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<document id="3F546681DB11758BD62BF06BB9A01359" ID-DOI="10.3853/j.2201-4349.72.2020.1765" ID-ISSN="2201-4349" ID-Zenodo-Dep="4654453" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617222158928" checkinUser="felipe" docAuthor="Popple, Lindsay W. &amp; Emery, David L." docDate="2020" docId="03B36A2341050A71FC6EFA1E651EFC4E" docLanguage="en" docName="RecAustMus.72.4.123-147.pdf" docOrigin="Records of the Australian Museum 72 (4)" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.3:RecAustMus.2019-.journal_article" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article" docStyleVersion="3" docTitle="Yoyetta loftyensis Popple &amp; Emery 2020, sp. nov." docType="treatment" docUuid="DFC5114D-A217-4281-9BD1-76A43C10D063" docUuidSource="ZooBank" docVersion="4" lastPageNumber="141" masterDocId="FF8A125B410D0A63FFA8FFCE6604FFF4" masterDocTitle="Four New Species of Cicadas in the Yoyetta abdominalis (Distant) Species Group (Hemiptera: Cicadidae: Cicadettinae) from Southeastern Australia" masterLastPageNumber="147" masterPageNumber="123" pageNumber="131" updateTime="1698898586388" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="012EC172255C50A2F4CFE1482217C337">Four New Species of Cicadas in the Yoyetta abdominalis (Distant) Species Group (Hemiptera: Cicadidae: Cicadettinae) from Southeastern Australia</mods:title>
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<mods:namePart id="31EFFB80C135EE153B7AC8789A317947">Popple, Lindsay W.</mods:namePart>
<mods:affiliation id="80DBB193A696CE75B0D63EDA0DFFC7AE">Biodiversity &amp; Geosciences, Queensland Museum, South Brisbane QLD 4101, Australia</mods:affiliation>
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<mods:namePart id="75B81D2657812AB5CB1AAB642CFA7B76">Emery, David L.</mods:namePart>
<mods:affiliation id="68D10F1E3BAE2067CD826FB60A16A1EB">Sydney School of Veterinary Science (SSVS), University of Sydney NSW 2006, Australia</mods:affiliation>
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<mods:title id="E441E4B99360038747AFEFB63FC06F51">Records of the Australian Museum</mods:title>
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<mods:date id="0D8389304841ABB82D396F7D77FADF72">2020</mods:date>
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<heading id="D0ED6C5941050A6BFC6EFA1E6338FA1B" bold="true" box="[966,1340,1487,1519]" centered="true" fontSize="12" level="2" pageId="8" pageNumber="131" reason="2">
<emphasis id="B96E072741050A6BFC6EFA1E6338FA1B" bold="true" box="[966,1340,1487,1519]" pageId="8" pageNumber="131">
<taxonomicName id="4C1AA0B641050A6BFC6EFA1E62C5FA1A" ID-CoL="8KLSL" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[966,1217,1487,1518]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="species" species="loftyensis" status="sp. nov.">
<emphasis id="B96E072741050A6BFC6EFA1E62C5FA1A" bold="true" box="[966,1217,1487,1518]" italics="true" pageId="8" pageNumber="131">Yoyetta loftyensis</emphasis>
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<subSubSection id="C30088BE41050A6BFC4CF9FE631AF9BF" box="[996,1310,1584,1611]" pageId="8" pageNumber="131" type="description">
<paragraph id="8BA5DB3541050A6BFC4CF9FE631AF9BF" blockId="8.[996,1310,1584,1611]" box="[996,1310,1584,1611]" pageId="8" pageNumber="131">
<figureCitation id="1321C7B041050A6BFC4CF9FE622AF9BF" box="[996,1070,1584,1611]" captionStart="Figure 1" captionStartId="2.[166,234,1164,1186]" captionTargetBox="[166,1463,151,1154]" captionTargetId="figure-483@2.[166,1466,151,1154]" captionTargetPageId="2" captionText="Figure 1. Map of south-eastern mainland Australia showing the geographical distribution of Yoyetta douglasi sp. nov. (grey circles), Y. enigmatica sp. nov. (grey triangles), Y. loftyensis sp. nov. (black triangles) and Y. ngarabal sp. nov. (black circles)." figureDoi="http://doi.org/10.5281/zenodo.4654455" httpUri="https://zenodo.org/record/4654455/files/figure.png" pageId="8" pageNumber="131">Figs 1</figureCitation>
,
<figureCitation id="1321C7B041050A6BFB95F9FF6266F9BF" box="[1085,1122,1585,1611]" captionStart="Figure 4" captionStartId="6.[166,234,1032,1054]" captionTargetBox="[167,1467,151,1018]" captionTargetId="figure-9@6.[167,1467,151,1018]" captionTargetPageId="6" captionText="Figure 4. Photos of male left opercula: (A) Yoyetta douglasi sp. nov.; (B) Y.enigmatica sp. nov.; (C) Y. loftyensis sp. nov.; (D) Y.ngarabal sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4654461" httpUri="https://zenodo.org/record/4654461/files/figure.png" pageId="8" pageNumber="131">4C</figureCitation>
,
<figureCitation id="1321C7B041050A6BFBC7F9FE6290F9BF" box="[1135,1172,1584,1611]" captionStart="Figure 5" captionStartId="7.[121,189,1038,1060]" captionTargetBox="[121,1421,151,1018]" captionTargetId="figure-10@7.[121,1421,151,1018]" captionTargetPageId="7" captionText="Figure 5. Photos of male left timbals, with dorsal edge at right and posterior margin at bottom: (A) Yoyetta douglasi sp. nov.; (B) Y. enigmatica sp. nov.; (C) Y. loftyensis sp. nov.; (D) Y. ngarabal sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4654463" httpUri="https://zenodo.org/record/4654463/files/figure.png" pageId="8" pageNumber="131">5C</figureCitation>
,
<figureCitation id="1321C7B041050A6BFB09F9FE62C3F9BE" box="[1185,1223,1584,1610]" captionStart="Figure 6" captionStartId="8.[166,234,1069,1091]" captionTargetBox="[167,1467,151,1050]" captionTargetId="figure-9@8.[167,1467,151,1050]" captionTargetPageId="8" captionText="Figure 6. Lateral views of the caudal abdomen of: (A) Yoyetta douglasi sp. nov.; (B) Y. enigmatica sp. nov.; (C) Y. loftyensis sp. nov.; (D) Y. ngarabal sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4654467" httpUri="https://zenodo.org/record/4654467/files/figure.png" pageId="8" pageNumber="131">6C</figureCitation>
,
<figureCitation id="1321C7B041050A6BFB7BF9FE631AF9BF" box="[1235,1310,1584,1611]" captionStart-0="Figure 12" captionStart-1="Figure 13" captionStart-2="Figure 14" captionStart-3="Figure 15" captionStartId-0="14.[166,234,1251,1273]" captionStartId-1="15.[121,189,1845,1867]" captionStartId-2="16.[167,235,1185,1207]" captionStartId-3="17.[121,189,1922,1944]" captionTargetBox-0="[166,1466,158,1236]" captionTargetBox-1="[122,1388,131,1838]" captionTargetBox-2="[167,1467,162,1163]" captionTargetBox-3="[134,1362,140,1901]" captionTargetId-0="figure-10@14.[166,1466,151,1236]" captionTargetId-1="figure-10@15.[121,1388,127,1838]" captionTargetId-2="figure-9@16.[167,1467,151,1163]" captionTargetId-3="figure-10@17.[121,1375,133,1910]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionText-0="Figure 12. Yoyetta loftyensis sp. nov.: (A) holotype male, 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E), dorsal habitus; (B) holotype male, ventral habitus; (C) female, 2.3 km S. of Birdwood (34°50'S 138°58'E) dorsal habitus; (D) female, 2.3 km S. of Birdwood, ventral habitus; (E) typical habitat, Adelaide; (F) typical habitat, Adelaide. Scale bars = 10 mm." captionText-1="Figure 13. Male calling song structure of Yoyetta loftyensis sp. nov. illustrated in waveform plots, including: (A) a single cueing phrase commencing with an extended dense syllable sequence/long echeme, followed by three normal phrases; (B) a series of four normal phrases, followed by a cueing phrase, then two sets of alternating normal and cueing phrases; (C) expanded section from (B) showing two normal phrases (demarcated by solid black bar above) and one cueing phrase (demarcated by dashed line above). The final subfigure (D) is a spectrogram displaying song frequency. This specimen was recorded in the field at 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E) by D. Marshall and K. Hill." captionText-2="Figure 14. Male calling song structure of Yoyetta loftyensis sp. nov. A comparison of phrase structures recorded from multiple sites, including: (A) Mt Lofty (34°59'S 138°43'E); (B) Hale Conservation Park (34°41'S 138°55'E); (C) 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E); and (D) 2.3 km S. of Birdwood (34°50'S 138°58'E). All examples are cueing phrases, apart from the one in (B) with a black scale bar above, which is a normal phrase. Recordings (A) and (B) were made in the field by LWP. Recordings (C) and (D) were made by D. Marshall and K. Hill." captionText-3="Figure 15. Yoyetta loftyensis sp. nov.: illustration of male genitalia; (a) pygofer, viewed laterally from the left; (b) pygofer, viewed ventrally; (c) aedeagus, viewed laterally from the left; (d) apex of theca. Characters depicted as in Fig. 7. Specimen from 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E)." figureDoi-0="http://doi.org/10.5281/zenodo.4654481" figureDoi-1="http://doi.org/10.5281/zenodo.4654483" figureDoi-2="http://doi.org/10.5281/zenodo.4654485" figureDoi-3="http://doi.org/10.5281/zenodo.4654487" httpUri-0="https://zenodo.org/record/4654481/files/figure.png" httpUri-1="https://zenodo.org/record/4654483/files/figure.png" httpUri-2="https://zenodo.org/record/4654485/files/figure.png" httpUri-3="https://zenodo.org/record/4654487/files/figure.png" pageId="8" pageNumber="131">12–15</figureCitation>
</paragraph>
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<materialsCitation id="3B72D16841050A6BFCEFF9AE6230F924" collectingDate="2006-11-30" collectionCode="SA, SAMA, SAM" collectorName="Marshall &amp; Hill &amp; Database No." elevation="229" latitude="-35.428883" location="Australia" longLatPrecision="1" longitude="138.5803" municipality="Victor Harbor Rd on Myponga Rd" pageId="8" pageNumber="131" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54A1659741050A6BFCEFF9AE65AAF98C" box="[839,942,1632,1656]" pageId="8" pageNumber="131" type="holotype">Holotype</typeStatus>
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, AU.
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.VHN,
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NW. of
<collectingMunicipality id="6BC1414F41050A6BFCEFF9B362CBF961" box="[839,1231,1661,1685]" pageId="8" pageNumber="131">Victor Harbor Rd on Myponga Rd</collectingMunicipality>
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&amp;
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<collectionCode id="ED0B43F041050A6BFAA3F9546352F946" box="[1291,1366,1690,1714]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="8" pageNumber="131" type="Museum">SAMA</collectionCode>
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20-017507 (
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.
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—
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, AU.
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<materialsCitation id="3B72D16841050A6BFB3FF91B6212F8B1" collectingDate="2006-11-29" collectionCode="SA, BIR" collectorName="S. Birdwood &amp; D. Marshall &amp; Hill &amp; Simon" elevation="391" latitude="-34.840984" location="Torrens" longLatPrecision="1" longitude="138.96146" pageId="8" pageNumber="131" specimenCount="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
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<collectorName id="26EFBEE341050A6BFB4CF91B636CF919" box="[1252,1384,1749,1773]" pageId="8" pageNumber="131">S. Birdwood</collectorName>
on road to
<location id="8EC58DEE41050A6BFCCCF93D65B3F8FE" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41050A6BFCCCF93D65B3F8FE" box="[868,951,1779,1802]" latitude="-34.840984" longLatPrecision="1" longitude="138.96146" name="Torrens" pageId="8" pageNumber="131" stateProvince="SOUTH AUSTRALIA">Torrens</location>
,
<quantity id="4CE276D041050A6BFC6CF93C6208F8FE" box="[964,1036,1778,1802]" metricMagnitude="2" metricUnit="m" metricValue="3.91" pageId="8" pageNumber="131" unit="m" value="391.0">
<elevation id="00373C0641050A6BFC6CF93C6208F8FE" box="[964,1036,1778,1802]" metricMagnitude="2" metricUnit="m" metricValue="3.91" pageId="8" pageNumber="131" unit="m" value="391.0">391 m</elevation>
</quantity>
,
<geoCoordinate id="EE2EBDF241050A6BFBB1F93C629EF8FE" box="[1049,1178,1778,1802]" degrees="34" direction="south" minutes="50.459" orientation="latitude" pageId="8" pageNumber="131" precision="1" value="-34.840984">34°50.459'S</geoCoordinate>
<geoCoordinate id="EE2EBDF241050A6BFB0BF93C6331F8FE" box="[1187,1333,1778,1802]" degrees="138" direction="east" minutes="57.687" orientation="longitude" pageId="8" pageNumber="131" precision="1" value="138.96146">138°57.687'E</geoCoordinate>
,
<date id="FFA4FDF541050A6BFAEBF93C63B3F8FE" box="[1347,1463,1778,1802]" pageId="8" pageNumber="131" value="2006-11-29">
<collectingDate id="EFE0041D41050A6BFAEBF93C63B3F8FE" box="[1347,1463,1778,1802]" pageId="8" pageNumber="131" value="2006-11-29">29.xi.2006</collectingDate>
</date>
,
<collectorName id="26EFBEE341050A6BFCEFF8DE65D0F8DC" box="[839,980,1808,1832]" pageId="8" pageNumber="131">D. Marshall</collectorName>
, K.
<collectorName id="26EFBEE341050A6BFBA0F8DE6230F8DC" box="[1032,1076,1808,1832]" pageId="8" pageNumber="131">Hill</collectorName>
(C.
<collectorName id="26EFBEE341050A6BFBC4F8DE62B1F8DC" box="[1132,1205,1808,1832]" pageId="8" pageNumber="131">Simon</collectorName>
lab voucher specimen 06.AU.
<collectionCode id="ED0B43F041050A6BFC3EF8E365BFF8B1" box="[918,955,1837,1861]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="8" pageNumber="131">SA</collectionCode>
.
<collectionCode id="ED0B43F041050A6BFC17F8E365F4F8B1" box="[959,1008,1837,1861]" pageId="8" pageNumber="131">BIR</collectionCode>
.01)
</materialsCitation>
;
<materialsCitation id="3B72D16841050A6BFB88F8E3659EF896" collectingDate="2006-11-29" collectionCode="MSM" collectorName="S. Birdwood &amp; D. Marshall &amp; Hill &amp; Simon" elevation="391" latitude="-34.840984" location="Torrens" longLatPrecision="1" longitude="138.96146" pageId="8" pageNumber="131" specimenCount="1" specimenCount-female="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41050A6BFB88F8E36246F8B1" box="[1056,1090,1837,1861]" pageId="8" pageNumber="131" type="female">1♀</specimenCount>
, same location and date as previous (
<collectionCode id="ED0B43F041050A6BFCE7F8846596F896" box="[847,914,1866,1890]" country="Japan" httpUri="http://grbio.org/cool/z37t-cwh9" name="Marine Science Museum, Tokai Univ." pageId="8" pageNumber="131">MSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72D16841050A6BFC0FF88462C9F896" box="[935,1229,1866,1890]" collectingDate="2006-11-30" collectionCode="SA, SAMA, SAM" collectorName="Marshall &amp; Hill &amp; Database No." elevation="229" latitude="-35.428883" location="Australia" longLatPrecision="1" longitude="138.5803" municipality="Victor Harbor Rd on Myponga Rd" pageId="8" pageNumber="131" specimenCount="1" specimenCount-female="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41050A6BFC0FF88465CCF896" box="[935,968,1866,1890]" pageId="8" pageNumber="131" type="female">1♀</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="3B72D16841050A6BFB72F884631AF84E" collectingDate="2011-01-08" collectionCode="SA, ADE" collectorName="Marshall &amp; Hill &amp; Simon" elevation="351" latitude="-35.004066" location="Balhannah on Jones Rd." longLatPrecision="1" longitude="138.825" pageId="8" pageNumber="131" specimenCount="1" specimenCount-male="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41050A6BFB72F88462F9F896" box="[1242,1277,1866,1890]" pageId="8" pageNumber="131" type="male">1♂</specimenCount>
, S. of
<location id="8EC58DEE41050A6BFAE1F88465D2F874" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41050A6BFAE1F88465D2F874" latitude="-35.004066" longLatPrecision="1" longitude="138.825" name="Balhannah on Jones Rd." pageId="8" pageNumber="131" stateProvince="SOUTH AUSTRALIA">Balhannah on Jones Rd.</location>
,
<locationDeviation id="9A17C21741050A6BFC4AF8A662DCF874" box="[994,1240,1896,1920]" location="Junction Rd." pageId="8" pageNumber="131">
<quantity id="4CE276D041050A6BFC4AF8A6622AF874" box="[994,1070,1896,1920]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="131" unit="km" value="1.2">1.2 km</quantity>
S Junction Rd.
</locationDeviation>
,
<quantity id="4CE276D041050A6BFB4DF8A96328F88B" box="[1253,1324,1895,1919]" metricMagnitude="2" metricUnit="m" metricValue="3.51" pageId="8" pageNumber="131" unit="m" value="351.0">
<elevation id="00373C0641050A6BFB4DF8A96328F88B" box="[1253,1324,1895,1919]" metricMagnitude="2" metricUnit="m" metricValue="3.51" pageId="8" pageNumber="131" unit="m" value="351.0">351 m</elevation>
</quantity>
,
<geoCoordinate id="EE2EBDF241050A6BFA91F8A963BEF88B" box="[1337,1466,1895,1919]" degrees="35" direction="south" minutes="00.244" orientation="latitude" pageId="8" pageNumber="131" precision="1" value="-35.004066">35°00.244'S</geoCoordinate>
<geoCoordinate id="EE2EBDF241050A6BFCEFF84B65E4F869" box="[839,992,1925,1949]" degrees="138" direction="east" minutes="49.500" orientation="longitude" pageId="8" pageNumber="131" precision="1" value="138.825">138°49.500'E</geoCoordinate>
,
<date id="FFA4FDF541050A6BFC46F84B624EF869" box="[1006,1098,1925,1949]" pageId="8" pageNumber="131" value="2011-01-08">
<collectingDate id="EFE0041D41050A6BFC46F84B624EF869" box="[1006,1098,1925,1949]" pageId="8" pageNumber="131" value="2011-01-08">8.i.2011</collectingDate>
</date>
,
<collectorName id="26EFBEE341050A6BFBF1F84B62BFF869" box="[1113,1211,1925,1949]" pageId="8" pageNumber="131">Marshall</collectorName>
&amp;
<collectorName id="26EFBEE341050A6BFB4DF84B6314F869" box="[1253,1296,1925,1949]" pageId="8" pageNumber="131">Hill</collectorName>
(C.
<collectorName id="26EFBEE341050A6BFAEFF84B638BF869" box="[1351,1423,1925,1949]" pageId="8" pageNumber="131">Simon</collectorName>
lab voucher specimen 11.AU.
<collectionCode id="ED0B43F041050A6BFBFDF86C627EF84E" box="[1109,1146,1954,1978]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="8" pageNumber="131">SA</collectionCode>
.
<collectionCode id="ED0B43F041050A6BFBD5F86C62BFF84E" box="[1149,1211,1954,1978]" pageId="8" pageNumber="131">ADE</collectionCode>
.01) (DE)
</materialsCitation>
;
<materialsCitation id="3B72D16841050A69FA80F86C66F4F96A" collectionCode="SAMA" collectorName="G. Rayson &amp; Belair &amp; National Parks &amp; Wildlife Service" lastPageId="10" lastPageNumber="133" location="Stack Jarra Stakes" pageId="8" pageNumber="131" specimenCount="1" specimenCount-male="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41050A6BFA80F86C634FF84E" box="[1320,1355,1954,1978]" pageId="8" pageNumber="131" type="male">1♂</specimenCount>
, Collected by
<collectorName id="26EFBEE341050A6BFCC3F80E65DBF82C" box="[875,991,1984,2008]" pageId="8" pageNumber="131">G. Rayson</collectorName>
, location
<location id="8EC58DEE41050A6BFBE2F80E630BF82C" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41050A6BFBE2F80E630BF82C" box="[1098,1295,1984,2008]" name="Stack Jarra Stakes" pageId="8" pageNumber="131" stateProvince="SOUTH AUSTRALIA">Stack Jarra Stakes</location>
f[ro]m W.A. to
<collectorName id="26EFBEE341070A69FF0EF9A666E3F974" box="[166,231,1640,1664]" pageId="10" pageNumber="133">Belair</collectorName>
RR, 23-11-77,
<collectorName id="26EFBEE341070A69FE30F9A6643DF974" box="[408,569,1640,1664]" pageId="10" pageNumber="133">National Parks</collectorName>
&amp;
<collectorName id="26EFBEE341070A69FDC9F9A66512F974" box="[609,790,1640,1664]" pageId="10" pageNumber="133">Wildlife Service</collectorName>
,
<collectionCode id="ED0B43F041070A69FF0EF94866F4F96A" box="[166,240,1670,1694]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="10" pageNumber="133" type="Museum">SAMA</collectionCode>
</materialsCitation>
<materialsCitation id="3B72D16841070A69FF5DF94867F2F969" box="[245,502,1669,1694]" location="Database No." pageId="10" pageNumber="133" specimenCount="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<location id="8EC58DEE41070A69FF5DF948677BF969" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FF5DF948677BF969" box="[245,383,1670,1694]" name="Database No." pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Database No.</location>
20-014465
</materialsCitation>
;
<materialsCitation id="3B72D16841070A69FE56F94867DBF94F" collectionCode="SAMA" collectorName="Mt. Barker S. A. &amp; Dec. &amp; N. McFarland" pageId="10" pageNumber="133" specimenCount="1" specimenCount-male="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41070A69FE56F9486424F96A" box="[510,544,1670,1694]" pageId="10" pageNumber="133" type="male">1♂</specimenCount>
,
<collectorName id="26EFBEE341070A69FD82F94864C9F96A" box="[554,717,1670,1694]" pageId="10" pageNumber="133">Mt. Barker S.A.</collectorName>
, 4.
<collectorName id="26EFBEE341070A69FD4FF948651EF96A" box="[743,794,1670,1694]" pageId="10" pageNumber="133">Dec.</collectorName>
[19]64,
<collectorName id="26EFBEE341070A69FF50F96D6788F94F" box="[248,396,1699,1723]" pageId="10" pageNumber="133">N. McFarland</collectorName>
,
<collectionCode id="ED0B43F041070A69FE3CF96D67DBF94F" box="[404,479,1699,1723]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="10" pageNumber="133" type="Museum">SAMA</collectionCode>
</materialsCitation>
<materialsCitation id="3B72D16841070A69FE4CF96D64EDF94F" box="[484,745,1699,1723]" location="Database No." pageId="10" pageNumber="133" specimenCount="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<location id="8EC58DEE41070A69FE4CF96D6474F94E" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FE4CF96D6474F94E" box="[484,624,1699,1723]" name="Database No." pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Database No.</location>
20-014439
</materialsCitation>
;
<materialsCitation id="3B72D16841070A69FD5AF96D64B5F92C" collectionCode="SAMA" collectorName="S. Aust &amp; Jny. &amp; A. White" location="Mt. Lofty" pageId="10" pageNumber="133" specimenCount="1" specimenCount-male="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41070A69FD5AF96D6511F94F" box="[754,789,1699,1723]" pageId="10" pageNumber="133" type="male">1♂</specimenCount>
,
<collectorName id="26EFBEE341070A69FF0EF90E66F3F92D" box="[166,247,1728,1753]" pageId="10" pageNumber="133">S. Aust</collectorName>
,
<location id="8EC58DEE41070A69FEABF90F6768F92C" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FEABF90F6768F92C" box="[259,364,1728,1752]" name="Mt. Lofty" pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Mt. Lofty</location>
,
<collectorName id="26EFBEE341070A69FED0F90E67A2F92C" box="[376,422,1728,1752]" pageId="10" pageNumber="133">Jny.</collectorName>
[18]85,
<collectorName id="26EFBEE341070A69FE5DF90F645DF92C" box="[501,601,1728,1753]" pageId="10" pageNumber="133">A. White</collectorName>
,
<collectionCode id="ED0B43F041070A69FDCEF90E64B5F92C" box="[614,689,1728,1752]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="10" pageNumber="133" type="Museum">SAMA</collectionCode>
</materialsCitation>
<materialsCitation id="3B72D16841070A69FD11F90E6742F902" location="Database No." pageId="10" pageNumber="133" specimenCount="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<location id="8EC58DEE41070A69FD11F90E66CAF901" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FD11F90E66CAF901" name="Database No." pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Database No.</location>
20-014440
</materialsCitation>
;
<materialsCitation id="3B72D16841070A69FEE7F9106744F8E7" collectionCode="SAMA" collectorName="Male &amp; W White" location="Reed Beds" municipality="Sth Aus" pageId="10" pageNumber="133" specimenCount="1" specimenCount-male="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41070A69FEE7F9106775F902" box="[335,369,1758,1782]" pageId="10" pageNumber="133" type="male">1♂</specimenCount>
,
<collectingMunicipality id="6BC1414F41070A69FED3F91067CBF902" box="[379,463,1758,1782]" pageId="10" pageNumber="133">Sth Aus</collectingMunicipality>
,
<location id="8EC58DEE41070A69FE7FF910644DF902" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FE7FF910644DF902" box="[471,585,1758,1782]" municipality="Sth Aus" name="Reed Beds" pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Reed Beds</location>
,
<collectorName id="26EFBEE341070A69FDFAF910648EF902" box="[594,650,1758,1782]" pageId="10" pageNumber="133">Male</collectorName>
, 9-Dec-70,
<collectorName id="26EFBEE341070A69FCA9F91066EDF8E7" pageId="10" pageNumber="133">W White</collectorName>
,
<collectionCode id="ED0B43F041070A69FF5DF9356744F8E7" box="[245,320,1787,1811]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="10" pageNumber="133" type="Museum">SAMA</collectionCode>
</materialsCitation>
<materialsCitation id="3B72D16841070A69FEEFF9356450F8E7" box="[327,596,1787,1811]" location="Database No." pageId="10" pageNumber="133" specimenCount="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<location id="8EC58DEE41070A69FEEFF93567D3F8E6" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FEEFF93567D3F8E6" box="[327,471,1787,1811]" name="Database No." pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Database No.</location>
20-017497
</materialsCitation>
;
<materialsCitation id="3B72D16841070A69FDC8F93567D8F8C4" collectingDate="1889-12" collectorName="J. W. Mellor" location="Morphett Vale" pageId="10" pageNumber="133" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41070A69FDC8F9356485F8E7" box="[608,641,1787,1811]" pageId="10" pageNumber="133" type="male">1♂</specimenCount>
<specimenCount id="9D1C10BC41070A69FD20F93564ADF8E7" box="[648,681,1787,1811]" pageId="10" pageNumber="133" type="female">1♀</specimenCount>
,
<location id="8EC58DEE41070A69FD1FF93566DCF8C4" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FD1FF93566DCF8C4" name="Morphett Vale" pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Morphett Vale</location>
,
<date id="FFA4FDF541070A69FF49F8D7674BF8C4" box="[225,335,1816,1840]" pageId="10" pageNumber="133" value="1889-12">
<collectingDate id="EFE0041D41070A69FF49F8D7674BF8C4" box="[225,335,1816,1840]" pageId="10" pageNumber="133" value="1889-12">Dec. 1889</collectingDate>
</date>
,
<collectorName id="26EFBEE341070A69FEF1F8D767D8F8C4" box="[345,476,1816,1840]" pageId="10" pageNumber="133">J. W. Mellor</collectorName>
</materialsCitation>
;
<materialsCitation id="3B72D16841070A69FE4DF8D66777F8BA" collectingDate="1958-11-22" collectionCode="SAMA" collectorName="Belair &amp; S. A., S. Dawson" pageId="10" pageNumber="133" specimenCount="1" specimenCount-male="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41070A69FE4DF8D6640CF8C4" box="[485,520,1816,1840]" pageId="10" pageNumber="133" type="male">1♂</specimenCount>
,
<date id="FFA4FDF541070A69FDBAF8D6648BF8C4" box="[530,655,1816,1840]" pageId="10" pageNumber="133" value="1958-11-22">
<collectingDate id="EFE0041D41070A69FDBAF8D6648BF8C4" box="[530,655,1816,1840]" pageId="10" pageNumber="133" value="1958-11-22">22-11-1958</collectingDate>
</date>
,
<collectorName id="26EFBEE341070A69FD31F8D664DFF8C4" box="[665,731,1816,1840]" pageId="10" pageNumber="133">Belair</collectorName>
,
<collectorName id="26EFBEE341070A69FD4CF8D6671AF8B9" pageId="10" pageNumber="133">S.A., S. Dawson</collectorName>
<collectionCode id="ED0B43F041070A69FE8FF8F86777F8BA" box="[295,371,1846,1870]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="10" pageNumber="133" type="Museum">SAMA</collectionCode>
</materialsCitation>
<materialsCitation id="3B72D16841070A69FED5F8F86512F8BA" box="[381,790,1845,1870]" location="Database No." pageId="10" pageNumber="133" specimenCount="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<location id="8EC58DEE41070A69FED5F8F86415F8B9" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FED5F8F86415F8B9" box="[381,529,1846,1870]" name="Database No." pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Database No.</location>
20-017501, 20-017499
</materialsCitation>
;
<materialsCitation id="3B72D16841070A69FF0EF89D64B1F89F" box="[166,693,1875,1899]" collectionCode="SAMA" collectorName="Rev. Burgess" location="Wilmington" pageId="10" pageNumber="133" specimenCount="2" specimenCount-female="2" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41070A69FF0EF89D66D8F89F" box="[166,220,1875,1899]" pageId="10" pageNumber="133" type="female">2♀♀</specimenCount>
,
<location id="8EC58DEE41070A69FF4EF89D6763F89F" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FF4EF89D6763F89F" box="[230,359,1875,1899]" name="Wilmington" pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Wilmington</location>
,
<collectorName id="26EFBEE341070A69FEC7F89D67FEF89F" box="[367,506,1875,1899]" pageId="10" pageNumber="133">Rev. Burgess</collectorName>
, 16.12.86,
<collectionCode id="ED0B43F041070A69FDC2F89D64B1F89F" box="[618,693,1875,1899]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="10" pageNumber="133" type="Museum">SAMA</collectionCode>
</materialsCitation>
<materialsCitation id="3B72D16841070A69FD11F89D67D4F87C" location="Database No." pageId="10" pageNumber="133" specimenCount="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<location id="8EC58DEE41070A69FD11F89D66CAF87C" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FD11F89D66CAF87C" name="Database No." pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Database No.</location>
20-014442, 20-014443
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;
<materialsCitation id="3B72D16841070A69FE75F8BE6770F852" collectionCode="SAMA" collectorName="S. Australia &amp; Rev. &amp; Burgess" pageId="10" pageNumber="133" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41070A69FE75F8BE67FBF87C" box="[477,511,1904,1928]" pageId="10" pageNumber="133" type="male">1♂</specimenCount>
<specimenCount id="9D1C10BC41070A69FDA0F8BE642DF87C" box="[520,553,1904,1928]" pageId="10" pageNumber="133" type="female">1♀</specimenCount>
,
<collectorName id="26EFBEE341070A69FD92F8BE64BFF87C" box="[570,699,1904,1928]" pageId="10" pageNumber="133">S. Australia</collectorName>
,
<collectorName id="26EFBEE341070A69FD60F8BF64FCF87C" box="[712,760,1905,1928]" pageId="10" pageNumber="133">Rev.</collectorName>
A. P.
<collectorName id="26EFBEE341070A69FF6BF840671FF852" box="[195,283,1934,1958]" pageId="10" pageNumber="133">Burgess</collectorName>
,
<collectionCode id="ED0B43F041070A69FE81F8406770F852" box="[297,372,1934,1958]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="10" pageNumber="133" type="Museum">SAMA</collectionCode>
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<materialsCitation id="3B72D16841070A69FED6F8406511F851" box="[382,789,1933,1958]" location="Database No." pageId="10" pageNumber="133" specimenCount="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<location id="8EC58DEE41070A69FED6F8406415F851" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FED6F8406415F851" box="[382,529,1934,1958]" name="Database No." pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Database No.</location>
20-014444, 20-014445
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;
<materialsCitation id="3B72D16841070A69FF0EF8656424F837" box="[166,544,1963,1987]" collectorName="S. H. Curnow" location="Mt Lofty Rge" pageId="10" pageNumber="133" specimenCount="2" specimenCount-female="2" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41070A69FF0EF86566D9F837" box="[166,221,1963,1987]" pageId="10" pageNumber="133" type="female">2♀♀</specimenCount>
,
<location id="8EC58DEE41070A69FF41F865677AF837" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FF41F865677AF837" box="[233,382,1963,1987]" name="Mt Lofty Rge" pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Mt Lofty Rge</location>
,
<collectorName id="26EFBEE341070A69FE21F8656424F837" box="[393,544,1963,1987]" pageId="10" pageNumber="133">S. H. Curnow</collectorName>
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;
<materialsCitation id="3B72D16841070A69FD82F8656215F974" box="[554,1041,1640,1987]" collectingDate="1891-12" collectionCode="SAMA" collectorName="S. Aust" location="Mt Lofty" pageId="10" pageNumber="133" specimenCount="1" specimenCount-female="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<specimenCount id="9D1C10BC41070A69FD82F8656448F837" box="[554,588,1963,1987]" pageId="10" pageNumber="133" type="female">1♀</specimenCount>
,
<collectorName id="26EFBEE341070A69FDF1F86564ADF837" box="[601,681,1963,1987]" pageId="10" pageNumber="133">S. Aust</collectorName>
,
<location id="8EC58DEE41070A69FD1CF8656512F837" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FD1CF8656512F837" box="[692,790,1963,1987]" name="Mt Lofty" pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Mt Lofty</location>
,
<date id="FFA4FDF541070A69FCEFF9A665B2F974" box="[839,950,1640,1664]" pageId="10" pageNumber="133" value="1891-12">
<collectingDate id="EFE0041D41070A69FCEFF9A665B2F974" box="[839,950,1640,1664]" pageId="10" pageNumber="133" value="1891-12">Dec-1891</collectingDate>
</date>
,
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</materialsCitation>
<materialsCitation id="3B72D16841070A69FBB3F9A66592F96A" collectionCode="SAM" location="Database No." pageId="10" pageNumber="133" specimenCount="1" stateProvince="SOUTH AUSTRALIA" typeStatus="paratype">
<location id="8EC58DEE41070A69FBB3F9A662B7F974" LSID="urn:lsid:plazi:treatment:03B36A2341050A71FC6EFA1E651EFC4E:8EC58DEE41070A69FBB3F9A662B7F974" box="[1051,1203,1640,1664]" name="Database No." pageId="10" pageNumber="133" stateProvince="SOUTH AUSTRALIA">Database No.</location>
20-017500, 20-017502 (
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF658BBD41040A6AFFD1F8A864ADF839" ID-DOI="http://doi.org/10.5281/zenodo.4654469" ID-Zenodo-Dep="4654469" httpUri="https://zenodo.org/record/4654469/files/figure.png" pageId="9" pageNumber="132" startId="9.[121,188,1894,1916]" targetBox="[131,1396,145,1868]" targetPageId="9">
<paragraph id="8BA5DB3541040A6AFFD1F8A864ADF839" blockId="9.[121,1422,1894,1997]" pageId="9" pageNumber="132">
<emphasis id="B96E072741040A6AFFD1F8A866C9F888" bold="true" box="[121,205,1894,1916]" pageId="9" pageNumber="132">Figure 7</emphasis>
.
<taxonomicName id="4C1AA0B641040A6AFF7FF8A96777F888" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[215,371,1895,1916]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="132" phylum="Arthropoda" rank="species" species="douglasi" status="sp. nov.">
<emphasis id="B96E072741040A6AFF7FF8A96777F888" box="[215,371,1895,1916]" italics="true" pageId="9" pageNumber="132">Yoyetta douglasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C41040A6AFED0F8A967C6F888" box="[376,450,1895,1916]" pageId="9" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
: illustration of male genitalia;
<emphasis id="B96E072741040A6AFD4BF8A964FBF888" box="[739,767,1895,1916]" italics="true" pageId="9" pageNumber="132">(a)</emphasis>
pygofer, viewed laterally from the left;
<emphasis id="B96E072741040A6AFBDBF8A96294F888" box="[1139,1168,1895,1916]" italics="true" pageId="9" pageNumber="132">(b)</emphasis>
pygofer, viewed ventrally;
<emphasis id="B96E072741040A6AFFD1F84C6690F863" box="[121,148,1922,1943]" italics="true" pageId="9" pageNumber="132">(c)</emphasis>
aedeagus, viewed laterally from the left;
<emphasis id="B96E072741040A6AFD98F84F6448F862" box="[560,588,1921,1942]" italics="true" pageId="9" pageNumber="132">(d)</emphasis>
apex of theca, viewed laterally from the left and
<emphasis id="B96E072741040A6AFB91F84C6250F863" box="[1081,1108,1922,1943]" italics="true" pageId="9" pageNumber="132">(e)</emphasis>
apex of theca, viewed dorsally. Characters include: aed, aedeagus; bpl, basal lobe of pygofer; cl, clasper; db, dorsal beak; ps, pseudoparameres; th, theca; upl, upper lobe of pygofer. Specimen from Halls Gap (37°14'S 142°32'E).
</paragraph>
</caption>
<caption id="DF658BBD41070A69FF0EFA036736F9ED" ID-DOI="http://doi.org/10.5281/zenodo.4654473" ID-Zenodo-Dep="4654473" httpUri="https://zenodo.org/record/4654473/files/figure.png" pageId="10" pageNumber="133" startId="10.[166,234,1485,1507]" targetBox="[166,1464,157,1472]" targetPageId="10">
<paragraph id="8BA5DB3541070A69FF0EFA036736F9ED" blockId="10.[166,1466,1485,1561]" pageId="10" pageNumber="133">
<emphasis id="B96E072741070A69FF0EFA0366FAFA17" bold="true" box="[166,254,1485,1507]" pageId="10" pageNumber="133">Figure 8</emphasis>
.
<taxonomicName id="4C1AA0B641070A69FEA4FA0067C1FA17" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[268,453,1486,1507]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="133" phylum="Arthropoda" rank="species" species="enigmatica" status="sp. nov.">
<emphasis id="B96E072741070A69FEA4FA0067C1FA17" box="[268,453,1486,1507]" italics="true" pageId="10" pageNumber="133">Yoyetta enigmatica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C41070A69FE65FA006419FA17" box="[461,541,1486,1507]" pageId="10" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="B96E072741070A69FD80FA006442FA17" box="[552,582,1486,1507]" italics="true" pageId="10" pageNumber="133">(A)</emphasis>
male, Lamington National Park (28°13'S 153°13'E), dorsal habitus;
<emphasis id="B96E072741070A69FB45FA00630FFA17" box="[1261,1291,1486,1507]" italics="true" pageId="10" pageNumber="133">(B)</emphasis>
male, Lamington National Park, ventral habitus;
<emphasis id="B96E072741070A69FE66FA2767EAFA0A" box="[462,494,1513,1534]" italics="true" pageId="10" pageNumber="133">(C)</emphasis>
typical habitat, Lamington National Park;
<emphasis id="B96E072741070A69FC2CFA2765A1FA0A" box="[900,933,1513,1534]" italics="true" pageId="10" pageNumber="133">(D)</emphasis>
typical habitat at Mt Kaputar (30°17'S 150°10'E). Scale bars = 10 mm.
</paragraph>
</caption>
<subSubSection id="C30088BE41070A69FCEFF97E63BFF912" pageId="10" pageNumber="133" type="etymology">
<paragraph id="8BA5DB3541070A69FCEFF97E63BFF912" blockId="10.[839,1467,1712,1766]" pageId="10" pageNumber="133">
<emphasis id="B96E072741070A69FCEFF97E65C6F93C" bold="true" box="[839,962,1712,1736]" pageId="10" pageNumber="133">Etymology</emphasis>
. Named after the Mt Lofty Ranges south-east of Adelaide,
<collectingRegion id="49DE15D741070A69FC19F9006257F912" box="[945,1107,1742,1766]" country="Australia" name="South Australia" pageId="10" pageNumber="133">South Australia</collectingRegion>
, where this species is most prolific.
</paragraph>
</subSubSection>
<subSubSection id="C30088BE41070A73FCEFF9356300F82C" lastPageId="16" lastPageNumber="139" pageId="10" pageNumber="133" type="distribution">
<paragraph id="8BA5DB3541070A68FCEFF9356490F8EA" blockId="10.[839,1467,1787,1987]" lastBlockId="11.[121,749,1710,1822]" lastPageId="11" lastPageNumber="134" pageId="10" pageNumber="133">
<emphasis id="B96E072741070A69FCEFF9356304F8E7" bold="true" box="[839,1280,1787,1811]" pageId="10" pageNumber="133">Distribution, habitat and seasonality</emphasis>
. This species is restricted to the Greater Adelaide region from near Victor Harbour north through the Mt Lofty Ranges to near Williamstown, with a potentially isolated population in the southern Flinders Ranges at Wilmington (
<figureCitation id="1321C7B041070A69FB56F8BE6338F87C" box="[1278,1340,1904,1928]" captionStart="Figure 1" captionStartId="2.[166,234,1164,1186]" captionTargetBox="[166,1463,151,1154]" captionTargetId="figure-483@2.[166,1466,151,1154]" captionTargetPageId="2" captionText="Figure 1. Map of south-eastern mainland Australia showing the geographical distribution of Yoyetta douglasi sp. nov. (grey circles), Y. enigmatica sp. nov. (grey triangles), Y. loftyensis sp. nov. (black triangles) and Y. ngarabal sp. nov. (black circles)." figureDoi="http://doi.org/10.5281/zenodo.4654455" httpUri="https://zenodo.org/record/4654455/files/figure.png" pageId="10" pageNumber="133">Fig. 1</figureCitation>
). It has also been heard calling in Adelaide Botanic Garden. Populations are found in open eucalypt forest and occasionally in parks with tall eucalypts (
<figureCitation id="1321C7B041060A68FEF1F96067DAF932" box="[345,478,1710,1734]" captionStart="Figure 12" captionStartId="14.[166,234,1251,1273]" captionTargetBox="[166,1466,158,1236]" captionTargetId="figure-10@14.[166,1466,151,1236]" captionTargetPageId="14" captionText="Figure 12. Yoyetta loftyensis sp. nov.: (A) holotype male, 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E), dorsal habitus; (B) holotype male, ventral habitus; (C) female, 2.3 km S. of Birdwood (34°50'S 138°58'E) dorsal habitus; (D) female, 2.3 km S. of Birdwood, ventral habitus; (E) typical habitat, Adelaide; (F) typical habitat, Adelaide. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4654481" httpUri="https://zenodo.org/record/4654481/files/figure.png" pageId="11" pageNumber="134">Fig. 12E, F</figureCitation>
). The adults occupy the upper and outer branches where they tend to remain fairly sedentary. Males move singing stations irregularly. Adults have been found during November and December.
</paragraph>
<caption id="DF658BBD41060A68FFD1FA326349F9B3" ID-DOI="http://doi.org/10.5281/zenodo.4654475" ID-Zenodo-Dep="4654475" httpUri="https://zenodo.org/record/4654475/files/figure.png" pageId="11" pageNumber="134" startId="11.[121,189,1532,1554]" targetBox="[122,1421,151,1515]" targetPageId="11">
<paragraph id="8BA5DB3541060A68FFD1FA326349F9B3" blockId="11.[121,1421,1532,1607]" pageId="11" pageNumber="134">
<emphasis id="B96E072741060A68FFD1FA3266D7F9E5" bold="true" box="[121,211,1532,1554]" pageId="11" pageNumber="134">Figure 9.</emphasis>
Male calling song structure of
<taxonomicName id="4C1AA0B641060A68FE5EFA3364AFF9E6" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[502,683,1533,1554]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="134" phylum="Arthropoda" rank="species" species="enigmatica" status="sp. nov.">
<emphasis id="B96E072741060A68FE5EFA3364AFF9E6" box="[502,683,1533,1554]" italics="true" pageId="11" pageNumber="134">Yoyetta enigmatica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C41060A68FD07FA3364FCF9E6" box="[687,760,1533,1554]" pageId="11" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
illustrated in waveform plots, including:
<emphasis id="B96E072741060A68FBD4FA32629EF9E5" box="[1148,1178,1532,1553]" italics="true" pageId="11" pageNumber="134">(A)</emphasis>
typical repeated echemes (in flight);
<emphasis id="B96E072741060A68FF4CF9D96706F9D8" box="[228,258,1559,1580]" italics="true" pageId="11" pageNumber="134">(B)</emphasis>
expanded section from
<emphasis id="B96E072741060A68FE47F9D96409F9D8" box="[495,525,1559,1580]" italics="true" pageId="11" pageNumber="134">(A)</emphasis>
showing the detailed structure of a single echeme. The final subfigure
<emphasis id="B96E072741060A68FB6DF9D962E1F9D8" box="[1221,1253,1559,1580]" italics="true" pageId="11" pageNumber="134">(C)</emphasis>
is a spectrogram displaying song frequency. This specimen was recorded in the field at Dawson Springs, Mt Kaputar (30°17'S 150°10'E) by LWP.
</paragraph>
</caption>
<paragraph id="8BA5DB3541060A6FFFD1F8F364A2F953" blockId="11.[121,749,1853,1995]" lastBlockId="12.[166,794,1415,1703]" lastPageId="12" lastPageNumber="135" pageId="11" pageNumber="134">
<emphasis id="B96E072741060A68FFD1F8F36703F8A2" bold="true" box="[121,263,1853,1878]" pageId="11" pageNumber="134">Calling song</emphasis>
. The calling song of
<taxonomicName id="4C1AA0B641060A68FE5DF8F06470F8A2" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[501,628,1854,1878]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="134" phylum="Arthropoda" rank="species" species="loftyensis" status="sp. nov.">
<emphasis id="B96E072741060A68FE5DF8F06470F8A2" box="[501,628,1854,1878]" italics="true" pageId="11" pageNumber="134">Y. loftyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C41060A68FDD6F8F064D6F8A1" box="[638,722,1854,1877]" pageId="11" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
is moderately high pitched, with a frequency plateau between approximately 10 and 15 kHz and dominant frequency of approximately 12 kHz. It contains repeated patterns of syllables (each
<geoCoordinate id="EE2EBDF241060A68FEB5F87D675CF83F" box="[285,344,1971,1995]" degrees="0.008" direction="north" orientation="latitude" pageId="11" pageNumber="134" precision="55" value="0.008">0.008</geoCoordinate>
<geoCoordinate id="EE2EBDF241060A68FEF0F87D67A5F83F" box="[344,417,1971,1995]" degrees="0.015" direction="west" orientation="longitude" pageId="11" pageNumber="134" precision="55" value="-0.015">–0.015</geoCoordinate>
s duration) that are grouped in phrases of two subtly different structures: a “normal phrase” and a “cueing phrase”. In both cases, each phrase commences with two (rarely one or three) preceding syllables, separated by a gap of 0.04–
<geoCoordinate id="EE2EBDF241060A68FC02F8C865E3F8E9" box="[938,999,1798,1822]" degrees="0.09" direction="south" orientation="latitude" pageId="11" pageNumber="134" precision="555" value="-0.09">0.09 s</geoCoordinate>
, followed by a shorter gap (0.01–
<geoCoordinate id="EE2EBDF241060A68FA97F8C8637AF8EA" box="[1343,1406,1798,1822]" degrees="0.07" direction="south" orientation="latitude" pageId="11" pageNumber="134" precision="555" value="-0.07">0.07 s</geoCoordinate>
), then an echeme (which could equivocally be interpreted as a dense syllable sequence; 0.26–
<geoCoordinate id="EE2EBDF241060A68FBF5F88E629EF8AC" box="[1117,1178,1856,1880]" degrees="0.71" direction="south" orientation="latitude" pageId="11" pageNumber="134" precision="555" value="-0.71">0.71 s</geoCoordinate>
duration) and then two (rarely three) subsequent syllables, separated by successively longer gaps (0.03–
<geoCoordinate id="EE2EBDF241060A68FC4DF8B5622DF866" box="[997,1065,1915,1939]" degrees="0.08" direction="south" orientation="latitude" pageId="11" pageNumber="134" precision="555" value="-0.08">0.08 s</geoCoordinate>
, increasing to 0.07–
<geoCoordinate id="EE2EBDF241060A68FAA1F8B56348F867" box="[1289,1356,1915,1939]" degrees="0.13" direction="south" orientation="latitude" pageId="11" pageNumber="134" precision="555" value="-0.13">0.13 s</geoCoordinate>
). The cueing phrase is characterized by the longer gap after the final syllable (0.09–
<geoCoordinate id="EE2EBDF241060A68FC45F878622AF83A" box="[1005,1070,1974,1998]" degrees="0.22" direction="south" orientation="latitude" pageId="11" pageNumber="134" precision="555" value="-0.22">0.22 s</geoCoordinate>
). In contrast, the normal phrases lack this longer gap, which makes the beginning and ending of each phrase less conspicuous (
<figureCitation id="1321C7B041010A6FFDB8FA6A6465FA49" box="[528,609,1444,1469]" captionStart="Figure 13" captionStartId="15.[121,189,1845,1867]" captionTargetBox="[122,1388,131,1838]" captionTargetId="figure-10@15.[121,1388,127,1838]" captionTargetPageId="15" captionText="Figure 13. Male calling song structure of Yoyetta loftyensis sp. nov. illustrated in waveform plots, including: (A) a single cueing phrase commencing with an extended dense syllable sequence/long echeme, followed by three normal phrases; (B) a series of four normal phrases, followed by a cueing phrase, then two sets of alternating normal and cueing phrases; (C) expanded section from (B) showing two normal phrases (demarcated by solid black bar above) and one cueing phrase (demarcated by dashed line above). The final subfigure (D) is a spectrogram displaying song frequency. This specimen was recorded in the field at 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E) by D. Marshall and K. Hill." figureDoi="http://doi.org/10.5281/zenodo.4654483" httpUri="https://zenodo.org/record/4654483/files/figure.png" pageId="12" pageNumber="135">Fig. 13</figureCitation>
). Occasionally, a one-off extended echeme (or dense syllable sequence; 3–19 s duration) is produced within the structure of a normal phrase or cueing phrase (
<figureCitation id="1321C7B041010A6FFECCFA3267C2F9E1" box="[356,454,1532,1557]" captionStart="Figure 13" captionStartId="15.[121,189,1845,1867]" captionTargetBox="[122,1388,131,1838]" captionTargetId="figure-10@15.[121,1388,127,1838]" captionTargetPageId="15" captionText="Figure 13. Male calling song structure of Yoyetta loftyensis sp. nov. illustrated in waveform plots, including: (A) a single cueing phrase commencing with an extended dense syllable sequence/long echeme, followed by three normal phrases; (B) a series of four normal phrases, followed by a cueing phrase, then two sets of alternating normal and cueing phrases; (C) expanded section from (B) showing two normal phrases (demarcated by solid black bar above) and one cueing phrase (demarcated by dashed line above). The final subfigure (D) is a spectrogram displaying song frequency. This specimen was recorded in the field at 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E) by D. Marshall and K. Hill." figureDoi="http://doi.org/10.5281/zenodo.4654483" httpUri="https://zenodo.org/record/4654483/files/figure.png" pageId="12" pageNumber="135">Fig. 13A</figureCitation>
). The two different phrase
<typeStatus id="54A1659741010A6FFD4AFA33651DF9E1" box="[738,793,1533,1557]" pageId="12" pageNumber="135">types</typeStatus>
may be produced interchangeably and there is little difference in overall duration between normal and cueing phrases (each lasting approximately 0.5–
<geoCoordinate id="EE2EBDF241010A6FFE61F99A67F8F999" box="[457,508,1620,1645]" degrees="1.1" direction="south" orientation="latitude" pageId="12" pageNumber="135" precision="5555" value="-1.1">1.1 s</geoCoordinate>
). A comparison of cueing phrase structures recorded from across the geographical distribution of this species is provided in
<figureCitation id="1321C7B041010A6FFDFEF94164A6F953" box="[598,674,1679,1703]" captionStart="Figure 14" captionStartId="16.[167,235,1185,1207]" captionTargetBox="[167,1467,162,1163]" captionTargetId="figure-9@16.[167,1467,151,1163]" captionTargetPageId="16" captionText="Figure 14. Male calling song structure of Yoyetta loftyensis sp. nov. A comparison of phrase structures recorded from multiple sites, including: (A) Mt Lofty (34°59'S 138°43'E); (B) Hale Conservation Park (34°41'S 138°55'E); (C) 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E); and (D) 2.3 km S. of Birdwood (34°50'S 138°58'E). All examples are cueing phrases, apart from the one in (B) with a black scale bar above, which is a normal phrase. Recordings (A) and (B) were made in the field by LWP. Recordings (C) and (D) were made by D. Marshall and K. Hill." figureDoi="http://doi.org/10.5281/zenodo.4654485" httpUri="https://zenodo.org/record/4654485/files/figure.png" pageId="12" pageNumber="135">Fig. 14</figureCitation>
.
</paragraph>
<caption id="DF658BBD41010A6FFF0FFB2B6445FAC4" ID-DOI="http://doi.org/10.5281/zenodo.4654477" ID-Zenodo-Dep="4654477" httpUri="https://zenodo.org/record/4654477/files/figure.png" pageId="12" pageNumber="135" startId="12.[167,234,1253,1275]" targetBox="[166,1466,164,1233]" targetPageId="12">
<paragraph id="8BA5DB3541010A6FFF0FFB2B6445FAC4" blockId="12.[167,1467,1253,1328]" pageId="12" pageNumber="135">
<emphasis id="B96E072741010A6FFF0FFB2B6702FB0E" bold="true" box="[167,262,1253,1275]" pageId="12" pageNumber="135">Figure 10</emphasis>
. Waveform plots comparing different recordings of the calling song of
<taxonomicName id="4C1AA0B641010A6FFC31FB28624FFB0F" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[921,1099,1254,1275]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="135" phylum="Arthropoda" rank="species" species="enigmatica" status="sp. nov.">
<emphasis id="B96E072741010A6FFC31FB28624FFB0F" box="[921,1099,1254,1275]" italics="true" pageId="12" pageNumber="135">Yoyetta enigmatica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C41010A6FFBF8FB28629CFB0F" box="[1104,1176,1254,1275]" pageId="12" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
from:
<emphasis id="B96E072741010A6FFB7CFB2B62F7FB0E" box="[1236,1267,1253,1274]" italics="true" pageId="12" pageNumber="135">(A)</emphasis>
Mt Kaputar (30°17'S 150°10'E);
<emphasis id="B96E072741010A6FFEBDFACE6737FAE1" box="[277,307,1280,1301]" italics="true" pageId="12" pageNumber="135">(B)</emphasis>
Mt Kaputar;
<emphasis id="B96E072741010A6FFE10FACE67D3FAE1" box="[440,471,1280,1301]" italics="true" pageId="12" pageNumber="135">(C)</emphasis>
Lamington National Park (28°13'S 153°13'E);
<emphasis id="B96E072741010A6FFC0BFACE65C0FAE1" box="[931,964,1280,1301]" italics="true" pageId="12" pageNumber="135">(D)</emphasis>
Springbrook National Park (28°13'S 153°16'E). All recordings were made in the field by LWP.
</paragraph>
</caption>
<paragraph id="8BA5DB3541010A6FFF0EF97464FFF8B3" blockId="12.[166,794,1722,1864]" pageId="12" pageNumber="135">Singing occurs almost exclusively when males are stationary. However, a phrase with an extended echeme may also be produced during short flights between singing stations. It is inferred that the female wing-flick response would be timed during these long gaps at the end of each cueing phrase.</paragraph>
<paragraph id="8BA5DB3541010A6FFF0EF89163BFF998" blockId="12.[166,794,1887,1999]" lastBlockId="12.[839,1467,1415,2000]" pageId="12" pageNumber="135">
<emphasis id="B96E072741010A6FFF0EF8916737F883" bold="true" box="[166,307,1887,1911]" pageId="12" pageNumber="135">Morphology</emphasis>
.
<emphasis id="B96E072741010A6FFEEAF8916778F883" bold="true" box="[322,380,1887,1911]" pageId="12" pageNumber="135">Male</emphasis>
(
<figureCitation id="1321C7B041010A6FFE24F89167E2F883" box="[396,486,1887,1911]" captionStart="Figure 4" captionStartId="6.[166,234,1032,1054]" captionTargetBox="[167,1467,151,1018]" captionTargetId="figure-9@6.[167,1467,151,1018]" captionTargetPageId="6" captionText="Figure 4. Photos of male left opercula: (A) Yoyetta douglasi sp. nov.; (B) Y.enigmatica sp. nov.; (C) Y. loftyensis sp. nov.; (D) Y.ngarabal sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4654461" httpUri="https://zenodo.org/record/4654461/files/figure.png" pageId="12" pageNumber="135">Figs 4C</figureCitation>
,
<figureCitation id="1321C7B041010A6FFE5BF8916411F883" box="[499,533,1887,1911]" captionStart="Figure 5" captionStartId="7.[121,189,1038,1060]" captionTargetBox="[121,1421,151,1018]" captionTargetId="figure-10@7.[121,1421,151,1018]" captionTargetPageId="7" captionText="Figure 5. Photos of male left timbals, with dorsal edge at right and posterior margin at bottom: (A) Yoyetta douglasi sp. nov.; (B) Y. enigmatica sp. nov.; (C) Y. loftyensis sp. nov.; (D) Y. ngarabal sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4654463" httpUri="https://zenodo.org/record/4654463/files/figure.png" pageId="12" pageNumber="135">5C</figureCitation>
,
<figureCitation id="1321C7B041010A6FFD8AF8916447F883" box="[546,579,1887,1911]" captionStart="Figure 6" captionStartId="8.[166,234,1069,1091]" captionTargetBox="[167,1467,151,1050]" captionTargetId="figure-9@8.[167,1467,151,1050]" captionTargetPageId="8" captionText="Figure 6. Lateral views of the caudal abdomen of: (A) Yoyetta douglasi sp. nov.; (B) Y. enigmatica sp. nov.; (C) Y. loftyensis sp. nov.; (D) Y. ngarabal sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4654467" httpUri="https://zenodo.org/record/4654467/files/figure.png" pageId="12" pageNumber="135">6C</figureCitation>
,
<figureCitation id="1321C7B041010A6FFDF8F891646EF883" box="[592,618,1887,1911]" captionStart="Figure 12" captionStartId="14.[166,234,1251,1273]" captionTargetBox="[166,1466,158,1236]" captionTargetId="figure-10@14.[166,1466,151,1236]" captionTargetPageId="14" captionText="Figure 12. Yoyetta loftyensis sp. nov.: (A) holotype male, 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E), dorsal habitus; (B) holotype male, ventral habitus; (C) female, 2.3 km S. of Birdwood (34°50'S 138°58'E) dorsal habitus; (D) female, 2.3 km S. of Birdwood, ventral habitus; (E) typical habitat, Adelaide; (F) typical habitat, Adelaide. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4654481" httpUri="https://zenodo.org/record/4654481/files/figure.png" pageId="12" pageNumber="135">12</figureCitation>
A–B, 15).
<emphasis id="B96E072741010A6FFD48F891651EF883" box="[736,794,1887,1911]" italics="true" pageId="12" pageNumber="135">Head</emphasis>
almost as wide as mesonotum, black; a central triangular yellow-brown fascia on midline posterior to ocelli, base on posterior margin, reducing anteriorly, ocelli pink to pale; dorsal postclypeus black, with a narrow yellow-brown fascia on midline, apex directly anteriorly, ventral surface black, ochraceous spot on anterior midline, with black transverse grooves, lateral margins brown-ochraceous; anteclypeus black, rostrum pale brown at base, black around apical third, brown centrally becoming dark brown to black at apex, reaching anterior of hind coxae; lorum black, gena black; eyes dull black; antennae black, supra-antennal plates black.
</paragraph>
<paragraph id="8BA5DB3541010A6FFCCDF9BC65E0F824" blockId="12.[839,1467,1415,2000]" pageId="12" pageNumber="135">
<emphasis id="B96E072741010A6FFCCDF9BC65B3F97E" box="[869,951,1650,1674]" italics="true" pageId="12" pageNumber="135">Thorax</emphasis>
mainly dull black. Pronotum black, with a central yellow-brown fascia extending from behind anterior margin almost to margin of pronotal collar; patchy dark brown markings over raised lateral areas of pronotum; paramedial fissures black, lateral fissures variably dark brown; pronotal collar dark brown to black, central third variably brown, margins of lateral angles brown, lateral areas black. Mesonotum black, parapsidal sutures black, scutal depressions and surrounds black; cruciform elevation arms black, lateral depressions light brown. Metanotum black, midline black, pale brown over central third, becoming black laterally.
</paragraph>
<caption id="DF658BBD41000A6EFFD1F8B36378F83D" ID-DOI="http://doi.org/10.5281/zenodo.4654479" ID-Zenodo-Dep="4654479" httpUri="https://zenodo.org/record/4654479/files/figure.png" pageId="13" pageNumber="136" startId="13.[121,187,1917,1939]" targetBox="[133,1409,163,1892]" targetPageId="13">
<paragraph id="8BA5DB3541000A6EFFD1F8B36378F83D" blockId="13.[121,1421,1917,1993]" pageId="13" pageNumber="136">
<emphasis id="B96E072741000A6EFFD1F8B366DFF866" bold="true" box="[121,219,1917,1939]" pageId="13" pageNumber="136">Figure 11.</emphasis>
<taxonomicName id="4C1AA0B641000A6EFF77F8B06794F867" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[223,400,1918,1939]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="136" phylum="Arthropoda" rank="species" species="enigmatica" status="sp. nov.">
<emphasis id="B96E072741000A6EFF77F8B06794F867" box="[223,400,1918,1939]" italics="true" pageId="13" pageNumber="136">Yoyetta enigmatica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C41000A6EFE3CF8B067D8F867" box="[404,476,1918,1939]" pageId="13" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
: illustration of male genitalia;
<emphasis id="B96E072741000A6EFD5CF8B0650BF867" box="[756,783,1918,1939]" italics="true" pageId="13" pageNumber="136">(a)</emphasis>
pygofer, viewed laterally from the left;
<emphasis id="B96E072741000A6EFBD2F8B06291F867" box="[1146,1173,1918,1939]" italics="true" pageId="13" pageNumber="136">(b)</emphasis>
pygofer, viewed ventrally;
<emphasis id="B96E072741000A6EFFD1F8576690F85A" box="[121,148,1945,1966]" italics="true" pageId="13" pageNumber="136">(c)</emphasis>
aedeagus viewed laterally from the left;
<emphasis id="B96E072741000A6EFDBFF8566436F859" box="[535,562,1944,1965]" italics="true" pageId="13" pageNumber="136">(d)</emphasis>
apical portion of theca, viewed laterally from the left;
<emphasis id="B96E072741000A6EFB90F8576257F85A" box="[1080,1107,1945,1966]" italics="true" pageId="13" pageNumber="136">(e)</emphasis>
terminal apex of theca;
<emphasis id="B96E072741000A6EFA90F856634BF859" box="[1336,1359,1944,1965]" italics="true" pageId="13" pageNumber="136">(f)</emphasis>
apical portion of theca, viewed ventrally. Characters depicted as in Fig. 7. Specimen from Dawson Springs, Mt Kaputar (30°17'S 150°10'E).
</paragraph>
</caption>
<caption id="DF658BBD41030A6DFF0EFB2D6238FADA" ID-DOI="http://doi.org/10.5281/zenodo.4654481" ID-Zenodo-Dep="4654481" httpUri="https://zenodo.org/record/4654481/files/figure.png" pageId="14" pageNumber="137" startId="14.[166,234,1251,1273]" targetBox="[166,1466,158,1236]" targetPageId="14">
<paragraph id="8BA5DB3541030A6DFF0EFB2D6238FADA" blockId="14.[166,1467,1251,1326]" pageId="14" pageNumber="137">
<emphasis id="B96E072741030A6DFF0EFB2D670BFB0C" bold="true" box="[166,271,1251,1273]" pageId="14" pageNumber="137">Figure 12.</emphasis>
<taxonomicName id="4C1AA0B641030A6DFEBDFB2A67B8FB0C" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[277,444,1251,1273]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="137" phylum="Arthropoda" rank="species" species="loftyensis" status="sp. nov.">
<emphasis id="B96E072741030A6DFEBDFB2A67B8FB0C" box="[277,444,1251,1273]" italics="true" pageId="14" pageNumber="137">Yoyetta loftyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C41030A6DFE6BFB2A6416FB0D" box="[451,530,1252,1273]" pageId="14" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="B96E072741030A6DFDB3FB2D643DFB0C" box="[539,569,1251,1272]" italics="true" pageId="14" pageNumber="137">(A)</emphasis>
holotype male, 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E), dorsal habitus;
<emphasis id="B96E072741030A6DFA33FB2D63BEFB0C" box="[1435,1466,1251,1272]" italics="true" pageId="14" pageNumber="137">(B)</emphasis>
holotype male, ventral habitus;
<emphasis id="B96E072741030A6DFE78FB3067F4FAE7" box="[464,496,1278,1299]" italics="true" pageId="14" pageNumber="137">(C)</emphasis>
female, 2.3 km S. of Birdwood (34°50'S 138°58'E) dorsal habitus;
<emphasis id="B96E072741030A6DFBC3FB306288FAE7" box="[1131,1164,1278,1299]" italics="true" pageId="14" pageNumber="137">(D)</emphasis>
female, 2.3 km S. of Birdwood, ventral habitus;
<emphasis id="B96E072741030A6DFEE9FAD7675BFADA" box="[321,351,1305,1326]" italics="true" pageId="14" pageNumber="137">(E)</emphasis>
typical habitat, Adelaide;
<emphasis id="B96E072741030A6DFDF4FAD7647EFADA" box="[604,634,1305,1326]" italics="true" pageId="14" pageNumber="137">(F)</emphasis>
typical habitat, Adelaide. Scale bars = 10 mm.
</paragraph>
</caption>
<paragraph id="8BA5DB3541030A6DFF6DFA5267FDF89F" blockId="14.[166,794,1435,1987]" pageId="14" pageNumber="137">
<emphasis id="B96E072741030A6DFF6DFA5266FAFA47" box="[197,254,1436,1459]" italics="true" pageId="14" pageNumber="137">Legs.</emphasis>
Coxae black, joints orange-red; basisterna black, bordered orange-red; trochanter black with diffuse brown areas; meracantha small, narrow, pale cream, black at base, pointed, marginally overlapping opercula; fore femora mainly orange-red, with black longitudinal markings on inner ventral sides and around base of femoral spines; femoral spines erect, black; mid femora black mainly orange red, with diffuse black areas on anterior sides, hind femora orange-brown with black markings on anterior apices, joints orange-red; fore tibia mainly black, dark reddish-brown on inner sides; mid tibia black, tending dark reddish-brown towards anterior apices; hind tibia dark reddish-brown with diffuse black areas; fore and mid tarsi dark reddish brown, tending black on outer sides; hind tarsi pale orange-brown, becoming dark brown towards claws; spines orange-brown; claws dark brown, black at tips.
</paragraph>
<paragraph id="8BA5DB3541030A6DFF6DF8BF65EFF9B2" blockId="14.[166,794,1435,1987]" lastBlockId="14.[839,1467,1435,1988]" pageId="14" pageNumber="137">
<emphasis id="B96E072741030A6DFF6DF8BF6701F87C" box="[197,261,1905,1928]" italics="true" pageId="14" pageNumber="137">Wings</emphasis>
with fore wing costal veins black anteriorly, orangebrown central rib, becoming brown distally, pterostigma mottled red, basal cell pale yellow, translucent, black anterior border, arculus black, basal membranes orange, other veins dark brown to black, proximal segment of vein CuA pale, becoming brown distally with eight apical cells; hind wing plaga white to whitish-cream over jugum and vein 3A, thin along vein 2A, subcostal vein pale, other veins brown, with six apical cells.
</paragraph>
<paragraph id="8BA5DB3541030A6DFCCDF985632DF94F" blockId="14.[839,1467,1435,1988]" pageId="14" pageNumber="137">
<emphasis id="B96E072741030A6DFCCDF98565CDF997" box="[869,969,1611,1635]" italics="true" pageId="14" pageNumber="137">Opercula</emphasis>
(
<figureCitation id="1321C7B041030A6DFC7EF9856222F990" box="[982,1062,1611,1636]" captionStart="Figure 4" captionStartId="6.[166,234,1032,1054]" captionTargetBox="[167,1467,151,1018]" captionTargetId="figure-9@6.[167,1467,151,1018]" captionTargetPageId="6" captionText="Figure 4. Photos of male left opercula: (A) Yoyetta douglasi sp. nov.; (B) Y.enigmatica sp. nov.; (C) Y. loftyensis sp. nov.; (D) Y.ngarabal sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4654461" httpUri="https://zenodo.org/record/4654461/files/figure.png" pageId="14" pageNumber="137">Fig. 4C</figureCitation>
) covering abdominal cavity, spatulate, following body axis ventrolaterally, depressed centrally, variably black at base, pale cream across remainder, tinged orange in some specimens, clearly separated.
</paragraph>
<paragraph id="8BA5DB3541030A6DFCCDF90F623BF87D" blockId="14.[839,1467,1435,1988]" pageId="14" pageNumber="137">
<emphasis id="B96E072741030A6DFCCDF90F65BDF92D" box="[869,953,1729,1753]" italics="true" pageId="14" pageNumber="137">Timbals</emphasis>
(
<figureCitation id="1321C7B041030A6DFC62F90F621BF92D" box="[970,1055,1729,1753]" captionStart="Figure 4" captionStartId="6.[166,234,1032,1054]" captionTargetBox="[167,1467,151,1018]" captionTargetId="figure-9@6.[167,1467,151,1018]" captionTargetPageId="6" captionText="Figure 4. Photos of male left opercula: (A) Yoyetta douglasi sp. nov.; (B) Y.enigmatica sp. nov.; (C) Y. loftyensis sp. nov.; (D) Y.ngarabal sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4654461" httpUri="https://zenodo.org/record/4654461/files/figure.png" pageId="14" pageNumber="137">Fig. 4D</figureCitation>
) with five distinct long ribs; long ribs 1–4 extending across surrounding membrane and fused dorsally along basal spur; long rib 5 independent of basal spur, comparatively shorter, extending ventrally across half of membrane; large ridged dome on posterior timbal plate extending across two-thirds of timbal; apodeme pit oval shaped, inconspicuous.
</paragraph>
<paragraph id="8BA5DB3541030A73FCCDF840672EF947" blockId="14.[839,1467,1435,1988]" lastBlockId="16.[166,794,1368,2008]" lastPageId="16" lastPageNumber="139" pageId="14" pageNumber="137">
<emphasis id="B96E072741030A6DFCCDF84065C8F852" box="[869,972,1934,1958]" italics="true" pageId="14" pageNumber="137">Abdomen</emphasis>
with tergite 1 black, membrane anterior to timbals orange-red, tergite 2 black, tergites 3–7 black with orange posterior margins, extending laterally to epipleurites, progressively increasing on either side of midline to reach anterior borders of tergites 4–7, posterior margins yellow; tergite 8 black, with red markings on lateral margins (
<figureCitation id="1321C7B0411D0A73FF06FA00670DFA12" box="[174,265,1486,1510]" captionStart="Figure 6" captionStartId="8.[166,234,1069,1091]" captionTargetBox="[167,1467,151,1050]" captionTargetId="figure-9@8.[167,1467,151,1050]" captionTargetPageId="8" captionText="Figure 6. Lateral views of the caudal abdomen of: (A) Yoyetta douglasi sp. nov.; (B) Y. enigmatica sp. nov.; (C) Y. loftyensis sp. nov.; (D) Y. ngarabal sp. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4654467" httpUri="https://zenodo.org/record/4654467/files/figure.png" pageId="16" pageNumber="139">Fig. 6C</figureCitation>
). Epipleurites 1–3 black anteriorly, orange red posteriorly, epipleurites 4–6 mainly orange. Sternite I black, sternite II black medially, orange-red over central third posterior to timbal cavity, black laterally, sternite III orange-red with central black spots on anterior and posterior margins, sternites IV–VI orange, sternite VII mainly orange, red posteriorly, sternite VIII fiery red, with pale brown pubescence.
</paragraph>
<caption id="DF658BBD41020A6CFFD1F8FB64D4F825" ID-DOI="http://doi.org/10.5281/zenodo.4654483" ID-Zenodo-Dep="4654483" httpUri="https://zenodo.org/record/4654483/files/figure.png" pageId="15" pageNumber="138" startId="15.[121,189,1845,1867]" targetBox="[122,1388,131,1838]" targetPageId="15">
<paragraph id="8BA5DB3541020A6CFFD1F8FB64D4F825" blockId="15.[121,1389,1845,2001]" pageId="15" pageNumber="138">
<emphasis id="B96E072741020A6CFFD1F8FB66E7F8BE" bold="true" box="[121,227,1845,1867]" pageId="15" pageNumber="138">Figure 13.</emphasis>
Male calling song structure of
<taxonomicName id="4C1AA0B641020A6CFDB3F8F864C0F8BE" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[539,708,1845,1867]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="138" phylum="Arthropoda" rank="species" species="loftyensis" status="sp. nov.">
<emphasis id="B96E072741020A6CFDB3F8F864C0F8BE" box="[539,708,1845,1867]" italics="true" pageId="15" pageNumber="138">Yoyetta loftyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C41020A6CFD63F8F8651CF8BF" box="[715,792,1846,1867]" pageId="15" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
illustrated in waveform plots, including:
<emphasis id="B96E072741020A6CFB1AF8F862D4F8BF" box="[1202,1232,1846,1867]" italics="true" pageId="15" pageNumber="138">(A)</emphasis>
a single cueing phrase commencing with an extended dense syllable sequence/long echeme, followed by three normal phrases;
<emphasis id="B96E072741020A6CFB1DF89E62D7F891" box="[1205,1235,1872,1893]" italics="true" pageId="15" pageNumber="138">(B)</emphasis>
a series of four normal phrases, followed by a cueing phrase, then two sets of alternating normal and cueing phrases;
<emphasis id="B96E072741020A6CFBCEF8A56282F874" box="[1126,1158,1899,1920]" italics="true" pageId="15" pageNumber="138">(C)</emphasis>
expanded section from
<emphasis id="B96E072741020A6CFFD1F8486693F86F" box="[121,151,1926,1947]" italics="true" pageId="15" pageNumber="138">(B)</emphasis>
showing two normal phrases (demarcated by solid black bar above) and one cueing phrase (demarcated by dashed line above). The final subfigure
<emphasis id="B96E072741020A6CFE9CF86E6751F841" box="[308,341,1952,1973]" italics="true" pageId="15" pageNumber="138">(D)</emphasis>
is a spectrogram displaying song frequency. This specimen was recorded in the field at 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E) by D. Marshall and K. Hill.
</paragraph>
</caption>
<caption id="DF658BBD411D0A73FF0FFB6F67EEFAD6" ID-DOI="http://doi.org/10.5281/zenodo.4654485" ID-Zenodo-Dep="4654485" httpUri="https://zenodo.org/record/4654485/files/figure.png" pageId="16" pageNumber="139" startId="16.[167,235,1185,1207]" targetBox="[167,1467,162,1163]" targetPageId="16">
<paragraph id="8BA5DB35411D0A73FF0FFB6F67EEFAD6" blockId="16.[167,1467,1185,1314]" pageId="16" pageNumber="139">
<emphasis id="B96E0727411D0A73FF0FFB6F6714FB42" bold="true" box="[167,272,1185,1207]" pageId="16" pageNumber="139">Figure 14.</emphasis>
Male calling song structure of
<taxonomicName id="4C1AA0B6411D0A73FDEFFB6C64EBFB42" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[583,751,1185,1207]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="loftyensis" status="sp. nov.">
<emphasis id="B96E0727411D0A73FDEFFB6C64EBFB42" box="[583,751,1185,1207]" italics="true" pageId="16" pageNumber="139">Yoyetta loftyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C411D0A73FD5FFB6F6547FB42" box="[759,835,1185,1206]" pageId="16" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
A comparison of phrase structures recorded from multiple sites, including:
<emphasis id="B96E0727411D0A73FEA6FB726729FB25" box="[270,301,1212,1233]" italics="true" pageId="16" pageNumber="139">(A)</emphasis>
Mt Lofty (34°59'S 138°43'E);
<emphasis id="B96E0727411D0A73FDF1FB72647CFB25" box="[601,632,1212,1233]" italics="true" pageId="16" pageNumber="139">(B)</emphasis>
Hale Conservation Park (34°41'S 138°55'E);
<emphasis id="B96E0727411D0A73FB9BFB726256FB25" box="[1075,1106,1212,1233]" italics="true" pageId="16" pageNumber="139">(C)</emphasis>
9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E); and
<emphasis id="B96E0727411D0A73FE3CFB1967B2FB18" box="[404,438,1239,1260]" italics="true" pageId="16" pageNumber="139">(D)</emphasis>
2.3 km S. of Birdwood (34°50'S 138°58'E). All examples are cueing phrases, apart from the one in
<emphasis id="B96E0727411D0A73FAC4FB19638EFB18" box="[1388,1418,1239,1260]" italics="true" pageId="16" pageNumber="139">(B)</emphasis>
with a black scale bar above, which is a normal phrase. Recordings
<emphasis id="B96E0727411D0A73FD57FB3F6519FAF2" box="[767,797,1265,1286]" italics="true" pageId="16" pageNumber="139">(A)</emphasis>
and
<emphasis id="B96E0727411D0A73FCE4FB3F656EFAF2" box="[844,874,1265,1286]" italics="true" pageId="16" pageNumber="139">(B)</emphasis>
were made in the field by LWP. Recordings
<emphasis id="B96E0727411D0A73FABEFB3F6332FAF2" box="[1302,1334,1265,1286]" italics="true" pageId="16" pageNumber="139">(C)</emphasis>
and
<emphasis id="B96E0727411D0A73FACDFB3F6382FAF2" box="[1381,1414,1265,1286]" italics="true" pageId="16" pageNumber="139">(D)</emphasis>
were made by D. Marshall and K. Hill.
</paragraph>
</caption>
<paragraph id="8BA5DB35411D0A73FF6DF976627DFA3C" blockId="16.[166,794,1368,2008]" lastBlockId="16.[839,1467,1368,1480]" pageId="16" pageNumber="139">
<emphasis id="B96E0727411D0A73FF6DF9766723F924" box="[197,295,1720,1744]" italics="true" pageId="16" pageNumber="139">Genitalia</emphasis>
(
<figureCitation id="1321C7B0411D0A73FE9BF9766779F924" box="[307,381,1720,1744]" captionStart="Figure 15" captionStartId="17.[121,189,1922,1944]" captionTargetBox="[134,1362,140,1901]" captionTargetId="figure-10@17.[121,1375,133,1910]" captionTargetPageId="17" captionText="Figure 15. Yoyetta loftyensis sp. nov.: illustration of male genitalia; (a) pygofer, viewed laterally from the left; (b) pygofer, viewed ventrally; (c) aedeagus, viewed laterally from the left; (d) apex of theca. Characters depicted as in Fig. 7. Specimen from 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E)." figureDoi="http://doi.org/10.5281/zenodo.4654487" httpUri="https://zenodo.org/record/4654487/files/figure.png" pageId="16" pageNumber="139">Fig. 15</figureCitation>
). Pygofer shiny black upper lobe mainly black with red posterior margin; basal lobe ochraceous; dorsal beak black, dark brown posterior margin, anal styles orange, red at tip. Uncus orange-brown; in lateral view beak-like and stumpy; lobes in ventral view bulbous, with rounded lateral termination; claspers clearly divided, triangular with apices gradually tapering laterally. Aedeagus with pseudoparameres extending around one third the length of theca; theca recurved ventrally at 180° towards apex, with transparent flange along outer margin of recurvature, broadly smooth, around 2× width of theca, terminating adjacent to apex of theca; apex short, tapered, sclerotized, angled 30° ventrally, with two lateral rows of prominent cornuti at base, numerous cornuti over apex.
</paragraph>
<paragraph id="8BA5DB35411D0A73FCEFFA256241F9F7" blockId="16.[839,1468,1515,2008]" box="[839,1093,1515,1539]" pageId="16" pageNumber="139">
<emphasis id="B96E0727411D0A73FCEFFA25659EF9F7" bold="true" box="[839,922,1515,1539]" pageId="16" pageNumber="139">Female</emphasis>
(
<figureCitation id="1321C7B0411D0A73FC1FFA256206F9F7" box="[951,1026,1515,1539]" captionStart="Figure 12" captionStartId="14.[166,234,1251,1273]" captionTargetBox="[166,1466,158,1236]" captionTargetId="figure-10@14.[166,1466,151,1236]" captionTargetPageId="14" captionText="Figure 12. Yoyetta loftyensis sp. nov.: (A) holotype male, 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E), dorsal habitus; (B) holotype male, ventral habitus; (C) female, 2.3 km S. of Birdwood (34°50'S 138°58'E) dorsal habitus; (D) female, 2.3 km S. of Birdwood, ventral habitus; (E) typical habitat, Adelaide; (F) typical habitat, Adelaide. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4654481" httpUri="https://zenodo.org/record/4654481/files/figure.png" pageId="16" pageNumber="139">Fig. 12</figureCitation>
C–D).
</paragraph>
<paragraph id="8BA5DB35411D0A73FCCDF9C662BEF9C9" blockId="16.[839,1468,1515,2008]" pageId="16" pageNumber="139">
<emphasis id="B96E0727411D0A73FCCDF9C665A4F9D4" box="[869,928,1544,1568]" italics="true" pageId="16" pageNumber="139">Head</emphasis>
and
<emphasis id="B96E0727411D0A73FC74F9C66227F9D4" box="[988,1059,1544,1568]" italics="true" pageId="16" pageNumber="139">thorax</emphasis>
similar to male, with darker brown markings along parapsidal sutures.
</paragraph>
<paragraph id="8BA5DB35411D0A73FCCDF98D6300F82C" blockId="16.[839,1468,1515,2008]" pageId="16" pageNumber="139">
<emphasis id="B96E0727411D0A73FCCDF98D65CEF9AF" box="[869,970,1603,1627]" italics="true" pageId="16" pageNumber="139">Abdomen</emphasis>
. Tergites similar to male. Sternite I–II black, posterior margins ochraceous, sternites III–VI orangered with midline black marking decreasing to sternite IV, sternite VII pale brown with anterior black spot either side of midline. Abdominal segment 9 with central black triangular marking, base at anterior margin, reducing over middle third, expanding over dorsal beak, flanked laterally by orange stripe and lateral black area, extending laterally over anterior third, remainder reddish-orange laterally with posterior black spot. Dorsal beak black, ovipositor dark brown, becoming black at tip, not extending beyond apex of abdominal segment 9. Anal styles orange-red; ovipositor sheath black, becoming paler brown ventrally, gonocoxites black laterally, ochraceous along midline.
</paragraph>
</subSubSection>
<caption id="DF658BBD411C0A72FFD1F84C640EF839" ID-DOI="http://doi.org/10.5281/zenodo.4654487" ID-Zenodo-Dep="4654487" httpUri="https://zenodo.org/record/4654487/files/figure.png" pageId="17" pageNumber="140" startId="17.[121,189,1922,1944]" targetBox="[134,1362,140,1901]" targetPageId="17">
<paragraph id="8BA5DB35411C0A72FFD1F84C640EF839" blockId="17.[121,1421,1922,1997]" pageId="17" pageNumber="140">
<emphasis id="B96E0727411C0A72FFD1F84C66E7F863" bold="true" box="[121,227,1922,1944]" pageId="17" pageNumber="140">Figure 15.</emphasis>
<taxonomicName id="4C1AA0B6411C0A72FF43F84D6790F863" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[235,404,1922,1944]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="140" phylum="Arthropoda" rank="species" species="loftyensis" status="sp. nov.">
<emphasis id="B96E0727411C0A72FF43F84D6790F863" box="[235,404,1922,1944]" italics="true" pageId="17" pageNumber="140">Yoyetta loftyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C411C0A72FE34F84C67EEF86C" box="[412,490,1922,1944]" pageId="17" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
: illustration of male genitalia;
<emphasis id="B96E0727411C0A72FCB7F84C653FF863" box="[799,827,1922,1943]" italics="true" pageId="17" pageNumber="140">(a)</emphasis>
pygofer, viewed laterally from the left;
<emphasis id="B96E0727411C0A72FB63F84C62ECF863" box="[1227,1256,1922,1943]" italics="true" pageId="17" pageNumber="140">(b)</emphasis>
pygofer, viewed ventrally;
<emphasis id="B96E0727411C0A72FF73F85366F2F846" box="[219,246,1949,1970]" italics="true" pageId="17" pageNumber="140">(c)</emphasis>
aedeagus, viewed laterally from the left;
<emphasis id="B96E0727411C0A72FD2AF8536499F846" box="[642,669,1949,1970]" italics="true" pageId="17" pageNumber="140">(d)</emphasis>
apex of theca. Characters depicted as in Fig. 7. Specimen from 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E).
</paragraph>
</caption>
<subSubSection id="C30088BE411F0A71FF0EFF56651EFC4E" pageId="18" pageNumber="141" type="description">
<paragraph id="8BA5DB35411F0A71FF0EFF566454FE89" blockId="18.[166,794,151,381]" pageId="18" pageNumber="141">
<emphasis id="B96E0727411F0A71FF0EFF566749FF44" bold="true" box="[166,333,152,176]" pageId="18" pageNumber="141">Measurements</emphasis>
(in mm;
<specimenCount id="9D1C10BC411F0A71FE04FF566409FF5B" box="[428,525,151,176]" pageId="18" pageNumber="141" type="male">10 males</specimenCount>
,
<specimenCount id="9D1C10BC411F0A71FDBEFF566485FF5B" box="[534,641,151,176]" pageId="18" pageNumber="141" type="female">8 females</specimenCount>
). Body length: male 20.9–24.7 (22.7); female 22.3–24.0 (28.5). Fore wing length: male 25.5–28.7 (26.9); female 28.3–31.2 (29.4). Fore wing width: male 9.2–10.6 (9.8); female 9.7–11.3 (10.5). Head width: male 6.0–7.0 (6.6); female 6.6–7.1 (6.9). Pronotum width: male 5.9–7.7 (6.9); female 6.4–8.0 (7.5). Abdomen width: male 6.1–7.2 (6.5); female 6.4–7.5 (7.0). Ovipositor length: female 6.0–8.4 (7.7).
</paragraph>
<paragraph id="8BA5DB35411F0A71FF0EFE5C66FDFD8C" blockId="18.[166,794,402,954]" pageId="18" pageNumber="141">
<emphasis id="B96E0727411F0A71FF0EFE5C67ACFE5E" bold="true" box="[166,424,402,426]" pageId="18" pageNumber="141">Distinguishing features</emphasis>
. Within the
<taxonomicName id="4C1AA0B6411F0A71FD8EFE5D64CDFE5E" box="[550,713,402,426]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B96E0727411F0A71FD8EFE5D64CDFE5E" box="[550,713,402,426]" italics="true" pageId="18" pageNumber="141">Y. abdominalis</emphasis>
</taxonomicName>
species group, the calling song of
<taxonomicName id="4C1AA0B6411F0A71FE12FE7E643CFE33" authorityName="Popple &amp; Emery" authorityYear="2020" box="[442,568,431,455]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="loftyensis">
<emphasis id="B96E0727411F0A71FE12FE7E643CFE33" box="[442,568,431,455]" italics="true" pageId="18" pageNumber="141">Y. loftyensis</emphasis>
</taxonomicName>
is most similar to the call of
<taxonomicName id="4C1AA0B6411F0A71FF44FE006455FE11" authority="Emery, Emery &amp; Popple" authorityName="Emery, Emery &amp; Popple" authorityYear="2019" box="[236,593,461,485]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="serrata">
<emphasis id="B96E0727411F0A71FF44FE00674BFE11" box="[236,335,462,485]" italics="true" pageId="18" pageNumber="141">Y. serrata</emphasis>
Emery, Emery &amp; Popple
</taxonomicName>
from south-eastern
<collectingCountry id="F30D9BA5411F0A71FF0EFE24670BFDF6" box="[166,271,490,514]" name="Australia" pageId="18" pageNumber="141">Australia</collectingCountry>
. The longer minimum gaps between syllables (&gt;
<geoCoordinate id="EE2EBDF2411F0A71FF14FDC96705FDEB" box="[188,257,519,543]" degrees="0.16" direction="south" orientation="latitude" pageId="18" pageNumber="141" precision="555" value="-0.16">0.16 s</geoCoordinate>
;
<bibRefCitation id="EF8BA6C4411F0A71FEA7FDC667D7FDD4" author="Emery, D. L. &amp; N. J. Emery &amp; L. W. Popple" box="[271,467,519,544]" pageId="18" pageNumber="141" pagination="277 - 347" refId="ref12949" refString="Emery, D. L., N. J. Emery, and L. W. Popple. 2019. A revision of the Yoyetta abdominalis (Distant) species group of cicadas (Hemiptera: Cicadidae: Cicadettinae), introducing eight new species. Records of the Australian Museum 71 (7): 277 - 347. https: // doi. org / 10.3853 / j. 2201 - 4349.71.2019.1720" type="journal article" year="2019">
Emery
<emphasis id="B96E0727411F0A71FEF6FDC66791FDEB" box="[350,405,519,543]" italics="true" pageId="18" pageNumber="141">et al.</emphasis>
2019
</bibRefCitation>
) distinguishes
<taxonomicName id="4C1AA0B6411F0A71FDDEFDC664DAFDEB" authorityName="Emery, Emery &amp; Popple" authorityYear="2019" box="[630,734,520,543]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="serrata">
<emphasis id="B96E0727411F0A71FDDEFDC664DAFDEB" box="[630,734,520,543]" italics="true" pageId="18" pageNumber="141">Y. serrata</emphasis>
</taxonomicName>
from
<taxonomicName id="4C1AA0B6411F0A71FF0EFDE8671BFDC9" authorityName="Popple &amp; Emery" authorityYear="2020" box="[166,287,549,573]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="loftyensis">
<emphasis id="B96E0727411F0A71FF0EFDE8671BFDC9" box="[166,287,549,573]" italics="true" pageId="18" pageNumber="141">Y. loftyensis</emphasis>
</taxonomicName>
. The former also lacks the characteristic structure of the cueing phrases produced by
<taxonomicName id="4C1AA0B6411F0A71FD84FD8D64AAFDAE" authorityName="Popple &amp; Emery" authorityYear="2020" box="[556,686,578,602]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="loftyensis">
<emphasis id="B96E0727411F0A71FD84FD8D64AAFDAE" box="[556,686,578,602]" italics="true" pageId="18" pageNumber="141">Y. loftyensis</emphasis>
</taxonomicName>
(see
<figureCitation id="1321C7B0411F0A71FD44FD8C66C7FD8C" captionStart="Figure 13" captionStartId="15.[121,189,1845,1867]" captionTargetBox="[122,1388,131,1838]" captionTargetId="figure-10@15.[121,1388,127,1838]" captionTargetPageId="15" captionText="Figure 13. Male calling song structure of Yoyetta loftyensis sp. nov. illustrated in waveform plots, including: (A) a single cueing phrase commencing with an extended dense syllable sequence/long echeme, followed by three normal phrases; (B) a series of four normal phrases, followed by a cueing phrase, then two sets of alternating normal and cueing phrases; (C) expanded section from (B) showing two normal phrases (demarcated by solid black bar above) and one cueing phrase (demarcated by dashed line above). The final subfigure (D) is a spectrogram displaying song frequency. This specimen was recorded in the field at 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E) by D. Marshall and K. Hill." figureDoi="http://doi.org/10.5281/zenodo.4654483" httpUri="https://zenodo.org/record/4654483/files/figure.png" pageId="18" pageNumber="141">Figs 13</figureCitation>
,
<figureCitation id="1321C7B0411F0A71FF66FDAE66E8FD8C" box="[206,236,608,632]" captionStart="Figure 14" captionStartId="16.[167,235,1185,1207]" captionTargetBox="[167,1467,162,1163]" captionTargetId="figure-9@16.[167,1467,151,1163]" captionTargetPageId="16" captionText="Figure 14. Male calling song structure of Yoyetta loftyensis sp. nov. A comparison of phrase structures recorded from multiple sites, including: (A) Mt Lofty (34°59'S 138°43'E); (B) Hale Conservation Park (34°41'S 138°55'E); (C) 9.8 km NW. of Victor Harbour Road (35°26'S 138°35'E); and (D) 2.3 km S. of Birdwood (34°50'S 138°58'E). All examples are cueing phrases, apart from the one in (B) with a black scale bar above, which is a normal phrase. Recordings (A) and (B) were made in the field by LWP. Recordings (C) and (D) were made by D. Marshall and K. Hill." figureDoi="http://doi.org/10.5281/zenodo.4654485" httpUri="https://zenodo.org/record/4654485/files/figure.png" pageId="18" pageNumber="141">14</figureCitation>
).
</paragraph>
<paragraph id="8BA5DB35411F0A71FF6DFDB3651EFC4E" blockId="18.[166,794,402,954]" pageId="18" pageNumber="141">
Morphologically,
<taxonomicName id="4C1AA0B6411F0A71FE3BFDB0641AFD61" authority="Popple &amp; Emery, 2020" authorityName="Popple &amp; Emery" authorityYear="2020" box="[403,542,637,661]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="loftyensis" status="sp. nov.">
<emphasis id="B96E0727411F0A71FE3BFDB0641AFD61" box="[403,542,637,661]" italics="true" pageId="18" pageNumber="141">Y. loftyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DBA5C411F0A71FD80FDB36480FD60" box="[552,644,637,660]" pageId="18" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
is closest to
<taxonomicName id="4C1AA0B6411F0A71FF0EFD5567ACFD46" authority="(Distant)" baseAuthorityName="Distant" box="[166,424,666,690]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B96E0727411F0A71FF0EFD556746FD46" box="[166,322,666,690]" italics="true" pageId="18" pageNumber="141">Y. abdominalis</emphasis>
(Distant)
</taxonomicName>
, which also occurs in south-eastern
<collectingRegion id="49DE15D7411F0A71FF0EFD796753FD3B" box="[166,343,695,719]" country="Australia" name="South Australia" pageId="18" pageNumber="141">South Australia</collectingRegion>
, and
<taxonomicName id="4C1AA0B6411F0A71FE30FD766402FD3B" authorityName="Emery, Emery &amp; Popple" authorityYear="2019" box="[408,518,696,719]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="serrata">
<emphasis id="B96E0727411F0A71FE30FD766402FD3B" box="[408,518,696,719]" italics="true" pageId="18" pageNumber="141">Y. serrata</emphasis>
</taxonomicName>
. It can be distinguished easily from
<taxonomicName id="4C1AA0B6411F0A71FE80FD18678BFD19" authorityName="Emery, Emery &amp; Popple" authorityYear="2019" box="[296,399,726,749]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="serrata">
<emphasis id="B96E0727411F0A71FE80FD18678BFD19" box="[296,399,726,749]" italics="true" pageId="18" pageNumber="141">Y. serrata</emphasis>
</taxonomicName>
by the colour of the fore wing basal membranes, which are orange (cf. pale whitish-grey). Males can be distinguished from
<taxonomicName id="4C1AA0B6411F0A71FE17FCDE6458FCD3" box="[447,604,783,807]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B96E0727411F0A71FE17FCDE6458FCD3" box="[447,604,783,807]" italics="true" pageId="18" pageNumber="141">Y. abdominalis</emphasis>
</taxonomicName>
by examining the apex of the theca, which is tapered and angled 30° ventrally (cf. short and club-like). Both sexes can be distinguished from fresh specimens of by
<taxonomicName id="4C1AA0B6411F0A71FE72FCA6647EFC8B" box="[474,634,871,895]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B96E0727411F0A71FE72FCA6647EFC8B" box="[474,634,871,895]" italics="true" pageId="18" pageNumber="141">Y. abdominalis</emphasis>
</taxonomicName>
by the lack of conspicuous golden hairs on the head and thorax. Otherwise, females cannot be distinguished reliably from
<taxonomicName id="4C1AA0B6411F0A71FDD4FC6D6517FC4E" box="[636,787,930,954]" class="Insecta" family="Cicadidae" genus="Yoyetta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="141" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B96E0727411F0A71FDD4FC6D6517FC4E" box="[636,787,930,954]" italics="true" pageId="18" pageNumber="141">Y. abdominalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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