311 lines
38 KiB
XML
311 lines
38 KiB
XML
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<mods:title id="A5918FDF9E95DC18449D0326E9F59070">A taxonomic review of the genus Ancyronyx Erichson, 1847 from Sulawesi (Insecta: Coleoptera: Elmidae)</mods:title>
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<mods:namePart id="05369032EA5D69D71D68DC7E845597B4">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="1452F805FFEDE8166168A593F5472AC7">Kodada, Ján</mods:namePart>
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<mods:title id="3CF23C72A000B600B2EF2F27BDADF8F5">Journal of Natural History</mods:title>
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<mods:part id="4B0D13F2922017DBDA6A5CA72D15552C">
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<mods:date id="AD9B3C0BACE53711482956291B0174D4">2017</mods:date>
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<mods:title id="74F1527A6E50422F5A8A650E47EA0BCB">J. Nat. Hist.</mods:title>
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<mods:number id="9ABF398B659A9613AB8840530F5AF9FE">2017-03-07</mods:number>
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<mods:number id="2002E94D0BB383BDFFA8AA9149967F49">51</mods:number>
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<mods:number id="DD3227E6BB1C0B6CB8D9C73FBCCD0B11">9 - 10</mods:number>
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<mods:start id="BEBBED8445336455FE450FD28A3E3B15">561</mods:start>
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<subSubSection id="69C26557FFDDFFE9AAF780C0C552B8FB" box="[441,875,1701,1727]" pageId="23" pageNumber="583" type="nomenclature">
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<paragraph id="216736DCFFDDFFE9AAF780C0C552B8FB" blockId="23.[441,875,1701,1727]" box="[441,875,1701,1727]" pageId="23" pageNumber="583">
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<heading id="7A2F81B0FFDDFFE9AAF780C0C552B8FB" box="[441,875,1701,1727]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="583" reason="2">
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<taxonomicName id="E6D84D5FFFDDFFE9AAF780C0C53CB8FB" ID-CoL="DPSP" authority="Freitag & Kodada, 2017" authorityName="Freitag & Kodada" authorityYear="2017" box="[441,773,1701,1727]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="583" phylum="Arthropoda" rank="species" species="longiparamerus" status="sp. nov.">
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<emphasis id="13ACEACEFFDDFFE9AAF780C0C53CB8FB" bold="true" box="[441,773,1701,1727]" italics="true" pageId="23" pageNumber="583">Ancyronyx longiparamerus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="089F57B5FFDDFFE9A84080C3C552B8FB" box="[782,875,1702,1727]" pageId="23" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="69C26557FFD2FFE6AA9686E8C573BEE2" box="[472,842,141,166]" pageId="24" pageNumber="584" type="description">
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<paragraph id="216736DCFFD2FFE6AA9686E8C573BEE2" blockId="24.[472,842,141,166]" box="[472,842,141,166]" pageId="24" pageNumber="584">
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(
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<figureCitation id="B9E32A59FFD2FFE6AAAE86E8C474BEE2" box="[480,589,141,166]" captionStart="Figure 2" captionStartId="8.[160,225,1580,1603]" captionTargetBox="[164,1145,142,1543]" captionTargetId="figure-1@8.[163,1153,142,1549]" captionTargetPageId="8" captionText="Figure 2. Habitus of Ancyronyx species from Sulawesi, (a) Ancyronyx longiparamerus (total length 1.80 mm), (b) Ancyronyx toraja (total length 1.75 mm), (c) Ancyronyx konjo (total length 1.95 mm), (d) Ancyronyx breviparamerus (total length 1.80 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746918" httpUri="https://zenodo.org/record/4746918/files/figure.png" pageId="24" pageNumber="584">Figures 2</figureCitation>
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(a); 10(a–c); 18; 19(c))
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</paragraph>
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</subSubSection>
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<subSubSection id="69C26557FFD2FFE7ABEE86B1C211BE8D" lastPageId="25" lastPageNumber="585" pageId="24" pageNumber="584" type="materials_examined">
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<paragraph id="216736DCFFD2FFE6ABEE86B1C70EBEA9" blockId="24.[160,1155,212,341]" box="[160,311,212,237]" pageId="24" pageNumber="584">
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<emphasis id="13ACEACEFFD2FFE6ABEE86B1C70EBEA9" bold="true" box="[160,311,212,237]" italics="true" pageId="24" pageNumber="584">Type locality</emphasis>
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</paragraph>
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<paragraph id="216736DCFFD2FFE6ABEE869DC705BF11" blockId="24.[160,1155,212,341]" pageId="24" pageNumber="584">
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<materialsCitation id="91B03C81FFD2FFE6ABEE869DC74CBF76" county="Rantepao" location="Mountain river" municipality="Salu Maiting'" pageId="24" pageNumber="584" specimenCount="1">
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<location id="24076007FFD2FFE6ABEE869DC771BF54" LSID="urn:lsid:plazi:treatment:A97187CAFFDDFFE4AAF780C0C59ABA71:24076007FFD2FFE6ABEE869DC771BF54" box="[160,328,248,272]" county="Rantepao" municipality="Salu Maiting'" name="Mountain river" pageId="24" pageNumber="584">Mountain river</location>
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‘
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||
<collectingMunicipality id="C103ACA6FFD2FFE6AA19869DC7D5BF54" box="[343,492,248,272]" pageId="24" pageNumber="584">Salu Maiting’</collectingMunicipality>
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||
flowing through pine forest/paddy fields,
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||
<quantity id="E6209B39FFD2FFE6A896869DC20DBF54" box="[984,1076,248,272]" metricMagnitude="4" metricUnit="m" metricValue="1.75" pageId="24" pageNumber="584" unit="km" value="17.5">17.5 km</quantity>
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||
northwest
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||
<collectingCounty id="C8064E50FFD2FFE6AB91877FC774BF76" box="[223,333,282,306]" pageId="24" pageNumber="584">Rantepao</collectingCounty>
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, ca
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</materialsCitation>
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.
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<geoCoordinate id="44EC501BFFD2FFE6AACD877FC433BF76" box="[387,522,282,306]" degrees="02" direction="south" minutes="49" orientation="latitude" pageId="24" pageNumber="584" precision="15" seconds="21" value="-2.8225">02°49′21′′ S</geoCoordinate>
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,
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<geoCoordinate id="44EC501BFFD2FFE6A959877FC492BF76" box="[535,683,282,306]" degrees="119" direction="east" minutes="51" orientation="longitude" pageId="24" pageNumber="584" precision="15" seconds="17" value="119.85472">119°51′17′′ E</geoCoordinate>
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,
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<quantity id="E6209B39FFD2FFE6A9F6877FC529BF76" box="[696,784,282,306]" metricMagnitude="3" metricUnit="m" metricValue="1.51" pageId="24" pageNumber="584" unit="m" value="1510.0">1510 m</quantity>
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a
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<materialsCitation id="91B03C81FFD2FFE6A862877FC506BF76" box="[812,831,282,306]" county="Rantepao" isTailEnumeration="true" location="Mountain river" municipality="Salu Maiting'" pageId="24" pageNumber="584" specimenCount="1">.s</materialsCitation>
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.l
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||
<materialsCitation id="91B03C81FFD2FFE6A809877FC70EBF11" country="Indonesia" location="South Sulawesi" pageId="24" pageNumber="584" specimenCount="1" stateProvince="Sulawesi Selatan">
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.,
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<collectingRegion id="E31CF83EFFD2FFE6A817877FC230BF76" box="[857,1033,282,306]" country="Indonesia" name="Sulawesi Selatan" pageId="24" pageNumber="584">South Sulawesi</collectingRegion>
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,
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<collectingCountry id="59CF764CFFD2FFE6AF58877FC2BABF76" box="[1046,1155,282,306]" name="Indonesia" pageId="24" pageNumber="584">Indonesia</collectingCountry>
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(
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<figureCitation id="B9E32A59FFD2FFE6ABE98758C72ABF11" box="[167,275,317,341]" captionStart="Figure 19" captionStartId="42.[160,225,1605,1628]" captionTargetBox="[163,1152,169,1575]" captionTargetId="figure-2@42.[163,1152,169,1575]" captionTargetPageId="42" captionText="Figure 19. Habitats of Ancyronyx species in Sulawesi, (a) the eastern slopes of the Pegunungan Fennema Mountains from where Ancyronyx breviparamerus, Ancyronyx henningi and Ancyronyx tobada were collected, (b) a small mountain creek in the Pegunungan Fennema Mountains (A), type locality of Ancyronyx breviparamerus and collection site of Ancyronyx tobada, (c) Salu Maiting River in North Toraja, type locality of Ancyronyx longiparamerus and collection site of Ancyronyx" figureDoi="http://doi.org/10.5281/zenodo.4746956" httpUri="https://zenodo.org/record/4746956/files/figure.png" pageId="24" pageNumber="584">Figure 19</figureCitation>
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||
(c))
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||
</materialsCitation>
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.
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||
</paragraph>
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<paragraph id="216736DCFFD2FFE6ABEE87E6C77CBFD8" blockId="24.[160,1156,387,585]" box="[160,325,387,412]" pageId="24" pageNumber="584">
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<emphasis id="13ACEACEFFD2FFE6ABEE87E6C77CBFD8" bold="true" box="[160,325,387,412]" italics="true" pageId="24" pageNumber="584">Type material</emphasis>
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</paragraph>
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<paragraph id="216736DCFFD2FFE7ABEE87C2C211BE8D" blockId="24.[160,1156,387,585]" lastBlockId="25.[160,1156,142,201]" lastPageId="25" lastPageNumber="585" pageId="24" pageNumber="584">
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<materialsCitation id="91B03C81FFD2FFE6ABEE87C2C5F7BC63" collectingDate="1997-12-31" collectionCode="NMW" collectorName="Freitag" country="Indonesia" elevation="1510" location="Salu Maiting'" longitude="119.85472" pageId="24" pageNumber="584" specimenCode="FR115" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
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<emphasis id="13ACEACEFFD2FFE6ABEE87C2C737BFFB" bold="true" box="[160,270,423,447]" pageId="24" pageNumber="584">
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<typeStatus id="FE63887EFFD2FFE6ABEE87C2C737BFFB" box="[160,270,423,447]" pageId="24" pageNumber="584" type="holotype">Holotype</typeStatus>
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</emphasis>
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<specimenCount id="37DEFD55FFD2FFE6AA5B87C3C710BFFA" box="[277,297,422,446]" pageId="24" pageNumber="584" type="male">♂</specimenCount>
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(
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<collectionCode id="47C9AE19FFD2FFE6AA7B87C2C74DBFFB" box="[309,372,423,447]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="584">NMW</collectionCode>
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[
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<specimenCode id="717E9EA7FFD2FFE6AACF87C2C7F7BFFB" box="[385,462,423,447]" collectionCode="NMW" country="Austria" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="584">FR115</specimenCode>
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]): ‘
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<collectingCountry id="59CF764CFFD2FFE6AAA787C2C452BFFB" box="[489,619,423,447]" name="Indonesia" pageId="24" pageNumber="584">INDONESIA</collectingCountry>
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: S
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<collectingRegion id="E31CF83EFFD2FFE6A9C687C2C4D5BFFB" box="[648,748,423,447]" country="Indonesia" name="Sulawesi Utara" pageId="24" pageNumber="584">Sulawesi</collectingRegion>
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,
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<locationDeviation id="30D52FFEFFD2FFE6A9BB87C2C5C6BFFB" box="[757,1023,423,447]" location="Rantepao" pageId="24" pageNumber="584">
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<quantity id="E6209B39FFD2FFE6A9BB87C2C574BFFB" box="[757,845,423,447]" metricMagnitude="4" metricUnit="m" metricValue="1.75" pageId="24" pageNumber="584" unit="km" value="17.5">17.5 km</quantity>
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NNW Rantepao
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</locationDeviation>
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<elevation id="AAF5D1EFFFD2FFE6AF4887C2C692BFA5" metricMagnitude="3" metricUnit="m" metricValue="1.51" pageId="24" pageNumber="584" unit="m" value="1510.0">
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<quantity id="E6209B39FFD2FFE6AF4887C2C263BFFB" box="[1030,1114,423,447]" metricMagnitude="3" metricUnit="m" metricValue="1.51" pageId="24" pageNumber="584" unit="m" value="1510.0">1510 m</quantity>
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a.s. l.
|
||
</elevation>
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||
, mount. riv. ’
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||
<location id="24076007FFD2FFE6AA0F87ACC7EABFA5" LSID="urn:lsid:plazi:treatment:A97187CAFFDDFFE4AAF780C0C59ABA71:24076007FFD2FFE6AA0F87ACC7EABFA5" box="[321,467,457,481]" country="Indonesia" longitude="119.85472" name="Salu Maiting'" pageId="24" pageNumber="584" stateProvince="Sulawesi Utara">Salu Maiting‘</location>
|
||
, pine forest/paddy fields, run; submerged wood, c. 02°49′21′ ′ S,
|
||
<geoCoordinate id="44EC501BFFD2FFE6AB858789C762BC40" box="[203,347,492,516]" degrees="119" direction="east" minutes="51" orientation="longitude" pageId="24" pageNumber="584" precision="15" seconds="17" value="119.85472">119°51′17′′ E</geoCoordinate>
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<date id="5566101CFFD2FFE6AA2A8789C7C9BC40" box="[356,496,492,516]" pageId="24" pageNumber="584" value="1997-12-31">
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||
<collectingDate id="4522E9F4FFD2FFE6AA2A8789C7C9BC40" box="[356,496,492,516]" pageId="24" pageNumber="584" value="1997-12-31">31 Dec.1997</collectingDate>
|
||
</date>
|
||
leg.
|
||
<collectorName id="8C2D530AFFD2FFE6A9658789C440BC40" box="[555,633,492,516]" pageId="24" pageNumber="584">Freitag</collectorName>
|
||
(Sul4f)’, terminal parts of abdomen including aedeagus and right antenna glued separately, right metatarsus absent
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="91B03C81FFD2FFE7A895846AC7EBBEE2" collectingDate="1997-12-31" collectionCode="CFM, CKB" collectorName="Freitag" country="Indonesia" elevation="1570" lastPageId="25" lastPageNumber="585" location="17.5 km NNW Rantepao" longitude="119.85472" pageId="24" pageNumber="584" specimenCode="FR113" specimenCount="2" specimenCount-male="2" stateProvince="Sulawesi Utara" typeStatus="paratype">
|
||
<emphasis id="13ACEACEFFD2FFE6A895846AC26BBC63" bold="true" box="[987,1106,527,551]" pageId="24" pageNumber="584">
|
||
<typeStatus id="FE63887EFFD2FFE6A895846AC26BBC63" box="[987,1106,527,551]" pageId="24" pageNumber="584" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="37DEFD55FFD2FFE6AF2C846AC2BABC62" box="[1122,1155,526,551]" pageId="24" pageNumber="584" type="male">1♂</specimenCount>
|
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(
|
||
<collectionCode id="47C9AE19FFD2FFE6ABE88454C6E3BC0D" box="[166,218,561,585]" pageId="24" pageNumber="584">CFM</collectionCode>
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[
|
||
<specimenCode id="717E9EA7FFD2FFE6ABA68454C70ABC0D" box="[232,307,561,585]" collectionCode="CFM" pageId="24" pageNumber="584">FR113</specimenCode>
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]) with same data as holotype;
|
||
<specimenCount id="37DEFD55FFD2FFE6A9D68454C483BC0D" box="[664,698,561,585]" pageId="24" pageNumber="584" type="male">1♂</specimenCount>
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(
|
||
<collectionCode id="47C9AE19FFD2FFE6A9878454C4C5BC0D" box="[713,764,561,585]" pageId="24" pageNumber="584">CKB</collectionCode>
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||
): ‘
|
||
<collectingCountry id="59CF764CFFD2FFE6A85D8454C5ACBC0D" box="[787,917,561,585]" name="Indonesia" pageId="24" pageNumber="584">INDONESIA</collectingCountry>
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: S
|
||
<collectingRegion id="E31CF83EFFD2FFE6A8F68454C225BC0D" box="[952,1052,561,585]" country="Indonesia" name="Sulawesi Utara" pageId="24" pageNumber="584">Sulawesi</collectingRegion>
|
||
,
|
||
<location id="24076007FFD2FFE7AF668454C769BEE2" LSID="urn:lsid:plazi:treatment:A97187CAFFDDFFE4AAF780C0C59ABA71:24076007FFD2FFE7AF668454C769BEE2" country="Indonesia" lastPageId="25" lastPageNumber="585" name="17.5 km NNW Rantepao" pageId="24" pageNumber="584" stateProvince="Sulawesi Utara">
|
||
<quantity id="E6209B39FFD2FFE6AF668454C2BABC0D" box="[1064,1155,561,585]" metricMagnitude="4" metricUnit="m" metricValue="1.75" pageId="24" pageNumber="584" unit="km" value="17.5">17.5 km</quantity>
|
||
NNW Rantepao
|
||
</location>
|
||
<quantity id="E6209B39FFD3FFE7AA1286EBC78DBEE2" box="[348,436,142,166]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="25" pageNumber="585" unit="m" value="1570.0">
|
||
<elevation id="AAF5D1EFFFD3FFE7AA1286EBC78DBEE2" box="[348,436,142,166]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="25" pageNumber="585" unit="m" value="1570.0">1570 m</elevation>
|
||
</quantity>
|
||
a
|
||
</materialsCitation>
|
||
<materialsCitation id="91B03C81FFD3FFE7AA9C86EBC7DCBEE2" box="[466,485,142,166]" collectingDate="1997-12-31" collectionCode="CFM, CKB" collectorName="Freitag" country="Indonesia" elevation="1570" isTailEnumeration="true" location="17.5 km NNW Rantepao" longitude="119.85472" pageId="25" pageNumber="585" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="paratype">.s</materialsCitation>
|
||
.l., small mount. river ’Salu Malarak‘, pine forest, run/riffle; submerged wood, c.
|
||
<geoCoordinate id="44EC501BFFD3FFE7AADF86D4C42FBE8D" box="[401,534,177,201]" degrees="02" direction="south" minutes="48" orientation="latitude" pageId="25" pageNumber="585" precision="15" seconds="51" value="-2.8141665">02°48′51′′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="44EC501BFFD3FFE7A96C86D4C488BE8D" box="[546,689,177,201]" degrees="119" direction="east" minutes="51" orientation="longitude" pageId="25" pageNumber="585" precision="15" seconds="21" value="119.855835">119°51′21′′ E</geoCoordinate>
|
||
<date id="5566101CFFD3FFE7A9F486D4C57CBE8D" box="[698,837,177,201]" pageId="25" pageNumber="585" value="1997-12-31">31 Dec.1997</date>
|
||
leg
|
||
<materialsCitation id="91B03C81FFD3FFE7A83D86D4C225BE8D" box="[883,1052,177,201]" collectingDate="1997-12-31" collectionCode="CFM, CKB" collectorName="Freitag & Sul" country="Indonesia" elevation="1570" isTailEnumeration="true" location="17.5 km NNW Rantepao" longitude="119.85472" pageId="25" pageNumber="585" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
|
||
.
|
||
<collectorName id="8C2D530AFFD3FFE7A83186D4C5F4BE8D" box="[895,973,177,201]" pageId="25" pageNumber="585">Freitag</collectorName>
|
||
(
|
||
<collectorName id="8C2D530AFFD3FFE7A89286D4C23BBE8D" box="[988,1026,177,201]" pageId="25" pageNumber="585">Sul</collectorName>
|
||
7f)
|
||
</materialsCitation>
|
||
’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="75A76654FFD2FFE6ABEE80C0C521B89D" ID-DOI="http://doi.org/10.5281/zenodo.4746934" ID-Zenodo-Dep="4746934" httpUri="https://zenodo.org/record/4746934/files/figure.png" pageId="24" pageNumber="584" startId="24.[160,225,1701,1724]" targetBox="[163,1151,628,1670]" targetPageId="24">
|
||
<paragraph id="216736DCFFD2FFE6ABEE80C0C521B89D" blockId="24.[160,1156,1700,1753]" pageId="24" pageNumber="584">
|
||
<emphasis id="13ACEACEFFD2FFE6ABEE80C0C736B8F8" bold="true" box="[160,271,1701,1724]" pageId="24" pageNumber="584">Figure 10.</emphasis>
|
||
<taxonomicName id="E6D84D5FFFD2FFE6AA5980C1C426B8FF" authorityName="Freitag & Kodada" authorityYear="2017" box="[279,543,1700,1723]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="584" phylum="Arthropoda" rank="species" species="longiparamerus">
|
||
<emphasis id="13ACEACEFFD2FFE6AA5980C1C426B8FF" box="[279,543,1700,1723]" italics="true" pageId="24" pageNumber="584">Ancyronyx longiparamerus</emphasis>
|
||
</taxonomicName>
|
||
from Rantepao (Sul7f); (a) male sternite IX, ventral; (b) aedeagus, lateral; (c) aedeagus, ventral; (Scale bars = 0.1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="69C26557FFD3FFE4ABEE8693C5F9BFDD" lastPageId="26" lastPageNumber="586" pageId="25" pageNumber="585" type="description">
|
||
<paragraph id="216736DCFFD3FFE7ABEE8693C71FBF4B" blockId="25.[160,1121,246,305]" box="[160,294,246,271]" pageId="25" pageNumber="585">
|
||
<emphasis id="13ACEACEFFD3FFE7ABEE8693C71FBF4B" bold="true" box="[160,294,246,271]" italics="true" pageId="25" pageNumber="585">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="216736DCFFD3FFE7ABEE877CC258BF75" blockId="25.[160,1121,246,305]" box="[160,1121,281,305]" pageId="25" pageNumber="585">
|
||
Body
|
||
<quantity id="E6209B39FFD3FFE7ABAC877CC7B8BF75" box="[226,385,281,305]" metricMagnitude="-3" metricUnit="m" metricValue="1.565" metricValueMax="1.58" metricValueMin="1.55" pageId="25" pageNumber="585" unit="mm" value="1.565" valueMax="1.58" valueMin="1.55">1.55–1.58 mm</quantity>
|
||
long, 2.2–2.3 times as long as wide (BL/EW), EW:
|
||
<quantity id="E6209B39FFD3FFE7A8F5877CC264BF75" box="[955,1117,281,305]" metricMagnitude="-4" metricUnit="m" metricValue="6.95" metricValueMax="7.1" metricValueMin="6.8" pageId="25" pageNumber="585" unit="mm" value="0.695" valueMax="0.71" valueMin="0.68">0.68–0.71 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="216736DCFFD3FFE7ABEE873BC736BC45" blockId="25.[160,1156,349,513]" pageId="25" pageNumber="585">
|
||
<emphasis id="13ACEACEFFD3FFE7ABEE873BC786BF33" bold="true" box="[160,447,349,375]" pageId="25" pageNumber="585">
|
||
<emphasis id="13ACEACEFFD3FFE7ABEE873BC715BF33" bold="true" box="[160,300,350,375]" italics="true" pageId="25" pageNumber="585">Colouration</emphasis>
|
||
<figureCitation id="B9E32A59FFD3FFE7AA7B8738C7A3BF33" box="[309,410,349,375]" captionStart="Figure 2" captionStartId="8.[160,225,1580,1603]" captionTargetBox="[164,1145,142,1543]" captionTargetId="figure-1@8.[163,1153,142,1549]" captionTargetPageId="8" captionText="Figure 2. Habitus of Ancyronyx species from Sulawesi, (a) Ancyronyx longiparamerus (total length 1.80 mm), (b) Ancyronyx toraja (total length 1.75 mm), (c) Ancyronyx konjo (total length 1.95 mm), (d) Ancyronyx breviparamerus (total length 1.80 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746918" httpUri="https://zenodo.org/record/4746918/files/figure.png" pageId="25" pageNumber="585">Figure 2</figureCitation>
|
||
(a).
|
||
</emphasis>
|
||
Predominantly very dark brown to black; elytra with conspicuous anterior pair of patches extending obliquely from humeri to first interval; posterior patches oblique and distinct. Pronotum and head black; entire femora dark brown; lighter areas of tarsomere 5, tibiae and antennal segments 1–8/9 paler brown to yellowish.
|
||
</paragraph>
|
||
<paragraph id="216736DCFFD3FFE7ABEE844BC5DFBCC8" blockId="25.[160,1156,558,652]" pageId="25" pageNumber="585">
|
||
<emphasis id="13ACEACEFFD3FFE7ABEE844BC6DDBC03" bold="true" box="[160,228,558,583]" italics="true" pageId="25" pageNumber="585">Head.</emphasis>
|
||
HW:
|
||
<quantity id="E6209B39FFD3FFE7AA63844AC7E8BC03" box="[301,465,559,583]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.7" metricValueMin="3.5" pageId="25" pageNumber="585" unit="mm" value="0.36" valueMax="0.37" valueMin="0.35">0.35–0.37 mm</quantity>
|
||
; ID:
|
||
<quantity id="E6209B39FFD3FFE7A949844AC492BC03" box="[519,683,559,583]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="25" pageNumber="585" unit="mm" value="0.2" valueMax="0.21" valueMin="0.19">0.19–0.21 mm</quantity>
|
||
; labrum and clypeus laterally with longitudinal granules; frons and vertex with moderately densely arranged short longitudinal granules. Gena and gula transversally microstriate; gular suture indistinct.
|
||
</paragraph>
|
||
<paragraph id="216736DCFFD3FFE7ABEE84DCC40BBAD0" blockId="25.[160,1156,697,1450]" pageId="25" pageNumber="585">
|
||
<emphasis id="13ACEACEFFD3FFE7ABEE84DCC6C0BC96" bold="true" box="[160,249,697,722]" italics="true" pageId="25" pageNumber="585">Thorax.</emphasis>
|
||
Pronotum slightly wider than long, widest at about posterior 0.3; posterolateral grooves moderately deep; posterior margin slightly trisinuate; PL:
|
||
<quantity id="E6209B39FFD3FFE7A8EE84B9C27DBCB0" box="[928,1092,732,756]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="25" pageNumber="585" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48 mm</quantity>
|
||
, MW:
|
||
<quantity id="E6209B39FFD3FFE7ABEE849AC77ABD53" box="[160,323,767,791]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="4.9" metricValueMin="4.8" pageId="25" pageNumber="585" unit="mm" value="0.485" valueMax="0.49" valueMin="0.48">0.48–0.49 mm</quantity>
|
||
; pronotal surface appears reticulated. Prosternum densely punctate and granulose; anterior portion narrow; prosternal process distinctly wider than long, broadly subpentagonal, anteriorly with a shallow transverse groove. Metascutellum subtriangular. Elytra 1.6–1.7 times as long as wide; EL:
|
||
<quantity id="E6209B39FFD3FFE7A9C58502C515BD3B" box="[651,812,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.16" metricValueMin="1.14" pageId="25" pageNumber="585" unit="mm" value="1.15" valueMax="1.16" valueMin="1.14">1.14–1.16 mm</quantity>
|
||
, EW:
|
||
<quantity id="E6209B39FFD3FFE7A8298502C231BD3B" box="[871,1032,871,895]" metricMagnitude="-4" metricUnit="m" metricValue="6.95" metricValueMax="7.1" metricValueMin="6.8" pageId="25" pageNumber="585" unit="mm" value="0.695" valueMax="0.71" valueMin="0.68">0.68–0.71 mm</quantity>
|
||
; punctures moderately large and moderately deeply impressed on disc, rather small and shallow on elytral declivity; intervals with small granules, latest less prominent posteriorly. Mesoventrite narrow, anteriorly transversely microstriate, remaining portion irregularly punctate and granulose. Metaventrite longer than combined length of pro- and mesoventrite, with transverse impression anteriorly; disc with moderately deep, almost glabrous longitudinal impression, surrounding area with irregular granules and dense punctures; anepisternum 3 narrow, glabrous, with one row of punctures. Hind wings present in all specimens examined.
|
||
</paragraph>
|
||
<paragraph id="216736DCFFD3FFE7AB8E82FAC440BBEE" blockId="25.[160,1156,697,1450]" pageId="25" pageNumber="585">Legs slightly longer than body; pro- and mesocoxae large, subglobular; metacoxae obtuse and moderately elevated at mesal portion, obliquely conoidal; only procoxae visible in dorsal view; trochanter short, broadly lanceolate, invisible in dorsal view, pointed at distal end. Femora and tibiae with elongate, dense granules (appear striated in low magnifications); tarsomeres with several moderately long hair-like setae, setae more conspicuous ventrally; combined length of tarsomeres 1–4 approximately as long as terminal tarsomere; claws long, moderately broad, strongly bent, base of each with three teeth; distal tooth distinctly largest.</paragraph>
|
||
<paragraph id="216736DCFFD3FFE7ABEE83B3C486B8FB" blockId="25.[160,1156,1494,1727]" pageId="25" pageNumber="585">
|
||
<emphasis id="13ACEACEFFD3FFE7ABEE83B3C721BBAB" bold="true" box="[160,280,1494,1519]" italics="true" pageId="25" pageNumber="585">Abdomen.</emphasis>
|
||
Ventrite 1 with intercoxal process short (about as long as remaining portion of ventrite 1), mesally almost glabrous. Ventrites 2–4 with surface similar as in ventrite 1; combined length of ventrites 3 and 4 as long as ventrite 1; ventrite 2 of intermediate size; ventrite 5 long, with cordiform granules; granules denser towards apex. Male sternite IX (
|
||
<figureCitation id="B9E32A59FFD3FFE7AA638007C7A4B83E" box="[301,413,1634,1658]" captionStart="Figure 10" captionStartId="24.[160,225,1701,1724]" captionTargetBox="[163,1151,628,1670]" captionTargetId="figure-139@24.[163,1152,628,1670]" captionTargetPageId="24" captionText="Figure 10. Ancyronyx longiparamerus from Rantepao (Sul7f); (a) male sternite IX, ventral; (b) aedeagus, lateral; (c) aedeagus, ventral; (Scale bars = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746934" httpUri="https://zenodo.org/record/4746934/files/figure.png" pageId="25" pageNumber="585">Figure 10</figureCitation>
|
||
(a)) 460 μm long; apical margin concave; with fewer than 10 moderately short setae on lateral portion; paraprocts short, not reaching apical margin; anterior strut moderately long, evenly tapering.
|
||
</paragraph>
|
||
<paragraph id="216736DCFFD0FFE4AB8E86EBC5F9BFDD" blockId="26.[160,1156,142,409]" pageId="26" pageNumber="586">
|
||
Aedeagus (
|
||
<figureCitation id="B9E32A59FFD0FFE4AA7286EBC79FBEE2" box="[316,422,142,166]" captionStart="Figure 10" captionStartId="24.[160,225,1701,1724]" captionTargetBox="[163,1151,628,1670]" captionTargetId="figure-139@24.[163,1152,628,1670]" captionTargetPageId="24" captionText="Figure 10. Ancyronyx longiparamerus from Rantepao (Sul7f); (a) male sternite IX, ventral; (b) aedeagus, lateral; (c) aedeagus, ventral; (Scale bars = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4746934" httpUri="https://zenodo.org/record/4746934/files/figure.png" pageId="26" pageNumber="586">Figure 10</figureCitation>
|
||
(b,c)) broad, elongate,
|
||
<quantity id="E6209B39FFD0FFE4A9D086EBC4CDBEE2" box="[670,756,142,166]" metricMagnitude="-1" metricUnit="m" metricValue="4.75" pageId="26" pageNumber="586" unit="mm" value="475.0">475 µm</quantity>
|
||
long. Median lobe moderately long and broad (
|
||
<quantity id="E6209B39FFD0FFE4AA5186D4C74EBE8D" box="[287,375,177,201]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="586" unit="mm" value="100.0">100 µm</quantity>
|
||
), slightly tapering towards base and almost regularly tapering in apical fifth, with small knob-like rounded tip, entirely slightly bent ventrad; ventral sac weakly sclerotized, simply plicate, reaching apical 0.08; fibula lightly sclerotized, apically enlarged; corona inconspicuous. Phallobase strongly sclerotized, rather simply shaped, asymmetrical, reaching 0.27 length of aedeagus. Parameres long and robust, regularly tapering in apical fourth; dorsolateral margin slightly concave; ventral margin slightly sinuous, with few very short basolateral setae and with more than 20 short setae in apical half.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="69C26557FFD0FFE4ABEE87ADC587BC40" box="[160,958,456,516]" pageId="26" pageNumber="586" type="distribution">
|
||
<paragraph id="216736DCFFD0FFE4ABEE87ADC715BFA5" blockId="26.[160,958,456,516]" box="[160,300,456,481]" pageId="26" pageNumber="586">
|
||
<emphasis id="13ACEACEFFD0FFE4ABEE87ADC715BFA5" bold="true" box="[160,300,456,481]" italics="true" pageId="26" pageNumber="586">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="216736DCFFD0FFE4ABEE8789C587BC40" blockId="26.[160,958,456,516]" box="[160,958,492,516]" pageId="26" pageNumber="586">
|
||
So far known from two localities in
|
||
<collectingRegion id="E31CF83EFFD0FFE4A9748789C4DFBC40" box="[570,742,492,516]" country="Indonesia" name="Sulawesi Selatan" pageId="26" pageNumber="586">South Sulawesi</collectingRegion>
|
||
(
|
||
<figureCitation id="B9E32A59FFD0FFE4A9B88789C552BC40" box="[758,875,492,516]" captionStart="Figure 18" captionStartId="41.[160,225,1188,1211]" captionTargetBox="[163,1151,142,1157]" captionTargetId="figure-116@41.[163,1152,142,1157]" captionTargetPageId="41" captionText="Figure 18. Map of Sulawesi with collecting site of Ancyronyx species based on their label data." figureDoi="http://doi.org/10.5281/zenodo.4746954" httpUri="https://zenodo.org/record/4746954/files/figure.png" pageId="26" pageNumber="586">Figures 18</figureCitation>
|
||
;
|
||
<figureCitation id="B9E32A59FFD0FFE4A8348789C5AEBC40" box="[890,919,492,516]" captionStart="Figure 19" captionStartId="42.[160,225,1605,1628]" captionTargetBox="[163,1152,169,1575]" captionTargetId="figure-2@42.[163,1152,169,1575]" captionTargetPageId="42" captionText="Figure 19. Habitats of Ancyronyx species in Sulawesi, (a) the eastern slopes of the Pegunungan Fennema Mountains from where Ancyronyx breviparamerus, Ancyronyx henningi and Ancyronyx tobada were collected, (b) a small mountain creek in the Pegunungan Fennema Mountains (A), type locality of Ancyronyx breviparamerus and collection site of Ancyronyx tobada, (c) Salu Maiting River in North Toraja, type locality of Ancyronyx longiparamerus and collection site of Ancyronyx" figureDoi="http://doi.org/10.5281/zenodo.4746956" httpUri="https://zenodo.org/record/4746956/files/figure.png" pageId="26" pageNumber="586">19</figureCitation>
|
||
(c)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="69C26557FFD0FFE4ABEE8456C7F0BCF0" pageId="26" pageNumber="586" type="etymology">
|
||
<paragraph id="216736DCFFD0FFE4ABEE8456C724BC08" blockId="26.[160,1155,563,692]" box="[160,285,563,588]" pageId="26" pageNumber="586">
|
||
<emphasis id="13ACEACEFFD0FFE4ABEE8456C724BC08" bold="true" box="[160,285,563,588]" italics="true" pageId="26" pageNumber="586">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="216736DCFFD0FFE4ABEE8432C7F0BCF0" blockId="26.[160,1155,563,692]" pageId="26" pageNumber="586">
|
||
The species epithet ‘
|
||
<taxonomicName id="E6D84D5FFFD0FFE4AADA8432C406BC2B" authorityName="Freitag & Kodada" authorityYear="2017" box="[404,575,599,623]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="586" phylum="Arthropoda" rank="species" species="longiparamerus">
|
||
<emphasis id="13ACEACEFFD0FFE4AADA8432C406BC2B" box="[404,575,599,623]" italics="true" pageId="26" pageNumber="586">longiparamerus</emphasis>
|
||
</taxonomicName>
|
||
’ refers to the comparably long parameres which allow us to distinguish the species most clearly from its congeners, and particularly from
|
||
<taxonomicName id="E6D84D5FFFD0FFE4ABEE84F9C7FBBCF0" authorityName="Freitag & Kodada" authorityYear="2017" box="[160,450,668,692]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="586" phylum="Arthropoda" rank="species" species="breviparamerus">
|
||
<emphasis id="13ACEACEFFD0FFE4ABEE84F9C7FBBCF0" box="[160,450,668,692]" italics="true" pageId="26" pageNumber="586">Ancyronyx breviparamerus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="69C26557FFD0FFE4ABEE8481C59ABA71" pageId="26" pageNumber="586" type="discussion">
|
||
<paragraph id="216736DCFFD0FFE4ABEE8481C727BCB9" blockId="26.[160,1156,740,1077]" box="[160,286,740,765]" pageId="26" pageNumber="586">
|
||
<emphasis id="13ACEACEFFD0FFE4ABEE8481C727BCB9" bold="true" box="[160,286,740,765]" italics="true" pageId="26" pageNumber="586">Comments</emphasis>
|
||
</paragraph>
|
||
<paragraph id="216736DCFFD0FFE4ABEE8562C757BD89" blockId="26.[160,1156,740,1077]" pageId="26" pageNumber="586">
|
||
The species is characterized by the very long and robust parameres, while it most resembles
|
||
<taxonomicName id="E6D84D5FFFD0FFE4AA55854FC407BD06" authorityName="Freitag & Kodada" authorityYear="2017" box="[283,574,810,834]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="586" phylum="Arthropoda" rank="species" species="breviparamerus">
|
||
<emphasis id="13ACEACEFFD0FFE4AA55854FC407BD06" box="[283,574,810,834]" italics="true" pageId="26" pageNumber="586">Ancyronyx breviparamerus</emphasis>
|
||
</taxonomicName>
|
||
in the external characters. These two species may be distinguished by the distinctly shorter parameres reaching only up to apical 0.6 of the median lobe in
|
||
<taxonomicName id="E6D84D5FFFD0FFE4AA18850AC424BDC3" authorityName="Freitag & Kodada" authorityYear="2017" box="[342,541,879,903]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="586" phylum="Arthropoda" rank="species" species="breviparamerus">
|
||
<emphasis id="13ACEACEFFD0FFE4AA18850AC424BDC3" box="[342,541,879,903]" italics="true" pageId="26" pageNumber="586">A. breviparamerus</emphasis>
|
||
</taxonomicName>
|
||
, as well as by the anterior elytral patches being faint and brownish in the latter species versus the more distinct, large and pale patches in
|
||
<taxonomicName id="E6D84D5FFFD0FFE4ABEE85D0C750BD89" authorityName="Freitag & Kodada" authorityYear="2017" box="[160,361,949,973]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="586" phylum="Arthropoda" rank="species" species="longiparamerus">
|
||
<emphasis id="13ACEACEFFD0FFE4ABEE85D0C750BD89" box="[160,361,949,973]" italics="true" pageId="26" pageNumber="586">A. longiparamerus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="216736DCFFD0FFE4AB8E85B2C59ABA71" blockId="26.[160,1156,740,1077]" pageId="26" pageNumber="586">
|
||
This species was exclusively found on submerged wood in small to medium-sized mountain rivers at altitudes at c.
|
||
<quantity id="E6209B39FFD0FFE4A952859FC44FBA56" box="[540,630,1018,1042]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="26" pageNumber="586" unit="m" value="1550.0">1550 m</quantity>
|
||
a.s.l. (
|
||
<figureCitation id="B9E32A59FFD0FFE4A98C859FC509BA56" box="[706,816,1018,1042]" captionStart="Figure 19" captionStartId="42.[160,225,1605,1628]" captionTargetBox="[163,1152,169,1575]" captionTargetId="figure-2@42.[163,1152,169,1575]" captionTargetPageId="42" captionText="Figure 19. Habitats of Ancyronyx species in Sulawesi, (a) the eastern slopes of the Pegunungan Fennema Mountains from where Ancyronyx breviparamerus, Ancyronyx henningi and Ancyronyx tobada were collected, (b) a small mountain creek in the Pegunungan Fennema Mountains (A), type locality of Ancyronyx breviparamerus and collection site of Ancyronyx tobada, (c) Salu Maiting River in North Toraja, type locality of Ancyronyx longiparamerus and collection site of Ancyronyx" figureDoi="http://doi.org/10.5281/zenodo.4746956" httpUri="https://zenodo.org/record/4746956/files/figure.png" pageId="26" pageNumber="586">Figure 19</figureCitation>
|
||
(c)). The collection sites were surrounded by secondary vegetation (pine forest) and paddy fields.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |