treatments-xml/data/D9/78/E8/D978E8BF0BBE5144B397F4690B165C75.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.97.65280" ID-GBIF-Dataset="f2891081-3c0d-4218-8196-8a99a766de15" ID-GBIF-Taxon="182239841" ID-Pensoft-Pub="1860-0743-1-281" ID-Pensoft-UUID="D784822D3E055039B543B7FC864ADF5C" ID-ZooBank="BCA8CCABC5784FEBB053F483D53922AD" LSID="urn:lsid:zoobank.org:act:05EFF40C-9238-49B8-9D79-7986979F674D" ModsDocID="1860-0743-97-1-281" checkinTime="1621457838280" checkinUser="pensoft" docAuthor="Vecchi, Matteo &amp; Stec, Daniel" docDate="2021" docId="D978E8BF0BBE5144B397F4690B165C75" docLanguage="en" docName="ZoosystEvol 97(1): 281-306" docOrigin="Zoosystematics and Evolution 97 (1)" docPubDate="2021-05-19" docSource="http://dx.doi.org/10.3897/zse.97.65280" docTitle="Macrobiotus annewintersae Vecchi &amp; Stec 2021, sp. nov." docType="treatment" docUuid="05EFF40C-9238-49B8-9D79-7986979F674D" docUuidSource="ZooBank" docVersion="4" id="D784822D3E055039B543B7FC864ADF5C" lastPageNumber="281" masterDocId="D784822D3E055039B543B7FC864ADF5C" masterDocTitle="Integrative descriptions of two new Macrobiotus species (Tardigrada, Eutardigrada, Macrobiotidae) from Mississippi (USA) and Crete (Greece)" masterLastPageNumber="306" masterPageNumber="281" pageId="0" pageNumber="281" updateTime="1634711172996" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Integrative descriptions of two new Macrobiotus species (Tardigrada, Eutardigrada, Macrobiotidae) from Mississippi (USA) and Crete (Greece)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vecchi, Matteo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7995-6827</mods:nameIdentifier>
<mods:affiliation>Department of Biological and Environmental Science, University of Jyvaskyla, PO Box 35, FI- 40014 Jyvaeskylae, Finland</mods:affiliation>
<mods:nameIdentifier type="email">matteo.m.vecchi@jyu.fi</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stec, Daniel</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6876-0717</mods:nameIdentifier>
<mods:affiliation>Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Faculty of Biology, Jagiellonian University, Gronostajowa 9, 30 - 387 Krakow, Poland</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-19</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>97</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:extent unit="page">
<mods:start>281</mods:start>
<mods:end>306</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.97.65280</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.65280</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-1-281</mods:identifier>
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<treatment ID-GBIF-Taxon="182239841" LSID="urn:lsid:zoobank.org:act:05EFF40C-9238-49B8-9D79-7986979F674D" httpUri="http://treatment.plazi.org/id/D978E8BF0BBE5144B397F4690B165C75" lastPageNumber="281" pageId="0" pageNumber="281">
<subSubSection pageId="0" pageNumber="281" type="nomenclature">
<paragraph pageId="0" pageNumber="281">
<taxonomicName LSID="http://zoobank.org/05EFF40C-9238-49B8-9D79-7986979F674D" authority="Vecchi &amp; Stec" authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae" status="sp. nov.">Macrobiotus annewintersae Vecchi &amp; Stec</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="281">sp. nov.</taxonomicNameLabel>
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Measurements [in µm] of selected morphological structures of individuals of Macrobiotus annewintersae sp. nov. mounted in Hoyer's medium (N-number of specimens / structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD-standard deviation)." httpUri="http://table.plazi.org/id/A24819DFF5ED4F5CB2328EA4473EC8D2" pageId="0" pageNumber="281" tableUuid="A24819DFF5ED4F5CB2328EA4473EC8D2">Tables 3</tableCitation>
, 
<figureCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Measurements [in µm] of selected morphological structures of the eggs of Macrobiotus annewintersae sp. nov. mounted in Hoyer's medium (N-number of eggs / structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD-standard deviation)." pageId="0" pageNumber="281">4</figureCitation>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Macrobiotus annewintersae sp. nov. - habitus and cuticular pores: A. Dorso-ventral view of the body (Holotype ♀;, PCM); B, C. Cuticular pores on the dorsal part of the body under PCM and under SEM, respectively. Arrowheads indicate pores and empty arrows indicate places on dorsal cuticle without pores. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure1" httpUri="https://binary.pensoft.net/fig/544821" pageId="0" pageNumber="281">, Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Macrobiotus annewintersae sp. nov. - cuticular structures on legs: A. External granulation on leg III under PCM; B. A cuticular bulge (pulvinus) on the internal surface of leg III under PCM; C. Granulation on leg IV under PCM; D. External granulation on leg III under SEM; E. A cuticular bulge (pulvinus) on the internal surface of leg III under SEM. Filled flat arrowheads indicate the granulation patch, empty flat arrowheads indicate pulvinus and filled indented arrowheads indicate muscle attachments. C assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure2" httpUri="https://binary.pensoft.net/fig/544822" pageId="0" pageNumber="281">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Macrobiotus annewintersae sp. nov. - claws: A, B. Claws III and IV, respectively, under PCM; C, D. Claws III and IV, respectively, under SEM. Filled indented arrowheads indicate double muscle attachments under the claws, empty indented arrowheads indicate a faintly visible divided cuticular bar. A and B assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure3" httpUri="https://binary.pensoft.net/fig/544823" pageId="0" pageNumber="281">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrobiotus annewintersae sp. nov. - buccal apparatus and the oral cavity armature under PCM: A. Dorso-ventral view of the entire buccal apparatus; B, C. Oral cavity armature in dorsal and ventral view, respectively; D, E. Placoid morphology in dorsal and ventral view, respectively. Empty flat arrowheads indicate the second band of teeth, filled indented arrowheads indicate the third band of teeth in the oral cavity, and empty indented arrowheads indicate central constriction in the first macroplacoid and subterminal constriction in the second macroplacoid. A, D and E assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure4" httpUri="https://binary.pensoft.net/fig/544824" pageId="0" pageNumber="281">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Macrobiotus annewintersae sp. nov. - anterior view of the mouth opening under SEM. Filled flat arrowhead indicates the first band of teeth. Scale bar in μm." figureDoi="10.3897/zse.97.65280.figure5" httpUri="https://binary.pensoft.net/fig/544825" pageId="0" pageNumber="281">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Macrobiotus annewintersae sp. nov. - egg chorion morphology under PCM: A, B. Egg surface; C, D. Midsection of the processes. Filled flat arrowheads indicate bubble-like structures within tentacular arms in the distal portion of the egg processes and empty flat arrowheads indicate dark thickenings / striae on the egg surface between processes. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure6" httpUri="https://binary.pensoft.net/fig/544826" pageId="0" pageNumber="281">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Macrobiotus annewintersae sp. nov. - egg chorion morphology under SEM: A, B. Entire egg; C-E. Details of the egg processes and egg surface between them; F. Details of the tentacular arms in the distal portion of each egg process. Filled indented arrowheads indicate micropores and empty indented arrowheads indicate lobes in tentacular arms covered by micro-granulation. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure7" httpUri="https://binary.pensoft.net/fig/544827" pageId="0" pageNumber="281">, 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Macrobiotus annewintersae sp. nov. - reproduction: A. Female under PCM; B. Male under PCM; C. Male under SEM. Filled indented arrowhead indicates spermathecae filled with spermatozoa, empty indented arrowhead indicates male's testis, arrows indicate lateral gibbosities on legs IV and filled flat arrowhead indicates cuticular pore on the ventral side of the body. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure8" httpUri="https://binary.pensoft.net/fig/544828" pageId="0" pageNumber="281">, 8</figureCitation>
, Suppl. material 1
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="281" type="etymology">
<paragraph pageId="0" pageNumber="281">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="281">We dedicate this species to MV friend and colleague Dr. Anne Winters, evolutionary ecologist, who collected the sample in which the new species was found.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="281" type="materials_examined">
<paragraph pageId="0" pageNumber="281">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="281">
146 animals and 
<specimenCount type="egg">56 eggs</specimenCount>
. Specimens mounted on microscope slides in 
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
medium (93 animals + 
<specimenCount type="egg">38 eggs</specimenCount>
), fixed on SEM stubs (51+18), and processed for DNA sequencing (2+0).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="281" type="type locality">
<paragraph pageId="0" pageNumber="281">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="281">
<geoCoordinate degrees="32" direction="north" minutes="21" orientation="latitude" precision="15" seconds="05" value="32.351387">32°21'05&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="89" direction="west" minutes="56" orientation="longitude" precision="15" seconds="30" value="-89.941666">89°56'30&quot;W</geoCoordinate>
; 106 m asl: suburban area of Jackson, Mississippi, USA; mixed leaf litter on ground; coll. December 2019 by Anne Winters.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="281" type="type depositories">
<paragraph pageId="0" pageNumber="281">Type depositories.</paragraph>
<paragraph pageId="0" pageNumber="281">
Holotype ♀ (slide US.084.01 with 10 paratypes) and 63 paratypes (slides: US.084.*, where the asterisk can be substituted by any of the following numbers: 02-05) and 20 eggs (slides US.084.*: 06-08) are deposited at the Institute of Zoology and Biomedical Research, Jagiellonian University (Gronostajowa 9, 30-387, 
<normalizedToken originalValue="Kraków">Krakow</normalizedToken>
, Poland). Additional paratypes (71 animals + 29 eggs) (slides: S207_SL*: 1-15; SEM stubs: S207_Stub*:1-4) are deposited at the Department of Biological and Environmental Sciences, University of 
<normalizedToken originalValue="Jyväskylä">Jyvaeskylae</normalizedToken>
(Survontie 9C, 40500, 
<normalizedToken originalValue="Jyväskylä">Jyvaeskylae</normalizedToken>
, Finland).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="281" type="description">
<paragraph pageId="0" pageNumber="281">Description of the new species.</paragraph>
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" italics="true" pageId="0" pageNumber="281">Animals</emphasis>
(measurements and statistics in Table 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Measurements [in µm] of selected morphological structures of individuals of Macrobiotus annewintersae sp. nov. mounted in Hoyer's medium (N-number of specimens / structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD-standard deviation)." httpUri="http://table.plazi.org/id/A24819DFF5ED4F5CB2328EA4473EC8D2" pageId="0" pageNumber="281" tableUuid="A24819DFF5ED4F5CB2328EA4473EC8D2">3</tableCitation>
):
</paragraph>
<caption ID-Table-UUID="A24819DFF5ED4F5CB2328EA4473EC8D2" httpUri="http://table.plazi.org/id/A24819DFF5ED4F5CB2328EA4473EC8D2" pageId="0" pageNumber="281" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">Table 3.</emphasis>
Measurements [in 
<normalizedToken originalValue="µm">µm</normalizedToken>
] of selected morphological structures of individuals of 
<taxonomicName authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus annewintersae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="281">sp. nov.</emphasis>
mounted in 
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
medium (N-number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD-standard deviation).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="281">
<table pageId="0" pageNumber="281">
<tr pageId="0" pageNumber="281">
<th colspan="1" pageId="0" pageNumber="281" rowspan="2">Character</th>
<th colspan="1" pageId="0" pageNumber="281" rowspan="2">N</th>
<th colspan="6" pageId="0" pageNumber="281" rowspan="1">Range</th>
<th colspan="2" pageId="0" pageNumber="281" rowspan="1">Mean</th>
<th colspan="2" pageId="0" pageNumber="281" rowspan="1">Sd</th>
<th colspan="2" pageId="0" pageNumber="281" rowspan="1">Holotype</th>
</tr>
<tr pageId="0" pageNumber="281">
<th colspan="3" pageId="0" pageNumber="281" rowspan="1">
<normalizedToken originalValue="µm">µm</normalizedToken>
</th>
<th colspan="3" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">pt</emphasis>
</th>
<th colspan="1" pageId="0" pageNumber="281" rowspan="1">
<normalizedToken originalValue="µm">µm</normalizedToken>
</th>
<th colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">pt</emphasis>
</th>
<th colspan="1" pageId="0" pageNumber="281" rowspan="1">
<normalizedToken originalValue="µm">µm</normalizedToken>
</th>
<th colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">pt</emphasis>
</th>
<th colspan="1" pageId="0" pageNumber="281" rowspan="1">
<normalizedToken originalValue="µm">µm</normalizedToken>
</th>
<th colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">Pt</emphasis>
</th>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Body length</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">29</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">287</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">441</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">934</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1226</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">371</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1074</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">46</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">84</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">434</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1226</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Buccal tube</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Buccal tube length</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">27.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">40.4</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">34.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">35.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Stylet support insertion point</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">21.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">32.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">76.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">81.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">27.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">79.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">27.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">77.7</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Buccal tube external width</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">29</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">6.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">12.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">17.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">4.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">13.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">15.3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Buccal tube internal width</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">29</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">4.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">6.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">11.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">9.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.1</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">9.3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Ventral lamina length</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">22</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">16.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">26.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">49.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">64.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">20.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">58.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">3.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">21.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">61.9</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Placoid lengths</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Macroplacoid 1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">6.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">10.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">20.9</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">28.9</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">24.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">1.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">26.6</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Macroplacoid 2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">3.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">6.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">12.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">18.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">15.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">15.8</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Microplacoid</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">4.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">4.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">11.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">2.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">7.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">2.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">8.2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Macroplacoid row</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">26</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">10.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">17.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">38.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">49.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">14.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">43.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">2.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">16.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">46.9</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Placoid row</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">26</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">13.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">22.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">48.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">62.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">18.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">54.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">3.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">20.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">58.5</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Claw 1 heights</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">External primary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">24</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">11.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">22.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">30.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">27.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">2.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">10.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">29.4</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">External secondary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">22</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">18.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">24.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">21.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">2.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">24.0</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Internal primary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">10.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">21.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">28.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">25.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.9</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">27.1</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Internal secondary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">23</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">16.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">22.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">20.1</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">21.2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Claw 2 heights</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">External primary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">26</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">11.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">25.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">32.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">10.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">29.1</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">1.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.9</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">11.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">31.1</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">External secondary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">6.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">18.9</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">26.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">23.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">2.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">26.3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Internal primary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">11.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">23.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">30.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">27.1</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.9</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">27.7</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Internal secondary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">26</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">15.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">24.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">20.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">2.1</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.6</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">24.3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Claw 3 heights</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">External primary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">11.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">25.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">31.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">28.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">10.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">30.8</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">External secondary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">24</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">19.1</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">27.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">22.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">1.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">2.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">26.3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Internal primary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">26</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">10.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">20.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">28.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.0</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">26.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">26.6</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Internal secondary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">24</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">16.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">23.1</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">20.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">1.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">21.8</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Claw 4 heights</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="281" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Anterior primary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">26</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">12.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">25.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">35.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">10.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">30.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">1.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">2.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">12.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">35.3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Anterior secondary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">14.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">26.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">22.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">2.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">26.3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Posterior primary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">9.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">14.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">29.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">37.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">11.5</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">33.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">1.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">2.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">12.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">35.9</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Posterior secondary branch</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">23</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">6.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">10.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">19.9</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">31.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">8.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">24.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="281">2.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">?</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">?</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="281">
In live animals, body translucent in smaller specimens and opaque whitish in larger animals; transparent after fixation in 
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
medium (Figure 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Macrobiotus annewintersae sp. nov. - habitus and cuticular pores: A. Dorso-ventral view of the body (Holotype ♀;, PCM); B, C. Cuticular pores on the dorsal part of the body under PCM and under SEM, respectively. Arrowheads indicate pores and empty arrows indicate places on dorsal cuticle without pores. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure1" httpUri="https://binary.pensoft.net/fig/544821" pageId="0" pageNumber="281">1</figureCitation>
). Eyes present in live animals and after fixation in 
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
medium. Small roundish cuticular pores on the dorsal and lateral cuticle, as well as on the external cuticle of all legs (0.2-0.6 
<normalizedToken originalValue="μm">μm</normalizedToken>
in diameter), visible under both PCM and SEM (Figures 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Macrobiotus annewintersae sp. nov. - habitus and cuticular pores: A. Dorso-ventral view of the body (Holotype ♀;, PCM); B, C. Cuticular pores on the dorsal part of the body under PCM and under SEM, respectively. Arrowheads indicate pores and empty arrows indicate places on dorsal cuticle without pores. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure1" httpUri="https://binary.pensoft.net/fig/544821" pageId="0" pageNumber="281">1B, C</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Macrobiotus annewintersae sp. nov. - cuticular structures on legs: A. External granulation on leg III under PCM; B. A cuticular bulge (pulvinus) on the internal surface of leg III under PCM; C. Granulation on leg IV under PCM; D. External granulation on leg III under SEM; E. A cuticular bulge (pulvinus) on the internal surface of leg III under SEM. Filled flat arrowheads indicate the granulation patch, empty flat arrowheads indicate pulvinus and filled indented arrowheads indicate muscle attachments. C assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure2" httpUri="https://binary.pensoft.net/fig/544822" pageId="0" pageNumber="281">2D</figureCitation>
). On the dorsal surface, pores are absent between cuticle folds and arranged in loose belts (Figure 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Macrobiotus annewintersae sp. nov. - habitus and cuticular pores: A. Dorso-ventral view of the body (Holotype ♀;, PCM); B, C. Cuticular pores on the dorsal part of the body under PCM and under SEM, respectively. Arrowheads indicate pores and empty arrows indicate places on dorsal cuticle without pores. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure1" httpUri="https://binary.pensoft.net/fig/544821" pageId="0" pageNumber="281">1C</figureCitation>
). Pores sparse on the ventral surface and visible only under SEM (Figure 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Macrobiotus annewintersae sp. nov. - reproduction: A. Female under PCM; B. Male under PCM; C. Male under SEM. Filled indented arrowhead indicates spermathecae filled with spermatozoa, empty indented arrowhead indicates male's testis, arrows indicate lateral gibbosities on legs IV and filled flat arrowhead indicates cuticular pore on the ventral side of the body. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure8" httpUri="https://binary.pensoft.net/fig/544828" pageId="0" pageNumber="281">8C</figureCitation>
). Patches of fine granulation, on the external surface of legs I-III as well as on the dorsal and dorso-lateral sides of legs IV, visible in PCM (Figure 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Macrobiotus annewintersae sp. nov. - cuticular structures on legs: A. External granulation on leg III under PCM; B. A cuticular bulge (pulvinus) on the internal surface of leg III under PCM; C. Granulation on leg IV under PCM; D. External granulation on leg III under SEM; E. A cuticular bulge (pulvinus) on the internal surface of leg III under SEM. Filled flat arrowheads indicate the granulation patch, empty flat arrowheads indicate pulvinus and filled indented arrowheads indicate muscle attachments. C assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure2" httpUri="https://binary.pensoft.net/fig/544822" pageId="0" pageNumber="281">2A, C</figureCitation>
) and SEM (Figure 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Macrobiotus annewintersae sp. nov. - cuticular structures on legs: A. External granulation on leg III under PCM; B. A cuticular bulge (pulvinus) on the internal surface of leg III under PCM; C. Granulation on leg IV under PCM; D. External granulation on leg III under SEM; E. A cuticular bulge (pulvinus) on the internal surface of leg III under SEM. Filled flat arrowheads indicate the granulation patch, empty flat arrowheads indicate pulvinus and filled indented arrowheads indicate muscle attachments. C assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure2" httpUri="https://binary.pensoft.net/fig/544822" pageId="0" pageNumber="281">2D</figureCitation>
). A pulvinus is present on the internal surface of legs I-III (Figure 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Macrobiotus annewintersae sp. nov. - cuticular structures on legs: A. External granulation on leg III under PCM; B. A cuticular bulge (pulvinus) on the internal surface of leg III under PCM; C. Granulation on leg IV under PCM; D. External granulation on leg III under SEM; E. A cuticular bulge (pulvinus) on the internal surface of leg III under SEM. Filled flat arrowheads indicate the granulation patch, empty flat arrowheads indicate pulvinus and filled indented arrowheads indicate muscle attachments. C assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure2" httpUri="https://binary.pensoft.net/fig/544822" pageId="0" pageNumber="281">2B, E</figureCitation>
).
</paragraph>
<caption ID-Table-UUID="C3BADBB570940027DBF5242597B5B48C" httpUri="http://table.plazi.org/id/C3BADBB570940027DBF5242597B5B48C" pageId="0" pageNumber="281" start="Table 4" startId="T4">
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">Table 4.</emphasis>
Measurements [in 
<normalizedToken originalValue="µm">µm</normalizedToken>
] of selected morphological structures of the eggs of 
<taxonomicName authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus annewintersae</emphasis>
</taxonomicName>
sp. nov. mounted in 
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
medium (N-number of eggs/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD-standard deviation).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="281">
<table pageId="0" pageNumber="281">
<tr pageId="0" pageNumber="281">
<th colspan="1" pageId="0" pageNumber="281" rowspan="1">Character</th>
<th colspan="1" pageId="0" pageNumber="281" rowspan="1">N</th>
<th colspan="3" pageId="0" pageNumber="281" rowspan="1">Range</th>
<th colspan="1" pageId="0" pageNumber="281" rowspan="1">Mean</th>
<th colspan="1" pageId="0" pageNumber="281" rowspan="1">Sd</th>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">egg bare diameter</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">59.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">76.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">66.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">3.7</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Egg full diameter</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">69.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">87.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">75.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">4.6</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Process height</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">63</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">4.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">7.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.7</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Process base width</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">63</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">5.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">4.1</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.7</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Process base/height ratio</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">63</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">52%</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">100%</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">71%</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">10%</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Terminal disc width</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">63</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">6.7</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">4.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.9</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Inter-process distance</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">63</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">6.9</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">4.2</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">0.9</td>
</tr>
<tr pageId="0" pageNumber="281">
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">Number of processes on the egg circumference</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">21</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">24.4</td>
<td colspan="1" pageId="0" pageNumber="281" rowspan="1">1.7</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/zse.97.65280.figure1" httpUri="https://binary.pensoft.net/fig/544821" pageId="0" pageNumber="281" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">Figure 1.</emphasis>
<taxonomicName authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus annewintersae</emphasis>
</taxonomicName>
sp. nov. - habitus and cuticular pores: 
<emphasis bold="true" pageId="0" pageNumber="281">A.</emphasis>
Dorso-ventral view of the body (Holotype ♀;, PCM); 
<emphasis bold="true" pageId="0" pageNumber="281">B, C.</emphasis>
Cuticular pores on the dorsal part of the body under PCM and under SEM, respectively. Arrowheads indicate pores and empty arrows indicate places on dorsal cuticle without pores. Scale bars in 
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<caption doi="10.3897/zse.97.65280.figure2" httpUri="https://binary.pensoft.net/fig/544822" pageId="0" pageNumber="281" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">Figure 2.</emphasis>
<taxonomicName authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus annewintersae</emphasis>
</taxonomicName>
sp. nov. - cuticular structures on legs: 
<emphasis bold="true" pageId="0" pageNumber="281">A.</emphasis>
External granulation on leg III under PCM; 
<emphasis bold="true" pageId="0" pageNumber="281">B.</emphasis>
A cuticular bulge (pulvinus) on the internal surface of leg III under PCM; 
<emphasis bold="true" pageId="0" pageNumber="281">C.</emphasis>
Granulation on leg IV under PCM; 
<emphasis bold="true" pageId="0" pageNumber="281">D.</emphasis>
External granulation on leg III under SEM; 
<emphasis bold="true" pageId="0" pageNumber="281">E.</emphasis>
A cuticular bulge (pulvinus) on the internal surface of leg III under SEM. Filled flat arrowheads indicate the granulation patch, empty flat arrowheads indicate pulvinus and filled indented arrowheads indicate muscle attachments. C assembled from several photos. Scale bars in 
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="281">
Claws Y-shaped, of the 
<taxonomicName class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus hufelandi" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus hufelandi</emphasis>
</taxonomicName>
type. Primary branches with distinct accessory points, a common tract, and an evident stalk connecting the claw to the lunula (Figure 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Macrobiotus annewintersae sp. nov. - claws: A, B. Claws III and IV, respectively, under PCM; C, D. Claws III and IV, respectively, under SEM. Filled indented arrowheads indicate double muscle attachments under the claws, empty indented arrowheads indicate a faintly visible divided cuticular bar. A and B assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure3" httpUri="https://binary.pensoft.net/fig/544823" pageId="0" pageNumber="281">3</figureCitation>
). The lunulae I-III are smooth (Figure 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Macrobiotus annewintersae sp. nov. - claws: A, B. Claws III and IV, respectively, under PCM; C, D. Claws III and IV, respectively, under SEM. Filled indented arrowheads indicate double muscle attachments under the claws, empty indented arrowheads indicate a faintly visible divided cuticular bar. A and B assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure3" httpUri="https://binary.pensoft.net/fig/544823" pageId="0" pageNumber="281">3A, C</figureCitation>
), whereas lunulae IV are dentate (Figure 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Macrobiotus annewintersae sp. nov. - claws: A, B. Claws III and IV, respectively, under PCM; C, D. Claws III and IV, respectively, under SEM. Filled indented arrowheads indicate double muscle attachments under the claws, empty indented arrowheads indicate a faintly visible divided cuticular bar. A and B assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure3" httpUri="https://binary.pensoft.net/fig/544823" pageId="0" pageNumber="281">3B, D</figureCitation>
). A divided cuticular bar with double muscle attachments are poorly visible under PCM (Figure 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Macrobiotus annewintersae sp. nov. - claws: A, B. Claws III and IV, respectively, under PCM; C, D. Claws III and IV, respectively, under SEM. Filled indented arrowheads indicate double muscle attachments under the claws, empty indented arrowheads indicate a faintly visible divided cuticular bar. A and B assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure3" httpUri="https://binary.pensoft.net/fig/544823" pageId="0" pageNumber="281">3A</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.97.65280.figure3" httpUri="https://binary.pensoft.net/fig/544823" pageId="0" pageNumber="281" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">Figure 3.</emphasis>
<taxonomicName authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus annewintersae</emphasis>
</taxonomicName>
sp. nov. - claws: 
<emphasis bold="true" pageId="0" pageNumber="281">A, B.</emphasis>
Claws III and IV, respectively, under PCM; 
<emphasis bold="true" pageId="0" pageNumber="281">C, D.</emphasis>
Claws III and IV, respectively, under SEM. Filled indented arrowheads indicate double muscle attachments under the claws, empty indented arrowheads indicate a faintly visible divided cuticular bar. A and B assembled from several photos. Scale bars in 
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="281">
Mouth antero-ventral. Bucco-pharyngeal apparatus of the 
<taxonomicName authorityName="C.A.S.Schultze" authorityYear="1834" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus</emphasis>
</taxonomicName>
type (Figure 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrobiotus annewintersae sp. nov. - buccal apparatus and the oral cavity armature under PCM: A. Dorso-ventral view of the entire buccal apparatus; B, C. Oral cavity armature in dorsal and ventral view, respectively; D, E. Placoid morphology in dorsal and ventral view, respectively. Empty flat arrowheads indicate the second band of teeth, filled indented arrowheads indicate the third band of teeth in the oral cavity, and empty indented arrowheads indicate central constriction in the first macroplacoid and subterminal constriction in the second macroplacoid. A, D and E assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure4" httpUri="https://binary.pensoft.net/fig/544824" pageId="0" pageNumber="281">4</figureCitation>
) with ventral lamina and ten peribuccal lamellae. The stylet furcae typically-shaped, the basal portion is enlarged and has two caudal branches with thickened, swollen, rounded apices. Under PCM, the oral cavity armature is of the 
<emphasis italics="true" pageId="0" pageNumber="281">patagonicus</emphasis>
type, 
<emphasis italics="true" pageId="0" pageNumber="281">i.e.</emphasis>
, with only the second and third bands of teeth visible (Figure 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrobiotus annewintersae sp. nov. - buccal apparatus and the oral cavity armature under PCM: A. Dorso-ventral view of the entire buccal apparatus; B, C. Oral cavity armature in dorsal and ventral view, respectively; D, E. Placoid morphology in dorsal and ventral view, respectively. Empty flat arrowheads indicate the second band of teeth, filled indented arrowheads indicate the third band of teeth in the oral cavity, and empty indented arrowheads indicate central constriction in the first macroplacoid and subterminal constriction in the second macroplacoid. A, D and E assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure4" httpUri="https://binary.pensoft.net/fig/544824" pageId="0" pageNumber="281">4B, C</figureCitation>
). However, under SEM the first band of teeth is visible and composed of one row of very small cones situated anteriorly in the oral cavity, just behind the bases of the peribuccal lamellae (Figure 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Macrobiotus annewintersae sp. nov. - anterior view of the mouth opening under SEM. Filled flat arrowhead indicates the first band of teeth. Scale bar in μm." figureDoi="10.3897/zse.97.65280.figure5" httpUri="https://binary.pensoft.net/fig/544825" pageId="0" pageNumber="281">5</figureCitation>
). The second band of teeth is situated between the ring fold and the third band of teeth and composed of 3-4 rows of teeth visible in PCM as granules (Figure 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrobiotus annewintersae sp. nov. - buccal apparatus and the oral cavity armature under PCM: A. Dorso-ventral view of the entire buccal apparatus; B, C. Oral cavity armature in dorsal and ventral view, respectively; D, E. Placoid morphology in dorsal and ventral view, respectively. Empty flat arrowheads indicate the second band of teeth, filled indented arrowheads indicate the third band of teeth in the oral cavity, and empty indented arrowheads indicate central constriction in the first macroplacoid and subterminal constriction in the second macroplacoid. A, D and E assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure4" httpUri="https://binary.pensoft.net/fig/544824" pageId="0" pageNumber="281">4B, C</figureCitation>
). The third band of teeth is divided into a dorsal (Figure 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrobiotus annewintersae sp. nov. - buccal apparatus and the oral cavity armature under PCM: A. Dorso-ventral view of the entire buccal apparatus; B, C. Oral cavity armature in dorsal and ventral view, respectively; D, E. Placoid morphology in dorsal and ventral view, respectively. Empty flat arrowheads indicate the second band of teeth, filled indented arrowheads indicate the third band of teeth in the oral cavity, and empty indented arrowheads indicate central constriction in the first macroplacoid and subterminal constriction in the second macroplacoid. A, D and E assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure4" httpUri="https://binary.pensoft.net/fig/544824" pageId="0" pageNumber="281">4B</figureCitation>
) and a ventral portion (Figure 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrobiotus annewintersae sp. nov. - buccal apparatus and the oral cavity armature under PCM: A. Dorso-ventral view of the entire buccal apparatus; B, C. Oral cavity armature in dorsal and ventral view, respectively; D, E. Placoid morphology in dorsal and ventral view, respectively. Empty flat arrowheads indicate the second band of teeth, filled indented arrowheads indicate the third band of teeth in the oral cavity, and empty indented arrowheads indicate central constriction in the first macroplacoid and subterminal constriction in the second macroplacoid. A, D and E assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure4" httpUri="https://binary.pensoft.net/fig/544824" pageId="0" pageNumber="281">4C</figureCitation>
). Under PCM, the dorsal teeth are seen as three distinct transverse ridges whereas the ventral teeth appear as two separate lateral transverse ridges between which one big tooth (sometimes circular in PCM) is visible (Figure 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrobiotus annewintersae sp. nov. - buccal apparatus and the oral cavity armature under PCM: A. Dorso-ventral view of the entire buccal apparatus; B, C. Oral cavity armature in dorsal and ventral view, respectively; D, E. Placoid morphology in dorsal and ventral view, respectively. Empty flat arrowheads indicate the second band of teeth, filled indented arrowheads indicate the third band of teeth in the oral cavity, and empty indented arrowheads indicate central constriction in the first macroplacoid and subterminal constriction in the second macroplacoid. A, D and E assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure4" httpUri="https://binary.pensoft.net/fig/544824" pageId="0" pageNumber="281">4B, C</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.97.65280.figure4" httpUri="https://binary.pensoft.net/fig/544824" pageId="0" pageNumber="281" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">Figure 4.</emphasis>
<taxonomicName authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus annewintersae</emphasis>
</taxonomicName>
sp. nov. - buccal apparatus and the oral cavity armature under PCM: 
<emphasis bold="true" pageId="0" pageNumber="281">A.</emphasis>
Dorso-ventral view of the entire buccal apparatus; 
<emphasis bold="true" pageId="0" pageNumber="281">B, C.</emphasis>
Oral cavity armature in dorsal and ventral view, respectively; 
<emphasis bold="true" pageId="0" pageNumber="281">D, E.</emphasis>
Placoid morphology in dorsal and ventral view, respectively. Empty flat arrowheads indicate the second band of teeth, filled indented arrowheads indicate the third band of teeth in the oral cavity, and empty indented arrowheads indicate central constriction in the first macroplacoid and subterminal constriction in the second macroplacoid. 
<emphasis bold="true" pageId="0" pageNumber="281">A, D</emphasis>
and 
<emphasis bold="true" pageId="0" pageNumber="281">E</emphasis>
assembled from several photos. Scale bars in 
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="281">
Pharyngeal bulb spherical, with triangular apophyses, two rod-shaped macroplacoids and a drop-shaped microplacoid (Figure 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrobiotus annewintersae sp. nov. - buccal apparatus and the oral cavity armature under PCM: A. Dorso-ventral view of the entire buccal apparatus; B, C. Oral cavity armature in dorsal and ventral view, respectively; D, E. Placoid morphology in dorsal and ventral view, respectively. Empty flat arrowheads indicate the second band of teeth, filled indented arrowheads indicate the third band of teeth in the oral cavity, and empty indented arrowheads indicate central constriction in the first macroplacoid and subterminal constriction in the second macroplacoid. A, D and E assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure4" httpUri="https://binary.pensoft.net/fig/544824" pageId="0" pageNumber="281">4A, D, E</figureCitation>
). The macroplacoid length sequence is 2&lt;1. The first and the second macroplacoid have a central and a subterminal constriction, respectively (Figure 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Macrobiotus annewintersae sp. nov. - buccal apparatus and the oral cavity armature under PCM: A. Dorso-ventral view of the entire buccal apparatus; B, C. Oral cavity armature in dorsal and ventral view, respectively; D, E. Placoid morphology in dorsal and ventral view, respectively. Empty flat arrowheads indicate the second band of teeth, filled indented arrowheads indicate the third band of teeth in the oral cavity, and empty indented arrowheads indicate central constriction in the first macroplacoid and subterminal constriction in the second macroplacoid. A, D and E assembled from several photos. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure4" httpUri="https://binary.pensoft.net/fig/544824" pageId="0" pageNumber="281">4D, E</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.97.65280.figure5" httpUri="https://binary.pensoft.net/fig/544825" pageId="0" pageNumber="281" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">Figure 5.</emphasis>
<taxonomicName authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus annewintersae</emphasis>
</taxonomicName>
sp. nov. - anterior view of the mouth opening under SEM. Filled flat arrowhead indicates the first band of teeth. Scale bar in 
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" italics="true" pageId="0" pageNumber="281">Eggs</emphasis>
(measurements and statistics in Table 
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Measurements [in µm] of selected morphological structures of the eggs of Macrobiotus annewintersae sp. nov. mounted in Hoyer's medium (N-number of eggs / structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD-standard deviation)." httpUri="http://table.plazi.org/id/C3BADBB570940027DBF5242597B5B48C" pageId="0" pageNumber="281" tableUuid="C3BADBB570940027DBF5242597B5B48C">4</tableCitation>
):
</paragraph>
<paragraph pageId="0" pageNumber="281">
The surface between processes is of the 
<taxonomicName authorityName="Binda &amp; Pilato" authorityYear="1972" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus persimilis" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="persimilis">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus persimilis</emphasis>
</taxonomicName>
type, 
<emphasis italics="true" pageId="0" pageNumber="281">i.e.</emphasis>
, with a continuous smooth chorion, never with pores or reticulum (Figures 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Macrobiotus annewintersae sp. nov. - egg chorion morphology under PCM: A, B. Egg surface; C, D. Midsection of the processes. Filled flat arrowheads indicate bubble-like structures within tentacular arms in the distal portion of the egg processes and empty flat arrowheads indicate dark thickenings / striae on the egg surface between processes. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure6" httpUri="https://binary.pensoft.net/fig/544826" pageId="0" pageNumber="281">6</figureCitation>
, 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Macrobiotus annewintersae sp. nov. - egg chorion morphology under SEM: A, B. Entire egg; C-E. Details of the egg processes and egg surface between them; F. Details of the tentacular arms in the distal portion of each egg process. Filled indented arrowheads indicate micropores and empty indented arrowheads indicate lobes in tentacular arms covered by micro-granulation. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure7" httpUri="https://binary.pensoft.net/fig/544827" pageId="0" pageNumber="281">7</figureCitation>
). Under PCM the surface between the processes is covered with wrinkles that appear as dark thickenings/striae, whereas under SEM the surface appears clearly wrinkled (Figures 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Macrobiotus annewintersae sp. nov. - egg chorion morphology under PCM: A, B. Egg surface; C, D. Midsection of the processes. Filled flat arrowheads indicate bubble-like structures within tentacular arms in the distal portion of the egg processes and empty flat arrowheads indicate dark thickenings / striae on the egg surface between processes. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure6" httpUri="https://binary.pensoft.net/fig/544826" pageId="0" pageNumber="281">6</figureCitation>
, 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Macrobiotus annewintersae sp. nov. - egg chorion morphology under SEM: A, B. Entire egg; C-E. Details of the egg processes and egg surface between them; F. Details of the tentacular arms in the distal portion of each egg process. Filled indented arrowheads indicate micropores and empty indented arrowheads indicate lobes in tentacular arms covered by micro-granulation. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure7" httpUri="https://binary.pensoft.net/fig/544827" pageId="0" pageNumber="281">7</figureCitation>
). Processes are of a modified 
<taxonomicName class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus hufelandi" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus hufelandi</emphasis>
</taxonomicName>
type (Figures 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Macrobiotus annewintersae sp. nov. - egg chorion morphology under PCM: A, B. Egg surface; C, D. Midsection of the processes. Filled flat arrowheads indicate bubble-like structures within tentacular arms in the distal portion of the egg processes and empty flat arrowheads indicate dark thickenings / striae on the egg surface between processes. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure6" httpUri="https://binary.pensoft.net/fig/544826" pageId="0" pageNumber="281">6</figureCitation>
, 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Macrobiotus annewintersae sp. nov. - egg chorion morphology under SEM: A, B. Entire egg; C-E. Details of the egg processes and egg surface between them; F. Details of the tentacular arms in the distal portion of each egg process. Filled indented arrowheads indicate micropores and empty indented arrowheads indicate lobes in tentacular arms covered by micro-granulation. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure7" httpUri="https://binary.pensoft.net/fig/544827" pageId="0" pageNumber="281">7</figureCitation>
). The proper terminal disc is absent and instead 2-8 thick tentacular arms (typically 5-6) are present in the distal part of the process (Figures 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Macrobiotus annewintersae sp. nov. - egg chorion morphology under PCM: A, B. Egg surface; C, D. Midsection of the processes. Filled flat arrowheads indicate bubble-like structures within tentacular arms in the distal portion of the egg processes and empty flat arrowheads indicate dark thickenings / striae on the egg surface between processes. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure6" httpUri="https://binary.pensoft.net/fig/544826" pageId="0" pageNumber="281">6</figureCitation>
, 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Macrobiotus annewintersae sp. nov. - egg chorion morphology under SEM: A, B. Entire egg; C-E. Details of the egg processes and egg surface between them; F. Details of the tentacular arms in the distal portion of each egg process. Filled indented arrowheads indicate micropores and empty indented arrowheads indicate lobes in tentacular arms covered by micro-granulation. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure7" httpUri="https://binary.pensoft.net/fig/544827" pageId="0" pageNumber="281">7</figureCitation>
). The tentacular arms present bubble-like structures (visible in PCM). Under SEM, each tentacular arm is distally divided into many irregular digitations that are sometime covered with micro-granulation (Figure 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Macrobiotus annewintersae sp. nov. - egg chorion morphology under SEM: A, B. Entire egg; C-E. Details of the egg processes and egg surface between them; F. Details of the tentacular arms in the distal portion of each egg process. Filled indented arrowheads indicate micropores and empty indented arrowheads indicate lobes in tentacular arms covered by micro-granulation. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure7" httpUri="https://binary.pensoft.net/fig/544827" pageId="0" pageNumber="281">7C-F</figureCitation>
). Also, under SEM micro-pores can be seen on the egg surface between the processes and around the process bases (Figure 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Macrobiotus annewintersae sp. nov. - egg chorion morphology under SEM: A, B. Entire egg; C-E. Details of the egg processes and egg surface between them; F. Details of the tentacular arms in the distal portion of each egg process. Filled indented arrowheads indicate micropores and empty indented arrowheads indicate lobes in tentacular arms covered by micro-granulation. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure7" httpUri="https://binary.pensoft.net/fig/544827" pageId="0" pageNumber="281">7C, E</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.97.65280.figure6" httpUri="https://binary.pensoft.net/fig/544826" pageId="0" pageNumber="281" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">Figure 6.</emphasis>
<taxonomicName authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus annewintersae</emphasis>
</taxonomicName>
sp. nov. - egg chorion morphology under PCM: 
<emphasis bold="true" pageId="0" pageNumber="281">A, B.</emphasis>
Egg surface; 
<emphasis bold="true" pageId="0" pageNumber="281">C, D.</emphasis>
Midsection of the processes. Filled flat arrowheads indicate bubble-like structures within tentacular arms in the distal portion of the egg processes and empty flat arrowheads indicate dark thickenings/striae on the egg surface between processes. Scale bars in 
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<caption doi="10.3897/zse.97.65280.figure7" httpUri="https://binary.pensoft.net/fig/544827" pageId="0" pageNumber="281" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">Figure 7.</emphasis>
<taxonomicName authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus annewintersae</emphasis>
</taxonomicName>
sp. nov. - egg chorion morphology under SEM: 
<emphasis bold="true" pageId="0" pageNumber="281">A, B.</emphasis>
Entire egg; 
<emphasis bold="true" pageId="0" pageNumber="281">C-E.</emphasis>
Details of the egg processes and egg surface between them; 
<emphasis bold="true" pageId="0" pageNumber="281">F.</emphasis>
Details of the tentacular arms in the distal portion of each egg process. Filled indented arrowheads indicate micropores and empty indented arrowheads indicate lobes in tentacular arms covered by micro-granulation. Scale bars in 
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">
<emphasis italics="true" pageId="0" pageNumber="281">Reproduction / Sexual dimorphism</emphasis>
.
</emphasis>
The species is dioecious. Spermathecae in females as well as testis in males, clearly visible under PCM up to 24 hours after mounting in 
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
medium, have been found to be filled with spermatozoa (Figure 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Macrobiotus annewintersae sp. nov. - reproduction: A. Female under PCM; B. Male under PCM; C. Male under SEM. Filled indented arrowhead indicates spermathecae filled with spermatozoa, empty indented arrowhead indicates male's testis, arrows indicate lateral gibbosities on legs IV and filled flat arrowhead indicates cuticular pore on the ventral side of the body. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure8" httpUri="https://binary.pensoft.net/fig/544828" pageId="0" pageNumber="281">8A, B</figureCitation>
). The species exhibits secondary sexual dimorphism in the form of clearly visible lateral gibbosities on the hind legs in males (Figure 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Macrobiotus annewintersae sp. nov. - reproduction: A. Female under PCM; B. Male under PCM; C. Male under SEM. Filled indented arrowhead indicates spermathecae filled with spermatozoa, empty indented arrowhead indicates male's testis, arrows indicate lateral gibbosities on legs IV and filled flat arrowhead indicates cuticular pore on the ventral side of the body. Scale bars in μm." figureDoi="10.3897/zse.97.65280.figure8" httpUri="https://binary.pensoft.net/fig/544828" pageId="0" pageNumber="281">8B, C</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.97.65280.figure8" httpUri="https://binary.pensoft.net/fig/544828" pageId="0" pageNumber="281" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">Figure 8.</emphasis>
<taxonomicName authorityName="Vecchi &amp; Stec" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus annewintersae" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus annewintersae</emphasis>
</taxonomicName>
sp. nov. - reproduction: 
<emphasis bold="true" pageId="0" pageNumber="281">A.</emphasis>
Female under PCM; 
<emphasis bold="true" pageId="0" pageNumber="281">B.</emphasis>
Male under PCM; 
<emphasis bold="true" pageId="0" pageNumber="281">C.</emphasis>
Male under SEM. Filled indented arrowhead indicates spermathecae filled with spermatozoa, empty indented arrowhead indicates 
<normalizedToken originalValue="male’s">male's</normalizedToken>
testis, arrows indicate lateral gibbosities on legs IV and filled flat arrowhead indicates cuticular pore on the ventral side of the body. Scale bars in 
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="281" type="dna sequences">
<paragraph pageId="0" pageNumber="281">DNA sequences.</paragraph>
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">18S rRNA</emphasis>
: GenBank: MW588024-MW588025; 659 and 664 bp long.
</paragraph>
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">28S rRNA</emphasis>
: GenBank: MW588030-MW588031; 679 and 703 bp long.
</paragraph>
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">ITS-2</emphasis>
: GenBank: MW588018-MW588019; 298 bp long.
</paragraph>
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">COI</emphasis>
: GenBank: MW593927-MW593928; 532 and 535 bp long.
</paragraph>
<paragraph pageId="0" pageNumber="281">
<emphasis bold="true" pageId="0" pageNumber="281">
<emphasis italics="true" pageId="0" pageNumber="281">Phenotypic differential diagnosis</emphasis>
.
</emphasis>
By having an egg chorion of the 
<emphasis italics="true" pageId="0" pageNumber="281">
<taxonomicName authorityName="Binda &amp; Pilato" authorityYear="1972" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus persimilis" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="persimilis">Macrobiotus persimilis</taxonomicName>
type
</emphasis>
(smooth or wrinkled chorion) and by having thick tentacular arms instead of a proper terminal disc on the distal part of egg processes, 
<taxonomicName class="Eutardigrada" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. annewintersae" order="Coleoptera" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis italics="true" pageId="0" pageNumber="281">M. annewintersae</emphasis>
</taxonomicName>
sp. nov. resembles only one species: 
<taxonomicName authorityName="Meyer, Domingue &amp; Hinton" authorityYear="2014" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" lsidName="Macrobiotus anemone" order="Parachela" pageId="0" pageNumber="281" phylum="Tardigrada" rank="species" species="anemone">
<emphasis italics="true" pageId="0" pageNumber="281">Macrobiotus anemone</emphasis>
</taxonomicName>
Meyer, Domingue &amp; Hinton, 2014 from USA. However, the new species differs specifically from:
</paragraph>
<paragraph pageId="0" pageNumber="281">• M. anemone by having eyes (absent in M. anemone), by the presence of granulation on all legs (absent in M. anemone), by having the oral cavity armature (OCA) of the patagonicus type (maculatus type - only the third band of teeth visible under light microscope - in M. anemone), by the presence of dentate lunulae in legs IV (smooth lunulae in legs IV in M. anemone), by having the thick tentacular arms in the distal part of the processes filled with bubble-like structures (tentacular arms solid in M. anemone, Figure 17) and by lacking a cavity between the process trunk and tentacular arms that appears in PCM as a clearly refracting dot (the cavity present in M. anemone, Figure 17).</paragraph>
</subSubSection>
</treatment>
</document>