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<document id="E7824A9A705F732763D16F86450015D4" ID-CLB-Dataset="8274" ID-DOI="10.11646/zootaxa.4860.3.4" ID-GBIF-Dataset="58ea28c2-03f1-4e50-b1f3-596aedd0d4e6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4414007" ID-ZooBank="5D5CA04F-E837-43F2-92B1-82B8C4B69F4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609704413559" checkinUser="plazi" docAuthor="Bayarsaikhan, Ulziijargal, Hirai, Norio, Černý, Karel, Kwon, Hyung-Wook &amp; Bae, Yang-Seop" docDate="2020" docId="9E18603A427F2F7FBCA28207FD931DD9" docLanguage="en" docName="zootaxa.4860.3.4.pdf" docOrigin="Zootaxa 4860 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diduga sphaeracephalus Bayarsaikhan &amp; Hirai &amp; Černý &amp; Kwon &amp; Bae 2020, n. sp." docType="treatment" docVersion="8" lastPageNumber="395" masterDocId="62211842427E2F7DBC358600FFC91D21" masterDocTitle="A new species and four new records of Diduga Moore (Lepidoptera, Erebidae Arctiinae) from Thailand" masterLastPageNumber="400" masterPageNumber="393" pageNumber="394" updateTime="1698851989482" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2C0CB7E765C3F79C0C4E1CDAD80CBCA1">A new species and four new records of Diduga Moore (Lepidoptera, Erebidae Arctiinae) from Thailand</mods:title>
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<mods:namePart id="471B474F734DDC239427FC594453C66F">Bayarsaikhan, Ulziijargal</mods:namePart>
<mods:nameIdentifier id="B389AA74CC8FBFC12248FE067AB53671" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="32A14590737293B88EED82840BA1CAF5">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Ko- rea. baeys @ inu. ac. kr; https: // orcid. org / 0000 - 0001 - 7356 - 5633</mods:affiliation>
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<mods:namePart id="9692D5C027310EBB83159C4F715A3701">Hirai, Norio</mods:namePart>
<mods:nameIdentifier id="14F5C8EFC1976DA37B449AAF021273F6" type="ORCID">0000-0003-1674-7443</mods:nameIdentifier>
<mods:affiliation id="F54E4609ADC24B41DEFB1307615CDEB7">Graduate School of Life and Environmental Sciences, Osaka Prefecture University, Osaka, 599 ˗ 8531, Japan. n _ hirai @ envi. osakafu-u. ac. jp; https: // orcid. org / 0000 - 0003 - 1674 - 7443</mods:affiliation>
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<mods:namePart id="69F92F738AA29078845D49D9D6D0E1ED">Černý, Karel</mods:namePart>
<mods:affiliation id="1FDFB2C59979F21207E6D116AF6FE19B">Tiergartenstrasse 27, A- 6020 Innsbruck, Austria.</mods:affiliation>
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<mods:namePart id="8DC0981ADDE142956C826E7F8CD58F17">Kwon, Hyung-Wook</mods:namePart>
<mods:affiliation id="8F77BC0035539C54AABD416CFCCAD99B">Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, South Korea.</mods:affiliation>
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<mods:namePart id="EEC5E5418CCD7DA7CF85BFD5B07FFC09">Bae, Yang-Seop</mods:namePart>
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<mods:affiliation id="46C0DC588D172BE4EC473A232B15ABFB">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Ko- rea. baeys @ inu. ac. kr; https: // orcid. org / 0000 - 0001 - 7356 - 5633 &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, South Korea.</mods:affiliation>
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<mods:date id="49788F2358BDB43B9D61F407FFB60F81">2020</mods:date>
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<treatment id="9E18603A427F2F7FBCA28207FD931DD9" ID-DOI="http://doi.org/10.5281/zenodo.4536024" ID-GBIF-Taxon="177319694" ID-Zenodo-Dep="4536024" LSID="urn:lsid:plazi:treatment:9E18603A427F2F7FBCA28207FD931DD9" httpUri="http://treatment.plazi.org/id/9E18603A427F2F7FBCA28207FD931DD9" lastPageId="2" lastPageNumber="395" pageId="1" pageNumber="394">
<subSubSection id="5EAB82A7427F2F7CBCA28207FCE81900" box="[151,801,1031,1058]" pageId="1" pageNumber="394" type="nomenclature">
<paragraph id="160ED12C427F2F7CBCA28207FCE81900" blockId="1.[151,801,1031,1094]" box="[151,801,1031,1058]" pageId="1" pageNumber="394">
<heading id="4D466640427F2F7CBCA28207FCE81900" bold="true" box="[151,801,1031,1058]" fontSize="11" level="1" pageId="1" pageNumber="394" reason="1">
<emphasis id="24C50D3E427F2F7CBCA28207FCE81900" bold="true" box="[151,801,1031,1058]" pageId="1" pageNumber="394">
<taxonomicName id="D1B1AAAF427F2F7CBCA28207FD1B1900" authority="Bayarsaikhan &amp; Bae" authorityName="Bayarsaikhan &amp; Hirai &amp; Černý &amp; Kwon &amp; Bae" authorityYear="2020" box="[151,722,1031,1058]" class="Insecta" family="Arctiidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="sphaeracephalus" status="sp. nov.">
<emphasis id="24C50D3E427F2F7CBCA28207FE0D1903" bold="true" box="[151,452,1031,1058]" italics="true" pageId="1" pageNumber="394">Diduga sphaeracephalus</emphasis>
Bayarsaikhan &amp; Bae
</taxonomicName>
,
<taxonomicNameLabel id="3FF6B045427F2F7CBEEA8207FCE81900" box="[735,801,1031,1057]" pageId="1" pageNumber="394" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="5EAB82A7427F2F7CBCA2822BFE6B1967" box="[151,418,1067,1094]" pageId="1" pageNumber="394" type="description">
<paragraph id="160ED12C427F2F7CBCA2822BFE6B1967" blockId="1.[151,801,1031,1094]" box="[151,418,1067,1094]" pageId="1" pageNumber="394">
(
<figureCitation id="8E8ACDA9427F2F7CBCAA822BFED41964" box="[159,285,1067,1093]" captionStart="FIGURES 17" captionStartId="3.[151,264,1840,1864]" captionTargetBox="[166,1434,185,1791]" captionTargetId="figure-19@3.[151,1436,179,1815]" captionTargetPageId="3" captionText="FIGURES 17. Adults of Diduga from Thailand (Coll. of OPU and CKC). 1a. D. sphaeracephalus n. sp., male (holotype, OPU˗037(INU˗10280T)). 1b. ditto, female (paratype, OPU˗028(INU˗10281T)). 2a. D. albicosta, male (INU(CKC)˗10369T). 2b. ditto, female (OPU˗036(INU˗10273T)). 3a. D. trichophora, male (INU(CKC)˗10372T). 3b. ditto, female (INU(CKC)˗10366T). 4. D. alternota, male (INU(CKC)˗10374T). 5. D. bispinosa, male (OPU˗037(INU˗10282T)). 6. D. dubatolovi, male (OPU˗029(INU˗10278T)). 7. D. allodubatolovi, male (OPU˗038(INU˗10275T))." figureDoi="http://doi.org/10.5281/zenodo.4414009" httpUri="https://zenodo.org/record/4414009/files/figure.png" pageId="1" pageNumber="394">Figs 1a, 1b</figureCitation>
,
<figureCitation id="8E8ACDA9427F2F7CBD1F822BFEB91964" box="[298,368,1067,1093]" captionStart="FIGURES 814" captionStartId="4.[151,264,1840,1864]" captionTargetBox="[162,1426,191,1795]" captionTargetId="figure-29@4.[151,1436,181,1816]" captionTargetPageId="4" captionText="FIGURES 814. Male genitalia of Diduga from Thailand (Coll. of OPU and CKC). 8a. D. sphaeracephalus n. sp., holotype (OPU˗037(INU˗10280T). 8b. Apical dentated plate of aedeagus vesica, holotype (OPU˗037(INU˗10280T)). 9. D. albicosta (INU(CKC)˗10369T). 10. D. trichophora (INU(CKC)˗10365T). 11. D. alternota (INU(CKC)˗10374T). 12. D. bispinosa (OPU˗027(INU˗10282T)). 13. D. dubatolovi (OPU˗029(INU˗10278T)). 14. D. allodubatolovi (OPU˗038(INU˗10275T))." figureDoi="http://doi.org/10.5281/zenodo.4414013" httpUri="https://zenodo.org/record/4414013/files/figure.png" pageId="1" pageNumber="394">8a, 8b</figureCitation>
,
<figureCitation id="8E8ACDA9427F2F7CBD48822BFE531967" box="[381,410,1067,1094]" captionStart="FIGURES 1517" captionStartId="6.[151,264,1187,1211]" captionTargetBox="[163,1420,199,1145]" captionTargetId="figure-29@6.[151,1436,181,1164]" captionTargetPageId="6" captionText="FIGURES 1517. Female genitalia of Diduga from Thailand (Coll. of OPU and CKC). 15. D. sphaeracephalus n. sp., paratype (OPU˗028(INU˗10281T)). 16. D. albicosta (OPU˗036(INU˗10273T)). 17. D. trichophora (INU(CKC)˗10371T)." figureDoi="http://doi.org/10.5281/zenodo.4414015" httpUri="https://zenodo.org/record/4414015/files/figure.png" pageId="1" pageNumber="394">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5EAB82A7427F2F7CBCA28273FBC519F4" pageId="1" pageNumber="394" type="materials_examined">
<paragraph id="160ED12C427F2F7CBCA28273FBC519F4" blockId="1.[151,1437,1139,2030]" pageId="1" pageNumber="394">
<emphasis id="24C50D3E427F2F7CBCA28273FE9B19AC" bold="true" box="[151,338,1139,1165]" pageId="1" pageNumber="394">Type materials.</emphasis>
<materialsCitation id="A6D9DB71427F2F7CBD6F8273FBFF1990" ID-GBIF-Occurrence="3034041301" collectingDate="1981-08-14" country="Thailand" location="Kaosoi Dao" pageId="1" pageNumber="394" specimenCount="1" specimenCount-male="1" stateProvince="Chanthaburi" typeStatus="holotype">
<emphasis id="24C50D3E427F2F7CBD6F8273FE1919AC" bold="true" box="[346,464,1139,1165]" pageId="1" pageNumber="394">
<typeStatus id="C90A6F8E427F2F7CBD6F8273FE0219AC" box="[346,459,1139,1165]" pageId="1" pageNumber="394" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="00B71AA5427F2F7CBDEC8273FE3719AC" box="[473,510,1139,1165]" pageId="1" pageNumber="394" type="male">1♂</specimenCount>
,
<collectingCountry id="6EA691BC427F2F7CBE3E8273FDB819AC" box="[523,625,1139,1165]" name="Thailand" pageId="1" pageNumber="394">Thailand</collectingCountry>
,
<collectingRegion id="D4751FCE427F2F7CBE4A8273FD3719AC" box="[639,766,1139,1165]" country="Thailand" name="Chanthaburi" pageId="1" pageNumber="394">Chantaburi</collectingRegion>
,
<location id="136E87F7427F2F7CBF3E8273FC5D19AF" LSID="urn:lsid:plazi:treatment:9E18603A427F2F7FBCA28207FD931DD9:136E87F7427F2F7CBF3E8273FC5D19AF" box="[779,916,1139,1166]" country="Thailand" name="Kaosoi Dao" pageId="1" pageNumber="394" stateProvince="Chanthaburi">Kaosoi Dao</location>
,
<date id="620FF7EC427F2F7CBF948273FBFD19AC" box="[929,1076,1139,1165]" pageId="1" pageNumber="394" value="1981-08-14">
<collectingDate id="724B0E04427F2F7CBF948273FBFD19AC" box="[929,1076,1139,1165]" pageId="1" pageNumber="394" value="1981-08-14">14.VIII.1981</collectingDate>
</date>
(leg.
<collectorName id="BB44B4FA427F2F7CB84D8274FB3D19AC" box="[1144,1268,1139,1166]" pageId="1" pageNumber="394">H. Kuroko</collectorName>
, S.
<collectorName id="BB44B4FA427F2F7CB9158273FA5019AF" box="[1312,1433,1139,1166]" pageId="1" pageNumber="394">Moriuti, Y</collectorName>
. Arita &amp;
<collectorName id="BB44B4FA427F2F7CBCC48298FE4F1993" box="[241,390,1175,1202]" pageId="1" pageNumber="394">Y. Yoshiyasu</collectorName>
),
<collectorName id="BB44B4FA427F2F7CBDAD8297FDF41993" box="[408,573,1175,1202]" pageId="1" pageNumber="394">Gen. Slide No.</collectorName>
OPU˗037(INU˗10280)
<collectingCountry id="6EA691BC427F2F7CBF0A8297FC6C1993" box="[831,933,1175,1202]" name="Thailand" pageId="1" pageNumber="394">Thailand</collectingCountry>
(
<collectorName id="BB44B4FA427F2F7CBF868297FC251990" box="[947,1004,1175,1201]" pageId="1" pageNumber="394">Coll.</collectorName>
OPU)
</materialsCitation>
.
<emphasis id="24C50D3E427F2F7CB8758297FB7D1990" bold="true" box="[1088,1204,1175,1201]" pageId="1" pageNumber="394">
<typeStatus id="C90A6F8E427F2F7CB8758297FB661990" box="[1088,1199,1175,1201]" pageId="1" pageNumber="394" type="paratype">Paratype</typeStatus>
.
</emphasis>
(
<specimenCount id="00B71AA5427F2F7CB8F08297FB231993" box="[1221,1258,1175,1202]" pageId="1" pageNumber="394" type="female">1 ♀</specimenCount>
)
<materialsCitation id="A6D9DB71427F2F7CB8CE8297FA5C1993" ID-GBIF-Occurrence="3034041312" box="[1275,1429,1175,1202]" country="Thailand" location="Thailand" pageId="1" pageNumber="394" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="6EA691BC427F2F7CB8CE8297FAAA1990" box="[1275,1379,1175,1201]" name="Thailand" pageId="1" pageNumber="394">Thailand</collectingCountry>
:
<specimenCount id="00B71AA5427F2F7CB9598297FA5C1993" box="[1388,1429,1175,1202]" pageId="1" pageNumber="394" type="female">1 ♀</specimenCount>
</materialsCitation>
, same to
<typeStatus id="C90A6F8E427F2F7CBCC082BBFE9319F4" box="[245,346,1211,1237]" pageId="1" pageNumber="394" type="holotype">holotype</typeStatus>
, Gen. Slide No. OPU˗028(INU˗10281)
<collectingCountry id="6EA691BC427F2F7CBF3B82BBFCBD19F7" box="[782,884,1211,1238]" name="Thailand" pageId="1" pageNumber="394">Thailand</collectingCountry>
(Coll. OPU).
</paragraph>
</subSubSection>
<subSubSection id="5EAB82A7427F2F7CBCF282DFFC14188F" pageId="1" pageNumber="394" type="diagnosis">
<paragraph id="160ED12C427F2F7CBCF282DFFC14188F" blockId="1.[151,1437,1139,2030]" pageId="1" pageNumber="394">
<emphasis id="24C50D3E427F2F7CBCF282DFFE8B19D8" bold="true" box="[199,322,1247,1273]" pageId="1" pageNumber="394">Diagnosis.</emphasis>
By the wing pattern of this species, it is hardly distinguishable from many others of the genus
<emphasis id="24C50D3E427F2F7CB94282E0FF1E183C" italics="true" pageId="1" pageNumber="394">
<taxonomicName id="D1B1AAAF427F2F7CB94282E0FF1B183C" authorityName="Moore (" authorityYear="1887" class="Insecta" family="Arctiidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">Diduga</taxonomicName>
.
</emphasis>
However, in male genitalia, membranous, rounded apex of valva with a upper angles spine, and aedeagus vesica with a large stout spine˗shaped and dentate plate˗shaped cornuti; in female genitalia, weakly sclerotized os- tial plate irregularly V˗shaped, and acorn˗shaped corpus bursae heavily covered with small spines wholly, and with diverse sized, strongly sclerotized spines based to strongly sclerotized two bands in thickened top of acorn˗shaped corpus bursae separates this species well from all the other known
<taxonomicName id="D1B1AAAF427F2F7CBFB78393FC1F188C" authorityName="Moore (" authorityYear="1887" box="[898,982,1427,1453]" class="Insecta" family="Arctiidae" genus="Diduga" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="24C50D3E427F2F7CBFB78393FC1F188C" box="[898,982,1427,1453]" italics="true" pageId="1" pageNumber="394">Diduga</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="5EAB82A7427F2F7CBCF283B7FA541ACC" pageId="1" pageNumber="394" type="description">
<paragraph id="160ED12C427F2F7CBCF283B7FA541ACC" blockId="1.[151,1437,1139,2030]" pageId="1" pageNumber="394">
<emphasis id="24C50D3E427F2F7CBCF283B7FE9018F0" bold="true" box="[199,345,1463,1489]" pageId="1" pageNumber="394">Description.</emphasis>
<emphasis id="24C50D3E427F2F7CBD5583B7FE5418F0" box="[352,413,1463,1489]" italics="true" pageId="1" pageNumber="394">Adult</emphasis>
(
<figureCitation id="8E8ACDA9427F2F7CBD9E83B7FDE018F3" box="[427,553,1463,1490]" captionStart="FIGURES 17" captionStartId="3.[151,264,1840,1864]" captionTargetBox="[166,1434,185,1791]" captionTargetId="figure-19@3.[151,1436,179,1815]" captionTargetPageId="3" captionText="FIGURES 17. Adults of Diduga from Thailand (Coll. of OPU and CKC). 1a. D. sphaeracephalus n. sp., male (holotype, OPU˗037(INU˗10280T)). 1b. ditto, female (paratype, OPU˗028(INU˗10281T)). 2a. D. albicosta, male (INU(CKC)˗10369T). 2b. ditto, female (OPU˗036(INU˗10273T)). 3a. D. trichophora, male (INU(CKC)˗10372T). 3b. ditto, female (INU(CKC)˗10366T). 4. D. alternota, male (INU(CKC)˗10374T). 5. D. bispinosa, male (OPU˗037(INU˗10282T)). 6. D. dubatolovi, male (OPU˗029(INU˗10278T)). 7. D. allodubatolovi, male (OPU˗038(INU˗10275T))." figureDoi="http://doi.org/10.5281/zenodo.4414009" httpUri="https://zenodo.org/record/4414009/files/figure.png" pageId="1" pageNumber="394">Figs 1a, 1b</figureCitation>
). Length of forewing 5˗5.5 mm in both sexes. Head, patagium and tegula deep yellow. Thorax dark brown. Forewing ground color dark brown, with broad, deep yellow costal margin, which hind border broadly waved; with row of small, dark brown dots in costal and terminal area; cilia deep yellow. Hindwing ground color pale brown; cilia pale brown. Abdomen brown, except deep yellow ventral, with deep yellow anal tuft in male.
<emphasis id="24C50D3E427F2F7CBCCE8047FE691B40" box="[251,416,1607,1633]" italics="true" pageId="1" pageNumber="394">Male genitalia</emphasis>
(
<figureCitation id="8E8ACDA9427F2F7CBD858047FDFB1B43" box="[432,562,1607,1634]" captionStart="FIGURES 814" captionStartId="4.[151,264,1840,1864]" captionTargetBox="[162,1426,191,1795]" captionTargetId="figure-29@4.[151,1436,181,1816]" captionTargetPageId="4" captionText="FIGURES 814. Male genitalia of Diduga from Thailand (Coll. of OPU and CKC). 8a. D. sphaeracephalus n. sp., holotype (OPU˗037(INU˗10280T). 8b. Apical dentated plate of aedeagus vesica, holotype (OPU˗037(INU˗10280T)). 9. D. albicosta (INU(CKC)˗10369T). 10. D. trichophora (INU(CKC)˗10365T). 11. D. alternota (INU(CKC)˗10374T). 12. D. bispinosa (OPU˗027(INU˗10282T)). 13. D. dubatolovi (OPU˗029(INU˗10278T)). 14. D. allodubatolovi (OPU˗038(INU˗10275T))." figureDoi="http://doi.org/10.5281/zenodo.4414013" httpUri="https://zenodo.org/record/4414013/files/figure.png" pageId="1" pageNumber="394">Figs 8a, 8b</figureCitation>
). Uncus waved, heavily covered with setae, apex strongly hook˗shaped, with small apical spine, almost same length with tegumen. Tegumen triangular, weakly sclerotized. Valva stout, symmetric; weakly sclerotized costal margin of valva with roundly broadened in median area, and apex of costal margin finished to a strongly sclerotized, large (1/3 length of costal margin), triangular plate, which plate with a sclerotized apical spine; apex of valva rounded, membranous, with a weakly sclerotized, small, semicircular spur in medial area of membranous apex, and heavily covered with setae. Apex of juxta broadly heart˗shaped. Saccus roundly U˗shaped. Aedeagus stout, and vesica with a large (almost same length with aedeagus), stout spine˗shaped cornutus in apical area and a band˗shaped plate covered with diverse sized spines in opposite side (near apex of aedeagus).
<emphasis id="24C50D3E427F2F7CBCA28167FE911AA0" box="[151,344,1895,1921]" italics="true" pageId="1" pageNumber="394">Female genitalia</emphasis>
(
<figureCitation id="8E8ACDA9427F2F7CBD5D8167FE7F1AA3" box="[360,438,1895,1922]" captionStart="FIGURES 1517" captionStartId="6.[151,264,1187,1211]" captionTargetBox="[163,1420,199,1145]" captionTargetId="figure-29@6.[151,1436,181,1164]" captionTargetPageId="6" captionText="FIGURES 1517. Female genitalia of Diduga from Thailand (Coll. of OPU and CKC). 15. D. sphaeracephalus n. sp., paratype (OPU˗028(INU˗10281T)). 16. D. albicosta (OPU˗036(INU˗10273T)). 17. D. trichophora (INU(CKC)˗10371T)." figureDoi="http://doi.org/10.5281/zenodo.4414015" httpUri="https://zenodo.org/record/4414015/files/figure.png" pageId="1" pageNumber="394">Fig 15</figureCitation>
). Papillae anales weakly covered with setae. Ostial plate weakly sclerotized, irregularly V˗shaped. Ductus bursae tubular, weakly sclerotized in first 1/3 of length, and strongly sclerotized near cervix of corpus bursae. Corpus bursae membranous, acorn˗shaped, heavily covered with small spines wholly, except top of acorn, which top thickened, with diverse sized, strongly sclerotized spines based to strongly sclerotized two bands.
</paragraph>
</subSubSection>
<subSubSection id="5EAB82A7427C2F7FBCF28696FC531D90" box="[199,922,150,177]" pageId="2" pageNumber="395" type="distribution">
<paragraph id="160ED12C427C2F7FBCF28696FC531D90" blockId="2.[151,1436,150,249]" box="[199,922,150,177]" pageId="2" pageNumber="395">
<emphasis id="24C50D3E427C2F7FBCF28696FEA81D91" bold="true" box="[199,353,150,176]" pageId="2" pageNumber="395">Distribution.</emphasis>
<collectingCountry id="6EA691BC427C2F7FBD528696FE021D91" box="[359,459,150,176]" name="Thailand" pageId="2" pageNumber="395">Thailand</collectingCountry>
(
<collectingRegion id="D4751FCE427C2F7FBDEC8696FD9E1D91" box="[473,599,150,176]" country="Thailand" name="Chanthaburi" pageId="2" pageNumber="395">Chantaburi</collectingRegion>
and Kaosoi Dao Provinces).
</paragraph>
</subSubSection>
<subSubSection id="5EAB82A7427C2F7FBCF286BAFD931DD9" pageId="2" pageNumber="395" type="etymology">
<paragraph id="160ED12C427C2F7FBCF286BAFD931DD9" blockId="2.[151,1436,150,249]" pageId="2" pageNumber="395">
<emphasis id="24C50D3E427C2F7FBCF286BAFE871DF5" bold="true" box="[199,334,186,212]" pageId="2" pageNumber="395">Etymology.</emphasis>
The species name is derived from the Greek
<emphasis id="24C50D3E427C2F7FBF7686BAFC541DF5" box="[835,925,186,212]" italics="true" pageId="2" pageNumber="395">sphaera</emphasis>
(= ball) and
<emphasis id="24C50D3E427C2F7FB81986BAFB4C1DF5" box="[1068,1157,186,212]" italics="true" pageId="2" pageNumber="395">cephala</emphasis>
(= head), referring to the rounded apex of valva in male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>