239 lines
36 KiB
XML
239 lines
36 KiB
XML
<document id="9B7BA5FB7CA00F39CF39B32B24FE6DBA" ID-DOI="10.11646/zootaxa.5026.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300792" ID-ZooBank="E712BC8E-3E75-422C-8587-69ED17B6CDD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630041845836" checkinUser="plazi" docAuthor="Fomichev, Alexander A." docDate="2021" docId="D220E4309B1EA407BFED8DEAFD64FAD6" docLanguage="en" docName="zootaxa.5026.4.7.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acantholycosa sergeevi Fomichev 2021, sp. n." docType="treatment" docVersion="4" lastPageNumber="579" masterDocId="2E199C489B12A40BBF7A8D7DFF9FFFAC" masterDocTitle="New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia" masterLastPageNumber="585" masterPageNumber="567" pageNumber="579" updateTime="1699139093203" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="02F07CB52A37E23DF59B91787381A59B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3EDA160A7927BAF84C8E926248656AF2">
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<mods:title id="99A70D19E97EA657CD2DAFC79AC64E3A">New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia</mods:title>
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<mods:name id="E0226368ADAC6C63FA03190507D118C9" type="personal">
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<mods:roleTerm id="9548A70CC8907DCAB786D3BE4E5B629F">Author</mods:roleTerm>
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<mods:namePart id="F24FE86BB5A6F650E5DFA6E595DB219B">Fomichev, Alexander A.</mods:namePart>
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<mods:affiliation id="5C2D421E7D3E106A3C6248DF233534E4">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. & Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="AA189D7123970A4C318AE6DD7A88C14F">text</mods:typeOfResource>
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<mods:title id="BC082D26386547B39DD4BA4CCBCF4136">Zootaxa</mods:title>
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<mods:part id="9B12B953CCCE92B3BEAD1D6AB9596227">
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<mods:date id="E6E883B8752FE68950B8889D7CD51850">2021</mods:date>
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<mods:number id="A137399C3223AE72CE9636F36AAF8C23">2021-08-27</mods:number>
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<mods:number id="F6960656D96384D44584D7B4615BEDDA">5026</mods:number>
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<mods:number id="902461A92F50F87DE80903AA8CD1663D">4</mods:number>
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<mods:classification id="8AE5E88AC7700A1DAA1F42A0378EB637">journal article</mods:classification>
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<mods:identifier id="965CF488FBCA552D44909664B6E26B2A" type="DOI">10.11646/zootaxa.5026.4.7</mods:identifier>
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<mods:identifier id="316769D571B9890C604FD7383A691126" type="ISSN">1175-5326</mods:identifier>
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<treatment id="D220E4309B1EA407BFED8DEAFD64FAD6" ID-DOI="http://doi.org/10.5281/zenodo.5307792" ID-Zenodo-Dep="5307792" LSID="urn:lsid:plazi:treatment:D220E4309B1EA407BFED8DEAFD64FAD6" httpUri="http://treatment.plazi.org/id/D220E4309B1EA407BFED8DEAFD64FAD6" lastPageNumber="579" pageId="12" pageNumber="579">
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<subSubSection id="129306AD9B1EA407BFED8DEAFE64FF1D" box="[151,507,151,178]" pageId="12" pageNumber="579" type="nomenclature">
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<paragraph id="5A3655269B1EA407BFED8DEAFE64FF1D" blockId="12.[151,584,151,214]" box="[151,507,151,178]" pageId="12" pageNumber="579">
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<heading id="017EE24A9B1EA407BFED8DEAFE64FF1D" bold="true" box="[151,507,151,178]" fontSize="11" level="1" pageId="12" pageNumber="579" reason="1">
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<emphasis id="68FD89349B1EA407BFED8DEAFE64FF1D" bold="true" box="[151,507,151,178]" pageId="12" pageNumber="579">
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<taxonomicName id="9D892EA59B1EA407BFED8DEAFE2DFF1D" ID-CoL="99CP9" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[151,434,151,178]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
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<emphasis id="68FD89349B1EA407BFED8DEAFE2DFF1D" bold="true" box="[151,434,151,178]" italics="true" pageId="12" pageNumber="579">Acantholycosa sergeevi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="73CE344F9B1EA407BEC38DEAFE64FF1D" box="[441,507,151,177]" pageId="12" pageNumber="579" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="129306AD9B1EA407BFED8DC6FDD7FF79" box="[151,584,187,214]" pageId="12" pageNumber="579" type="description">
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<paragraph id="5A3655269B1EA407BFED8DC6FDD7FF79" blockId="12.[151,584,151,214]" box="[151,584,187,214]" pageId="12" pageNumber="579">
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<figureCitation id="C2B249A39B1EA407BFED8DC6FF64FF79" box="[151,251,187,213]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="12" pageNumber="579">Figs 6–7</figureCitation>
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,
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<figureCitation id="C2B249A39B1EA407BE7C8DC6FED1FF79" box="[262,334,187,213]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="12" pageNumber="579">10–11</figureCitation>
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,
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<figureCitation id="C2B249A39B1EA407BE238DC6FE3DFF79" box="[345,418,187,213]" captionStart="FIGURES 32–43" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[154,1435,189,1719]" captionTargetId="figure-33@6.[151,1437,181,1719]" captionTargetPageId="6" captionText="FIGURES 32–43. Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of Acantholycosa sayanensis (32–33), A. sidorovi sp. n. (34–35), A. levinae (36–37) and A. sergeevi sp. n. (38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pe extension of the palea, Pl palea, Ta terminal apophysis, Tg tegular apophysis, Ts triangle-shaped projection of the palea." figureDoi="http://doi.org/10.5281/zenodo.5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="12" pageNumber="579">38–43</figureCitation>
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,
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<figureCitation id="C2B249A39B1EA407BED78DC6FE69FF7A" box="[429,502,187,214]" captionStart="FIGURES 44–53" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[163,1427,188,1770]" captionTargetId="figure-15@7.[151,1436,181,1770]" captionTargetPageId="7" captionText="FIGURES 44–53. Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of Acantholycosa sergeevi sp. n. (44–45) and Mongolicosa lizae sp. n. (46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pl palea, Sd sperm duct, Ta terminal apophysis, Tg tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="12" pageNumber="579">44–45</figureCitation>
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,
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<figureCitation id="C2B249A39B1EA407BD7B8DC6FDD7FF79" box="[513,584,187,213]" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="12" pageNumber="579">74–76</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="129306AD9B1EA407BFED8C7EFADDFEED" pageId="12" pageNumber="579" type="materials_examined">
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<paragraph id="5A3655269B1EA407BFED8C7EFADDFEED" blockId="12.[151,1437,259,1402]" pageId="12" pageNumber="579">
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<emphasis id="68FD89349B1EA407BFED8C7EFF46FEB1" bold="true" box="[151,217,259,285]" pageId="12" pageNumber="579">Type.</emphasis>
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<materialsCitation id="EAE15F7B9B1EA407BF9B8C7EFAA1FEED" collectingDate="2020-07-16" collectionCode="ISEA" collectorName="A. A. Fomichev" country="Russia" elevation="1900" location="Borus Mt. Range" pageId="12" pageNumber="579" specimenCount="1" specimenCount-male="1" stateProvince="Krasnoyarsk" typeStatus="holotype">
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<collectingCountry id="229E15B69B1EA407BF9B8C7EFED6FEB1" box="[225,329,259,285]" name="Russia" pageId="12" pageNumber="579">RUSSIA</collectingCountry>
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:
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<emphasis id="68FD89349B1EA407BE2F8C7EFDAEFEB1" bold="true" box="[341,561,259,285]" pageId="12" pageNumber="579">
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||
<collectingRegion id="984D9BC49B1EA407BE2F8C7EFDAEFEB1" box="[341,561,259,285]" country="Russia" name="Krasnoyarsk" pageId="12" pageNumber="579">Krasnoyarsk Krai</collectingRegion>
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</emphasis>
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:
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<typeStatus id="8532EB849B1EA407BD388C7EFD3BFEB1" box="[578,676,259,285]" pageId="12" pageNumber="579" type="holotype">holotype</typeStatus>
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<specimenCount id="4C8F9EAF9B1EA407BDD68C79FD5EFEB2" box="[684,705,260,286]" pageId="12" pageNumber="579" type="male">♂</specimenCount>
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(
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<collectionCode id="3C98CDE39B1EA407BDA88C7EFC8BFEB1" box="[722,788,259,285]" pageId="12" pageNumber="579">ISEA</collectionCode>
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, 001.8660),
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<location id="5F5603FD9B1EA407BCD98C79FBF7FEB2" LSID="urn:lsid:plazi:treatment:D220E4309B1EA407BFED8DEAFD64FAD6:5F5603FD9B1EA407BCD98C79FBF7FEB2" box="[931,1128,260,286]" country="Russia" name="Borus Mt. Range" pageId="12" pageNumber="579" stateProvince="Krasnoyarsk">Borus Mt. Range</location>
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, Malyi Borus and Bol’shoi Borus (=Poilova) Mountains, 52°47–48’N, 91°31–33’E, scree,
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<quantity id="9D71F8C39B1EA407BC268C5AFC66FEEE" box="[860,1017,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="12" pageNumber="579" unit="m" value="1900.0" valueMax="2300.0" valueMin="1500.0">
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<elevation id="D1A4B2159B1EA407BC268C5AFC66FEEE" box="[860,1017,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="12" pageNumber="579" unit="m" value="1900.0" valueMax="2300.0" valueMin="1500.0">1500–2300 m</elevation>
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</quantity>
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,
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<date id="2E3773E69B1EA407BB7F8C5AFB1AFEED" box="[1029,1157,295,321]" pageId="12" pageNumber="579" value="2020-07-16">
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<collectingDate id="3E738A0E9B1EA407BB7F8C5AFB1AFEED" box="[1029,1157,295,321]" pageId="12" pageNumber="579" value="2020-07-16">16.07.2020</collectingDate>
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</date>
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,
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<collectorName id="F77C30F09B1EA407BBF58C5AFAA1FEED" box="[1167,1342,295,321]" pageId="12" pageNumber="579">A.A. Fomichev</collectorName>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="129306AD9B1EA407BFBD8C36FD8DFE01" pageId="12" pageNumber="579" type="etymology">
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<paragraph id="5A3655269B1EA407BFBD8C36FD8DFE01" blockId="12.[151,1437,259,1402]" pageId="12" pageNumber="579">
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<emphasis id="68FD89349B1EA407BFBD8C36FED1FEC9" bold="true" box="[199,334,331,357]" pageId="12" pageNumber="579">Etymology.</emphasis>
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The specific name is a patronym given in honor of my friend, Vasiliy V. Sergeev (
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<collectingRegion id="984D9BC49B1EA407BA718C36FA06FEC9" box="[1291,1433,331,357]" country="Russia" name="Novosibirsk" pageId="12" pageNumber="579">Novosibirsk</collectingRegion>
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,
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<collectingCountry id="229E15B69B1EA407BFED8C12FF79FE26" box="[151,230,367,394]" name="Russia" pageId="12" pageNumber="579">Russia</collectingCountry>
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), who helped in spider collection and in organization of the field trip to Borus Mt. Range, where the
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<typeStatus id="8532EB849B1EA407BA408C12FA03FE25" box="[1338,1436,367,393]" pageId="12" pageNumber="579" type="holotype">holotype</typeStatus>
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of this new species was collected.
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</paragraph>
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</subSubSection>
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<subSubSection id="129306AD9B1EA407BFBD8CCAFD45FC95" pageId="12" pageNumber="579" type="diagnosis">
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<paragraph id="5A3655269B1EA407BFBD8CCAFD45FC95" blockId="12.[151,1437,259,1402]" pageId="12" pageNumber="579">
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<emphasis id="68FD89349B1EA407BFBD8CCAFEDDFE7D" bold="true" box="[199,322,439,465]" pageId="12" pageNumber="579">Diagnosis.</emphasis>
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The new species is similar to the Siberian species
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<taxonomicName id="9D892EA59B1EA407BC0C8CCAFC73FE7D" baseAuthorityName="Marusik" baseAuthorityYear="1993" box="[886,1004,439,465]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sterneri">
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<emphasis id="68FD89349B1EA407BC0C8CCAFC73FE7D" box="[886,1004,439,465]" italics="true" pageId="12" pageNumber="579">A. sterneri</emphasis>
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</taxonomicName>
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as well as
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<taxonomicName id="9D892EA59B1EA407BB1C8CCAFB40FE7D" baseAuthorityName="Levi & Levi" baseAuthorityYear="1951" box="[1126,1247,439,465]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="solituda">
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<emphasis id="68FD89349B1EA407BB1C8CCAFB40FE7D" box="[1126,1247,439,465]" italics="true" pageId="12" pageNumber="579">A. solituda</emphasis>
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</taxonomicName>
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, from the Rocky Mountains (
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<collectingCountry id="229E15B69B1EA407BE5B8CA6FEC0FE59" box="[289,351,475,501]" name="United States of America" pageId="12" pageNumber="579">USA</collectingCountry>
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). All three species share the following features: palea (
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<emphasis id="68FD89349B1EA407BCA78CA6FC69FE59" box="[989,1014,475,501]" italics="true" pageId="12" pageNumber="579">Pl</emphasis>
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) with a prominent extension (
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<emphasis id="68FD89349B1EA407BA2C8CA1FAEBFE59" box="[1366,1396,476,501]" italics="true" pageId="12" pageNumber="579">Pe</emphasis>
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) in the terminal part and the triangle shaped projection (
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<emphasis id="68FD89349B1EA407BD918F7DFC9CFDB5" box="[747,771,512,537]" italics="true" pageId="12" pageNumber="579">Ts</emphasis>
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) on its posterior edge, strong terminal apophysis (
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<emphasis id="68FD89349B1EA407BA448F7DFAC6FDB5" box="[1342,1369,512,537]" italics="true" pageId="12" pageNumber="579">Ta</emphasis>
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) with bifurcate tip and thick, cotton-like pubescence on legs I and II.
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<taxonomicName id="9D892EA59B1EA407BCF98F5EFB11FD91" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[899,1166,547,573]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
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<emphasis id="68FD89349B1EA407BCF98F5EFB11FD91" box="[899,1166,547,573]" italics="true" pageId="12" pageNumber="579">Acantholycosa sergeevi</emphasis>
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</taxonomicName>
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<emphasis id="68FD89349B1EA407BBE38F5EFB43FD91" bold="true" box="[1177,1244,547,573]" pageId="12" pageNumber="579">
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<taxonomicNameLabel id="73CE344F9B1EA407BBE38F5EFB43FD91" box="[1177,1244,547,573]" pageId="12" pageNumber="579" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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differs from the both aforementioned species by the more developed and pointed triangle shaped projection on the posterior edge of the palea (cf.
|
||
<figureCitation id="C2B249A39B1EA407BE2B8F16FE50FD29" box="[337,463,619,645]" captionStart="FIGURES 32–43" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[154,1435,189,1719]" captionTargetId="figure-33@6.[151,1437,181,1719]" captionTargetPageId="6" captionText="FIGURES 32–43. Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of Acantholycosa sayanensis (32–33), A. sidorovi sp. n. (34–35), A. levinae (36–37) and A. sergeevi sp. n. (38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pe extension of the palea, Pl palea, Ta terminal apophysis, Tg tegular apophysis, Ts triangle-shaped projection of the palea." figureDoi="http://doi.org/10.5281/zenodo.5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="12" pageNumber="579">Figs 41–42</figureCitation>
|
||
and fig 3, 7 in
|
||
<bibRefCitation id="3E1828D79B1EA407BD048F16FC5FFD29" author="Kronestedt, T. & Marusik, Y. M." box="[638,960,619,645]" pageId="12" pageNumber="579" pagination="63 - 71" refId="ref10150" refString="Kronestedt, T. & Marusik, Y. M. (2002) On Acantholycosa solituda (Levi & Levi) and A. sterneri (Marusik) (Araneae: Lycosidae), a pair of geographically distant allied species. Acta Arachnologica, 51 (1), 63 - 71. https: // doi. org / 10.2476 / asjaa. 51.63" type="journal article" year="2002">Kronestedt & Marusik 2002</bibRefCitation>
|
||
). The new species can be differed from
|
||
<taxonomicName id="9D892EA59B1EA407BAFE8F16FF71FD05" baseAuthorityName="Marusik" baseAuthorityYear="1993" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sterneri">
|
||
<emphasis id="68FD89349B1EA407BAFE8F16FF71FD05" italics="true" pageId="12" pageNumber="579">A. sterneri</emphasis>
|
||
</taxonomicName>
|
||
by the terminal part of the palea less extended, the embolus (
|
||
<emphasis id="68FD89349B1EA407BCEF8FEDFC25FD05" box="[917,954,656,681]" italics="true" pageId="12" pageNumber="579">Em</emphasis>
|
||
) directed at 2:30 o’clock (vs. 3:30 o’clock) and by the almost straight anterior edge of the tegular apophysis (
|
||
<emphasis id="68FD89349B1EA407BCF18FC9FC36FD61" box="[907,937,692,717]" italics="true" pageId="12" pageNumber="579">Tg</emphasis>
|
||
) (vs. triangular, cf.
|
||
<figureCitation id="C2B249A39B1EA407BBF58FCEFA8DFD61" box="[1167,1298,691,717]" captionStart="FIGURES 32–43" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[154,1435,189,1719]" captionTargetId="figure-33@6.[151,1437,181,1719]" captionTargetPageId="6" captionText="FIGURES 32–43. Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of Acantholycosa sayanensis (32–33), A. sidorovi sp. n. (34–35), A. levinae (36–37) and A. sergeevi sp. n. (38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pe extension of the palea, Pl palea, Ta terminal apophysis, Tg tegular apophysis, Ts triangle-shaped projection of the palea." figureDoi="http://doi.org/10.5281/zenodo.5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="12" pageNumber="579">Figs 38, 41</figureCitation>
|
||
,
|
||
<figureCitation id="C2B249A39B1EA407BA5A8FCEFAA3FD61" box="[1312,1340,691,717]" captionStart="FIGURES 44–53" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[163,1427,188,1770]" captionTargetId="figure-15@7.[151,1436,181,1770]" captionTargetPageId="7" captionText="FIGURES 44–53. Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of Acantholycosa sergeevi sp. n. (44–45) and Mongolicosa lizae sp. n. (46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pl palea, Sd sperm duct, Ta terminal apophysis, Tg tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="12" pageNumber="579">44</figureCitation>
|
||
and fig.
|
||
<quantity id="9D71F8C39B1EA407BFED8FA5FF5CFD5D" box="[151,195,727,754]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="12" pageNumber="579" unit="in" value="7.0">7 in</quantity>
|
||
<bibRefCitation id="3E1828D79B1EA407BFB18FAAFD91FD5D" author="Kronestedt, T. & Marusik, Y. M." box="[203,526,727,753]" pageId="12" pageNumber="579" pagination="63 - 71" refId="ref10150" refString="Kronestedt, T. & Marusik, Y. M. (2002) On Acantholycosa solituda (Levi & Levi) and A. sterneri (Marusik) (Araneae: Lycosidae), a pair of geographically distant allied species. Acta Arachnologica, 51 (1), 63 - 71. https: // doi. org / 10.2476 / asjaa. 51.63" type="journal article" year="2002">Kronestedt & Marusik 2002</bibRefCitation>
|
||
).
|
||
<taxonomicName id="9D892EA59B1EA407BD5B8FAAFCB5FD5D" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[545,810,727,753]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
|
||
<emphasis id="68FD89349B1EA407BD5B8FAAFCB5FD5D" box="[545,810,727,753]" italics="true" pageId="12" pageNumber="579">Acantholycosa sergeevi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="68FD89349B1EA407BC488FAAFCEDFD5D" bold="true" box="[818,882,727,753]" pageId="12" pageNumber="579">
|
||
<taxonomicNameLabel id="73CE344F9B1EA407BC488FAAFCEDFD5D" box="[818,882,727,753]" pageId="12" pageNumber="579" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="9D892EA59B1EA407BBDD8FAAFABEFD5D" baseAuthorityName="Levi & Levi" baseAuthorityYear="1951" box="[1191,1313,727,753]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="solituda">
|
||
<emphasis id="68FD89349B1EA407BBDD8FAAFABEFD5D" box="[1191,1313,727,753]" italics="true" pageId="12" pageNumber="579">A. solituda</emphasis>
|
||
</taxonomicName>
|
||
by thinner embolus (in anterior view) and by a straight basal arm (
|
||
<emphasis id="68FD89349B1EA407BC758F81FCB1FCB9" box="[783,814,764,789]" italics="true" pageId="12" pageNumber="579">Ba</emphasis>
|
||
) of the tegular apophysis (vs. bent posteriorly, cf.
|
||
<figureCitation id="C2B249A39B1EA407BA168F86FF29FC95" captionStart="FIGURES 32–43" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[154,1435,189,1719]" captionTargetId="figure-33@6.[151,1437,181,1719]" captionTargetPageId="6" captionText="FIGURES 32–43. Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of Acantholycosa sayanensis (32–33), A. sidorovi sp. n. (34–35), A. levinae (36–37) and A. sergeevi sp. n. (38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pe extension of the palea, Pl palea, Ta terminal apophysis, Tg tegular apophysis, Ts triangle-shaped projection of the palea." figureDoi="http://doi.org/10.5281/zenodo.5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="12" pageNumber="579">Figs 42</figureCitation>
|
||
,
|
||
<figureCitation id="C2B249A39B1EA407BFBB8E62FF42FC95" box="[193,221,799,825]" captionStart="FIGURES 44–53" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[163,1427,188,1770]" captionTargetId="figure-15@7.[151,1436,181,1770]" captionTargetPageId="7" captionText="FIGURES 44–53. Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of Acantholycosa sergeevi sp. n. (44–45) and Mongolicosa lizae sp. n. (46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pl palea, Sd sperm duct, Ta terminal apophysis, Tg tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="12" pageNumber="579">44</figureCitation>
|
||
and fig.
|
||
<quantity id="9D71F8C39B1EA407BE3B8E62FE17FC95" box="[321,392,799,826]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="12" pageNumber="579" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4–5 in</quantity>
|
||
<bibRefCitation id="3E1828D79B1EA407BEF58E62FD51FC95" author="Kronestedt, T. & Marusik, Y. M." box="[399,718,799,825]" pageId="12" pageNumber="579" pagination="63 - 71" refId="ref10150" refString="Kronestedt, T. & Marusik, Y. M. (2002) On Acantholycosa solituda (Levi & Levi) and A. sterneri (Marusik) (Araneae: Lycosidae), a pair of geographically distant allied species. Acta Arachnologica, 51 (1), 63 - 71. https: // doi. org / 10.2476 / asjaa. 51.63" type="journal article" year="2002">Kronestedt & Marusik 2002</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="129306AD9B1EA407BFBD8E3EFE0EFB45" pageId="12" pageNumber="579" type="description">
|
||
<paragraph id="5A3655269B1EA407BFBD8E3EFB0EFB9A" blockId="12.[151,1437,259,1402]" pageId="12" pageNumber="579">
|
||
<emphasis id="68FD89349B1EA407BFBD8E3EFEC6FCF1" bold="true" box="[199,345,835,861]" pageId="12" pageNumber="579">Description.</emphasis>
|
||
Male (
|
||
<typeStatus id="8532EB849B1EA407BECA8E3EFD86FCF2" box="[432,537,835,862]" pageId="12" pageNumber="579" type="holotype">holotype</typeStatus>
|
||
). Total length 10.3. Carapace: 5.1 long, 4.25 wide. General appearance as in
|
||
<figureCitation id="C2B249A39B1EA407BFED8E1AFF40FC2D" box="[151,223,871,897]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="12" pageNumber="579">Fig. 6</figureCitation>
|
||
. Coloration. Carapace and chelicerae black. Sternum and labium dark-brown. Endites yellow-gray. Palps black. Coxae yellow-gray with yellow spots. Legs dark brown, almost black. Legs I–III have a thick cotton-like pubescence of curled hairs. Abdomen dark gray dorsally, almost black. Cardiac mark dark brown. Venter of the abdomen yellow-gray. Spinnerets black. Spination of leg I: Fe d1-1-1 p0-0-3 r0-1-1; Pa p1 r1; Ti d1-0-1 p1-0-0 v2- 2-2-2-2-2-2-2; Mt p1-1-0 r1-1-0 v2-2-0. Leg measurements: I 4.6, 2.2, 4.35, 4.15, 1.85 (17.15); II 4.55, 2.15, 4.2, 4.25, 1.95 (17.1); III 4.6, 2.0, 3.85, 5.45, 2.05 (17.95); IV 5.35, 2.05, 4.8, 7.85, 2.55 (22.6).
|
||
</paragraph>
|
||
<paragraph id="5A3655269B1EA407BFBD8942FC43FB69" blockId="12.[151,1437,259,1402]" pageId="12" pageNumber="579">
|
||
Palp as in
|
||
<figureCitation id="C2B249A39B1EA407BE478942FE22FBF5" box="[317,445,1087,1113]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="12" pageNumber="579">Figs 10–11</figureCitation>
|
||
,
|
||
<figureCitation id="C2B249A39B1EA407BEB38942FD8DFBF5" box="[457,530,1087,1113]" captionStart="FIGURES 32–43" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[154,1435,189,1719]" captionTargetId="figure-33@6.[151,1437,181,1719]" captionTargetPageId="6" captionText="FIGURES 32–43. Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of Acantholycosa sayanensis (32–33), A. sidorovi sp. n. (34–35), A. levinae (36–37) and A. sergeevi sp. n. (38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pe extension of the palea, Pl palea, Ta terminal apophysis, Tg tegular apophysis, Ts triangle-shaped projection of the palea." figureDoi="http://doi.org/10.5281/zenodo.5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="12" pageNumber="579">38–43</figureCitation>
|
||
,
|
||
<figureCitation id="C2B249A39B1EA407BD648942FDF7FBF6" box="[542,616,1087,1114]" captionStart="FIGURES 44–53" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[163,1427,188,1770]" captionTargetId="figure-15@7.[151,1436,181,1770]" captionTargetPageId="7" captionText="FIGURES 44–53. Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of Acantholycosa sergeevi sp. n. (44–45) and Mongolicosa lizae sp. n. (46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pl palea, Sd sperm duct, Ta terminal apophysis, Tg tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="12" pageNumber="579">44–45</figureCitation>
|
||
. Tegular apophysis hammer-shaped, with reduced apical arm. Palea with abrupt retrolateral margin. Triangle-shaped projection straight and sharply pointed, bearing conical tubercle on the retrolateral edge. Terminal apophysis massive, subdivided into two lobes. Embolus completely straight. Distal part of the embolus bears a rectangular laminar outgrowth on the anterior edge.
|
||
</paragraph>
|
||
<paragraph id="5A3655269B1EA407BFBD89B2FE0EFB45" blockId="12.[151,1437,259,1402]" box="[199,401,1231,1257]" pageId="12" pageNumber="579">Female unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="129306AD9B1EA407BFBD898EFB63FAA2" box="[199,1276,1267,1294]" pageId="12" pageNumber="579" type="distribution">
|
||
<paragraph id="5A3655269B1EA407BFBD898EFB63FAA2" blockId="12.[151,1437,259,1402]" box="[199,1276,1267,1294]" pageId="12" pageNumber="579">
|
||
<emphasis id="68FD89349B1EA407BFBD898EFEFEFAA1" bold="true" box="[199,353,1267,1293]" pageId="12" pageNumber="579">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="8532EB849B1EA407BD1A8989FD0FFAA2" box="[608,656,1268,1294]" pageId="12" pageNumber="579">type</typeStatus>
|
||
locality in the Western Sayan Mountains (
|
||
<figureCitation id="C2B249A39B1EA407BB16898EFB6FFAA2" box="[1132,1264,1267,1294]" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="12" pageNumber="579">Figs 74–76</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="129306AD9B1EA407BFBD886AFD64FAD6" pageId="12" pageNumber="579" type="discussion">
|
||
<paragraph id="5A3655269B1EA407BFBD886AFD64FAD6" blockId="12.[151,1437,259,1402]" pageId="12" pageNumber="579">
|
||
<emphasis id="68FD89349B1EA407BFBD886AFED0FA9D" bold="true" box="[199,335,1303,1329]" pageId="12" pageNumber="579">Comments.</emphasis>
|
||
This species was found in the same locality and in the same scree patches as
|
||
<taxonomicName id="9D892EA59B1EA407BB8D886AFE8FFAF9" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sayanensis">
|
||
<emphasis id="68FD89349B1EA407BB8D886AFE8FFAF9" italics="true" pageId="12" pageNumber="579">Acantholycosa sayanensis</emphasis>
|
||
</taxonomicName>
|
||
(see above). Interestingly,
|
||
<taxonomicName id="9D892EA59B1EA407BD3F8846FD5DFAF9" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[581,706,1339,1365]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
|
||
<emphasis id="68FD89349B1EA407BD3F8846FD5DFAF9" box="[581,706,1339,1365]" italics="true" pageId="12" pageNumber="579">A. sergeevi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="68FD89349B1EA407BDB08846FC94FAF9" bold="true" box="[714,779,1339,1365]" pageId="12" pageNumber="579">
|
||
<taxonomicNameLabel id="73CE344F9B1EA407BDB08846FC94FAF9" box="[714,779,1339,1365]" pageId="12" pageNumber="579" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is much rarer than
|
||
<taxonomicName id="9D892EA59B1EA407BC918846FB1BFAF9" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[1003,1156,1339,1365]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sayanensis">
|
||
<emphasis id="68FD89349B1EA407BC918846FB1BFAF9" box="[1003,1156,1339,1365]" italics="true" pageId="12" pageNumber="579">A. sayanensis</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="8532EB849B1EA407BBBD8846FAB6FAF9" box="[1223,1321,1339,1365]" pageId="12" pageNumber="579" type="holotype">holotype</typeStatus>
|
||
male was collected together with
|
||
<specimenCount id="4C8F9EAF9B1EA407BEE58822FDA8FAD5" box="[415,567,1375,1401]" pageId="12" pageNumber="579" type="generic">30 specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="9D892EA59B1EA407BD268822FD6BFAD5" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[604,756,1375,1401]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sayanensis">
|
||
<emphasis id="68FD89349B1EA407BD268822FD6BFAD5" box="[604,756,1375,1401]" italics="true" pageId="12" pageNumber="579">A. sayanensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |