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<document id="310BB1DD8FCDB182D3550A82C22A1672" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier &amp; Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FFFE615BFF026310FE2CFEB6" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madecorphnus cuccodoroi Frolov 2010" docType="treatment" docVersion="15" lastPageNumber="50" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="48" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:title id="19E978C1EB79EACC5B9434C3F2A42E09">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<mods:namePart id="40796631B4AB69169F744ECE1FBCFF8E">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="B0222393D5EBDE7C40B1AB3AB4F8F332">Montreuil, Olivier</mods:namePart>
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<mods:namePart id="10C739EFB07E6E8E91A19AE837AA8C01">Akhmetova, Lilia A.</mods:namePart>
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<mods:title id="DB62105DEAE4761E070A38DC420B1AB8">Zootaxa</mods:title>
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<mods:date id="85FB9FBC97D78A139F4C52729933E2DB">2016</mods:date>
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<subSubSection id="9BFC5F9DFFFE6145FF026310FEBCFE72" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="D3590C16FFFE6145FF026310FDF6FE93" blockId="47.[151,636,332,391]" box="[151,636,332,358]" pageId="47" pageNumber="48">
<heading id="8811BB7AFFFE6145FF026310FDF6FE93" bold="true" box="[151,636,332,358]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<taxonomicName id="14E67795FFFE6145FF026310FDF6FE93" authority="Frolov, 2010" authorityName="Frolov" authorityYear="2010" box="[151,636,332,358]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="E192D004FFFE6145FF026310FDF6FE93" bold="true" box="[151,636,332,358]" pageId="47" pageNumber="48">
<emphasis id="E192D004FFFE6145FF026310FE50FE93" bold="true" box="[151,474,332,358]" italics="true" pageId="47" pageNumber="48">Madecorphnus cuccodoroi</emphasis>
Frolov, 2010
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D3590C16FFFE6145FF026332FEBCFE72" blockId="47.[151,636,332,391]" box="[151,310,366,391]" pageId="47" pageNumber="48">
(
<figureCitation id="4BDD1093FFFE6145FF0A6332FF70FE72" box="[159,250,366,391]" captionStart="FIGURE 24" captionStartId="48.[151,250,1930,1952]" captionTargetBox="[226,1348,468,1875]" captionTargetId="figure@48.[202,1385,409,1909]" captionTargetPageId="48" captionText="FIGURE 24. Madecorphnus cuccodoroi, holotype male (A, C, D, E), paratype female (B). Habitus in dorsal view, internal sac of aedeagus (C), aedeagus in lateral view and parameres in dorsal view (D), labels (E), distributional records (F)." httpUri="https://zenodo.org/record/200467/files/figure.png" pageId="47" pageNumber="48">Figs. 24</figureCitation>
AF)
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFE6145FF0263EAFDECFE39" box="[151,614,438,460]" pageId="47" pageNumber="48" type="reference_group">
<paragraph id="D3590C16FFFE6145FF0263EAFDECFE39" blockId="47.[151,614,438,460]" box="[151,614,438,460]" pageId="47" pageNumber="48">
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<taxonomicName id="14E67795FFFE6145FF0263EAFDE8FE39" authority="Frolov, 2010a: 64" authorityName="Frolov" authorityPageNumber="64" authorityYear="2010" box="[151,610,438,460]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="E192D004FFFE6145FF0263EAFE2CFE39" box="[151,422,438,460]" italics="true" pageId="47" pageNumber="48">Madecorphnus cuccodoroi</emphasis>
<treatmentCitation id="52472A07FFFE6145FE3963EAFDE8FE39" ID-DOI="http://doi.org/10.5281/zenodo.3505135" ID-Zenodo-Dep="3505135" LSID="urn:lsid:zoobank.org:act:CF051797-A4EE-4172-A537-8731D86D92D1" author="Frolov" box="[428,610,438,460]" httpUri="http://treatment.plazi.org/id/039687DBFFE3FFF0FE906FEBFAF5FCBE" page="64" pageId="47" pageNumber="48" year="2010">
<bibRefCitation id="B77771E7FFFE6145FE3963EAFDB6FE39" author="Frolov" box="[428,572,438,460]" pageId="47" pageNumber="48" refString="Frolov, A. V. (2010 a) A new species of scarab beetle, Madecorphnus cuccodoroi (Coleoptera: Scarabaeidae: Orphninae), from northern Madagascar. Zootaxa, 2531, 64 - 68." type="journal article" year="2010" yearSuffix="a">Frolov, 2010a</bibRefCitation>
: 64
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFE6145FF0263A5FE37FC13" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="D3590C16FFFE6145FF0263A5FE8DFDAC" blockId="47.[151,1437,505,2006]" pageId="47" pageNumber="48">
<materialsCitation id="638E064BFFFE6145FF0263A5FDCAFDC0" ID-GBIF-Occurrence="1416610277" country="Madagascar" elevation="1300" latitude="-12.5525" location="Mt Ambre" longLatPrecision="21" longitude="49.161114" pageId="47" pageNumber="48" specimenCount="2" typeStatus="holotype">
<emphasis id="E192D004FFFE6145FF0263A5FE4BFDE7" bold="true" box="[151,449,505,530]" pageId="47" pageNumber="48">
<typeStatus id="0C5DB2B4FFFE6145FF0263A5FF58FDE7" box="[151,210,505,530]" pageId="47" pageNumber="48">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="0C5DB2B4FFFE6145FE5C63A5FDBBFDE7" box="[457,561,505,530]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
(
<figureCitation id="4BDD1093FFFE6145FDD463A5FD15FDE4" box="[577,671,505,530]" captionStart="FIGURE 24" captionStartId="48.[151,250,1930,1952]" captionTargetBox="[226,1348,468,1875]" captionTargetId="figure@48.[202,1385,409,1909]" captionTargetPageId="48" captionText="FIGURE 24. Madecorphnus cuccodoroi, holotype male (A, C, D, E), paratype female (B). Habitus in dorsal view, internal sac of aedeagus (C), aedeagus in lateral view and parameres in dorsal view (D), labels (E), distributional records (F)." httpUri="https://zenodo.org/record/200467/files/figure.png" pageId="47" pageNumber="48">Figs. 24</figureCitation>
A, CE), male, “
<collectingCountry id="ABF14C86FFFE6145FCCE63A5FBADFDE4" box="[859,1063,505,529]" name="Madagascar" pageId="47" pageNumber="48">MADAGASCAR</collectingCountry>
,
<location id="D6395ACDFFFE6145FBA663A6FB27FDE7" LSID="urn:lsid:plazi:treatment:5B4FBD00FFFE615BFF026310FE2CFEB6:D6395ACDFFFE6145FBA663A6FB27FDE7" box="[1075,1197,505,530]" country="Madagascar" latitude="-12.5525" longLatPrecision="21" longitude="49.161114" name="Mt Ambre" pageId="47" pageNumber="48">Mt Ambre</location>
,
<quantity id="141EA1F3FFFE6145FB2C63A5FB79FDE7" box="[1209,1267,505,530]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="47" pageNumber="48" unit="km" value="3.0">3 km</quantity>
NE N.P. camp site,
<geoCoordinate id="B6D26AD1FFFE6145FF586042FEC6FDC3" box="[205,332,542,566]" direction="south" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="-12.5525">12°33'09''S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FFFE6145FECC6042FE51FDC3" box="[345,475,542,566]" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="15" value="49.161114">49°09'40''E</geoCoordinate>
,
<elevation id="58CBEB25FFFE6145FE726042FDCAFDC0" box="[487,576,542,566]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="47" pageNumber="48" unit="m" value="1300.0">
<quantity id="141EA1F3FFFE6145FE726042FDCAFDC0" box="[487,576,542,566]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="47" pageNumber="48" unit="m" value="1300.0">1300 m</quantity>
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.
<date id="A7582AD6FFFE6145FDDE6042FD4AFDC3" box="[587,704,542,566]" pageId="47" pageNumber="48" value="2003-02-28">28.II.2003</date>
#3a G. Cuccodoro / Holotypus
<emphasis id="E192D004FFFE6145FBB06041FBFBFDC3" box="[1061,1137,541,566]" italics="true" pageId="47" pageNumber="48">Madec</emphasis>
.
<taxonomicName id="14E67795FFFE6145FB156041FA1BFDC3" authority="Frolov 2010" authorityName="Frolov" authorityYear="2010" box="[1152,1425,541,566]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="E192D004FFFE6145FB156041FB77FDC3" box="[1152,1277,541,566]" italics="true" pageId="47" pageNumber="48">cuccodoroi</emphasis>
Frolov 2010
</taxonomicName>
” (MHNG).
</paragraph>
<paragraph id="D3590C16FFFE6145FF52603AFE44FCA3" blockId="47.[151,1437,505,2006]" pageId="47" pageNumber="48">
<materialsCitation id="638E064BFFFE6145FF52603AFA12FD88" ID-GBIF-Occurrence="1416610307" box="[199,1432,613,638]" country="Madagascar" elevation="1050" latitude="49.195" location="Mt Ambre" pageId="47" pageNumber="48" specimenCount="1" typeStatus="paratype">
<typeStatus id="0C5DB2B4FFFE6145FF52603AFEB3FD8B" box="[199,313,614,638]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
: one male, “
<collectingCountry id="ABF14C86FFFE6145FE526039FDD8FD8B" box="[455,594,613,638]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
,
<location id="D6395ACDFFFE6145FDCA603AFD53FD8B" LSID="urn:lsid:plazi:treatment:5B4FBD00FFFE615BFF026310FE2CFEB6:D6395ACDFFFE6145FDCA603AFD53FD8B" box="[607,729,613,638]" country="Madagascar" latitude="49.195" name="Mt Ambre" pageId="47" pageNumber="48">Mt Ambre</location>
, 2,5 km SE N.P. camp site,
<geoCoordinate id="B6D26AD1FFFE6145FB8A603AFB15FD8B" box="[1055,1183,614,638]" direction="south" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="-12.5199995">12°31'12''S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FFFE6145FB38603AFABBFD8B" box="[1197,1329,614,638]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="49.195">49°11'42''N</geoCoordinate>
,
<quantity id="141EA1F3FFFE6145FAAB6039FA12FD88" box="[1342,1432,613,638]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="47" pageNumber="48" unit="m" value="1050.0">
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</quantity>
</materialsCitation>
.
<materialsCitation id="638E064BFFFE6145FF0260D6FE0CFD30" ID-GBIF-Occurrence="1416610317" collectingDate="2003-03-11" collectionCode="MHNG" collectorName="G. Cuccodoro" country="Madagascar" elevation="1000" latitude="49.175556" location="Mt Ambre" pageId="47" pageNumber="48" specimenCount="1" typeStatus="paratype">
<collectingDate id="B71CD33EFFFE6145FF0260D6FE9CFD57" box="[151,278,650,674]" pageId="47" pageNumber="48" value="2003-03-11">
<date id="A7582AD6FFFE6145FF0260D6FE9CFD57" box="[151,278,650,674]" pageId="47" pageNumber="48" value="2003-03-11">11.III.2003</date>
</collectingDate>
,
<collectorName id="7E1369C0FFFE6145FEB660D6FE4BFD57" box="[291,449,649,674]" pageId="47" pageNumber="48">G. Cuccodoro</collectorName>
” (
<collectionCode id="B5F794D3FFFE6145FE4E60D6FDBDFD57" box="[475,567,650,674]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="47" pageNumber="48">MHNG</collectionCode>
); one male, “
<collectingCountry id="ABF14C86FFFE6145FD5960D5FCD2FD57" box="[716,856,649,674]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
,
<location id="D6395ACDFFFE6145FCF060D6FC55FD57" LSID="urn:lsid:plazi:treatment:5B4FBD00FFFE615BFF026310FE2CFEB6:D6395ACDFFFE6145FCF060D6FC55FD57" box="[869,991,649,674]" country="Madagascar" latitude="49.175556" name="Mt Ambre" pageId="47" pageNumber="48">Mt Ambre</location>
,
<quantity id="141EA1F3FFFE6145FC7E60D6FBAFFD57" box="[1003,1061,649,674]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="km" value="1.0">1 km</quantity>
NW N.P. camp site,
<geoCoordinate id="B6D26AD1FFFE6145FA8260D5FA1DFD57" box="[1303,1431,649,674]" direction="south" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="-12.531944">12°31'55''S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FFFE6145FF0260F2FE96FD33" box="[151,284,686,710]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="49.175556">49°10'32''N</geoCoordinate>
,
<elevation id="58CBEB25FFFE6145FEBE60F2FE0CFD30" box="[299,390,686,710]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1000.0">
<quantity id="141EA1F3FFFE6145FEBE60F2FE0CFD30" box="[299,390,686,710]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1000.0">1000 m</quantity>
</elevation>
</materialsCitation>
.
<materialsCitation id="638E064BFFFE6145FE0160F2FD0EFD1C" ID-GBIF-Occurrence="1416610313" collectingDate="2003-03-02" collectionCode="MHNG" collectorName="G. Cuccodoro" country="Madagascar" elevation="1250" latitude="49.167225" location="Mt Ambre" pageId="47" pageNumber="48" specimenCount="1" typeStatus="paratype">
<collectingDate id="B71CD33EFFFE6145FE0160F2FD8DFD33" box="[404,519,686,710]" pageId="47" pageNumber="48" value="2003-03-02">
<date id="A7582AD6FFFE6145FE0160F2FD8DFD33" box="[404,519,686,710]" pageId="47" pageNumber="48" value="2003-03-02">2.III.2003</date>
</collectingDate>
,
<collectorName id="7E1369C0FFFE6145FD8060F2FD3EFD33" box="[533,692,685,710]" pageId="47" pageNumber="48">G. Cuccodoro</collectorName>
” (
<collectionCode id="B5F794D3FFFE6145FD4460F2FCA7FD33" box="[721,813,686,710]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="47" pageNumber="48">MHNG</collectionCode>
); two males, “
<collectingCountry id="ABF14C86FFFE6145FC4160F1FBEAFD33" box="[980,1120,685,710]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
,
<location id="D6395ACDFFFE6145FBFB60F2FB63FD33" LSID="urn:lsid:plazi:treatment:5B4FBD00FFFE615BFF026310FE2CFEB6:D6395ACDFFFE6145FBFB60F2FB63FD33" box="[1134,1257,685,710]" country="Madagascar" latitude="49.167225" name="Mt Ambre" pageId="47" pageNumber="48">Mt Ambre</location>
,
<quantity id="141EA1F3FFFE6145FB6D60F2FABEFD33" box="[1272,1332,685,710]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="47" pageNumber="48" unit="km" value="3.0">3 km</quantity>
NE N.P. camp site,
<geoCoordinate id="B6D26AD1FFFE6145FE9A608EFE1AFD1F" box="[271,400,722,746]" direction="south" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="-12.541667">12°32'30''S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FFFE6145FE09608EFDAAFD1F" box="[412,544,722,746]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="49.167225">49°10'02''N</geoCoordinate>
,
<elevation id="58CBEB25FFFE6145FDB9608DFD0EFD1C" box="[556,644,721,746]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="47" pageNumber="48" unit="m" value="1250.0">
<quantity id="141EA1F3FFFE6145FDB9608DFD0EFD1C" box="[556,644,721,746]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="47" pageNumber="48" unit="m" value="1250.0">1250 m</quantity>
</elevation>
</materialsCitation>
.
<materialsCitation id="638E064BFFFE6145FD1A608DFC39FCF8" ID-GBIF-Occurrence="1416610309" collectingDate="2003-03-15" collectionCode="MHNG, ZIN" collectorName="G. Cuccodoro" country="Madagascar" elevation="950" latitude="49.194447" location="Mt Ambre" pageId="47" pageNumber="48" specimenCount="1" typeStatus="paratype">
<collectingDate id="B71CD33EFFFE6145FD1A608DFC85FD1F" box="[655,783,721,746]" pageId="47" pageNumber="48" value="2003-03-15">
<date id="A7582AD6FFFE6145FD1A608DFC85FD1F" box="[655,783,721,746]" pageId="47" pageNumber="48" value="2003-03-15">15.III.2003</date>
</collectingDate>
,
<collectorName id="7E1369C0FFFE6145FC8E608EFC32FD1F" box="[795,952,721,746]" pageId="47" pageNumber="48">G. Cuccodoro</collectorName>
” (
<collectionCode id="B5F794D3FFFE6145FC45608EFBADFD1F" box="[976,1063,722,746]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="47" pageNumber="48">MHNG</collectionCode>
,
<collectionCode id="B5F794D3FFFE6145FBA7608EFBECFD1F" box="[1074,1126,722,746]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="47" pageNumber="48">ZIN</collectionCode>
); one female, “
<collectingCountry id="ABF14C86FFFE6145FA99608DFA12FD1F" box="[1292,1432,721,746]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
,
<location id="D6395ACDFFFE6145FF0260AAFE9AFCFB" LSID="urn:lsid:plazi:treatment:5B4FBD00FFFE615BFF026310FE2CFEB6:D6395ACDFFFE6145FF0260AAFE9AFCFB" box="[151,272,757,782]" country="Madagascar" latitude="49.194447" name="Mt Ambre" pageId="47" pageNumber="48">Mt Ambre</location>
, 2,5 km SE N.P. camp site,
<geoCoordinate id="B6D26AD1FFFE6145FDDE60A9FD41FCFB" box="[587,715,757,782]" direction="south" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="-12.520833">12°31'15''S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FFFE6145FD4D60AAFCD6FCFB" box="[728,860,758,782]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="49.194447">49°11'40''N</geoCoordinate>
,
<elevation id="58CBEB25FFFE6145FCFD60A9FC39FCF8" box="[872,947,757,782]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="47" pageNumber="48" unit="m" value="950.0">
<specimenCount id="C5E0C79FFFFE6145FCFD60A9FC39FCF8" box="[872,947,757,782]" pageId="47" pageNumber="48" type="generic">
<quantity id="141EA1F3FFFE6145FCFD60A9FC39FCF8" box="[872,947,757,782]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="47" pageNumber="48" unit="m" value="950.0">950 m</quantity>
</specimenCount>
</elevation>
</materialsCitation>
.
<materialsCitation id="638E064BFFFE6145FC2A60AAFB62FCC4" ID-GBIF-Occurrence="1416610302" collectingDate="2003-03-03" collectionCode="MHNG" collectorName="G. Cuccodoro" country="Madagascar" elevation="1000" latitude="49.17361" location="Mt Ambre" pageId="47" pageNumber="48" specimenCount="1" typeStatus="paratype">
<collectingDate id="B71CD33EFFFE6145FC2A60AAFBB5FCFB" box="[959,1087,758,782]" pageId="47" pageNumber="48" value="2003-03-03">
<date id="A7582AD6FFFE6145FC2A60AAFBB5FCFB" box="[959,1087,758,782]" pageId="47" pageNumber="48" value="2003-03-03">03.III.2003</date>
</collectingDate>
,
<collectorName id="7E1369C0FFFE6145FBD960AAFB62FCFB" box="[1100,1256,757,782]" pageId="47" pageNumber="48">G. Cuccodoro</collectorName>
” (
<collectionCode id="B5F794D3FFFE6145FA9760AAFAD4FCFB" box="[1282,1374,758,782]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="47" pageNumber="48">MHNG</collectionCode>
); two females, “
<collectingCountry id="ABF14C86FFFE6145FE9C6145FE1FFCC7" box="[265,405,793,818]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
,
<location id="D6395ACDFFFE6145FE376146FD94FCC7" LSID="urn:lsid:plazi:treatment:5B4FBD00FFFE615BFF026310FE2CFEB6:D6395ACDFFFE6145FE376146FD94FCC7" box="[418,542,793,818]" country="Madagascar" latitude="49.17361" name="Mt Ambre" pageId="47" pageNumber="48">Mt Ambre</location>
, 0,5 km NE N.P. camp site,
<geoCoordinate id="B6D26AD1FFFE6145FCFE6145FC61FCC7" box="[875,1003,793,818]" direction="south" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="-12.530556">12°31'50''S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FFFE6145FC6F6145FBF5FCC7" box="[1018,1151,793,818]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="49.17361">49°10'25''N</geoCoordinate>
,
<elevation id="58CBEB25FFFE6145FB186146FB62FCC4" box="[1165,1256,794,818]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1000.0">
<quantity id="141EA1F3FFFE6145FB186146FB62FCC4" box="[1165,1256,794,818]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1000.0">1000 m</quantity>
</elevation>
</materialsCitation>
. 0
<date id="A7582AD6FFFE6145FA9D6146FAFCFCC7" box="[1288,1398,794,818]" pageId="47" pageNumber="48" value="2003-03-01">1.III.2003</date>
, G. Cuccodoro” (MHNG, ZIN).
</paragraph>
<paragraph id="D3590C16FFFE6145FF52613DFE37FC13" blockId="47.[151,1437,505,2006]" pageId="47" pageNumber="48">
<materialsCitation id="638E064BFFFE6145FF52613DFD5CFC37" ID-GBIF-Occurrence="1416610314" collectingDate="2001-01-20" collectingDateMax="2001-01-26" collectingDateMin="2001-01-20" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="925" latitude="-12.534445" location="Parc National Montagne d'Ambre" longLatPrecision="21" longitude="49.179447" municipality="Province d'Antsiranana" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Antsiranana">
<emphasis id="E192D004FFFE6145FF52613DFC4DFC8F" bold="true" box="[199,967,865,890]" pageId="47" pageNumber="48">
Additional material examined.
<collectingCountry id="ABF14C86FFFE6145FDD2613DFC93FC8F" box="[583,793,865,890]" name="Madagascar" pageId="47" pageNumber="48">MADAGASCAR</collectingCountry>
:
<collectingRegion id="1122C2F4FFFE6145FCBC613DFC49FC8F" box="[809,963,865,890]" country="Madagascar" name="Antsiranana" pageId="47" pageNumber="48">Antsiranana</collectingRegion>
:
</emphasis>
six males, “
<collectingCountry id="ABF14C86FFFE6145FBC9613EFAA0FC8F" box="[1116,1322,866,890]" name="Madagascar" pageId="47" pageNumber="48">MADAGASCAR</collectingCountry>
:
<collectingMunicipality id="333D966CFFFE6145FAAD613DFEBEFC6B" pageId="47" pageNumber="48">Province d'Antsiranana</collectingMunicipality>
,
<location id="D6395ACDFFFE6145FED061DAFD45FC6B" LSID="urn:lsid:plazi:treatment:5B4FBD00FFFE615BFF026310FE2CFEB6:D6395ACDFFFE6145FED061DAFD45FC6B" box="[325,719,901,926]" country="Madagascar" latitude="-12.534445" longLatPrecision="21" longitude="49.179447" municipality="Province d'Antsiranana" name="Parc National Montagne d'Ambre" pageId="47" pageNumber="48" stateProvince="Antsiranana">Parc National Montagne d'Ambre</location>
<locationDeviation id="C2EB1534FFFE6145FD4E61D9FBECFC6B" box="[731,1126,901,926]" pageId="47" pageNumber="48">
<quantity id="141EA1F3FFFE6145FD4E61D9FCA5FC6B" box="[731,815,901,926]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="47" pageNumber="48" unit="km" value="3.6">3.6 km</quantity>
235° SW Joffreville Elev
</locationDeviation>
<elevation id="58CBEB25FFFE6145FBE661D9FB35FC68" box="[1139,1215,901,926]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="47" pageNumber="48" unit="m" value="925.0">
<quantity id="141EA1F3FFFE6145FBE661D9FB35FC68" box="[1139,1215,901,926]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="47" pageNumber="48" unit="m" value="925.0">925 m</quantity>
</elevation>
<collectingDate id="B71CD33EFFFE6145FB5961D9FA02FC6B" box="[1228,1416,901,926]" pageId="47" pageNumber="48" value="2001-01-20" valueMax="2001-01-26" valueMin="2001-01-20">
<date id="A7582AD6FFFE6145FB5961D9FA02FC6B" box="[1228,1416,901,926]" pageId="47" pageNumber="48" value="2001-01-20" valueMax="2001-01-26" valueMin="2001-01-20">2026 Jan 2001</date>
</collectingDate>
/
<geoCoordinate id="B6D26AD1FFFE6145FF0261F6FE86FC37" box="[151,268,937,962]" direction="south" orientation="latitude" pageId="47" pageNumber="48" precision="15" value="-12.534445">12°324″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FFFE6145FE8C61F6FE11FC37" box="[281,411,937,962]" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="15" value="49.179447">49°1046″E</geoCoordinate>
coll.
<collectorName id="7E1369C0FFFE6145FE4961F5FDAEFC37" box="[476,548,937,962]" pageId="47" pageNumber="48">Fisher</collectorName>
,
<collectorName id="7E1369C0FFFE6145FDA561F5FD12FC37" box="[560,664,937,962]" pageId="47" pageNumber="48">Griswold</collectorName>
<emphasis id="E192D004FFFE6145FD3461F6FD5CFC37" box="[673,726,937,962]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
</materialsCitation>
.
<materialsCitation id="638E064BFFFE6145FD7661F5FE33FC13" ID-GBIF-Occurrence="1416610300" collectionCode="CASC" country="Madagascar" location="Sciences" pageId="47" pageNumber="48" specimenCount="1" stateProvince="California">
<collectingRegion id="1122C2F4FFFE6145FD7661F5FCDEFC37" box="[739,852,937,962]" country="United States of America" name="California" pageId="47" pageNumber="48">California</collectingRegion>
Acad. of
<location id="D6395ACDFFFE6145FC5361F5FBA1FC37" LSID="urn:lsid:plazi:treatment:5B4FBD00FFFE615BFF026310FE2CFEB6:D6395ACDFFFE6145FC5361F5FBA1FC37" box="[966,1067,937,962]" country="Madagascar" name="Sciences" pageId="47" pageNumber="48" stateProvince="California">Sciences</location>
, sifted litter, montane rainforest, code: BLF2564” (
<collectionCode id="B5F794D3FFFE6145FEF46192FE3AFC13" box="[353,432,974,998]" pageId="47" pageNumber="48">CASC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFE6145FF5261ADFC81FADF" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="D3590C16FFFE6145FF5261ADFC81FADF" blockId="47.[151,1437,505,2006]" pageId="47" pageNumber="48">
<emphasis id="E192D004FFFE6145FF5261ADFECBFBFF" bold="true" box="[199,321,1009,1034]" pageId="47" pageNumber="48">Diagnosis.</emphasis>
This species is externally similar to
<taxonomicName id="14E67795FFFE6145FD7D61AEFB90FBFF" authority="Frolov, 2010" authorityName="Frolov" authorityYear="2010" box="[744,1050,1009,1034]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="E192D004FFFE6145FD7D61AEFC0BFBFC" box="[744,897,1010,1033]" italics="true" pageId="47" pageNumber="48">M. peyrierasi</emphasis>
Frolov, 2010
</taxonomicName>
and
<taxonomicName id="14E67795FFFE6145FBC161AEFA16FBFF" authority="Frolov, 2010" authorityName="Frolov" authorityYear="2010" box="[1108,1436,1009,1034]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="perinetensis">
<emphasis id="E192D004FFFE6145FBC161AEFA88FBFC" box="[1108,1282,1010,1033]" italics="true" pageId="47" pageNumber="48">M. perinetensis</emphasis>
Frolov, 2010
</taxonomicName>
and can reliably be separated from them only by the shape of the parameres and, especially, the internal sac armature. In
<taxonomicName id="14E67795FFFE6145FEBE6666FE4EFBA4" box="[299,452,1082,1105]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="E192D004FFFE6145FEBE6666FE4EFBA4" box="[299,452,1082,1105]" italics="true" pageId="47" pageNumber="48">M. peyrierasi</emphasis>
</taxonomicName>
and
<taxonomicName id="14E67795FFFE6145FD956666FD24FBA4" box="[512,686,1082,1105]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="perinetensis">
<emphasis id="E192D004FFFE6145FD956666FD24FBA4" box="[512,686,1082,1105]" italics="true" pageId="47" pageNumber="48">M. perinetensis</emphasis>
</taxonomicName>
, the internal sac armature consists of 2 larger sclerites basally; 2 smaller, comma-shaped sclerites apically; and an area of numerous minute spinules situated between the sclerites (
<figureCitation id="4BDD1093FFFE6145FF0A66DDFF76FB6F" box="[159,252,1153,1178]" captionStart="FIGURE 26" captionStartId="51.[151,250,1889,1911]" captionTargetBox="[216,1373,283,1854]" captionTargetId="figure@51.[202,1384,265,1868]" captionTargetPageId="51" captionText="FIGURE 26. Madecorphnus perinetensis, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200469/files/figure.png" pageId="47" pageNumber="48">Figs. 26</figureCitation>
DE, 27 DE). In
<taxonomicName id="14E67795FFFE6145FE5966DEFDE5FB6F" box="[460,623,1153,1178]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="E192D004FFFE6145FE5966DEFDE5FB6F" box="[460,623,1153,1178]" italics="true" pageId="47" pageNumber="48">M. cuccodoroi</emphasis>
</taxonomicName>
, the 2 larger sclerites are probably homologous to those in the previous species, but they differ slightly in shape. The area composed of numerous minute spinules is larger in
<taxonomicName id="14E67795FFFE6145FAE866FAFE9EFB17" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="E192D004FFFE6145FAE866FAFE9EFB17" italics="true" pageId="47" pageNumber="48">M. cuccodoroi</emphasis>
</taxonomicName>
, the spinules are somewhat coarser, and a part of the apical spinules is more robust and they are almost tooth shaped (
<figureCitation id="4BDD1093FFFE6145FEA166B1FE0DFAF3" box="[308,391,1261,1286]" captionStart="FIGURE 24" captionStartId="48.[151,250,1930,1952]" captionTargetBox="[226,1348,468,1875]" captionTargetId="figure@48.[202,1385,409,1909]" captionTargetPageId="48" captionText="FIGURE 24. Madecorphnus cuccodoroi, holotype male (A, C, D, E), paratype female (B). Habitus in dorsal view, internal sac of aedeagus (C), aedeagus in lateral view and parameres in dorsal view (D), labels (E), distributional records (F)." httpUri="https://zenodo.org/record/200467/files/figure.png" pageId="47" pageNumber="48">Fig. 24</figureCitation>
C).
<taxonomicName id="14E67795FFFE6145FE3866B1FD53FAF3" box="[429,729,1261,1286]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="E192D004FFFE6145FE3866B1FD53FAF3" box="[429,729,1261,1286]" italics="true" pageId="47" pageNumber="48">Madecorphnus cuccodoroi</emphasis>
</taxonomicName>
also differs in the shape of the parameres, which are somewhat wider and more curved at apices (
<figureCitation id="4BDD1093FFFE6145FD84674DFDE4FADF" box="[529,622,1297,1322]" captionStart="FIGURE 24" captionStartId="48.[151,250,1930,1952]" captionTargetBox="[226,1348,468,1875]" captionTargetId="figure@48.[202,1385,409,1909]" captionTargetPageId="48" captionText="FIGURE 24. Madecorphnus cuccodoroi, holotype male (A, C, D, E), paratype female (B). Habitus in dorsal view, internal sac of aedeagus (C), aedeagus in lateral view and parameres in dorsal view (D), labels (E), distributional records (F)." httpUri="https://zenodo.org/record/200467/files/figure.png" pageId="47" pageNumber="48">Figs. 24</figureCitation>
D, 26B, 27B).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFE615AFF526769FCECFE90" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="description">
<paragraph id="D3590C16FFFE6145FF526769FE9FFA87" blockId="47.[151,1437,505,2006]" pageId="47" pageNumber="48">
<emphasis id="E192D004FFFE6145FF526769FE29FABB" bold="true" box="[199,419,1333,1358]" pageId="47" pageNumber="48">Description. Male.</emphasis>
Body length 5.0
<quantity id="141EA1F3FFFE6145FDF96769FD4EFAB8" box="[620,708,1333,1358]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="47" pageNumber="48" unit="mm" value="5.5">5.5 mm</quantity>
. Color uniformly dark brown, almost black, legs somewhat paler (
<figureCitation id="4BDD1093FFFE6145FF0A6705FF78FA87" box="[159,242,1369,1394]" captionStart="FIGURE 24" captionStartId="48.[151,250,1930,1952]" captionTargetBox="[226,1348,468,1875]" captionTargetId="figure@48.[202,1385,409,1909]" captionTargetPageId="48" captionText="FIGURE 24. Madecorphnus cuccodoroi, holotype male (A, C, D, E), paratype female (B). Habitus in dorsal view, internal sac of aedeagus (C), aedeagus in lateral view and parameres in dorsal view (D), labels (E), distributional records (F)." httpUri="https://zenodo.org/record/200467/files/figure.png" pageId="47" pageNumber="48">Fig. 24</figureCitation>
A).
</paragraph>
<paragraph id="D3590C16FFFE6145FF526721FCF1F9F7" blockId="47.[151,1437,505,2006]" pageId="47" pageNumber="48">Right mandible slightly longer than left, without tooth behind apex. Labrum trapezoidal, with slightly rounded sides, length about 1/6 width (in dorsal view). Frontoclypeus slightly asymmetrical, apically obtuse, with 2 long and 2 shorter setae on the apical margin. Frontoclypeus slightly depressed apicomedially. Head finely punctate with minute punctures separated by greater than 4 times their diameter.</paragraph>
<paragraph id="D3590C16FFFE6145FF526452FD37F967" blockId="47.[151,1437,505,2006]" pageId="47" pageNumber="48">Pronotum approximately 1.5 times wider than long, widest medially. Disc of pronotum convex, without any depressions, tubercles, or ridges. Punctation on pronotum finer than on head, almost indistinct. Margins with relatively wide border, lateral margins with 4 long setae: 1 seta on basal angle, 1 seta approximately in the middle of lateral margin, and 2 setae on the apical angle.</paragraph>
<paragraph id="D3590C16FFFE6145FF5264C1FC3FF943" blockId="47.[151,1437,505,2006]" box="[199,949,1693,1718]" pageId="47" pageNumber="48">Scutellum triangular, angulate apically, about 1/12 length of elytra.</paragraph>
<paragraph id="D3590C16FFFE6145FF52649DFBFEF8D7" blockId="47.[151,1437,505,2006]" pageId="47" pageNumber="48">Elytra convex, with distinct humeral and apical, widest at basal third. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra sparsely punctate with relatively large punctures. Epipleura with long, sparse, brown setae. Base of elytron with border from scutellum to humeral callus.</paragraph>
<paragraph id="D3590C16FFFE6145FF526571FF46F847" blockId="47.[151,1437,505,2006]" pageId="47" pageNumber="48">Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex of protibia with robust, spur-like seta and a few smaller setae basally. Mesothoracic and metathoracic legs similar in shape to each other. Longer tibial spur shorter than 2 basal tarsomeres in mesothoracic legs and as long as 2 tarsomeres in metathoracic legs.</paragraph>
<paragraph id="D3590C16FFFE615AFF5265E2FC0AFF21" blockId="47.[151,1437,505,2006]" lastBlockId="48.[151,1436,151,357]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
Parameres with small teeth laterally, narrowly rounded apically in dorsal view and curved downwards in lateral view (
<figureCitation id="4BDD1093FFE1615AFF4962CBFEA5FF45" box="[220,303,151,176]" captionStart="FIGURE 24" captionStartId="48.[151,250,1930,1952]" captionTargetBox="[226,1348,468,1875]" captionTargetId="figure@48.[202,1385,409,1909]" captionTargetPageId="48" captionText="FIGURE 24. Madecorphnus cuccodoroi, holotype male (A, C, D, E), paratype female (B). Habitus in dorsal view, internal sac of aedeagus (C), aedeagus in lateral view and parameres in dorsal view (D), labels (E), distributional records (F)." httpUri="https://zenodo.org/record/200467/files/figure.png" pageId="48" pageNumber="49">Fig. 24</figureCitation>
D). Internal sac with 2 long sclerites and a large number of small spinules; some spinules more robust and form 2 indistinctly separated clusters in apical part (
<figureCitation id="4BDD1093FFE1615AFC9862E0FCEAFF21" box="[781,864,188,213]" captionStart="FIGURE 24" captionStartId="48.[151,250,1930,1952]" captionTargetBox="[226,1348,468,1875]" captionTargetId="figure@48.[202,1385,409,1909]" captionTargetPageId="48" captionText="FIGURE 24. Madecorphnus cuccodoroi, holotype male (A, C, D, E), paratype female (B). Habitus in dorsal view, internal sac of aedeagus (C), aedeagus in lateral view and parameres in dorsal view (D), labels (E), distributional records (F)." httpUri="https://zenodo.org/record/200467/files/figure.png" pageId="48" pageNumber="49">Fig. 24</figureCitation>
C).
</paragraph>
<paragraph id="D3590C16FFE1615AFF526283FDE7FEE8" blockId="48.[151,1436,151,357]" pageId="48" pageNumber="49">
<emphasis id="E192D004FFE1615AFF526283FEAFFF0D" bold="true" box="[199,293,223,248]" pageId="48" pageNumber="49">Female.</emphasis>
Females (
<figureCitation id="4BDD1093FFE1615AFE0F6283FE67FF0D" box="[410,493,223,248]" captionStart="FIGURE 24" captionStartId="48.[151,250,1930,1952]" captionTargetBox="[226,1348,468,1875]" captionTargetId="figure@48.[202,1385,409,1909]" captionTargetPageId="48" captionText="FIGURE 24. Madecorphnus cuccodoroi, holotype male (A, C, D, E), paratype female (B). Habitus in dorsal view, internal sac of aedeagus (C), aedeagus in lateral view and parameres in dorsal view (D), labels (E), distributional records (F)." httpUri="https://zenodo.org/record/200467/files/figure.png" pageId="48" pageNumber="49">Fig. 24</figureCitation>
B) differ from males in having the apical spur of the protibiae that is slightly longer and more robust than apical seta in males.
</paragraph>
<paragraph id="D3590C16FFE1615AFF52637BFCECFE90" blockId="48.[151,1436,151,357]" pageId="48" pageNumber="49">
<emphasis id="E192D004FFE1615AFF52637BFEB5FEB5" bold="true" box="[199,319,295,320]" pageId="48" pageNumber="49">Variation.</emphasis>
Body length of the examined specimens varies from 5.0
<quantity id="141EA1F3FFE1615AFC53637BFB94FECA" box="[966,1054,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="48" pageNumber="49" unit="mm" value="5.5">5.5 mm</quantity>
. Left and right mandibles of some males are almost equal in length and similar to those in females.
</paragraph>
</subSubSection>
<caption id="87995C9EFFE1615AFF0265D6FA84F84A" httpUri="https://zenodo.org/record/200467/files/figure.png" pageId="48" pageNumber="49" targetBox="[226,1348,468,1875]" targetPageId="48">
<paragraph id="D3590C16FFE1615AFF0265D6FA84F84A" blockId="48.[151,1435,1930,1983]" pageId="48" pageNumber="49">
<emphasis id="E192D004FFE1615AFF0265D6FEAAF855" bold="true" box="[151,288,1930,1952]" pageId="48" pageNumber="49">FIGURE 24.</emphasis>
<taxonomicName id="14E67795FFE1615AFEB365D6FDBFF855" box="[294,565,1930,1952]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="E192D004FFE1615AFEB365D6FDBFF855" box="[294,565,1930,1952]" italics="true" pageId="48" pageNumber="49">Madecorphnus cuccodoroi</emphasis>
</taxonomicName>
, holotype male (A, C, D, E), paratype female (B). Habitus in dorsal view, internal sac of aedeagus (C), aedeagus in lateral view and parameres in dorsal view (D), labels (E), distributional records (F).
</paragraph>
</caption>
<subSubSection id="9BFC5F9DFFE0615BFF5262CBFE2CFEB6" pageId="49" pageNumber="50" type="distribution">
<paragraph id="D3590C16FFE0615BFF5262CBFE2CFEB6" blockId="49.[151,1436,151,323]" pageId="49" pageNumber="50">
<emphasis id="E192D004FFE0615BFF5262CBFE7BFF45" bold="true" box="[199,497,151,176]" pageId="49" pageNumber="50">Distribution and habitat.</emphasis>
All specimens were collected in the Amber Mountain National Park in
<collectingRegion id="1122C2F4FFE0615BFA8062CBFF71FF20" country="Madagascar" name="Antsiranana" pageId="49" pageNumber="50">Antsiranana Province</collectingRegion>
(northern
<collectingCountry id="ABF14C86FFE0615BFEFA62E0FE72FF20" box="[367,504,188,213]" name="Madagascar" pageId="49" pageNumber="50">Madagascar</collectingCountry>
,
<figureCitation id="4BDD1093FFE0615BFD9662E0FDDFFF21" box="[515,597,188,213]" captionStart="FIGURE 24" captionStartId="48.[151,250,1930,1952]" captionTargetBox="[226,1348,468,1875]" captionTargetId="figure@48.[202,1385,409,1909]" captionTargetPageId="48" captionText="FIGURE 24. Madecorphnus cuccodoroi, holotype male (A, C, D, E), paratype female (B). Habitus in dorsal view, internal sac of aedeagus (C), aedeagus in lateral view and parameres in dorsal view (D), labels (E), distributional records (F)." httpUri="https://zenodo.org/record/200467/files/figure.png" pageId="49" pageNumber="50">Fig. 24</figureCitation>
F). The park occupies the Madagascan northernmost remnant of indigenous humid forest covering about
<quantity id="141EA1F3FFE0615BFE7C62BFFDC8FF0E" box="[489,578,226,251]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="49" pageNumber="50" unit="km" value="200.0">200 km</quantity>
2. All specimens were collected by sifting forest litter. The number of the collected specimens suggests that the population density of
<taxonomicName id="14E67795FFE0615BFCAB6354FC68FEEA" box="[830,994,262,287]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="E192D004FFE0615BFCAB6354FC68FEEA" box="[830,994,262,287]" italics="true" pageId="49" pageNumber="50">M. cuccodoroi</emphasis>
</taxonomicName>
is low at least during the season when the collecting was done.
</paragraph>
</subSubSection>
</treatment>
</document>