treatments-xml/data/9E/B3/C6/9EB3C65E7B5E5E9C9CDD06471CD72582.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.972.52853" ID-GBIF-Dataset="b5be8a15-c690-495c-9b35-a347dd965f7e" ID-PMC="PMC7544732" ID-Pensoft-Pub="1313-2970-972-1" ID-Pensoft-UUID="2751774FF66A5579AE66145BE5055C8E" ID-PubMed="33071542" ID-ZooBank="791674942E9242C38D1FD4DE7264D7B7" ModsDocID="1313-2970-972-1" checkinTime="1601578722686" checkinUser="pensoft" docAuthor="Dayrat, Benoit, Goulding, Tricia C., Apte, Deepak, Aslam, Sadar, Bourke, Adam, Comendador, Joseph, Khalil, Munawar, Ngo, Xuan Qu ảng, Tan, Siong Kiat &amp; Tan, Shau Hwai" docDate="2020" docId="9EB3C65E7B5E5E9C9CDD06471CD72582" docLanguage="en" docName="ZooKeys 972: 1-224" docOrigin="ZooKeys 972" docSource="http://dx.doi.org/10.3897/zookeys.972.52853" docTitle="Peronia madagascariensis" docType="treatment" docVersion="5" id="2751774FF66A5579AE66145BE5055C8E" lastPageNumber="1" masterDocId="2751774FF66A5579AE66145BE5055C8E" masterDocTitle="Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae)" masterLastPageNumber="224" masterPageNumber="1" pageNumber="1" updateTime="1668169548897" updateUser="ExternalLinkService">
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<mods:title>Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dayrat, Benoit</mods:namePart>
<mods:affiliation>Department of Biology, Pennsylvania State University, University Park, PA 16802, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-1514-4854</mods:nameIdentifier>
<mods:nameIdentifier type="email">bdayrat@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Goulding, Tricia C.</mods:namePart>
<mods:affiliation>Department of Biology, Pennsylvania State University, University Park, PA 16802, USA</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Apte, Deepak</mods:namePart>
<mods:affiliation>Bombay Natural History Society, Hornbill House, Opp. Lion Gate, Shaheed Bhagat Singh Road, Mumbai 400 001, Maharashtra, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Aslam, Sadar</mods:namePart>
<mods:affiliation>Centre of Excellence in Marine Biology, University of Karachi, Karachi 75270, Pakistan</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4340-7885</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bourke, Adam</mods:namePart>
<mods:affiliation>College of Engineering, Information Technology and the Environment, Charles Darwin University, Ellengowan Dr, Casuarina, NT 0810, Australia</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Comendador, Joseph</mods:namePart>
<mods:affiliation>National Museum of the Philippines, Taft Ave, Ermita, Manila, 1000, Metro Manila, Philippines</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Khalil, Munawar</mods:namePart>
<mods:affiliation>Department of Marine Science, Universitas Malikussaleh, Reuleut Main Campus, Kecamatan Muara Batu, North Aceh, Aceh, 24355, Indonesia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8264-5317</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ngo, Xuan Qu ảng</mods:namePart>
<mods:affiliation>Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart>Tan, Siong Kiat</mods:namePart>
<mods:affiliation>Lee Kong Chian Natural History Museum, 2 Conservatory Dr, National University of Singapore, 117377, Singapore</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tan, Shau Hwai</mods:namePart>
<mods:affiliation>Centre for Marine and Coastal Studies, Universiti Sains Malaysia, 11800, Minden Penang, Malaysia &amp; Marine Science Laboratory, School of Biological Sciences, Universiti Sains Malaysia, 11800, Minden Penang, Malaysia</mods:affiliation>
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<mods:number>972</mods:number>
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<treatment ID-GBIF-Taxon="168209566" LSID="urn:lsid:plazi:treatment:9EB3C65E7B5E5E9C9CDD06471CD72582" httpUri="http://treatment.plazi.org/id/9EB3C65E7B5E5E9C9CDD06471CD72582" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="9EB3C65E-7B5E-5E9C-9CDD-06471CD72582" authority="(Labbe, 1934 a)" baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
Peronia madagascariensis (
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a)
</taxonomicName>
<figureCitation captionStart="Figure 21" captionStartId="F23" captionText="Figure 21. Peronia madagascariensis A-D digestive system, dorsal view, with intestinal loops of type V E posterior, hermaphroditic (female) reproductive system A holotype, Paraperonia madagascariensis, Madagascar (MNHN-IM- 2000 - 33680) B paralectotype, Paraperonia gondwanae, Mumbai, western India (MNHN-IM- 2000 - 33682) C paralectotype, Paraperonia gondwanae, Red Sea (MNHN-IM- 2000 - 33683) D Madagascar, [5501] (MNHN-IM- 2009 - 16392) E same as D. Scale bars: 5 mm (A-D), 2 mm (E). Abbreviations: dd deferent duct, ddg dorsal digestive gland, fgm female gland mass, hg hermaphroditic gland, i intestine, ov oviduct, pdg posterior digestive gland, rs receptaculum seminis, sp spermatheca, st stomach." figureDoi="10.3897/zookeys.972.52853.figure21" httpUri="https://binary.pensoft.net/fig/458321" pageId="0" pageNumber="1">Figs 21</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F24" captionText="Figure 22. Radula, Peronia madagascariensis, South Africa, [5841] (NMSA W 7547) A right half rows of teeth B rachidian and innermost lateral teeth C rachidian and innermost lateral teeth D lateral teeth E lateral teeth, frontal view F outermost lateral teeth. Scale bars: 200 μm (A), 20 μm (B, E, F), 40 μm (C), 100 μm (D)." figureDoi="10.3897/zookeys.972.52853.figure22" httpUri="https://binary.pensoft.net/fig/458322" pageId="0" pageNumber="1">, 22</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F25" captionText="Figure 23. Anterior, male, copulatory apparatus, Peronia madagascariensis A Madagascar, [5501] (MNHN-IM- 2009 - 16392) B South Africa, [5841] (NMSA W 7547). Scale bars: 5 mm (A), 2 mm (B). Abbreviations: ag accessory penial gland, dd deferent duct, ms muscular sac, ps penial sheath, rm retractor muscle, v vestibule." figureDoi="10.3897/zookeys.972.52853.figure23" httpUri="https://binary.pensoft.net/fig/458323" pageId="0" pageNumber="1">, 23</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F26" captionText="Figure 24. Penial hooks, Peronia madagascariensis, Madagascar A, C [5500] (MNHN-IM- 2009 - 16391) B, D [5504] (MNHN-IM- 2009 - 16412) E [5506] (MNHN-IM- 2009 - 16418) F [5501] (MNHN-IM- 2009 - 16392). Scale bars: 60 μm (A), 100 μm (B), 20 μm (C, D), 40 μm (E), 10 μm (F)." figureDoi="10.3897/zookeys.972.52853.figure24" httpUri="https://binary.pensoft.net/fig/458324" pageId="0" pageNumber="1">, 24</figureCitation>
<figureCitation captionStart="Figure 25" captionStartId="F27" captionText="Figure 25. Accessory penial gland spine, Peronia madagascariensis, Madagascar A, E [5500] (MNHN-IM- 2009 - 16391) B, F [5502] (MNHN-IM- 2009 - 16393) C, G [5504] (MNHN-IM- 2009 - 16412) D, H [5506] (MNHN-IM- 2009 - 16418). Scale bars: 400 μm (A-D), 20 μm (E-H)." figureDoi="10.3897/zookeys.972.52853.figure25" httpUri="https://binary.pensoft.net/fig/458325" pageId="0" pageNumber="1">, 25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">Paraperonia madagascariensis</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a: 199, fig. 15.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia jousseaumei" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">Paraperonia jousseaumei</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a: 198, figs 12-14. Syn. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
(
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia madagascariensis</emphasis>
</taxonomicName>
). Madagascar • holotype, by monotypy, 40/40 mm; Fort Dauphin [Taolagnaro]; 1932; 
<normalizedToken originalValue="Décary">Decary</normalizedToken>
leg.; MNHN-IM-2000-33680. Originally, no jar clearly labeled as the type material of 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
was found at the MNHN, but the holotype could be traced back. The original description of 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
is based on a single individual (40/38 mm) from Fort-Dauphin collected by 
<normalizedToken originalValue="Décary">Decary</normalizedToken>
(the French botanist Raymond 
<normalizedToken originalValue="Décary">Decary</normalizedToken>
[1891-1973]) in 1932. Only one old jar was found at the MNHN with a specimen collected from Fort-Dauphin (MNHN-IM-2000-33680). The information on the label (specimen collected by 
<normalizedToken originalValue="Décary">Decary</normalizedToken>
in 1932) matches the information provided in 
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
original description of 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
, and even the specimen size matches. Therefore, that specimen is considered to be the holotype by monotypy of 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
. The holotype was dissected by 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
. The radula, the posterior (hermaphroditic) reproductive parts, and the male parts are all missing. The intestinal loops are of type V (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F23" captionText="Figure 21. Peronia madagascariensis A-D digestive system, dorsal view, with intestinal loops of type V E posterior, hermaphroditic (female) reproductive system A holotype, Paraperonia madagascariensis, Madagascar (MNHN-IM- 2000 - 33680) B paralectotype, Paraperonia gondwanae, Mumbai, western India (MNHN-IM- 2000 - 33682) C paralectotype, Paraperonia gondwanae, Red Sea (MNHN-IM- 2000 - 33683) D Madagascar, [5501] (MNHN-IM- 2009 - 16392) E same as D. Scale bars: 5 mm (A-D), 2 mm (E). Abbreviations: dd deferent duct, ddg dorsal digestive gland, fgm female gland mass, hg hermaphroditic gland, i intestine, ov oviduct, pdg posterior digestive gland, rs receptaculum seminis, sp spermatheca, st stomach." figureDoi="10.3897/zookeys.972.52853.figure21" httpUri="https://binary.pensoft.net/fig/458321" pageId="0" pageNumber="1">21A</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.972.52853.figure21" httpUri="https://binary.pensoft.net/fig/458321" pageId="0" pageNumber="1" start="Figure 21" startId="F23">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 21.</emphasis>
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
digestive system, dorsal view, with intestinal loops of type V 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
posterior, hermaphroditic (female) reproductive system 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
holotype, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia madagascariensis</emphasis>
</taxonomicName>
, Madagascar (MNHN-IM-2000-33680) 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
paralectotype, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia gondwanae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia gondwanae</emphasis>
</taxonomicName>
, Mumbai, western India (MNHN-IM-2000-33682) 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
paralectotype, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia gondwanae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia gondwanae</emphasis>
</taxonomicName>
, Red Sea (MNHN-IM-2000-33683) 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
Madagascar, [5501] (MNHN-IM-2009-16392) 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
same as D. Scale bars: 5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
), 2 mm (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
). Abbreviations: dd deferent duct, ddg dorsal digestive gland, fgm female gland mass, hg hermaphroditic gland, i intestine, ov oviduct, pdg posterior digestive gland, rs receptaculum seminis, sp spermatheca, st stomach.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
(
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia jousseaumei" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia jousseaumei</emphasis>
</taxonomicName>
). The type material of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia jousseaumei" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia jousseaumei</emphasis>
</taxonomicName>
could not be located at the MNHN. The original description of 
<taxonomicName lsidName="P. jousseaumei" pageId="0" pageNumber="1" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">P. jousseaumei</emphasis>
</taxonomicName>
was based on ten individuals (45/38 to 40/30 mm) from the Red Sea (&quot;Mer Rouge&quot;) collected by Jousseaume in 1892. Only two old jars were found at the MNHN with that collecting information. One of them contains specimens that are part of the type series of 
<taxonomicName lsidName="P. gondwanae" pageId="0" pageNumber="1" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">P. gondwanae</emphasis>
</taxonomicName>
because the specific name &quot; 
<taxonomicName lsidName="gondwanae" pageId="0" pageNumber="1" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">gondwanae</emphasis>
</taxonomicName>
&quot; is written on an old label (MNHN-IM-2000-33683). The three labels of the other jar (MNHN-IM-2014-7993) say: &quot;Peronia Mer Rouge Mr Jousseaume n°15, 1892,&quot; &quot;Oncidium [written over 
<normalizedToken originalValue="“Oncidiella”">&quot;Oncidiella&quot;</normalizedToken>
] peronii Cuvier Mer Rouge M. Jousseaume n°15-1892,&quot; and, for unknown reasons, 
<normalizedToken originalValue="“60.”">&quot;60.&quot;</normalizedToken>
This jar contains six specimens of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
, from 60/45 to 25/15 mm, two of which were dissected, possibly by 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
. The intestinal loops of the two dissected specimens are of type I and thus are not in agreement with 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a
</bibRefCitation>
: fig. 12) original illustration of the intestinal loops of type V in 
<taxonomicName lsidName="P. jousseaumei" pageId="0" pageNumber="1" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">P. jousseaumei</emphasis>
</taxonomicName>
. Also, the sizes and the number of individuals do not match the original description of 
<taxonomicName lsidName="P. jousseaumei" pageId="0" pageNumber="1" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">P. jousseaumei</emphasis>
</taxonomicName>
. Those specimens could possibly be some of the eight non-type specimens that 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 190) mentioned in his re-description of 
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
collected by Jousseaume from the Red Sea (&quot;Mer Rouge&quot;) in 
<normalizedToken originalValue="“1852”">&quot;1852&quot;</normalizedToken>
(likely a mistake for 1892). Given that 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
does not specify their size, it is not possible to know to what species 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
thought those specimens belong exactly (MNHN-IM-2014-7993).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
South Africa • 2 specimens 35/23 mm [5841] and 18/13 mm [5842]; KwaZulu-Natal, Durban, Treasure Beach; 
<geoCoordinate degrees="29" direction="south" minutes="57.294" orientation="latitude" precision="1" value="-29.9549">29°57.294'S</geoCoordinate>
, 
<geoCoordinate degrees="30" direction="east" minutes="59.514" orientation="longitude" precision="1" value="30.9919">30°59.514'E</geoCoordinate>
; 18 Nov 2010; D Herbert and L Davis leg.; rocky intertidal zone; NMSA W7547.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Mozambique • 1 specimen 42/37 mm [735]; Cabo Delgado Province, Pemba, Wimbi Beach, Pemba Beach Hotel; 
<geoCoordinate degrees="12" direction="south" minutes="58" orientation="latitude" precision="925" value="-12.966666">12°58'S</geoCoordinate>
, 
<geoCoordinate degrees="40" direction="east" minutes="32" orientation="longitude" precision="925" value="40.533333">40°32'E</geoCoordinate>
; 14 Jul 2006; DG Reid leg.; on shady rock at base of limestone cliff, in upper eulittoral behind intertidal platform; NHMUK 20060414.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Madagascar • 1 specimen 55/40 mm [5500]; Ambatobe, 
<normalizedToken originalValue="près">pres</normalizedToken>
Soamanitse; 
<geoCoordinate degrees="25" direction="south" minutes="27.4" orientation="latitude" precision="92" value="-25.456667">25°27.4'S</geoCoordinate>
, 
<geoCoordinate degrees="44" direction="east" minutes="57.4" orientation="longitude" precision="92" value="44.956665">44°57.4'E</geoCoordinate>
; 24 May &amp; 7 Jun 2010; MNHN Expedition Atimo Vatae leg.; st BM02, 0-1 m; MNHN-IM-2009-16391. • 1 specimen 40/35 mm [5504]; same collection data as for the preceding; MNHN-IM-2009-16412. • 1 specimen 40/30 mm [5501]; Ambatomainty; 
<geoCoordinate degrees="25" direction="south" minutes="26.3" orientation="latitude" precision="92" value="-25.438334">25°26.3'S</geoCoordinate>
, 
<geoCoordinate degrees="44" direction="east" minutes="56.5" orientation="longitude" precision="92" value="44.941666">44°56.5'E</geoCoordinate>
; 25 May 2010; MNHN Expedition Atimo Vatae leg.; st BM03, 0-1 m; MNHN-IM-2009-16392. • 1 specimen 55/40 mm [5502]; same collection data as for the preceding; MNHN-IM-2009-16393. • 1 specimen 55/40 mm [5503]; Ambatobe, Bavarama; 
<geoCoordinate degrees="25" direction="south" minutes="27.9" orientation="latitude" precision="92" value="-25.465">25°27.9'S</geoCoordinate>
, 
<geoCoordinate degrees="44" direction="east" minutes="57.6" orientation="longitude" precision="92" value="44.96">44°57.6'E</geoCoordinate>
; 28 &amp; 29 May 2010; MNHN Expedition Atimo Vatae leg.; st BM06, 0-1 m; MNHN-IM-2009-16396. • 1 specimen 40/35 mm [5506]; same collection data as for the preceding; MNHN-IM-2009-16418.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Oman • 1 specimen 10/7 [703]; Muscat, Cemetery Bay; 
<geoCoordinate degrees="23" direction="north" minutes="37.250" orientation="latitude" precision="1" value="23.620832">23°37.250'N</geoCoordinate>
, 
<geoCoordinate degrees="58" direction="east" minutes="36.016" orientation="longitude" precision="1" value="58.600266">58°36.016'E</geoCoordinate>
; 9 Feb 2004; G Paulay &amp; M Claereboudt leg.; coral community, reef slope, on ophiolitic bedrock and rubble; UF 332088.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material examined</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">(historical museum collections).</emphasis>
Oman • 3 specimens 80/60 mm; Qurm Beach, near Muscat; 
<geoCoordinate degrees="23" direction="north" minutes="37.56" orientation="latitude" precision="9" value="23.626">23°37.56'N</geoCoordinate>
, 
<geoCoordinate degrees="58" direction="east" minutes="28.86" orientation="longitude" precision="9" value="58.481">58°28.86'E</geoCoordinate>
; 26 Jan 2005; V Bonito, M Claereboudt &amp; G Paulay leg.; intertidal rocky shore; UF 368019.
</paragraph>
<paragraph pageId="0" pageNumber="1">Iran • 3 specimens 80/65 mm to 75/65 mm; Persian Gulf, Strait of Hormuz, Qeshm Island; 18 Apr 1937; G Thorson leg.; st 69; NHMD 635302.</paragraph>
<paragraph pageId="0" pageNumber="1">
Yemen • 1 specimen 55/55 mm; Socotra, off Quadub; 
<geoCoordinate degrees="12" direction="north" minutes="39.015" orientation="latitude" precision="1" value="12.65025">12°39.015'N</geoCoordinate>
, 
<geoCoordinate degrees="53" direction="east" minutes="55.730" orientation="longitude" precision="1" value="53.928833">53°55.730'E</geoCoordinate>
; 18 Mar 1999; Salim Al-Moghrabi (from N 
<normalizedToken originalValue="Yonow’s">Yonow's</normalizedToken>
personal collection) leg.; intertidal, ST-064 SAM-1; SMF 358305.
</paragraph>
<paragraph pageId="0" pageNumber="1">South Africa • 1 specimen 70/45 mm; Port Natal, Durban; 30S, 31E; Wahlberg leg.; littoral rocky bottom; SMNH 180711.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="genbank and bold sequences">
<paragraph pageId="0" pageNumber="1">GenBank and BOLD sequences.</paragraph>
<paragraph pageId="0" pageNumber="1">
One COI sequence was obtained from BOLD (LGEN099-14) for an individual identified as 
<taxonomicName authorityName="Cuvier" authorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium verruculatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium verruculatum</emphasis>
</taxonomicName>
and collected from Dwarka, Gujarat, on the western coast of India (ca. 22°N), which is the easternmost known locality for 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
. A second COI sequence was obtained from GenBank (LC027608) for an individual identified as 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
sp. and collected from the coast of Iran in the Persian Gulf. Both sequences were unpublished.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1">
(Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
). From South Africa to the Red Sea and western India (ca. 22°N): South Africa, Mozambique, Madagascar (type locality of 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
), Gulf of Oman, Iran (Strait of Hormuz), Yemen (Socotra), India (Mumbai, Gujarat), Red Sea (type locality of 
<taxonomicName lsidName="P. jousseaumei" pageId="0" pageNumber="1" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">P. jousseaumei</emphasis>
</taxonomicName>
). All records are new except for the type locality in Madagascar. 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
is, so far, not present in Mauritius.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
was named after its type locality, Madagascar. 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia jousseaumei" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia jousseaumei</emphasis>
</taxonomicName>
was named after 
<normalizedToken originalValue="Félix">Felix</normalizedToken>
Pierre Jousseaume [1835-1921], a medical doctor and malacologist who collected many specimens from the Red Sea preserved at the MNHN and which 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a)
</bibRefCitation>
studied for his monograph on onchidiids.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
is found in the rocky intertidal, like most other 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
slugs.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="color and morphology">
<paragraph pageId="0" pageNumber="1">Color and morphology.</paragraph>
<paragraph pageId="0" pageNumber="1">No picture of live animals was available. The color of preserved specimens is not different from other species (greyish brown and mottled with darker and lighter areas dorsally, and light brown greyish ventrally). The dorsal notum of live animals is covered by dozens of papillae of various sizes. In large individuals, dorsal papillae can be particularly tall (easily up to 4 mm), even in preserved specimens, and are evenly distributed over the entire notum. Preserved, they are very difficult to distinguish from retracted dorsal gills in the posterior half of the notum, but they are regular papillae with or without eyes. Some papillae bear black dorsal eyes at their tip. The number of papillae with dorsal eyes is variable (from 12 to 18). Dorsal gills seem taller and denser than in other species. The largest specimens in our fresh material are 55 mm long but two additional museum specimens are much longer (80 mm).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="digestive system">
<paragraph pageId="0" pageNumber="1">Digestive system</paragraph>
<paragraph pageId="0" pageNumber="1">
(Figs 
<figureCitation captionStart="Figure 21" captionStartId="F23" captionText="Figure 21. Peronia madagascariensis A-D digestive system, dorsal view, with intestinal loops of type V E posterior, hermaphroditic (female) reproductive system A holotype, Paraperonia madagascariensis, Madagascar (MNHN-IM- 2000 - 33680) B paralectotype, Paraperonia gondwanae, Mumbai, western India (MNHN-IM- 2000 - 33682) C paralectotype, Paraperonia gondwanae, Red Sea (MNHN-IM- 2000 - 33683) D Madagascar, [5501] (MNHN-IM- 2009 - 16392) E same as D. Scale bars: 5 mm (A-D), 2 mm (E). Abbreviations: dd deferent duct, ddg dorsal digestive gland, fgm female gland mass, hg hermaphroditic gland, i intestine, ov oviduct, pdg posterior digestive gland, rs receptaculum seminis, sp spermatheca, st stomach." figureDoi="10.3897/zookeys.972.52853.figure21" httpUri="https://binary.pensoft.net/fig/458321" pageId="0" pageNumber="1">21A-D</figureCitation>
, 
<figureCitation captionStart="Figure 22" captionStartId="F24" captionText="Figure 22. Radula, Peronia madagascariensis, South Africa, [5841] (NMSA W 7547) A right half rows of teeth B rachidian and innermost lateral teeth C rachidian and innermost lateral teeth D lateral teeth E lateral teeth, frontal view F outermost lateral teeth. Scale bars: 200 μm (A), 20 μm (B, E, F), 40 μm (C), 100 μm (D)." figureDoi="10.3897/zookeys.972.52853.figure22" httpUri="https://binary.pensoft.net/fig/458322" pageId="0" pageNumber="1">22</figureCitation>
). Examples of radular formulae are presented in Table 
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Radular formulae in Peronia species. Each formula follows the same format: number of rows x number of lateral teeth per left half row - 1 (rachidian tooth) - number of lateral teeth per right half row. Each DNA extraction number corresponds to one individual. The letter H next to an extraction number indicates a holotype." httpUri="http://table.plazi.org/id/F6A603768F442AB5CA201CDCEC46CA6A" pageId="0" pageNumber="1" tableUuid="F6A603768F442AB5CA201CDCEC46CA6A">5</tableCitation>
. The median cusp of the rachidian teeth is approximately 55 
<normalizedToken originalValue="μm">μm</normalizedToken>
long. The hook of the lateral teeth is approximately 100 to 130 
<normalizedToken originalValue="μm">μm</normalizedToken>
long. The intestinal loops are of type V.
</paragraph>
<caption doi="10.3897/zookeys.972.52853.figure22" httpUri="https://binary.pensoft.net/fig/458322" pageId="0" pageNumber="1" start="Figure 22" startId="F24">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 22.</emphasis>
Radula, 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
, South Africa, [5841] (NMSA W7547) 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
right half rows of teeth 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
rachidian and innermost lateral teeth 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
rachidian and innermost lateral teeth 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
lateral teeth 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
lateral teeth, frontal view 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
outermost lateral teeth. Scale bars: 200 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), 20 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">B, E, F</emphasis>
), 40 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), 100 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reproductive system">
<paragraph pageId="0" pageNumber="1">Reproductive system</paragraph>
<paragraph pageId="0" pageNumber="1">
(Figs 
<figureCitation captionStart="Figure 21" captionStartId="F23" captionText="Figure 21. Peronia madagascariensis A-D digestive system, dorsal view, with intestinal loops of type V E posterior, hermaphroditic (female) reproductive system A holotype, Paraperonia madagascariensis, Madagascar (MNHN-IM- 2000 - 33680) B paralectotype, Paraperonia gondwanae, Mumbai, western India (MNHN-IM- 2000 - 33682) C paralectotype, Paraperonia gondwanae, Red Sea (MNHN-IM- 2000 - 33683) D Madagascar, [5501] (MNHN-IM- 2009 - 16392) E same as D. Scale bars: 5 mm (A-D), 2 mm (E). Abbreviations: dd deferent duct, ddg dorsal digestive gland, fgm female gland mass, hg hermaphroditic gland, i intestine, ov oviduct, pdg posterior digestive gland, rs receptaculum seminis, sp spermatheca, st stomach." figureDoi="10.3897/zookeys.972.52853.figure21" httpUri="https://binary.pensoft.net/fig/458321" pageId="0" pageNumber="1">21E</figureCitation>
, 
<figureCitation captionStart="Figure 23" captionStartId="F25" captionText="Figure 23. Anterior, male, copulatory apparatus, Peronia madagascariensis A Madagascar, [5501] (MNHN-IM- 2009 - 16392) B South Africa, [5841] (NMSA W 7547). Scale bars: 5 mm (A), 2 mm (B). Abbreviations: ag accessory penial gland, dd deferent duct, ms muscular sac, ps penial sheath, rm retractor muscle, v vestibule." figureDoi="10.3897/zookeys.972.52853.figure23" httpUri="https://binary.pensoft.net/fig/458323" pageId="0" pageNumber="1">23</figureCitation>
- 
<figureCitation captionStart="Figure 25" captionStartId="F27" captionText="Figure 25. Accessory penial gland spine, Peronia madagascariensis, Madagascar A, E [5500] (MNHN-IM- 2009 - 16391) B, F [5502] (MNHN-IM- 2009 - 16393) C, G [5504] (MNHN-IM- 2009 - 16412) D, H [5506] (MNHN-IM- 2009 - 16418). Scale bars: 400 μm (A-D), 20 μm (E-H)." figureDoi="10.3897/zookeys.972.52853.figure25" httpUri="https://binary.pensoft.net/fig/458325" pageId="0" pageNumber="1">25</figureCitation>
). In the anterior (male) parts, the muscular sac of the accessory penial gland is less than 15 mm long. The hollow spine of the accessory penial gland is narrow, elongated, and straight or slightly curved, and its shape (including at its tip) varies between individuals. Its length ranges from 2 mm ([5502] MNHN-IM-2009-16393) to 2.4 mm ([5500] MNHN-IM-2009-16391). Its diameter at the conical base ranges from 200 to 230 
<normalizedToken originalValue="μm">μm</normalizedToken>
. Its diameter at the tip ranges from 70 to 80 
<normalizedToken originalValue="μm">μm</normalizedToken>
. The retractor muscle is shorter or longer than the penial sheath and inserts near the heart. Inside the penial sheath, the penis is a narrow, elongated, soft, hollow tube. Its distal end bears conical hooks which are less than 100 
<normalizedToken originalValue="μm">μm</normalizedToken>
long.
</paragraph>
<caption doi="10.3897/zookeys.972.52853.figure23" httpUri="https://binary.pensoft.net/fig/458323" pageId="0" pageNumber="1" start="Figure 23" startId="F25">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 23.</emphasis>
Anterior, male, copulatory apparatus, 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Madagascar, [5501] (MNHN-IM-2009-16392) 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
South Africa, [5841] (NMSA W7547). Scale bars: 5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), 2 mm (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
). Abbreviations: ag accessory penial gland, dd deferent duct, ms muscular sac, ps penial sheath, rm retractor muscle, v vestibule.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.972.52853.figure24" httpUri="https://binary.pensoft.net/fig/458324" pageId="0" pageNumber="1" start="Figure 24" startId="F26">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 24.</emphasis>
Penial hooks, 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
, Madagascar 
<emphasis bold="true" pageId="0" pageNumber="1">A, C</emphasis>
[5500] (MNHN-IM-2009-16391) 
<emphasis bold="true" pageId="0" pageNumber="1">B, D</emphasis>
[5504] (MNHN-IM-2009-16412) 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
[5506] (MNHN-IM-2009-16418) 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
[5501] (MNHN-IM-2009-16392). Scale bars: 60 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), 100 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), 20 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
), 40 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), 10 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.972.52853.figure25" httpUri="https://binary.pensoft.net/fig/458325" pageId="0" pageNumber="1" start="Figure 25" startId="F27">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 25.</emphasis>
Accessory penial gland spine, 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
, Madagascar 
<emphasis bold="true" pageId="0" pageNumber="1">A, E</emphasis>
[5500] (MNHN-IM-2009-16391) 
<emphasis bold="true" pageId="0" pageNumber="1">B, F</emphasis>
[5502] (MNHN-IM-2009-16393) 
<emphasis bold="true" pageId="0" pageNumber="1">C, G</emphasis>
[5504] (MNHN-IM-2009-16412) 
<emphasis bold="true" pageId="0" pageNumber="1">D, H</emphasis>
[5506] (MNHN-IM-2009-16418). Scale bars: 400 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
), 20 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">E-H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnostic features">
<paragraph pageId="0" pageNumber="1">Diagnostic features</paragraph>
<paragraph pageId="0" pageNumber="1">
(Table 
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
). 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
is characterized by a unique combination of two anatomical traits: intestinal loops of type V and a spine of the accessory penial gland longer than 2 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The name 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia madagascariensis</emphasis>
</taxonomicName>
clearly applies to a 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species because of the dorsal gills on the notum of the holotype. The holotype was entirely dissected by 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
. The radula, the posterior (hermaphroditic) reproductive parts, and the anterior copulatory apparatus are missing. The intestinal loops are of type V (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F23" captionText="Figure 21. Peronia madagascariensis A-D digestive system, dorsal view, with intestinal loops of type V E posterior, hermaphroditic (female) reproductive system A holotype, Paraperonia madagascariensis, Madagascar (MNHN-IM- 2000 - 33680) B paralectotype, Paraperonia gondwanae, Mumbai, western India (MNHN-IM- 2000 - 33682) C paralectotype, Paraperonia gondwanae, Red Sea (MNHN-IM- 2000 - 33683) D Madagascar, [5501] (MNHN-IM- 2009 - 16392) E same as D. Scale bars: 5 mm (A-D), 2 mm (E). Abbreviations: dd deferent duct, ddg dorsal digestive gland, fgm female gland mass, hg hermaphroditic gland, i intestine, ov oviduct, pdg posterior digestive gland, rs receptaculum seminis, sp spermatheca, st stomach." figureDoi="10.3897/zookeys.972.52853.figure21" httpUri="https://binary.pensoft.net/fig/458321" pageId="0" pageNumber="1">21A</figureCitation>
), as illustrated by 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: fig. 17). The name 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
applies to the species described here because it is, according to our molecular data, the only 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species with intestinal loops of type V along the eastern African coast, from South Africa to the Persian Gulf and western India, including Madagascar. Note that some of our fresh material was collected only 150 km east of the type locality in southern Madagascar. Some internal characters described by 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 199) could not be verified on the holotype because most internal parts are missing, but they are similar to the species described here. In particular, the length of the spine of the accessory penial gland (2 mm) is compatible with what was observed in our material.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Additional, non-type specimens were found in historical museum collections which could be identified as 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
due to the presence of intestinal loops of type V, from Oman (UF 368019), the Strait of Hormuz (NHMD 635302), and Socotra (SMF 358305). Those localities, however, are all already included within the known distribution of 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
based on our DNA sequences, as the Strait of Hormuz is very close to the Gulf of Oman. Finally, one of the &quot; 
<emphasis italics="true" pageId="0" pageNumber="1">a</emphasis>
&quot; paralectotypes of 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a
</bibRefCitation>
: 199) 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia gondwanae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia gondwanae</emphasis>
</taxonomicName>
from Bombay (MNHN-IM-2000-33682), with intestinal loops of type V (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F23" captionText="Figure 21. Peronia madagascariensis A-D digestive system, dorsal view, with intestinal loops of type V E posterior, hermaphroditic (female) reproductive system A holotype, Paraperonia madagascariensis, Madagascar (MNHN-IM- 2000 - 33680) B paralectotype, Paraperonia gondwanae, Mumbai, western India (MNHN-IM- 2000 - 33682) C paralectotype, Paraperonia gondwanae, Red Sea (MNHN-IM- 2000 - 33683) D Madagascar, [5501] (MNHN-IM- 2009 - 16392) E same as D. Scale bars: 5 mm (A-D), 2 mm (E). Abbreviations: dd deferent duct, ddg dorsal digestive gland, fgm female gland mass, hg hermaphroditic gland, i intestine, ov oviduct, pdg posterior digestive gland, rs receptaculum seminis, sp spermatheca, st stomach." figureDoi="10.3897/zookeys.972.52853.figure21" httpUri="https://binary.pensoft.net/fig/458321" pageId="0" pageNumber="1">21B</figureCitation>
), belongs to 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
. Note that two of those museum specimens are longer (80 mm) than our fresh material (less than 55 mm).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
slugs with intestinal loops of type V are without doubt present in the Red Sea. For instance, one of the &quot; 
<emphasis italics="true" pageId="0" pageNumber="1">c</emphasis>
&quot; paralectotypes of 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a
</bibRefCitation>
: 200) 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia gondwanae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia gondwanae</emphasis>
</taxonomicName>
from Suez (MNHN-IM-2000-33683) is characterized by intestinal loops of type V (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F23" captionText="Figure 21. Peronia madagascariensis A-D digestive system, dorsal view, with intestinal loops of type V E posterior, hermaphroditic (female) reproductive system A holotype, Paraperonia madagascariensis, Madagascar (MNHN-IM- 2000 - 33680) B paralectotype, Paraperonia gondwanae, Mumbai, western India (MNHN-IM- 2000 - 33682) C paralectotype, Paraperonia gondwanae, Red Sea (MNHN-IM- 2000 - 33683) D Madagascar, [5501] (MNHN-IM- 2009 - 16392) E same as D. Scale bars: 5 mm (A-D), 2 mm (E). Abbreviations: dd deferent duct, ddg dorsal digestive gland, fgm female gland mass, hg hermaphroditic gland, i intestine, ov oviduct, pdg posterior digestive gland, rs receptaculum seminis, sp spermatheca, st stomach." figureDoi="10.3897/zookeys.972.52853.figure21" httpUri="https://binary.pensoft.net/fig/458321" pageId="0" pageNumber="1">21C</figureCitation>
), which means that it does not belong to 
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(characterized by intestinal loops of type I). 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a)
</bibRefCitation>
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia jousseaumei" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia jousseaumei</emphasis>
</taxonomicName>
, with the Red Sea as type locality, is also characterized by intestinal loops of type V. Even though the type material of 
<taxonomicName lsidName="P. jousseaumei" pageId="0" pageNumber="1" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">P. jousseaumei</emphasis>
</taxonomicName>
could not be located at the MNHN, 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a
</bibRefCitation>
: fig. 12) drawing of the internal anatomy of 
<taxonomicName lsidName="P. jousseaumei" pageId="0" pageNumber="1" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">P. jousseaumei</emphasis>
</taxonomicName>
clearly illustrates intestinal loops of type V. Given that 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
is widespread from South Africa all the way to western India, including the Strait of Hormuz, it is accepted here that it also is distributed in the Red Sea. That, however, will still need to be confirmed with fresh material from both the Red Sea and the Gulf of Aden. If it appears that the populations of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
slugs with intestinal loops of type V from the Red Sea are a distinct species, then the name 
<taxonomicName lsidName="P. jousseaumei" pageId="0" pageNumber="1" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">P. jousseaumei</emphasis>
</taxonomicName>
could apply to them and be valid. Finally, given that 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
is present in the Strait of Hormuz, it most likely also is distributed in the rest of the Persian Gulf, which hopefully will be confirmed at some point with fresh material.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Even though the names 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia jousseaumei" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia jousseaumei</emphasis>
</taxonomicName>
were never used prior to the present contribution, they are not regarded as new combinations because 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
has already been regarded as a synonym of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
by 
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a065863" author="Britton, KM" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="1" pagination="179 - 191" refId="B27" refString="Britton, KM, 1984. The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera. Journal of Molluscan Studies 50: 179 - 191, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a065863" title="The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera." url="https://doi.org/10.1093/oxfordjournals.mollus.a065863" volume="50" year="1984">Britton (1984</bibRefCitation>
: 182) and because it has also been made clear that the genus 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
included all species of slugs with dorsal gills (e.g., 
<bibRefCitation DOI="https://doi.org/10.1080/00222933.2017.1347297" author="Dayrat, B" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="1" pagination="1851 - 1897" refId="B53" refString="Dayrat, B, Goulding, TC, Apte, D, Bhave, V, Ngo, XQ, 2017. A new genus and four new species of onchidiid slugs from South-East Asia (Mollusca: Gastropoda: Pulmonata: Onchidiidae). Journal of Natural History 51: 1851 - 1897, DOI: https://doi.org/10.1080/00222933.2017.1347297" title="A new genus and four new species of onchidiid slugs from South-East Asia (Mollusca: Gastropoda: Pulmonata: Onchidiidae)." url="https://doi.org/10.1080/00222933.2017.1347297" volume="51" year="2017">Dayrat et al. 2017</bibRefCitation>
: 1861).
</paragraph>
<paragraph pageId="0" pageNumber="1">
The specimen [703] from Oman was tentatively identified as 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
sp. 2 by 
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2011.02.014" author="Dayrat, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="1" pagination="425 - 437" refId="B50" refString="Dayrat, B, Conrad, M, Balayan, S, White, TR, Albrecht, C, Golding, R, Gomes, S, Harasewych, MG, Frias Martins, AM de, 2011. Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling. Molecular Phylogenetics and Evolution 59: 425 - 437, DOI: https://doi.org/10.1016/j.ympev.2011.02.014" title="Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling." url="https://doi.org/10.1016/j.ympev.2011.02.014" volume="59" year="2011">Dayrat et al. (2011)</bibRefCitation>
but it clearly belongs to 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic relationships between Peronia species based on concatenated mitochondrial COI and 16 S DNA sequences. Numbers by the branches are the bootstrap values (maximum likelihood analysis) and the posterior probabilities (Bayesian analysis). Only the values&gt; 50 % (ML) and&gt; 0.9 (Bayesian) are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 3 - 6." figureDoi="10.3897/zookeys.972.52853.figure2" httpUri="https://binary.pensoft.net/fig/458300" pageId="0" pageNumber="1">2</figureCitation>
). Also, note that its COI sequence was resubmitted to GenBank because the old one (GenBank HQ660044) was inaccurate. The specimen [735] from Mozambique was tentatively identified as 
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">Peronia cf. peronii</taxonomicName>
by 
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2011.02.014" author="Dayrat, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="1" pagination="425 - 437" refId="B50" refString="Dayrat, B, Conrad, M, Balayan, S, White, TR, Albrecht, C, Golding, R, Gomes, S, Harasewych, MG, Frias Martins, AM de, 2011. Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling. Molecular Phylogenetics and Evolution 59: 425 - 437, DOI: https://doi.org/10.1016/j.ympev.2011.02.014" title="Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling." url="https://doi.org/10.1016/j.ympev.2011.02.014" volume="59" year="2011">Dayrat et al. (2011)</bibRefCitation>
. This identification should be disregarded because the specimen [735] belongs to 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic relationships between Peronia species based on concatenated mitochondrial COI and 16 S DNA sequences. Numbers by the branches are the bootstrap values (maximum likelihood analysis) and the posterior probabilities (Bayesian analysis). Only the values&gt; 50 % (ML) and&gt; 0.9 (Bayesian) are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 3 - 6." figureDoi="10.3897/zookeys.972.52853.figure2" httpUri="https://binary.pensoft.net/fig/458300" pageId="0" pageNumber="1">2</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
A specimen from Durban (30°S), South Africa, preserved in Stockholm (SMNH 180711) identified as 
<taxonomicName lsidName="O. verruculatum" pageId="0" pageNumber="1" rank="species" species="verruculatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. verruculatum</emphasis>
</taxonomicName>
by 
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928</bibRefCitation>
: 44, 73) is identified here as 
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
because of its intestinal loops of type V (Table 
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
). Various records of 
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="O. savignyi" pageId="0" pageNumber="1" rank="species" species="savignyi">
<emphasis italics="true" pageId="0" pageNumber="1">O. savignyi</emphasis>
</taxonomicName>
, and 
<taxonomicName authorityName="Cuvier" authorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium verruculatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium verruculatum</emphasis>
</taxonomicName>
from Natal, South Africa (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.13936" author="Krauss, F" journalOrPublisher="Ebner &amp; Seubert, Stuttgart" pageId="0" pageNumber="1" refId="B106" refString="Krauss, F, 1848. Die Suedafrikanischen Mollusken. Ein Beitrag zur Kenntniss der Mollusken des Kap- und Natallandes und zur Geographischen Verbreitung derselben, mit Beschreibung und Abbildung der neuen Arten. Ebner &amp; Seubert, Stuttgart, DOI: https://doi.org/10.5962/bhl.title.13936" title="Die Suedafrikanischen Mollusken. Ein Beitrag zur Kenntniss der Mollusken des Kap- und Natallandes und zur Geographischen Verbreitung derselben, mit Beschreibung und Abbildung der neuen Arten." url="https://doi.org/10.5962/bhl.title.13936" year="1848">Krauss 1848</bibRefCitation>
: 72; 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12875" author="Sturany, R" journalOrPublisher="Kaiserlich-Koeniglichen Hof- und Staatsdruckerei, Wien" pageId="0" pageNumber="1" refId="B174" refString="Sturany, R, 1898. Catalog der bisher bekannt gewordenen Suedafrikanischen Land- und Suesswasser-Mollusken mit besonderer Beruecksichtigung des von Dr Penther gesammelten Materiales. Kaiserlich-Koeniglichen Hof- und Staatsdruckerei, Wien, DOI: https://doi.org/10.5962/bhl.title.12875" title="Catalog der bisher bekannt gewordenen Suedafrikanischen Land- und Suesswasser-Mollusken mit besonderer Beruecksichtigung des von Dr Penther gesammelten Materiales." url="https://doi.org/10.5962/bhl.title.12875" year="1898">Sturany 1898</bibRefCitation>
: 73; 
<bibRefCitation author="Collinge, WE" journalOrPublisher="Annals of the Natal Museum" pageId="0" pageNumber="1" pagination="159 - 174" publicationUrl="https://www.biodiversitylibrary.org/page/6322951" refId="B36" refString="Collinge, WE, 1910. The slugs of Natal. Annals of the Natal Museum 2: 159 - 174, https://www.biodiversitylibrary.org/page/6322951" title="The slugs of Natal." url="https://www.biodiversitylibrary.org/page/6322951" volume="2" year="1910">Collinge 1910</bibRefCitation>
: 171-172; 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11477" author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" pagination="59 - 306" refId="B37" refString="Connolly, M, 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306, DOI: https://doi.org/10.5962/bhl.title.11477" title="A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum." url="https://doi.org/10.5962/bhl.title.11477" volume="11" year="1912">Connolly 1912</bibRefCitation>
: 224-225, 
<bibRefCitation author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/40865351" refId="B38" refString="Connolly, M, 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1-660. [pls I-XIX] https://www.biodiversitylibrary.org/page/40865351" title="A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1 - 660. [pls I-XIX]" url="https://www.biodiversitylibrary.org/page/40865351" year="1939">1939</bibRefCitation>
: 454; Webb 1969) most likely are records of 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia madagascariensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia madagascariensis</emphasis>
</taxonomicName>
, although 
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #5) could also be present in northeastern South Africa because it is known in Maputo, southern Mozambique (ca. 26°S).
</paragraph>
</subSubSection>
</treatment>
</document>