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<document id="711F57EF043F6739979ED7D3483DCCEC" ID-DOI="10.11646/zootaxa.3920.1.3" ID-GBIF-Dataset="9f10093d-31b9-45c3-9ec3-62ee8078ad74" ID-ISSN="1175-5326" ID-Zenodo-Dep="287903" ID-ZooBank="5A9E6C27-4E20-4AE5-8287-4E4762BFCD6A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994510914" checkinUser="plazi" docAuthor="Molineri, C., Salles, F. F. &amp; Emmerich, D." docDate="2015" docId="D62C004AFF965A2CFF7283E0FB17FC13" docLanguage="en" docName="zt03920p068.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Campsurus molinai Molineri, Salles &amp; Emmerich, 2015, sp. nov." docType="treatment" docVersion="6" lastPageNumber="64" masterDocId="2A157832FF9B5A21FFE58377FFE6FFD5" masterDocTitle="Revision of Campsurus violaceus species group (Ephemeroptera: Polymitarcyidae) with new synonymies and nomina dubia in Campsurus Eaton, 1868" masterLastPageNumber="68" masterPageNumber="51" pageNumber="64" updateTime="1698562500379" updateUser="plazi">
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<mods:title id="F5D0E402EBD6B574C8F72977EAAEFCDD">Revision of Campsurus violaceus species group (Ephemeroptera: Polymitarcyidae) with new synonymies and nomina dubia in Campsurus Eaton, 1868</mods:title>
</mods:titleInfo>
<mods:name id="A18E3D41765BAA480BC3B9B7CEAB2A08" type="personal">
<mods:role id="5511B4360225A74BCB7602DD9A3DD3E6">
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<mods:namePart id="7B360ED8F464ADC6EBB28ABA9F1255A7">Molineri, C.</mods:namePart>
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<mods:namePart id="F9A97C888E81F43D57312D6CF252D81C">Salles, F. F.</mods:namePart>
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<mods:name id="EA4377FC5BDF3FDD04B8A93123CF330A" type="personal">
<mods:role id="0295EF6D33530005B8D90F4134026DDF">
<mods:roleTerm id="66AC1AB0698B367C211A9C53DE39E6F3">Author</mods:roleTerm>
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<mods:namePart id="4BBBC1CB71825289E3E982F181E1C7C8">Emmerich, D.</mods:namePart>
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<mods:typeOfResource id="4B93A77D06E273096B8361383E241699">text</mods:typeOfResource>
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<mods:title id="75787D748E425681E62079735304B10C">Zootaxa</mods:title>
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<mods:part id="050AECCF2DB6FF45303DBA9F4A8AAFB2">
<mods:date id="CD1DD3A2E1C2830A9BEF392D459415DF">2015</mods:date>
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<mods:number id="72CF7795DCA46B1638F62B1BBDEEDF36">3920</mods:number>
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<mods:identifier id="72B838815056A0022BA58C6660B92075" type="DOI">10.11646/zootaxa.3920.1.3</mods:identifier>
<mods:identifier id="70F0DF8F44BD8000BC761F3A761E4D67" type="GBIF-Dataset">9f10093d-31b9-45c3-9ec3-62ee8078ad74</mods:identifier>
<mods:identifier id="08D79EC661E68598E32C95558CCBB69C" type="ISSN">1175-5326</mods:identifier>
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<treatment id="D62C004AFF965A2CFF7283E0FB17FC13" ID-DOI="http://doi.org/10.5281/zenodo.5659652" ID-GBIF-Taxon="127671673" ID-Zenodo-Dep="5659652" LSID="urn:lsid:plazi:treatment:D62C004AFF965A2CFF7283E0FB17FC13" httpUri="http://treatment.plazi.org/id/D62C004AFF965A2CFF7283E0FB17FC13" lastPageNumber="64" pageId="13" pageNumber="64">
<subSubSection id="169FE2D7FF965A2CFF7283E0FEC9FF06" pageId="13" pageNumber="64" type="nomenclature">
<paragraph id="5E3AB15CFF965A2CFF7283E0FE0CFF67" blockId="13.[151,490,151,211]" box="[151,490,151,178]" pageId="13" pageNumber="64">
<heading id="05720630FF965A2CFF7283E0FE0CFF67" bold="true" box="[151,490,151,178]" fontSize="11" level="1" pageId="13" pageNumber="64" reason="1">
<emphasis id="6CF16D4EFF965A2CFF7283E0FE0CFF67" bold="true" box="[151,490,151,178]" pageId="13" pageNumber="64">
<taxonomicName id="9985CADFFF965A2CFF7283E0FE60FF64" box="[151,390,151,177]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="64" phylum="Arthropoda" rank="species" species="molinai" status="sp. nov.">
<emphasis id="6CF16D4EFF965A2CFF7283E0FE60FF64" bold="true" box="[151,390,151,177]" italics="true" pageId="13" pageNumber="64">Campsurus molinai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77C2D035FF965A2CFE6983EFFE0CFF67" box="[396,490,152,178]" pageId="13" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5E3AB15CFF965A2CFF7283CDFEC9FF06" blockId="13.[151,490,151,211]" box="[151,303,186,211]" pageId="13" pageNumber="64">
(
<figureCitation id="C6BEADD9FF965A2CFF7A83CDFECEFF06" box="[159,296,186,211]" captionStart="FIGURES 38 40. C" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[163,1436,1036,1707]" captionTargetId="figure@13.[150,1436,1022,1716]" captionTargetPageId="13" captionText="FIGURES 38 40. C. molinai sp. nov., male adult. 38, male genitalia, v. v.; 39, pedestal, detail, v. v.; 40, penes and pedestals, v. v. (photograph). Abbreviations: ip = inner projection (pedestal); op = outer projection (pedestal); p = penes; pb = pyramidal base (penes); t = &quot; thumb &quot; (penes)." httpUri="https://zenodo.org/record/287909/files/figure.png" pageId="13" pageNumber="64">Figs. 3840</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="169FE2D7FF965A2CFF728275FC52FEEA" pageId="13" pageNumber="64" type="materials_examined">
<paragraph id="5E3AB15CFF965A2CFF728275FC52FEEA" blockId="13.[151,1437,258,967]" pageId="13" pageNumber="64">
<emphasis id="6CF16D4EFF965A2CFF728275FEAFFECE" bold="true" box="[151,329,258,283]" pageId="13" pageNumber="64">
<typeStatus id="813E0FFEFF965A2CFF728275FF34FECE" box="[151,210,258,283]" pageId="13" pageNumber="64">Type</typeStatus>
material.
</emphasis>
<typeStatus id="813E0FFEFF965A2CFEB18275FE5AFECE" box="[340,444,258,283]" pageId="13" pageNumber="64" type="holotype">Holotype</typeStatus>
male imago from
<collectingCountry id="2692F1CCFF965A2CFD798275FD17FECE" box="[668,753,258,283]" name="Bolivia" pageId="13" pageNumber="64">Bolivia</collectingCountry>
, Beni, La Granja lake,
<date id="2A3B979CFF965A2CFBEC8275FB8AFECE" box="[1033,1132,258,283]" pageId="13" pageNumber="64" value="2005-05-02">2.v.2005</date>
, C. Molina col. (
<collectionCode id="38942999FF965A2CFADC8274FA76FECE" box="[1337,1424,259,283]" httpUri="http://grbio.org/cool/jxh3-1jar" name="Instituto de Ecologia" pageId="13" pageNumber="64">UMSA</collectionCode>
).
<typeStatus id="813E0FFEFF965A2CFF728250FF1FFEEA" box="[151,249,295,319]" pageId="13" pageNumber="64" type="paratype">Paratype</typeStatus>
male imago (slide IBN344CM), same data as
<typeStatus id="813E0FFEFF965A2CFCE48251FC85FEEA" box="[769,867,294,319]" pageId="13" pageNumber="64" type="holotype">holotype</typeStatus>
(IBN).
</paragraph>
</subSubSection>
<subSubSection id="169FE2D7FF965A2CFF22823DFDE9FEB6" box="[199,527,330,355]" pageId="13" pageNumber="64" type="distribution">
<paragraph id="5E3AB15CFF965A2CFF22823DFDE9FEB6" blockId="13.[151,1437,258,967]" box="[199,527,330,355]" pageId="13" pageNumber="64">
<emphasis id="6CF16D4EFF965A2CFF22823DFE86FEB6" bold="true" box="[199,352,330,355]" pageId="13" pageNumber="64">Distribution.</emphasis>
<collectingCountry id="2692F1CCFF965A2CFE82823DFE5CFEB6" box="[359,442,330,355]" name="Bolivia" pageId="13" pageNumber="64">Bolivia</collectingCountry>
(Beni).
</paragraph>
</subSubSection>
<subSubSection id="169FE2D7FF965A2CFF228219FDFCFE7E" pageId="13" pageNumber="64" type="etymology">
<paragraph id="5E3AB15CFF965A2CFF228219FDFCFE7E" blockId="13.[151,1437,258,967]" pageId="13" pageNumber="64">
<emphasis id="6CF16D4EFF965A2CFF228219FEA8FE52" bold="true" box="[199,334,366,391]" pageId="13" pageNumber="64">Etymology.</emphasis>
The species is dedicated to Dr. Carlos
<taxonomicName id="9985CADFFF965A2CFCC78219FC00FE52" authority="Bolivian" authorityName="Bolivian" box="[802,998,366,391]" class="Magnoliopsida" family="Asteraceae" genus="Molina" higherTaxonomySource="CoL" kingdom="Plantae" order="Asterales" pageId="13" pageNumber="64" phylum="Tracheophyta" rank="genus">Molina, Bolivian</taxonomicName>
limnologist that collected and kindly offered the material of the species.
</paragraph>
</subSubSection>
<subSubSection id="169FE2D7FF965A2CFF2282C1FE6BFC8E" pageId="13" pageNumber="64" type="diagnosis">
<paragraph id="5E3AB15CFF965A2CFF2282C1FC8EFDEE" blockId="13.[151,1437,258,967]" pageId="13" pageNumber="64">
<emphasis id="6CF16D4EFF965A2CFF2282C1FEA7FE1A" bold="true" box="[199,321,438,463]" pageId="13" pageNumber="64">Diagnosis.</emphasis>
1) pedestals stout, subquadrate, inner distal corner larger and roundly expanded, outer corner thinner and acute (
<figureCitation id="C6BEADD9FF965A2CFE8F82ADFE10FE26" box="[362,502,474,499]" captionStart="FIGURES 38 40. C" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[163,1436,1036,1707]" captionTargetId="figure@13.[150,1436,1022,1716]" captionTargetPageId="13" captionText="FIGURES 38 40. C. molinai sp. nov., male adult. 38, male genitalia, v. v.; 39, pedestal, detail, v. v.; 40, penes and pedestals, v. v. (photograph). Abbreviations: ip = inner projection (pedestal); op = outer projection (pedestal); p = penes; pb = pyramidal base (penes); t = &quot; thumb &quot; (penes)." httpUri="https://zenodo.org/record/287909/files/figure.png" pageId="13" pageNumber="64">Figs. 3840</figureCitation>
); 2) penis strongly developed and sclerotized, with a small spine-like projection at apex, basal 1/2 of penis strongly expanded dorsally (
<figureCitation id="C6BEADD9FF965A2CFD198289FC6DFDC2" box="[764,907,510,535]" captionStart="FIGURES 38 40. C" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[163,1436,1036,1707]" captionTargetId="figure@13.[150,1436,1022,1716]" captionTargetPageId="13" captionText="FIGURES 38 40. C. molinai sp. nov., male adult. 38, male genitalia, v. v.; 39, pedestal, detail, v. v.; 40, penes and pedestals, v. v. (photograph). Abbreviations: ip = inner projection (pedestal); op = outer projection (pedestal); p = penes; pb = pyramidal base (penes); t = &quot; thumb &quot; (penes)." httpUri="https://zenodo.org/record/287909/files/figure.png" pageId="13" pageNumber="64">Figs. 38, 40</figureCitation>
); 3) thumb not finger-like, reduced to a basal membrane on inner ventral margin of each penis (&quot;t&quot; in
<figureCitation id="C6BEADD9FF965A2CFCED8155FCBAFDEE" box="[776,860,546,571]" captionStart="FIGURES 38 40. C" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[163,1436,1036,1707]" captionTargetId="figure@13.[150,1436,1022,1716]" captionTargetPageId="13" captionText="FIGURES 38 40. C. molinai sp. nov., male adult. 38, male genitalia, v. v.; 39, pedestal, detail, v. v.; 40, penes and pedestals, v. v. (photograph). Abbreviations: ip = inner projection (pedestal); op = outer projection (pedestal); p = penes; pb = pyramidal base (penes); t = &quot; thumb &quot; (penes)." httpUri="https://zenodo.org/record/287909/files/figure.png" pageId="13" pageNumber="64">Fig. 40</figureCitation>
).
</paragraph>
<paragraph id="5E3AB15CFF965A2CFF228131FE6BFC8E" blockId="13.[151,1437,258,967]" pageId="13" pageNumber="64">
Male imago. Length (mm): body, 13.013.5; FW, 11.011.2; HW, 5.05.2; cerci, 31.0; foreleg, 5.55.7. Body coloration completely faded. Foreleg ratio of segments in relation to femur (
<quantity id="997D1CB9FF965A2CFBC0811DFB61FD56" box="[1061,1159,618,643]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="64" unit="mm" value="1.6">1.6 mm</quantity>
): femur (1), tibia (1.0), tarsomere 1 (0.1); tarsomere 2(0.3); tarsomere 3 (0.3), tarsomere 4 (0.3), tarsomere 5 (0.3), short claw (0.2), long claw (0.3). Tarsal segments 25 subequal, tarsomere 1 very short. Genitalia (
<figureCitation id="C6BEADD9FF965A2CFC1481C5FB9AFD1E" box="[1009,1148,690,715]" captionStart="FIGURES 38 40. C" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[163,1436,1036,1707]" captionTargetId="figure@13.[150,1436,1022,1716]" captionTargetPageId="13" captionText="FIGURES 38 40. C. molinai sp. nov., male adult. 38, male genitalia, v. v.; 39, pedestal, detail, v. v.; 40, penes and pedestals, v. v. (photograph). Abbreviations: ip = inner projection (pedestal); op = outer projection (pedestal); p = penes; pb = pyramidal base (penes); t = &quot; thumb &quot; (penes)." httpUri="https://zenodo.org/record/287909/files/figure.png" pageId="13" pageNumber="64">Figs. 3840</figureCitation>
): sternum IX subquadrate with rounded apical margin; pedestal stout, slightly flattened dorsoventrally, distally with large rounded inner corner and short acute outer corner (
<figureCitation id="C6BEADD9FF965A2CFDAC818DFD47FCC6" box="[585,673,762,787]" captionStart="FIGURES 38 40. C" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[163,1436,1036,1707]" captionTargetId="figure@13.[150,1436,1022,1716]" captionTargetPageId="13" captionText="FIGURES 38 40. C. molinai sp. nov., male adult. 38, male genitalia, v. v.; 39, pedestal, detail, v. v.; 40, penes and pedestals, v. v. (photograph). Abbreviations: ip = inner projection (pedestal); op = outer projection (pedestal); p = penes; pb = pyramidal base (penes); t = &quot; thumb &quot; (penes)." httpUri="https://zenodo.org/record/287909/files/figure.png" pageId="13" pageNumber="64">Fig. 39</figureCitation>
); penes strongly developed and sclerotized, curved ventrally and medially, finger-like on distal half, with small acute projection at apex; basal 1/2 of penes strongly projected dorsally (
<figureCitation id="C6BEADD9FF965A2CFEE58035FE67FC8E" box="[256,385,834,859]" captionStart="FIGURES 38 40. C" captionStartId="13.[151,264,1737,1760]" captionTargetBox="[163,1436,1036,1707]" captionTargetId="figure@13.[150,1436,1022,1716]" captionTargetPageId="13" captionText="FIGURES 38 40. C. molinai sp. nov., male adult. 38, male genitalia, v. v.; 39, pedestal, detail, v. v.; 40, penes and pedestals, v. v. (photograph). Abbreviations: ip = inner projection (pedestal); op = outer projection (pedestal); p = penes; pb = pyramidal base (penes); t = &quot; thumb &quot; (penes)." httpUri="https://zenodo.org/record/287909/files/figure.png" pageId="13" pageNumber="64">Fig. 3840</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="169FE2D7FF965A2CFF228011FB17FC13" pageId="13" pageNumber="64" type="discussion">
<paragraph id="5E3AB15CFF965A2CFF228011FB17FC13" blockId="13.[151,1437,258,967]" pageId="13" pageNumber="64">
<emphasis id="6CF16D4EFF965A2CFF228011FEA3FCAA" bold="true" box="[199,325,870,895]" pageId="13" pageNumber="64">Discussion</emphasis>
. This new species is similar to
<taxonomicName id="9985CADFFF965A2CFD518010FCA7FCAB" box="[692,833,871,894]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="64" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="6CF16D4EFF965A2CFD518010FCA7FCAB" box="[692,833,871,894]" italics="true" pageId="13" pageNumber="64">C. truncatus</emphasis>
</taxonomicName>
in genital morphology, but pedestals are stouter and penes much more developed (each lobe is very wide and large) and sclerotized. The same differences apply when compared with
<taxonomicName id="9985CADFFF965A2CFEA080D8FE32FC13" box="[325,468,942,966]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="64" phylum="Arthropoda" rank="species" species="mahunkai">
<emphasis id="6CF16D4EFF965A2CFEA080D8FE32FC13" box="[325,468,942,966]" italics="true" pageId="13" pageNumber="64">C. mahunkai</emphasis>
</taxonomicName>
, known also from
<collectingCountry id="2692F1CCFF965A2CFD4380D9FD1FFC12" box="[678,761,942,967]" name="Bolivia" pageId="13" pageNumber="64">Bolivia</collectingCountry>
but here treated as synonym of
<taxonomicName id="9985CADFFF965A2CFB8580D8FB0CFC13" box="[1120,1258,943,966]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="64" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="6CF16D4EFF965A2CFB8580D8FB0CFC13" box="[1120,1258,943,966]" italics="true" pageId="13" pageNumber="64">C. truncatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>