378 lines
41 KiB
XML
378 lines
41 KiB
XML
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<mods:title id="ED5E8FEF659FEDEE2BFE39CFC3707957">A taxonomic review of the genus Ancyronyx Erichson, 1847 from Sulawesi (Insecta: Coleoptera: Elmidae)</mods:title>
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<mods:namePart id="1CB87457A6A5FEFD6042892DABA64342">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="425CC68EFCD308FFDEBC80ECA67455CA">Kodada, Ján</mods:namePart>
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<mods:title id="781598831A1E0294F847883376A8E763">Journal of Natural History</mods:title>
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<mods:part id="F56AA26F507867E58E516C99E91AAEC0">
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<mods:date id="7AAB061F3B5C19D00F149D7BCDFC632F">2017</mods:date>
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<mods:title id="779CA733F02EFBF38D5ACC066FC87ED0">J. Nat. Hist.</mods:title>
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<mods:number id="832B31516CC239FAA8082F3BBC489448">2017-03-07</mods:number>
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<subSubSection id="69C26557FFEDFFD9AAD184E6C5BDBCD9" box="[415,900,643,669]" pageId="39" pageNumber="599" type="nomenclature">
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<paragraph id="216736DCFFEDFFD9AAD184E6C5BDBCD9" blockId="39.[415,900,643,704]" box="[415,900,643,669]" pageId="39" pageNumber="599">
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<heading id="7A2F81B0FFEDFFD9AAD184E6C5BDBCD9" box="[415,900,643,669]" centered="true" fontSize="11" level="2" pageId="39" pageNumber="599" reason="2">
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<taxonomicName id="E6D84D5FFFEDFFD9AAD184E6C5BDBCD9" ID-CoL="DPT2" authority="Jach, 2004" authorityName="Jach" authorityYear="2004" box="[415,900,643,669]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="599" phylum="Arthropoda" rank="species" species="raffaelacatharina">
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<emphasis id="13ACEACEFFEDFFD9AAD184E6C4C7BCD9" bold="true" box="[415,766,643,669]" italics="true" pageId="39" pageNumber="599">Ancyronyx raffaelacatharina</emphasis>
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<bibRefCitation id="45494B2DFFEDFFD9A84684E1C5BDBCD9" author="Jach MA" box="[776,900,644,669]" pageId="39" pageNumber="599" pagination="389 - 395" refId="ref18593" refString="Jach MA. 2004. Descriptions of two new species of Ancyronyx Erichson (Insecta: coleoptera: elmidae). Ann Naturhist Mus Wien B. 105: 389 - 395." type="journal article" year="2004">Jäch, 2004</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="69C26557FFEDFFD9AAB484C2C510BC84" box="[506,809,678,704]" pageId="39" pageNumber="599" type="description">
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<paragraph id="216736DCFFEDFFD9AAB484C2C510BC84" blockId="39.[415,900,643,704]" box="[506,809,678,704]" pageId="39" pageNumber="599">
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(
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<figureCitation id="B9E32A59FFEDFFD9A94C84C2C448BC84" box="[514,625,679,704]" captionStart="Figure 3" captionStartId="9.[160,225,1528,1551]" captionTargetBox="[164,1152,142,1496]" captionTargetId="figure-1@9.[163,1153,142,1497]" captionTargetPageId="9" captionText="Figure 3. Habitus of Ancyronyx species from Sulawesi, (a) Ancyronyx henningi from Gintu (Sul13f) (total length 1.61 mm; BL: 1.45 mm!), (b) Ancyronyx henningi from Tentena (Sul18 g) (total length 1.58 mm; BL: 1.33 mm!), (c) Ancyronyx vincentarnoldi (total length 1.80 mm), (d) Ancyronyx raffaelacatharina (total length 2.15 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746920" httpUri="https://zenodo.org/record/4746920/files/figure.png" pageId="39" pageNumber="599">Figures 3</figureCitation>
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(d), 16(a–d), 18)
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</paragraph>
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</subSubSection>
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<caption id="75A76654FFEDFFD9ABEF80FDC5AAB888" ID-DOI="http://doi.org/10.5281/zenodo.4746946" ID-Zenodo-Dep="4746946" httpUri="https://zenodo.org/record/4746946/files/figure.png" pageId="39" pageNumber="599" startId="39.[161,226,1688,1711]" targetBox="[164,1152,812,1657]" targetPageId="39">
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<paragraph id="216736DCFFEDFFD9ABEF80FDC5AAB888" blockId="39.[161,1158,1687,1740]" pageId="39" pageNumber="599">
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<emphasis id="13ACEACEFFEDFFD9ABEF80FDC735B8EB" bold="true" box="[161,268,1688,1711]" pageId="39" pageNumber="599">Figure 16.</emphasis>
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<taxonomicName id="E6D84D5FFFEDFFD9AA5B80F2C411B8EA" authorityName="Jach" authorityYear="2004" box="[277,552,1687,1711]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="599" phylum="Arthropoda" rank="species" species="raffaelacatharina">
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<emphasis id="13ACEACEFFEDFFD9AA5B80F2C411B8EA" box="[277,552,1687,1711]" italics="true" pageId="39" pageNumber="599">Ancyronyx raffaelacatharina</emphasis>
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</taxonomicName>
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from Dumoga; (a) male sternite IX, ventral; (b) aedeagus, lateral; (c) aedeagus, ventral; (d) ovipositor, ventral; (Scale bars = 0.1 mm).
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</paragraph>
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</caption>
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<caption id="75A76654FFE2FFD6ABEE843AC22FBC8A" ID-DOI="http://doi.org/10.5281/zenodo.4746950" ID-Zenodo-Dep="4746950" httpUri="https://zenodo.org/record/4746950/files/figure.png" pageId="40" pageNumber="600" startId="40.[160,225,607,630]" targetBox="[164,1152,143,576]" targetPageId="40">
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<paragraph id="216736DCFFE2FFD6ABEE843AC22FBC8A" blockId="40.[160,1155,606,718]" pageId="40" pageNumber="600">
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<emphasis id="13ACEACEFFE2FFD6ABEE843AC73EBC32" bold="true" box="[160,263,607,630]" pageId="40" pageNumber="600">Figure 17.</emphasis>
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Statistical parsimony tree of mtDNA haplotypes produced by TCS based on
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<emphasis id="13ACEACEFFE2FFD6A8B8843BC21BBC31" box="[1014,1058,606,629]" italics="true" pageId="40" pageNumber="600">cox1</emphasis>
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fragment sequences. The tree is scaled in horizontal direction by number of substitutions,
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<emphasis id="13ACEACEFFE2FFD6A8F58419C5F9BCD7" box="[955,960,636,659]" italics="true" pageId="40" pageNumber="600">‘</emphasis>
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missing haplotypes
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<emphasis id="13ACEACEFFE2FFD6AF308419C2BABCD7" box="[1150,1155,636,659]" italics="true" pageId="40" pageNumber="600">’</emphasis>
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||
are not indicated. Circles represent existing haplotypes from the sample set, with circle size indicating the number of specimens with identical haplotype at the respective position.
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</paragraph>
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</caption>
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<subSubSection id="69C26557FFE2FFD6ABEE8569C4AABD60" box="[160,659,780,804]" pageId="40" pageNumber="600" type="reference_group">
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<paragraph id="216736DCFFE2FFD6ABEE8569C4AABD60" blockId="40.[160,659,780,804]" box="[160,659,780,804]" pageId="40" pageNumber="600">
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<treatmentCitationGroup id="01C811F2FFE2FFD6ABEE8569C4AABD60" box="[160,659,780,804]" pageId="40" pageNumber="600">
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<treatmentCitation id="A07910CDFFE2FFD6ABEE8569C4B6BD60" author="Jach MA" box="[160,655,780,804]" page="392" pageId="40" pageNumber="600" year="2004">
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<taxonomicName id="E6D84D5FFFE2FFD6ABEE8569C4B6BD60" ID-CoL="DPT2" authority="Jach, 2004: 392" authorityName="Jach" authorityPageNumber="392" authorityYear="2004" box="[160,655,780,804]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="600" phylum="Arthropoda" rank="species" species="raffaelacatharina">
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<emphasis id="13ACEACEFFE2FFD6ABEE8569C7ECBD60" box="[160,469,780,804]" italics="true" pageId="40" pageNumber="600">Ancyronyx raffaelacatharina</emphasis>
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<bibRefCitation id="45494B2DFFE2FFD6AA908569C4B6BD60" author="Jach MA" box="[478,655,780,804]" pageId="40" pageNumber="600" pagination="389 - 395" refId="ref18593" refString="Jach MA. 2004. Descriptions of two new species of Ancyronyx Erichson (Insecta: coleoptera: elmidae). Ann Naturhist Mus Wien B. 105: 389 - 395." type="journal article" year="2004">Jäch, 2004: 392</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="69C26557FFE2FFD7ABEE853DC58DBB70" lastPageId="41" lastPageNumber="601" pageId="40" pageNumber="600" type="materials_examined">
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<paragraph id="216736DCFFE2FFD6ABEE853DC42CBA05" blockId="40.[160,1157,856,1089]" pageId="40" pageNumber="600">
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<emphasis id="13ACEACEFFE2FFD6ABEE853DC7B3BD35" bold="true" box="[160,394,856,881]" italics="true" pageId="40" pageNumber="600">Material examined.</emphasis>
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<materialsCitation id="91B03C81FFE2FFD6AAD5853CC7B9BDD7" collectionCode="NMW" country="Indonesia" location="mountain Riv" pageId="40" pageNumber="600" specimenCode="FR109" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara">
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<specimenCount id="37DEFD55FFE2FFD6AAD5853CC7F1BD34" box="[411,456,856,881]" pageId="40" pageNumber="600" type="male">1 ♂</specimenCount>
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(
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<collectionCode id="47C9AE19FFE2FFD6AA97853CC421BD35" box="[473,536,857,881]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="600">NMW</collectionCode>
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[
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<specimenCode id="717E9EA7FFE2FFD6A967853CC44FBD35" box="[553,630,857,881]" collectionCode="NMW" country="Austria" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="600">FR109</specimenCode>
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]): ‘
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<collectingCountry id="59CF764CFFE2FFD6A9DB853CC52EBD35" box="[661,791,857,881]" name="Indonesia" pageId="40" pageNumber="600">INDONESIA</collectingCountry>
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: SE
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<collectingRegion id="E31CF83EFFE2FFD6A805853CC596BD35" box="[843,943,857,881]" country="Indonesia" name="Sulawesi Utara" pageId="40" pageNumber="600">Sulawesi</collectingRegion>
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,
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N Kolaka
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</locationDeviation>
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, small
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<location id="24076007FFE2FFD6ABAA851EC7B9BDD7" LSID="urn:lsid:plazi:treatment:A97187CAFFEDFFD5AAD184E6C7CDBF32:24076007FFE2FFD6ABAA851EC7B9BDD7" box="[228,384,891,915]" country="Indonesia" name="mountain Riv" pageId="40" pageNumber="600" stateProvince="Sulawesi Utara">mountain Riv</location>
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</materialsCitation>
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.; secondary forest; run, submerged wood;
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a
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<materialsCitation id="91B03C81FFE2FFD6A8F0851EC5E8BDD7" box="[958,977,891,915]" collectionCode="NMW" country="Indonesia" isTailEnumeration="true" location="mountain Riv" pageId="40" pageNumber="600" specimenCount="1" stateProvince="Sulawesi Utara">.s</materialsCitation>
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.l., c.
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04°01
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<emphasis id="13ACEACEFFE2FFD6AF74851EC279BDD7" bold="true" box="[1082,1088,891,915]" italics="true" pageId="40" pageNumber="600">′</emphasis>
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28
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<emphasis id="13ACEACEFFE2FFD6AF15851EC25EBDD7" bold="true" box="[1115,1127,891,915]" italics="true" pageId="40" pageNumber="600">′′</emphasis>
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S
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</geoCoordinate>
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,
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<geoCoordinate id="44EC501BFFE2FFD6ABEE85FBC717BDF2" box="[160,302,926,950]" degrees="121" direction="east" minutes="36" orientation="longitude" pageId="40" pageNumber="600" precision="15" seconds="16" value="121.60445">
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121°36
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<emphasis id="13ACEACEFFE2FFD6ABA385FBC6CABDF2" bold="true" box="[237,243,926,950]" italics="true" pageId="40" pageNumber="600">′</emphasis>
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16
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<emphasis id="13ACEACEFFE2FFD6AA4085FBC723BDF2" bold="true" box="[270,282,926,950]" italics="true" pageId="40" pageNumber="600">′′</emphasis>
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E
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</geoCoordinate>
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<date id="5566101CFFE2FFD6AA7B85FBC784BDF2" box="[309,445,926,950]" pageId="40" pageNumber="600" value="1997-12-28">28 Dec.1997</date>
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leg
|
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<materialsCitation id="91B03C81FFE2FFD6AAA785FBC7E6BD9D" collectionCode="CFM" collectorName="Freitag & Sul" country="Indonesia" location="mountain Riv" pageId="40" pageNumber="600" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara">
|
||
.
|
||
<collectorName id="8C2D530AFFE2FFD6AABD85FBC478BDF2" box="[499,577,926,950]" pageId="40" pageNumber="600">Freitag</collectorName>
|
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(
|
||
<collectorName id="8C2D530AFFE2FFD6A90085FBC44ABDF2" box="[590,627,926,950]" pageId="40" pageNumber="600">Sul</collectorName>
|
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2f)’;
|
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<specimenCount id="37DEFD55FFE2FFD6A9EE85FBC4F1BDF2" box="[672,712,926,950]" pageId="40" pageNumber="600" type="female">1 ♀</specimenCount>
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(
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<collectionCode id="47C9AE19FFE2FFD6A99A85FBC535BDF2" box="[724,780,926,950]" pageId="40" pageNumber="600">CFM</collectionCode>
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): ‘
|
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<collectingCountry id="59CF764CFFE2FFD6A86F85FBC59ABDF2" box="[801,931,926,950]" name="Indonesia" pageId="40" pageNumber="600">INDONESIA</collectingCountry>
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||
: SE
|
||
<collectingRegion id="E31CF83EFFE2FFD6A88285FBC209BDF2" box="[972,1072,926,950]" country="Indonesia" name="Sulawesi Utara" pageId="40" pageNumber="600">Sulawesi</collectingRegion>
|
||
,
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||
<locationDeviation id="30D52FFEFFE2FFD6AF7785FBC711BD9D" location="Kolaka" pageId="40" pageNumber="600">
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<quantity id="E6209B39FFE2FFD6AF7785FBC2BABDF2" box="[1081,1155,926,950]" metricMagnitude="3" metricUnit="m" metricValue="8.1" pageId="40" pageNumber="600" unit="km" value="8.1">8.1 km</quantity>
|
||
ENE Kolaka
|
||
</locationDeviation>
|
||
,
|
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<location id="24076007FFE2FFD6AA7585A4C7E6BD9D" LSID="urn:lsid:plazi:treatment:A97187CAFFEDFFD5AAD184E6C7CDBF32:24076007FFE2FFD6AA7585A4C7E6BD9D" box="[315,479,961,985]" country="Indonesia" name="mountain Riv" pageId="40" pageNumber="600" stateProvince="Sulawesi Utara">mountain Riv</location>
|
||
</materialsCitation>
|
||
.; secondary forest; run, submerged wood;
|
||
<quantity id="E6209B39FFE2FFD6A8BE85A4C206BD9D" box="[1008,1087,961,985]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="40" pageNumber="600" unit="m" value="155.0">155 m</quantity>
|
||
a
|
||
<materialsCitation id="91B03C81FFE2FFD6AF2F85A4C24DBD9D" box="[1121,1140,961,985]" collectionCode="CFM" collectorName="Freitag & Sul" country="Indonesia" isTailEnumeration="true" location="mountain Riv" pageId="40" pageNumber="600" specimenCount="1" stateProvince="Sulawesi Utara">.s</materialsCitation>
|
||
.l., c.
|
||
<geoCoordinate id="44EC501BFFE2FFD6ABE18586C702BDBF" box="[175,315,995,1019]" degrees="04" direction="south" minutes="02" orientation="latitude" pageId="40" pageNumber="600" precision="15" seconds="06" value="-4.035">
|
||
04°02
|
||
<emphasis id="13ACEACEFFE2FFD6ABBF8586C6CEBDBF" bold="true" box="[241,247,995,1019]" italics="true" pageId="40" pageNumber="600">′</emphasis>
|
||
06
|
||
<emphasis id="13ACEACEFFE2FFD6AA5C8586C727BDBF" bold="true" box="[274,286,995,1019]" italics="true" pageId="40" pageNumber="600">′′</emphasis>
|
||
S
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="44EC501BFFE2FFD6AA058586C7D9BDBF" box="[331,480,995,1019]" degrees="121" direction="east" minutes="40" orientation="longitude" pageId="40" pageNumber="600" precision="15" seconds="04" value="121.66778">
|
||
121°40
|
||
<emphasis id="13ACEACEFFE2FFD6AAD68586C7A7BDBF" bold="true" box="[408,414,995,1019]" italics="true" pageId="40" pageNumber="600">′</emphasis>
|
||
04
|
||
<emphasis id="13ACEACEFFE2FFD6AAF78586C7FCBDBF" bold="true" box="[441,453,995,1019]" italics="true" pageId="40" pageNumber="600">′′</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
<date id="5566101CFFE2FFD6AAA38586C445BDBF" box="[493,636,995,1019]" pageId="40" pageNumber="600" value="1997-12-29">29 Dec.1997</date>
|
||
leg
|
||
<materialsCitation id="91B03C81FFE2FFD6A9E08586C501BA5A" collectionCode="CSH, CKB" collectorName="Freitag & Sul" country="Indonesia" location="Dumoga Bone" pageId="40" pageNumber="600" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Sulawesi Utara">
|
||
.
|
||
<collectorName id="8C2D530AFFE2FFD6A9F08586C535BDBF" box="[702,780,995,1019]" pageId="40" pageNumber="600">Freitag</collectorName>
|
||
(
|
||
<collectorName id="8C2D530AFFE2FFD6A8518586C57CBDBF" box="[799,837,995,1019]" pageId="40" pageNumber="600">Sul</collectorName>
|
||
3f)’;
|
||
<specimenCount id="37DEFD55FFE2FFD6A8368586C595BDBF" box="[888,940,995,1019]" pageId="40" pageNumber="600" type="male">3♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="37DEFD55FFE2FFD6A88E8586C5CDBDBF" box="[960,1012,995,1019]" pageId="40" pageNumber="600" type="female">2♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="47C9AE19FFE2FFD6AF498586C203BDBF" box="[1031,1082,995,1019]" country="China" name="Chenshan Botanical Garden" pageId="40" pageNumber="600" type="Herbarium">CSH</collectionCode>
|
||
,
|
||
<collectionCode id="47C9AE19FFE2FFD6AF048586C242BDBF" box="[1098,1147,995,1019]" pageId="40" pageNumber="600">CKB</collectionCode>
|
||
): ‘
|
||
<collectingCountry id="59CF764CFFE2FFD6ABE88263C71CBA5A" box="[166,293,1030,1054]" name="Indonesia" pageId="40" pageNumber="600">INDONESIA</collectingCountry>
|
||
N
|
||
<collectingRegion id="E31CF83EFFE2FFD6AA068263C790BA5A" box="[328,425,1030,1054]" country="Indonesia" name="Sulawesi Utara" pageId="40" pageNumber="600">Sulawesi</collectingRegion>
|
||
<locationDeviation id="30D52FFEFFE2FFD6AAFF8263C466BA5A" box="[433,607,1030,1054]" location="Toraut" pageId="40" pageNumber="600">
|
||
<quantity id="E6209B39FFE2FFD6AAFF8263C7D2BA5A" box="[433,491,1030,1054]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="600" unit="km" value="1.0">1 km</quantity>
|
||
W Toraut
|
||
</locationDeviation>
|
||
,
|
||
<location id="24076007FFE2FFD6A9258263C537BA5A" LSID="urn:lsid:plazi:treatment:A97187CAFFEDFFD5AAD184E6C7CDBF32:24076007FFE2FFD6A9258263C537BA5A" box="[619,782,1030,1054]" country="Indonesia" name="Dumoga Bone" pageId="40" pageNumber="600" stateProvince="Sulawesi Utara">Dumoga Bone</location>
|
||
NP
|
||
</materialsCitation>
|
||
.
|
||
<quantity id="E6209B39FFE2FFD6A8758263C584BA5A" box="[827,957,1030,1054]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="40" pageNumber="600" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200–300 m</quantity>
|
||
,
|
||
<geoCoordinate id="44EC501BFFE2FFD6A8878263C27CBA5A" box="[969,1093,1030,1054]" degrees="0" direction="north" minutes="34" orientation="latitude" pageId="40" pageNumber="600" precision="15" seconds="17" value="0.5713889">
|
||
0°34
|
||
<emphasis id="13ACEACEFFE2FFD6A8B58263C238BA5A" bold="true" box="[1019,1025,1030,1054]" italics="true" pageId="40" pageNumber="600">′</emphasis>
|
||
17
|
||
<emphasis id="13ACEACEFFE2FFD6AF528263C211BA5A" bold="true" box="[1052,1064,1030,1054]" italics="true" pageId="40" pageNumber="600">′′</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="44EC501BFFE2FFD6AF1C8263C6C4BA05" degrees="123" direction="east" minutes="54" orientation="longitude" pageId="40" pageNumber="600" precision="15" seconds="19" value="123.90528">
|
||
123° 54
|
||
<emphasis id="13ACEACEFFE2FFD6ABF5824DC6F8BA04" bold="true" box="[187,193,1064,1088]" italics="true" pageId="40" pageNumber="600">′</emphasis>
|
||
19
|
||
<emphasis id="13ACEACEFFE2FFD6AB93824DC6D0BA04" bold="true" box="[221,233,1064,1088]" italics="true" pageId="40" pageNumber="600">′′</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
<date id="5566101CFFE2FFD6AA4B824CC744BA05" box="[261,381,1065,1089]" pageId="40" pageNumber="600" value="2006-02-01" valueMax="2006-02-02" valueMin="2006-02-01">
|
||
1
|
||
<materialsCitation id="91B03C81FFE2FFD6AA42824CC71FBA05" box="[268,294,1065,1089]" collectionCode="CSH, CKB" collectorName="Freitag & Sul" country="Indonesia" isTailEnumeration="true" location="Dumoga Bone" pageId="40" pageNumber="600" specimenCount="1" stateProvince="Sulawesi Utara">.-2</materialsCitation>
|
||
.II.2006
|
||
</date>
|
||
leg
|
||
<materialsCitation id="91B03C81FFE2FFD6AAE6824CC7FFBA05" box="[424,454,1065,1089]" collectionCode="CSH, CKB" collectorName="Freitag & Sul" country="Indonesia" isTailEnumeration="true" location="Dumoga Bone" pageId="40" pageNumber="600" specimenCount="1" stateProvince="Sulawesi Utara">. A</materialsCitation>
|
||
. Skale’.
|
||
</paragraph>
|
||
<paragraph id="216736DCFFE2FFD6ABEE8211C787BAC9" blockId="40.[160,1156,1140,1373]" box="[160,446,1140,1165]" pageId="40" pageNumber="600">
|
||
<emphasis id="13ACEACEFFE2FFD6ABEE8211C787BAC9" bold="true" box="[160,446,1140,1165]" italics="true" pageId="40" pageNumber="600">Additions to description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="216736DCFFE2FFD6ABEE82FDC4C3BA97" blockId="40.[160,1156,1140,1373]" pageId="40" pageNumber="600">
|
||
In addition to the original species description based on
|
||
<specimenCount id="37DEFD55FFE2FFD6A80982FDC5E1BAF4" box="[839,984,1176,1200]" pageId="40" pageNumber="600" type="female">two females</specimenCount>
|
||
, the following characters are of extended species-level significance:
|
||
</paragraph>
|
||
<paragraph id="216736DCFFE2FFD6AB8E82B8C7C1BB19" blockId="40.[160,1156,1140,1373]" pageId="40" pageNumber="600">
|
||
Body
|
||
<quantity id="E6209B39FFE2FFD6AA4E82B8C7BABAB1" box="[256,387,1245,1269]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="40" pageNumber="600" unit="mm" value="1.9" valueMax="2.1" valueMin="1.7">1.7–2.1 mm</quantity>
|
||
long, 2.2–2.5 times as long as wide (
|
||
<collectionCode id="47C9AE19FFE2FFD6A86E82B8C507BAB1" box="[800,830,1245,1269]" pageId="40" pageNumber="600">BL</collectionCode>
|
||
/
|
||
<collectionCode id="47C9AE19FFE2FFD6A80882B8C557BAB1" box="[838,878,1245,1269]" pageId="40" pageNumber="600">EW</collectionCode>
|
||
),
|
||
<collectionCode id="47C9AE19FFE2FFD6A83082B8C59DBAB1" box="[894,932,1245,1269]" pageId="40" pageNumber="600">EW</collectionCode>
|
||
:
|
||
<quantity id="E6209B39FFE2FFD6A8E182B8C276BAB1" box="[943,1103,1245,1269]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="40" pageNumber="600" unit="mm" value="0.8" valueMax="0.85" valueMin="0.75">0.75–0.85 mm</quantity>
|
||
. The dark elytral portions, however, are slightly extended, and the anterior yellowish pattern is broadly V-shaped (
|
||
<figureCitation id="B9E32A59FFE2FFD6AAC08346C7D5BB7F" box="[398,492,1315,1339]" captionStart="Figure 3" captionStartId="9.[160,225,1528,1551]" captionTargetBox="[164,1152,142,1496]" captionTargetId="figure-1@9.[163,1153,142,1497]" captionTargetPageId="9" captionText="Figure 3. Habitus of Ancyronyx species from Sulawesi, (a) Ancyronyx henningi from Gintu (Sul13f) (total length 1.61 mm; BL: 1.45 mm!), (b) Ancyronyx henningi from Tentena (Sul18 g) (total length 1.58 mm; BL: 1.33 mm!), (c) Ancyronyx vincentarnoldi (total length 1.80 mm), (d) Ancyronyx raffaelacatharina (total length 2.15 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746920" httpUri="https://zenodo.org/record/4746920/files/figure.png" pageId="40" pageNumber="600">Figure 3</figureCitation>
|
||
(d)). Most of the specimens bear entirely dark brown head, without any pale dorsal areas.
|
||
</paragraph>
|
||
<paragraph id="216736DCFFE2FFD6ABEE83F4C422BBAB" blockId="40.[160,1156,1425,1727]" pageId="40" pageNumber="600">
|
||
<emphasis id="13ACEACEFFE2FFD6ABEE83F4C721BBEE" bold="true" box="[160,280,1425,1450]" italics="true" pageId="40" pageNumber="600">Abdomen.</emphasis>
|
||
Male sternite IX (
|
||
<figureCitation id="B9E32A59FFE2FFD6AAA483F7C46CBBEE" box="[490,597,1426,1450]" captionStart="Figure 16" captionStartId="39.[161,226,1688,1711]" captionTargetBox="[164,1152,812,1657]" captionTargetId="figure-198@39.[164,1153,811,1657]" captionTargetPageId="39" captionText="Figure 16. Ancyronyx raffaelacatharina from Dumoga; (a) male sternite IX, ventral; (b) aedeagus, lateral; (c) aedeagus, ventral; (d) ovipositor, ventral; (Scale bars = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746946" httpUri="https://zenodo.org/record/4746946/files/figure.png" pageId="40" pageNumber="600">Figure 16</figureCitation>
|
||
(a)) deeply excised apically; medioapical rim with fine pubescence; lateroapical portions with c.10 small spines; paraprocts moderately long, not reaching apical margin.
|
||
</paragraph>
|
||
<paragraph id="216736DCFFE2FFD7AB8E839FC58DBB70" blockId="40.[160,1156,1425,1727]" lastBlockId="41.[160,1156,1274,1332]" lastPageId="41" lastPageNumber="601" pageId="40" pageNumber="600">
|
||
Aedeagus (
|
||
<figureCitation id="B9E32A59FFE2FFD6AA70839FC790B856" box="[318,425,1530,1554]" captionStart="Figure 16" captionStartId="39.[161,226,1688,1711]" captionTargetBox="[164,1152,812,1657]" captionTargetId="figure-198@39.[164,1153,811,1657]" captionTargetPageId="39" captionText="Figure 16. Ancyronyx raffaelacatharina from Dumoga; (a) male sternite IX, ventral; (b) aedeagus, lateral; (c) aedeagus, ventral; (d) ovipositor, ventral; (Scale bars = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746946" httpUri="https://zenodo.org/record/4746946/files/figure.png" pageId="40" pageNumber="600">Figure 16</figureCitation>
|
||
(b,c))
|
||
<quantity id="E6209B39FFE2FFD6AAA8839FC402B856" box="[486,571,1530,1554]" metricMagnitude="-1" metricUnit="m" metricValue="3.65" pageId="40" pageNumber="600" unit="mm" value="365.0">365 µm</quantity>
|
||
long. Median lobe
|
||
<quantity id="E6209B39FFE2FFD6A858839FC564B856" box="[790,861,1530,1554]" metricMagnitude="-2" metricUnit="m" metricValue="7.8" pageId="40" pageNumber="600" unit="mm" value="78.0">78 µm</quantity>
|
||
wide and evenly tapering posteriad; apex broadly rounded; subapical portion abruptly bent ventrad (lateral view); lateral portion with few short spines; ventral sac and fibula as in previous species; corona conspicuous. Phallobase asymmetrical; basal margin obliquely concave, moderately sclerotized. Parameres with about six setae of varying size (three moderately short ones at apex and three moderately to very long ones situated more towards the base); surface additionally with scattered minute, spine-like setae; parameres short, reaching about 0.76 of length of aedeagus, not contiguous ventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="75A76654FFE3FFD7ABEE82C1C267BAFF" ID-DOI="http://doi.org/10.5281/zenodo.4746954" ID-Zenodo-Dep="4746954" box="[160,1118,1188,1211]" httpUri="https://zenodo.org/record/4746954/files/figure.png" pageId="41" pageNumber="601" startId="41.[160,225,1188,1211]" targetBox="[163,1151,142,1157]" targetPageId="41">
|
||
<paragraph id="216736DCFFE3FFD7ABEE82C1C267BAFF" blockId="41.[160,1118,1188,1211]" box="[160,1118,1188,1211]" pageId="41" pageNumber="601">
|
||
<emphasis id="13ACEACEFFE3FFD7ABEE82C1C730BAFF" bold="true" box="[160,265,1188,1211]" pageId="41" pageNumber="601">Figure 18.</emphasis>
|
||
Map of Sulawesi with collecting site of
|
||
<taxonomicName id="E6D84D5FFFE3FFD7A9EE82C1C56BBAFF" box="[672,850,1188,1211]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="601" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="13ACEACEFFE3FFD7A9EE82C1C53ABAFF" box="[672,771,1188,1211]" italics="true" pageId="41" pageNumber="601">Ancyronyx</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
based on their label data.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="69C26557FFE3FFD7ABEE830DC7B0BB82" pageId="41" pageNumber="601" type="distribution">
|
||
<paragraph id="216736DCFFE3FFD7ABEE830DC715BBC5" blockId="41.[160,1155,1384,1478]" box="[160,300,1384,1409]" pageId="41" pageNumber="601">
|
||
<emphasis id="13ACEACEFFE3FFD7ABEE830DC715BBC5" bold="true" box="[160,300,1384,1409]" italics="true" pageId="41" pageNumber="601">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="216736DCFFE3FFD7ABEE83EEC7B0BB82" blockId="41.[160,1155,1384,1478]" pageId="41" pageNumber="601">
|
||
The species is known from several localities on southeastern, central, and northern
|
||
<collectingRegion id="E31CF83EFFE3FFD7ABEE83CBC738BB82" box="[160,257,1454,1478]" country="Indonesia" name="Sulawesi Utara" pageId="41" pageNumber="601">Sulawesi</collectingRegion>
|
||
(
|
||
<figureCitation id="B9E32A59FFE3FFD7AA5F83CBC745BB82" box="[273,380,1454,1478]" captionStart="Figure 18" captionStartId="41.[160,225,1188,1211]" captionTargetBox="[163,1151,142,1157]" captionTargetId="figure-116@41.[163,1152,142,1157]" captionTargetPageId="41" captionText="Figure 18. Map of Sulawesi with collecting site of Ancyronyx species based on their label data." figureDoi="http://doi.org/10.5281/zenodo.4746954" httpUri="https://zenodo.org/record/4746954/files/figure.png" pageId="41" pageNumber="601">Figure 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="69C26557FFE3FFD5ABEE839CC7CDBF32" lastPageId="43" lastPageNumber="603" pageId="41" pageNumber="601" type="discussion">
|
||
<paragraph id="216736DCFFE3FFD7ABEE839CC727B856" blockId="41.[160,1155,1529,1727]" box="[160,286,1529,1554]" pageId="41" pageNumber="601">
|
||
<emphasis id="13ACEACEFFE3FFD7ABEE839CC727B856" bold="true" box="[160,286,1529,1554]" italics="true" pageId="41" pageNumber="601">Comments</emphasis>
|
||
</paragraph>
|
||
<paragraph id="216736DCFFE3FFD5ABEE8078C5BABF75" blockId="41.[160,1155,1529,1727]" lastBlockId="43.[160,1156,142,374]" lastPageId="43" lastPageNumber="603" pageId="41" pageNumber="601">
|
||
The specimens examined agree quite well with the description and illustrations of the previously known
|
||
<specimenCount id="37DEFD55FFE3FFD7AA3C805AC7C6B813" box="[370,511,1599,1623]" pageId="41" pageNumber="601" type="female">two females</specimenCount>
|
||
. The ovipositor (
|
||
<figureCitation id="B9E32A59FFE3FFD7A9F1805AC515B813" box="[703,812,1599,1623]" captionStart="Figure 16" captionStartId="39.[161,226,1688,1711]" captionTargetBox="[164,1152,812,1657]" captionTargetId="figure-198@39.[164,1153,811,1657]" captionTargetPageId="39" captionText="Figure 16. Ancyronyx raffaelacatharina from Dumoga; (a) male sternite IX, ventral; (b) aedeagus, lateral; (c) aedeagus, ventral; (d) ovipositor, ventral; (Scale bars = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746946" httpUri="https://zenodo.org/record/4746946/files/figure.png" pageId="41" pageNumber="601">Figure 16</figureCitation>
|
||
(d)) corresponds well with the description (comp.
|
||
<bibRefCitation id="45494B2DFFE3FFD7AA378007C7D1B83E" author="Jach MA" box="[377,488,1634,1658]" pageId="41" pageNumber="601" pagination="389 - 395" refId="ref18593" refString="Jach MA. 2004. Descriptions of two new species of Ancyronyx Erichson (Insecta: coleoptera: elmidae). Ann Naturhist Mus Wien B. 105: 389 - 395." type="journal article" year="2004">Jäch 2004</bibRefCitation>
|
||
: p. 395 and
|
||
<figureCitation id="B9E32A59FFE3FFD7A93D8007C4F6B83E" box="[627,719,1634,1658]" captionStart="Figure 7" captionStartId="17.[160,225,1383,1406]" captionTargetBox="[163,1153,142,1352]" captionTargetId="figure-72@17.[163,1153,142,1352]" captionTargetPageId="17" captionText="Figure 7. Ancyronyx skalei from Dumoga; (a) male sternite IX, ventral; (b) aedeagus, lateral; (c) aedeagus, ventral; (d) ovipositor, ventral; (Scale bars = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746928" httpUri="https://zenodo.org/record/4746928/files/figure.png" pageId="41" pageNumber="601">Figure 7</figureCitation>
|
||
), although in the specimens examined herein, it is slightly larger (
|
||
<quantity id="E6209B39FFE3FFD7AA8880E0C424B8D9" box="[454,541,1669,1693]" metricMagnitude="-1" metricUnit="m" metricValue="3.95" pageId="41" pageNumber="601" unit="mm" value="395.0">395 µm</quantity>
|
||
long) and the number of setae as well as the extent of strioles at the coxite vary. However, these minor differences are considered as intraspecific variation only. This species may be readily distinguished from other members of the
|
||
<taxonomicName id="E6D84D5FFFE1FFD5AA6886D4C439BE8D" box="[294,512,177,201]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="603" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="13ACEACEFFE1FFD5AA6886D4C439BE8D" box="[294,512,177,201]" italics="true" pageId="43" pageNumber="603">Ancyronyx variabilis</emphasis>
|
||
</taxonomicName>
|
||
group from
|
||
<collectingRegion id="E31CF83EFFE1FFD5A9DA86D4C4CCBE8D" box="[660,757,177,201]" country="Indonesia" name="Sulawesi Utara" pageId="43" pageNumber="603">Sulawesi</collectingRegion>
|
||
by the unique extensive yellowish colour pattern of the elytra and by the form of the pronotum. It shares with
|
||
<taxonomicName id="E6D84D5FFFE1FFD5ABEE8693C765BF4A" authorityName="Freitag & Kodada" authorityYear="2017" box="[160,348,246,270]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="603" phylum="Arthropoda" rank="species" species="vincentarnoldi">
|
||
<emphasis id="13ACEACEFFE1FFD5ABEE8693C765BF4A" box="[160,348,246,270]" italics="true" pageId="43" pageNumber="603">A. vincentarnoldi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B9E32A59FFE1FFD5AA3E8693C7D9BF4A" box="[368,480,246,270]" captionStart="Figure 15" captionStartId="36.[160,225,1596,1619]" captionTargetBox="[164,1152,648,1565]" captionTargetId="figure-120@36.[163,1153,648,1565]" captionTargetPageId="36" captionText="Figure 15. Ancyronyx vincentarnoldi from Kolaka (male: Sul2f) and (female: Sul3f); (a) male sternite IX, ventral, mutilated; (b) aedeagus, lateral; (c) aedeagus, ventral; D: ovipositor, ventral; (Scale bars = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746944" httpUri="https://zenodo.org/record/4746944/files/figure.png" pageId="43" pageNumber="603">Figure 15</figureCitation>
|
||
(b,c)) the similar shape and proportions of the aedeagus; for differences see key and comments under that species above.
|
||
</paragraph>
|
||
<caption id="75A76654FFE0FFD4ABEE8020C2BDB895" ID-DOI="http://doi.org/10.5281/zenodo.4746956" ID-Zenodo-Dep="4746956" httpUri="https://zenodo.org/record/4746956/files/figure.png" pageId="42" pageNumber="602" startId="42.[160,225,1605,1628]" targetBox="[163,1152,169,1575]" targetPageId="42">
|
||
<paragraph id="216736DCFFE0FFD4ABEE8020C2BDB895" blockId="42.[160,1156,1605,1746]" pageId="42" pageNumber="602">
|
||
<emphasis id="13ACEACEFFE0FFD4ABEE8020C732B818" bold="true" box="[160,267,1605,1628]" pageId="42" pageNumber="602">Figure 19.</emphasis>
|
||
Habitats of
|
||
<taxonomicName id="E6D84D5FFFE0FFD4AAC18020C47AB818" box="[399,579,1605,1628]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="602" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="13ACEACEFFE0FFD4AAC18020C7CBB818" box="[399,498,1605,1628]" italics="true" pageId="42" pageNumber="602">Ancyronyx</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in Sulawesi, (a) the eastern slopes of the Pegunungan Fennema Mountains from where
|
||
<taxonomicName id="E6D84D5FFFE0FFD4A94B8007C533B83D" authorityName="Freitag & Kodada" authorityYear="2017" box="[517,778,1634,1657]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="602" phylum="Arthropoda" rank="species" species="breviparamerus">
|
||
<emphasis id="13ACEACEFFE0FFD4A94B8007C533B83D" box="[517,778,1634,1657]" italics="true" pageId="42" pageNumber="602">Ancyronyx breviparamerus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E6D84D5FFFE0FFD4A8538007C5D9B83D" authorityName="Freitag & Kodada" authorityYear="2017" box="[797,992,1634,1657]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="602" phylum="Arthropoda" rank="species" species="henningi">
|
||
<emphasis id="13ACEACEFFE0FFD4A8538007C5D9B83D" box="[797,992,1634,1657]" italics="true" pageId="42" pageNumber="602">Ancyronyx henningi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E6D84D5FFFE0FFD4AF6F8007C6DDB8D2" authorityName="Freitag & Kodada" authorityYear="2017" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="602" phylum="Arthropoda" rank="species" species="tobada">
|
||
<emphasis id="13ACEACEFFE0FFD4AF6F8007C6DDB8D2" italics="true" pageId="42" pageNumber="602">Ancyronyx tobada</emphasis>
|
||
</taxonomicName>
|
||
were collected, (b) a small mountain creek in the Pegunungan Fennema Mountains (A), type locality of
|
||
<taxonomicName id="E6D84D5FFFE0FFD4AA0E80F8C478B8F0" authorityName="Freitag & Kodada" authorityYear="2017" box="[320,577,1693,1716]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="602" phylum="Arthropoda" rank="species" species="breviparamerus">
|
||
<emphasis id="13ACEACEFFE0FFD4AA0E80F8C478B8F0" box="[320,577,1693,1716]" italics="true" pageId="42" pageNumber="602">Ancyronyx breviparamerus</emphasis>
|
||
</taxonomicName>
|
||
and collection site of
|
||
<taxonomicName id="E6D84D5FFFE0FFD4A86980F8C5EEB8F0" authorityName="Freitag & Kodada" authorityYear="2017" box="[807,983,1693,1716]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="602" phylum="Arthropoda" rank="species" species="tobada">
|
||
<emphasis id="13ACEACEFFE0FFD4A86980F8C5EEB8F0" box="[807,983,1693,1716]" italics="true" pageId="42" pageNumber="602">Ancyronyx tobada</emphasis>
|
||
</taxonomicName>
|
||
, (c) Salu Maiting River in North Toraja, type locality of
|
||
<taxonomicName id="E6D84D5FFFE0FFD4A96380DFC508B895" authorityName="Freitag & Kodada" authorityYear="2017" box="[557,817,1722,1745]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="602" phylum="Arthropoda" rank="species" species="longiparamerus">
|
||
<emphasis id="13ACEACEFFE0FFD4A96380DFC508B895" box="[557,817,1722,1745]" italics="true" pageId="42" pageNumber="602">Ancyronyx longiparamerus</emphasis>
|
||
</taxonomicName>
|
||
and collection site of
|
||
<taxonomicName id="E6D84D5FFFE0FFD4AF6F80DFC2BDB895" authorityName="Erichson" authorityYear="1847" box="[1057,1156,1722,1745]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="602" phylum="Arthropoda" rank="genus">Ancyronyx</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="216736DCFFE1FFD5AB8E8759C7CDBF32" blockId="43.[160,1156,142,374]" pageId="43" pageNumber="603">
|
||
The newly collected specimens were found syntopically with
|
||
<taxonomicName id="E6D84D5FFFE1FFD5A83E8759C2BABF10" authorityName="Freitag & Kodada" authorityYear="2017" box="[880,1155,316,340]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="603" phylum="Arthropoda" rank="species" species="vincentarnoldi">
|
||
<emphasis id="13ACEACEFFE1FFD5A83E8759C2BABF10" box="[880,1155,316,340]" italics="true" pageId="43" pageNumber="603">Ancyronyx vincentarnoldi</emphasis>
|
||
</taxonomicName>
|
||
(see comments for the latter).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |