350 lines
29 KiB
XML
350 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.151.50538" ID-GBIF-Dataset="a62873a2-c4e1-4c6d-9baa-5c5fb94e7348" ID-PMC="PMC7329918" ID-Pensoft-Pub="1314-2003-151-107" ID-Pensoft-UUID="42D1EB2DF0545744B92BFCCCB395C45B" ID-PubMed="32655282" ModsDocID="1314-2003-151-107" checkinTime="1593036331120" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Cuta-Alarcon, Lia E., Bravo-Pedraza, William J. & Soreng, Robert J." docDate="2020" docId="DED1619CE72D57139424FFF7019A50FD" docLanguage="en" docName="PhytoKeys 151: 107-160" docOrigin="PhytoKeys 151" docSource="http://dx.doi.org/10.3897/phytokeys.151.50538" docTitle="Agrostis mertensii Trin., Linnaea 10 (3): 302. 1836" docType="treatment" docVersion="4" id="42D1EB2DF0545744B92BFCCCB395C45B" lastPageNumber="107" masterDocId="42D1EB2DF0545744B92BFCCCB395C45B" masterDocTitle="Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species" masterLastPageNumber="160" masterPageNumber="107" pageNumber="107" updateTime="1668139837814" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Sylvester, Steven P.</mods:namePart>
|
||
<mods:affiliation>College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China & Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
|
||
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
|
||
<mods:nameIdentifier type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Cuta-Alarcon, Lia E.</mods:namePart>
|
||
<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Bravo-Pedraza, William J.</mods:namePart>
|
||
<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Soreng, Robert J.</mods:namePart>
|
||
<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
|
||
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>PhytoKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2020</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>151</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>107</mods:start>
|
||
<mods:end>160</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/phytokeys.151.50538</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1314-2003-151-107</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">42D1EB2DF0545744B92BFCCCB395C45B</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="164730846" LSID="urn:lsid:plazi:treatment:DED1619CE72D57139424FFF7019A50FD" httpUri="http://treatment.plazi.org/id/DED1619CE72D57139424FFF7019A50FD" lastPageNumber="107" pageId="0" pageNumber="107">
|
||
<subSubSection pageId="0" pageNumber="107" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<taxonomicName LSID="DED1619C-E72D-5713-9424-FFF7019A50FD" authority="Trin., Linnaea 10 (3): 302. 1836" authorityName="Trin., Linnaea 10 (3): 302." authorityYear="1836" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis mertensii" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="mertensii">Agrostis mertensii Trin., Linnaea 10 (3): 302. 1836</taxonomicName>
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Agrostis mertensii A spikelet, lateral view, with floret detached and raised above the glumes B whole plant. Images of Cuta-Alarcon 365 (FMB)." figureDoi="10.3897/phytokeys.151.50538.figure7" httpUri="https://binary.pensoft.net/fig/424845" pageId="0" pageNumber="107">Fig. 7</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="107" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis laxiflora subsp. var. var. mertensii" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="variety" species="laxiflora" subSpecies="var." variety="mertensii">Agrostis laxiflora var. mertensii</taxonomicName>
|
||
(Trin.) Griseb., Fl. Ross. 4 (13): 442. 1852.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis canina subsp. var. var. mertensii" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="variety" species="canina" subSpecies="var." variety="mertensii">Agrostis canina var. mertensii</taxonomicName>
|
||
(Trin.) Kuntze, Revis. Gen. Pl. 3[3]: 338. 1898.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
=
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis boliviana" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boliviana">Agrostis boliviana</taxonomicName>
|
||
Mez, Repert. Spec. Nov. Regni Veg. 18 (1-3): 1. 1922. Type: Bolivia. Pinos bei Tarija, 3000 m alt., 22 Jan. 1904, K.A.G. Fiebrig 2821 (lectotype, designated by
|
||
<bibRefCitation author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
||
<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
|
||
de Agrasar (2012
|
||
</bibRefCitation>
|
||
: 115): BAA (BAA00000014 [image!]); isolectotypes: BAA (BAA00000211 [image!] fragm. ex K), G (G00099216 [image!]), K (K000308377 [image!]), L (L0819974 [image!]; L0819973 [image!]); syntypes: Bolivia. Calderillo, 3000 m alt., Mar. 1904, K.A.G. Fiebrig 2905, BAA (BAA00000013 [image!]; BAA00000210 [image!]), E (E00373832 [image!]), G (G00099217 [image!]), GH (GH00221373 [image!]; GH00221374 [image!]), K (K000308376 [image!]), S (S05-10054 [image!])).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
=
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis gelida" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="gelida">Agrostis gelida</taxonomicName>
|
||
Trin.,
|
||
<normalizedToken originalValue="Mém">Mem</normalizedToken>
|
||
. Acad. Imp. Sci.
|
||
<normalizedToken originalValue="Saint-Pétersbourg">Saint-Petersbourg</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="Sér">Ser</normalizedToken>
|
||
. 6, Sci. Math., Seconde Pt. Sci. Nat. 6 (2, Bot.): 343. 1841. Type: Peru. Andibus de Pasco, [ad nives aeternas], E.F. Poeppig s.n. (holotype: LE-TRIN (LE-TRIN1613.01); isotype: US (US75321 fragm.)). Many other heterotypic synonyms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="107" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="107">Type.</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">USA. Alaska, 1829, D. Mertens s.n. (lectotype, designated by Widen (1971: 52): LE-TRIN (LE-TRIN1622.01, plant 1); isolectotypes: BAA (BAA00001355 [image!] fragm. ex LE-TRIN), S (S-G-263 [image!] fragm. ex LE-TRIN)).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="107" type="description">
|
||
<paragraph pageId="0" pageNumber="107">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<emphasis bold="true" pageId="0" pageNumber="107">Perennial herbs</emphasis>
|
||
, tufted, sometimes with short lateral tending rhizomes present or stooling with pseudostolons present.
|
||
<emphasis bold="true" pageId="0" pageNumber="107">Tillers</emphasis>
|
||
extravaginal, with cataphylls present.
|
||
<emphasis bold="true" pageId="0" pageNumber="107">Culms</emphasis>
|
||
20-65 cm tall, erect, ascendant, or geniculate at the base, delicate and slender, with 0-2(-3) nodes exerted at flowering, smooth.
|
||
<emphasis bold="true" pageId="0" pageNumber="107">Leaves</emphasis>
|
||
mostly cauline but with basal leaves present, glabrous, smooth throughout or finely scaberulous on the blade adaxial surface and margins;
|
||
<emphasis bold="true" pageId="0" pageNumber="107">ligules</emphasis>
|
||
(0.6-)2-6 mm long, membranous or scarious, obtuse to acuminate, not or slightly to moderately decurrent with the sheath, abaxial surface smooth or scaberulous;
|
||
<emphasis bold="true" pageId="0" pageNumber="107">blades</emphasis>
|
||
10-13 cm long, 1-3 mm wide when opened out, usually flat, soft and lax, less often involute and semi-rigid, tiller blades tending to be more involute and thin, semi-rigid, smooth throughout or finely scaberulous on the adaxial surface and/or margins, apices acute to acuminate.
|
||
<emphasis bold="true" pageId="0" pageNumber="107">Panicles</emphasis>
|
||
5-15 cm long, c. 4-16 cm wide, open when flowering and mature, congested when immature, lax, ovoid to pyramidal, slightly to usually greatly exerted from the basal foliage, lateral branches without spikelets in the lower
|
||
<normalizedToken originalValue="½">1/2</normalizedToken>
|
||
, long, ascending, spreading, to somewhat divergent and not held close to the central inflorescence axis at flowering or maturity, adpressed when young, central axis and panicle branches scabrous or smooth;
|
||
<emphasis bold="true" pageId="0" pageNumber="107">pedicels</emphasis>
|
||
1-6(-15) mm long, sometimes shorter to usually much longer than their spikelets, not or slightly dilated at their apex, smooth or scaberulous.
|
||
<emphasis bold="true" pageId="0" pageNumber="107">Spikelets</emphasis>
|
||
(not including awn) (2.2-)2.5-3(-4) mm long;
|
||
<emphasis bold="true" pageId="0" pageNumber="107">glumes</emphasis>
|
||
unequal, the lower slightly longer than the upper by up to c. 0.5 mm, 1-veined, keels smooth to scabrous throughout or more commonly scabrous in the upper half, apices acute;
|
||
<emphasis bold="true" pageId="0" pageNumber="107">floret</emphasis>
|
||
usually 2/3-3/4 the length of the glumes;
|
||
<emphasis bold="true" pageId="0" pageNumber="107">calluses</emphasis>
|
||
lightly pilose with 2 sparse lateral tufts of short hairs;
|
||
<emphasis bold="true" pageId="0" pageNumber="107">lemmas</emphasis>
|
||
1.7-2 mm long, glabrous, smooth, 5-veined, apex obtuse to acute, 2-dentate, 2-4 mucronate, awned, awn (1.8-)2.5-4.5 mm long, inserted dorsally in the middle or lower third, exerted from the glumes, flexuose to geniculate, twisted, persistant;
|
||
<emphasis bold="true" pageId="0" pageNumber="107">paleas</emphasis>
|
||
absent or 0.2-0.5 mm long, <
|
||
<normalizedToken originalValue="¼">1/4</normalizedToken>
|
||
the length of the lemma;
|
||
<emphasis bold="true" pageId="0" pageNumber="107">rachilla</emphasis>
|
||
absent;
|
||
<emphasis bold="true" pageId="0" pageNumber="107">anthers</emphasis>
|
||
0.5-1 mm long.
|
||
</paragraph>
|
||
<caption doi="10.3897/phytokeys.151.50538.figure7" httpUri="https://binary.pensoft.net/fig/424845" pageId="0" pageNumber="107" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<emphasis bold="true" pageId="0" pageNumber="107">Figure 7.</emphasis>
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis mertensii" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="mertensii">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis mertensii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="107">A</emphasis>
|
||
spikelet, lateral view, with floret detached and raised above the glumes
|
||
<emphasis bold="true" pageId="0" pageNumber="107">B</emphasis>
|
||
whole plant. Images of Cuta-Alarcon 365 (FMB).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="107" type="distribution">
|
||
<paragraph pageId="0" pageNumber="107">Distribution and ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
Exhibits a disjunct distribution, being found in very cold Arctic and sub-Arctic areas of the Northern Hemisphere (i.e. North America, Europe and Asia), and also in the high Andes of Venezuela, Colombia, Peru, Bolivia, Argentina and Chile (
|
||
<bibRefCitation author="Soreng, RJ" journalOrPublisher="Contributions from the United States National Herbarium" pageId="0" pageNumber="107" publicationUrl="http://www.tropicos.org/project/" refId="B34" refString="Soreng, RJ, Davidse, G, Peterson, PM, Zuloaga, FO, Judziewicz, EJ, Filgueiras, TS, Morrone, O, 2003 and onwards. . Catalogue of New World Grasses. http://www.tropicos.org/project/" title=". Catalogue of New World Grasses." url="http://www.tropicos.org/project/" year="2003 and onwards">Soreng et al. 2003 and onwards</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
||
<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
|
||
de Agrasar 2012
|
||
</bibRefCitation>
|
||
). It has most likely been introduced and naturalized in South America.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="107" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="107">Other specimens examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<emphasis bold="true" pageId="0" pageNumber="107">Colombia</emphasis>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="107">
|
||
<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
|
||
</emphasis>
|
||
: Munic. Arcabuco,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
El Valle, Vereda el Carmen,
|
||
<normalizedToken originalValue="páramo">paramo</normalizedToken>
|
||
muy
|
||
<normalizedToken originalValue="húmedo">humedo</normalizedToken>
|
||
dominado por
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chusquea" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Chusquea</emphasis>
|
||
</taxonomicName>
|
||
, pastoreo natural,
|
||
<geoCoordinate degrees="5.75425" direction="north" orientation="latitude" precision="1" value="5.75425">5.75425N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.3830278" direction="west" orientation="longitude" precision="1" value="-73.383026">73.3830278W</geoCoordinate>
|
||
, 3430 m alt., 15 Nov. 2017, S.P. Sylvester 3067b (COL, FMB, K, US); Munic. Duitama,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de Agueros,
|
||
<normalizedToken originalValue="vía">via</normalizedToken>
|
||
que conduce a vereda de Avendanos,
|
||
<geoCoordinate degrees="5.91464" direction="north" orientation="latitude" precision="1" value="5.91464">5.91464N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.07114" direction="west" orientation="longitude" precision="1" value="-73.07114">73.07114W</geoCoordinate>
|
||
, 3445 m alt., 28 Oct. 2017, S.P. Sylvester 3047 (FMB, K, US); Munic. Duitama,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de Agueros, semi disturbed
|
||
<normalizedToken originalValue="páramo">paramo</normalizedToken>
|
||
in Agueros reserve above ridge- along path running through reserve,
|
||
<geoCoordinate degrees="5.91653" direction="north" orientation="latitude" precision="1" value="5.91653">5.91653N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.07164" direction="west" orientation="longitude" precision="1" value="-73.07164">73.07164W</geoCoordinate>
|
||
, 3445 m alt., 29 Oct. 2017, S.P. Sylvester 3052 (FMB, K, UPTC, US); Munic. Duitama,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de La Rusia, Guanenta Alto
|
||
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
||
Fonce National Park, top of the ridge
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
|
||
Negra just below military base, ridge along the top of a steep rocky landscape, open
|
||
<normalizedToken originalValue="páramo">paramo</normalizedToken>
|
||
with
|
||
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia cachaluensis" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="cachaluensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Espeletia cachaluensis</emphasis>
|
||
</taxonomicName>
|
||
, no signs of grazing,
|
||
<geoCoordinate degrees="5.58389" direction="north" orientation="latitude" precision="1" value="5.58389">5.58389N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.053263" direction="west" orientation="longitude" precision="1" value="-73.05326">73.053263W</geoCoordinate>
|
||
, 3970 m alt., 21 Nov. 2017, M. Vorontsova 2198 (FMB, K, US); Munic. Duitama,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de La Rusia, Fina Betania, vereda El Carmen on other side of river,
|
||
<geoCoordinate degrees="5.95333" direction="north" orientation="latitude" precision="1" value="5.95333">5.95333N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.10864" direction="west" orientation="longitude" precision="1" value="-73.10864">73.10864W</geoCoordinate>
|
||
, 3445 m alt., 30 Oct. 2017, S.P. Sylvester 3060 (FMB, K, US); Munic. Duitama,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de La Rusia,
|
||
<normalizedToken originalValue="vía">via</normalizedToken>
|
||
que conduce a Vereda de Avendanos,
|
||
<normalizedToken originalValue="páramo">paramo</normalizedToken>
|
||
semi-perturbado, pastado por cabras y quemado regularmente hasta hace 1
|
||
<normalizedToken originalValue="año">ano</normalizedToken>
|
||
,
|
||
<geoCoordinate degrees="5.95011" direction="north" orientation="latitude" precision="1" value="5.95011">5.95011N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.09097" direction="west" orientation="longitude" precision="1" value="-73.09097">73.09097W</geoCoordinate>
|
||
, 3795 m alt., 1 Oct. 2017, S.P. Sylvester 3004 (COL, FMB, K, US); Munic. Duitama, valley between Guantiva and La Rusia, between Susacon and Onsaga, 2 km towards Onzaga,
|
||
<geoCoordinate degrees="6.10091" direction="north" orientation="latitude" precision="1" value="6.10091">6.10091N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="72.48263" direction="west" orientation="longitude" precision="1" value="-72.48263">72.48263W</geoCoordinate>
|
||
, 3292 m alt., 24 Nov. 2017, M. Vorontsova 2304a (FMB, K, US); Munic. Mongua,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de
|
||
<normalizedToken originalValue="Ocetá">Oceta</normalizedToken>
|
||
, Valle de Laguna Negra, pajonal frailejonal, terreno inclinado en medio del valle,
|
||
<geoCoordinate degrees="5.70664" direction="north" orientation="latitude" precision="1" value="5.70664">5.70664N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="72.80361" direction="west" orientation="longitude" precision="1" value="-72.80361">72.80361W</geoCoordinate>
|
||
, 3699 m alt., 29 Nov. 2017, L.E.
|
||
<normalizedToken originalValue="Cuta-Alarcón">Cuta-Alarcon</normalizedToken>
|
||
365 (FMB, K, UPTC, US); Munic. Mongui,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de
|
||
<normalizedToken originalValue="Ocetá">Oceta</normalizedToken>
|
||
, Sector La Pedrisca, Vereda Vallado, potrero con 20
|
||
<normalizedToken originalValue="años">anos</normalizedToken>
|
||
de abandono dominado por
|
||
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Senecio" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Senecio" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Senecio</emphasis>
|
||
</taxonomicName>
|
||
y
|
||
<taxonomicName authorityName="Cuatrec" authorityYear="1973" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia boyacensis" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Espeletia boyacensis</emphasis>
|
||
</taxonomicName>
|
||
, y
|
||
<normalizedToken originalValue="gramíneas">gramineas</normalizedToken>
|
||
<normalizedToken originalValue="exóticas">exoticas</normalizedToken>
|
||
,
|
||
<geoCoordinate degrees="5.69969" direction="north" orientation="latitude" precision="1" value="5.69969">5.69969N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="72.80892" direction="west" orientation="longitude" precision="1" value="-72.80892">72.80892W</geoCoordinate>
|
||
, 3751 m alt., 30 Nov. 2017, L.E.
|
||
<normalizedToken originalValue="Cuta-Alarcón">Cuta-Alarcon</normalizedToken>
|
||
377 (FMB, K, SI, US); Munic. Sotaquira, protected area of
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
El Valle, day 2 East ridge walk, somewhat disturbed patch of open
|
||
<normalizedToken originalValue="páramo">paramo</normalizedToken>
|
||
between patches of
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chusquea tessellata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tessellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Chusquea tessellata</emphasis>
|
||
</taxonomicName>
|
||
[Munro], dominated by
|
||
<taxonomicName authorityName="Cuatrec" authorityYear="1973" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia boyacensis" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Espeletia boyacensis</emphasis>
|
||
</taxonomicName>
|
||
, limited natural grazing,
|
||
<geoCoordinate degrees="5.44436" direction="north" orientation="latitude" precision="1" value="5.44436">5.44436N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.22089" direction="west" orientation="longitude" precision="1" value="-73.22089">73.22089W</geoCoordinate>
|
||
, 3717 m alt., 16 Nov. 2017, M. Vorontsova 2155 (FMB, K, US).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="107" type="notes">
|
||
<paragraph pageId="0" pageNumber="107">Notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">This species is highly variable in terms of its habit, the form of its leaf blades (flat or involute) and ligule (short and obtuse to long and acuminate), and the form of the panicle (panicle branches adpressed and congested when young while open when mature). The combination of open panicle (when mature), lemma with a dorsally inserted awn, and a minute or absent palea are diagnostic for this species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="107" type="similar species">
|
||
<paragraph pageId="0" pageNumber="107">Similar species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis perennans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis perennans</emphasis>
|
||
</taxonomicName>
|
||
s.l., which can be principally differentiated by the absence of an awn, or if awn present, this being inserted in the upper half of the dorsal surface of the lemma or subapically and to 0.5 mm long.
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis pittieri" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="pittieri">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis pittieri</emphasis>
|
||
</taxonomicName>
|
||
Hack., considered endemic to Costa Rica by
|
||
<bibRefCitation author="Pohl, RW" editor="Burger, WC" journalOrPublisher="Fieldiana Botany" pageId="0" pageNumber="107" pagination="1 - 608" refId="B27" refString="Pohl, RW, 1980. Family 15. Gramineae. In: Burger, WC, Ed., Flora Costaricensis. Fieldiana Botany 4: 1 - 608" title="Family 15. Gramineae." volume="4" volumeTitle="Flora Costaricensis." year="1980">Pohl (1980)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Morales-Quiros, JF" editor="Hammel, BE" journalOrPublisher="Monographs in Systematic Botany from the Missouri Botanical Garden 93 (3)" pageId="0" pageNumber="107" pagination="598 - 821" refId="B22" refString="Morales-Quiros, JF, 2003. Poaceae. In: Hammel, BE, Grayum, MH, Herrera-Mora, C, Zamora-Villalobos, N, Eds., Manual of Plants of Costa Rica. Monographs in Systematic Botany from the Missouri Botanical Garden 93(3): 598 - 821" title="Poaceae." volumeTitle="Manual of Plants of Costa Rica." year="2003">
|
||
<normalizedToken originalValue="Morales-Quirós">Morales-Quiros</normalizedToken>
|
||
(2003)
|
||
</bibRefCitation>
|
||
, also bears close similarity to
|
||
<taxonomicName lsidName="A. mertensii" pageId="0" pageNumber="107" rank="species" species="mertensii">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. mertensii</emphasis>
|
||
</taxonomicName>
|
||
and may be placed as a synonym of the latter in future research.
|
||
<bibRefCitation author="Hokche, O" journalOrPublisher="Fundacion Instituto Botanico de Venezuela, Caracas" pageId="0" pageNumber="107" refId="B16" refString="Hokche, O, Berry, PE, Huber, O [Eds], 2008. Nuevo Catalogo de la Flora Vascular de Venezuela. Fundacion Instituto Botanico de Venezuela, Caracas" title="Nuevo Catalogo de la Flora Vascular de Venezuela." year="2008">Hokche et al. (2008)</bibRefCitation>
|
||
and
|
||
<bibRefCitation DOI="https://doi.org/10.5479/si.19382812.100" author="Dorr, LJ" journalOrPublisher="Smithsonian Contributions to Botany" pageId="0" pageNumber="107" pagination="1 - 289" refId="B6" refString="Dorr, LJ, 2014. Flora of Guaramacal (Venezuela): Monocotyledons. Smithsonian Contributions to Botany 100: 1 - 289, DOI: https://doi.org/10.5479/si.19382812.100" title="Flora of Guaramacal (Venezuela): Monocotyledons." url="https://doi.org/10.5479/si.19382812.100" volume="100" year="2014">Dorr (2014)</bibRefCitation>
|
||
record
|
||
<taxonomicName lsidName="A. pittieri" pageId="0" pageNumber="107" rank="species" species="pittieri">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. pittieri</emphasis>
|
||
</taxonomicName>
|
||
for Venezuela, with
|
||
<bibRefCitation DOI="https://doi.org/10.5479/si.19382812.100" author="Dorr, LJ" journalOrPublisher="Smithsonian Contributions to Botany" pageId="0" pageNumber="107" pagination="1 - 289" refId="B6" refString="Dorr, LJ, 2014. Flora of Guaramacal (Venezuela): Monocotyledons. Smithsonian Contributions to Botany 100: 1 - 289, DOI: https://doi.org/10.5479/si.19382812.100" title="Flora of Guaramacal (Venezuela): Monocotyledons." url="https://doi.org/10.5479/si.19382812.100" volume="100" year="2014">Dorr (2014)</bibRefCitation>
|
||
stating
|
||
<taxonomicName lsidName="A. pittieri" pageId="0" pageNumber="107" rank="species" species="pittieri">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. pittieri</emphasis>
|
||
</taxonomicName>
|
||
can be tentatively differentiated by having linear and narrow panicles with green spikelets and lemmas 1.9-2 mm long, while
|
||
<taxonomicName lsidName="A. mertensii" pageId="0" pageNumber="107" rank="species" species="mertensii">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. mertensii</emphasis>
|
||
</taxonomicName>
|
||
has broadly ovate to lanceolate panicles with purple spikelets and lemmas 2-2.5 mm long. However, these
|
||
<normalizedToken originalValue="characters’">characters'</normalizedToken>
|
||
overlap in specimens studied from throughout the range of
|
||
<taxonomicName lsidName="A. mertensii" pageId="0" pageNumber="107" rank="species" species="mertensii">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. mertensii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |