown center, without transverse veinlets.
<emphasis id="B96EEADEFFBE1F2EFB43F7ABFABD8037" bold="true" box="[1179,1348,2066,2092]" pageId="65" pageNumber="66">Inflorescences</emphasis>
curving, arching, or pendulous amongst the leaves, with few to numerous partial inflorescences; rachis bracts narrow, closely sheathing, sparsely tomentose, usually without hairs at the apex; partial inflorescences 56; proximalmost rachillae straight, 4.6(4.05.2) cm long and 1.0(0.91.2) mm diameter in fruit; rachillae uneven at or near anthesis with lines of warty outgrowths, these often becoming more pronounced as fruits develop; stamens 8; fruit pedicels
<quantity id="4CE29B29FFBC1F2CFAFCFEB3FA55893E" box="[1316,1452,266,293]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="67" pageNumber="68" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.20.3 mm</quantity>
long;
<emphasis id="B96EEADEFFBC1F2CFF12FE97FEF28953" bold="true" box="[202,267,302,328]" pageId="67" pageNumber="68">fruits</emphasis>
3.9(3.74.2) mm long and 4.3(4.04.7) mm diameter, tan; fruit surfaces densely muricate; seed surfaces deeply lobed, the lobes running from base of seeds approximately to equator.
</paragraph>
</subSubSection>
<caption id="DF656644FFBD1F2DFF50F8C4FAD68F88" ID-DOI="http://doi.org/10.5281/zenodo.8389399" ID-Zenodo-Dep="8389399" box="[136,1327,1917,1939]" httpUri="https://zenodo.org/record/8389399/files/figure.png" pageId="66" pageNumber="67" startId="66.[136,214,1917,1938]" targetBox="[151,1433,194,1891]" targetPageId="66" targetType="plate">
<paragraph id="8BA536CCFFBD1F2DFF50F8C4FAD68F88" blockId="66.[136,1327,1917,1939]" box="[136,1327,1917,1939]" pageId="66" pageNumber="67">
<emphasis id="B96EEADEFFBD1F2DFF50F8C4FF0D8F89" bold="true" box="[136,244,1917,1938]" pageId="66" pageNumber="67">PLATE 19</emphasis>
.
<taxonomicName id="4C1A4D4FFFBD1F2DFED8F8C4FDDC8F88" box="[256,549,1917,1939]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="montgomeryana">
<emphasis id="B96EEADEFFBD1F2DFED8F8C4FDDC8F88" box="[256,549,1917,1939]" italics="true" pageId="66" pageNumber="67">Coccothrinax montgomeryana</emphasis>
</taxonomicName>
, Dominican Republic, with slender, curved stems. (Image by Eladio Fernández).
</paragraph>
</caption>
<subSubSection id="C3006547FFBC1F2CFF65FECFFE5289AF" pageId="67" pageNumber="68" type="distribution">
<paragraph id="8BA536CCFFBC1F2CFF65FECFFE5289AF" blockId="67.[136,1452,158,581]" pageId="67" pageNumber="68">
<emphasis id="B96EEADEFFBC1F2CFF65FECFFDFF898B" bold="true" box="[189,518,374,400]" pageId="67" pageNumber="68">Distribution and habitat:—</emphasis>
<collectingCountry id="F30D765CFFBC1F2CFDDEFECFFDB8898B" box="[518,577,374,400]" name="Cuba" pageId="67" pageNumber="68">Cuba</collectingCountry>
(
<collectingRegion id="49DEF82EFFBC1F2CFD88FECFFD1A898B" box="[592,739,374,400]" country="Cuba" name="Guantanamo" pageId="67" pageNumber="68">Guantánamo</collectingRegion>
) (
<figureCitation id="13212A49FFBC1F2CFD22FECFFCB2898A" box="[762,843,374,401]" captionStart="FIGURE 15" captionStartId="67.[136,229,1957,1979]" captionTargetBox="[155,1429,649,1928]" captionTargetId="figure-194@67.[150,1434,644,1933]" captionTargetPageId="67" captionText="FIGURE 15. Distribution of Coccothrinax montgomeryana, C. munizii, C. orientalis, and C. pauciramosa." figureDoi="http://doi.org/10.5281/zenodo.8389401" httpUri="https://zenodo.org/record/8389401/files/figure.png" pageId="67" pageNumber="68">Fig. 15</figureCitation>
) in dry, scrub forest on limestone cliffs near the sea at 163(25300) m elevation.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFBC1F2CFF65FE07FC1B8A5F" pageId="67" pageNumber="68" type="discussion">
<paragraph id="8BA536CCFFBC1F2CFF65FE07FC1B8A5F" blockId="67.[136,1452,158,581]" pageId="67" pageNumber="68">
<emphasis id="B96EEADEFFBC1F2CFF65FE07FE5689C3" bold="true" box="[189,431,446,472]" pageId="67" pageNumber="68">Taxonomic notes:—</emphasis>
As a preliminary species,
<taxonomicName id="4C1A4D4FFFBC1F2CFD07FE07FC2D89C3" box="[735,980,446,472]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="67" pageNumber="68" phylum="Tracheophyta" rank="species" species="munizii">
<emphasis id="B96EEADEFFBC1F2CFD07FE07FC2D89C3" box="[735,980,446,472]" italics="true" pageId="67" pageNumber="68">Coccothrinax munizii</emphasis>
</taxonomicName>
has a unique combination of qualitative character states and is recognized as a phylogenetic species. It is one of only four species with small, muricate fruits, the other three (
<emphasis id="B96EEADEFFBC1F2CFEE6FDBEFDA78A3B" box="[318,606,518,545]" italics="true" pageId="67" pageNumber="68">
<taxonomicName id="4C1A4D4FFFBC1F2CFEE6FDBEFE2F8A3B" authorityName="Mejia &amp; Garcia" authorityYear="1997" box="[318,470,518,545]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="67" pageNumber="68" phylum="Tracheophyta" rank="species" species="boschiana">C. boschiana</taxonomicName>
,
<taxonomicName id="4C1A4D4FFFBC1F2CFE3BFDBEFDA78A3B" box="[483,606,518,545]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="67" pageNumber="68" phylum="Tracheophyta" rank="species" species="ekmanii">C. ekmanii</taxonomicName>
</emphasis>
,
<taxonomicName id="4C1A4D4FFFBC1F2CFDB5FDBEFCE98A3B" box="[621,784,519,545]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="67" pageNumber="68" phylum="Tracheophyta" rank="species" species="samanensis">
<emphasis id="B96EEADEFFBC1F2CFDB5FDBEFCE98A3B" box="[621,784,519,545]" italics="true" pageId="67" pageNumber="68">C. samanensis</emphasis>
</taxonomicName>
) are endemic to Hispaniola.
<taxonomicName id="4C1A4D4FFFBC1F2CFB82FDBFFAB48A3B" box="[1114,1357,518,544]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="67" pageNumber="68" phylum="Tracheophyta" rank="species" species="munizii">
<emphasis id="B96EEADEFFBC1F2CFB82FDBFFAB48A3B" box="[1114,1357,518,544]" italics="true" pageId="67" pageNumber="68">Coccothrinax munizii</emphasis>
</taxonomicName>
appears most similar morphologically to
<taxonomicName id="4C1A4D4FFFBC1F2CFE2FFD92FD728A5F" authorityName="Mejia &amp; Garcia" authorityYear="1997" box="[503,651,554,581]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="67" pageNumber="68" phylum="Tracheophyta" rank="species" species="boschiana">
<emphasis id="B96EEADEFFBC1F2CFE2FFD92FD728A5F" box="[503,651,554,581]" italics="true" pageId="67" pageNumber="68">C. boschiana</emphasis>
</taxonomicName>
(see notes under that species).
</paragraph>
</subSubSection>
</treatment>
</document>