treatments-xml/data/C5/65/7B/C5657B52FE9FB21744D1F8F3CCB80F32.xml
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<mods:title id="C5689415FFBF52BCB7DE77EF188192FF">Decapod Crustacea of the Californian and Oregonian Zoogeographic Provinces 3371</mods:title>
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<paragraph id="4D73CA44FE9FB21044D1F8F3CAA90C78" blockId="259.[151,687,1850,1909]" box="[151,687,1850,1876]" pageId="259" pageNumber="260">
<heading id="163B7D28FE9FB21044D1F8F3CAA90C78" bold="true" box="[151,687,1850,1876]" fontSize="11" level="1" pageId="259" pageNumber="260" reason="1">
<taxonomicName id="8ACCB1C7FE9FB21044D1F8F3CAA90C78" ID-CoL="7DDQV" authority="crenulata (Lockington, 1877)" authorityName="crenulata (Lockington" authorityYear="1877" baseAuthorityName="Lockington" baseAuthorityYear="1877" box="[151,687,1850,1876]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="259" pageNumber="260" phylum="Arthropoda" rank="subSpecies" species="crenulata" subSpecies="crenulata">
<emphasis id="7FB81656FE9FB21044D1F8F3CAA90C78" bold="true" box="[151,687,1850,1876]" pageId="259" pageNumber="260">
<emphasis id="7FB81656FE9FB21044D1F8F3C9BA0C78" bold="true" box="[151,444,1850,1876]" italics="true" pageId="259" pageNumber="260">Uca crenulata crenulata</emphasis>
(Lockington, 1877)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="05D699CFFE9FB21044D1F894C9690C59" box="[151,367,1885,1909]" pageId="259" pageNumber="260" type="description">
<paragraph id="4D73CA44FE9FB21044D1F894C9690C59" blockId="259.[151,687,1850,1909]" box="[151,367,1885,1909]" pageId="259" pageNumber="260">
(
<figureCitation id="D5F7D6C1FE9FB21044D9F894C9020C59" box="[159,260,1885,1909]" captionStart="FIGURE 60" captionStartId="258.[151,250,1838,1861]" captionTargetBox="[154,1411,198,1816]" captionTargetId="figure-112@258.[151,1436,195,1816]" captionTargetPageId="258" captionText="FIGURE 60. Families Pinnotheridae, Ocypodidae, Palicidae, Grapsidae. A, B, Pinnotheres pugettensis Holmes, 1900; A, entire crab; B, chela. C, Uca crenulata crenulata (Lockington, 1877). D, Palicus cortezi (Crane, 1937). E, Palicus lucasii Rathbun, 1898. F, Planes major (MacLeay, 1838). G, H, Planes marinus Rathbun, 1915; G, chela; H, crab in dorsal view. Scales: A, C = 5 mm, DF, H = 10 m. A, B, G, H from Hart 1982, C drawn from crab from Santa Catalina I., D from Montagne &amp; Cadien 2001, E after Rathbun 1917, F drawn from crab from western Panama." pageId="259" pageNumber="260">Fig. 60C</figureCitation>
, Pl. 15G)
</paragraph>
</subSubSection>
<subSubSection id="05D699CFFE9FB21744D1F86CC9B40A25" lastPageId="260" lastPageNumber="261" pageId="259" pageNumber="260" type="reference_group">
<paragraph id="4D73CA44FE9FB21044D1F86CCA610C97" blockId="259.[151,1436,1957,2010]" box="[151,615,1957,1979]" pageId="259" pageNumber="260">
<taxonomicName id="8ACCB1C7FE9FB21044D1F86CCA650C97" ID-CoL="6JYKT" authority="Lockington, 1877: 149" authorityName="Lockington" authorityPageNumber="149" authorityYear="1877" box="[151,611,1957,1979]" class="Malacostraca" family="Ocypodidae" genus="Gelasimus" kingdom="Animalia" order="Decapoda" pageId="259" pageNumber="260" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="7FB81656FE9FB21044D1F86CC9740C97" box="[151,370,1957,1979]" italics="true" pageId="259" pageNumber="260">Gelasimus crenulatus</emphasis>
Lockington, 1877: 149
</taxonomicName>
.
</paragraph>
<paragraph id="4D73CA44FE9FB21744D1F80DC9B90BE0" blockId="259.[151,1436,1957,2010]" lastBlockId="260.[151,1436,152,266]" lastPageId="260" lastPageNumber="261" pageId="259" pageNumber="260">
<treatmentCitationGroup id="6DDCED6AFE9FB21744D1F80DC9B90BE0" lastPageId="260" lastPageNumber="261" pageId="259" pageNumber="260">
<taxonomicName id="8ACCB1C7FE9FB21044D1F80DC9210CF6" ID-CoL="7DDQV" baseAuthorityName="Lockington" baseAuthorityYear="1876" box="[151,295,1988,2010]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="259" pageNumber="260" phylum="Arthropoda" rank="species" species="crenulata">
<emphasis id="7FB81656FE9FB21044D1F80DC9210CF6" box="[151,295,1988,2010]" italics="true" pageId="259" pageNumber="260">Uca crenulata</emphasis>
</taxonomicName>
. —
<treatmentCitation id="CC6DEC55FE9FB2104515F80ACA050CF5" author="Holmes, S. J." box="[339,515,1987,2009]" page="75" pageId="259" pageNumber="260" year="1900">
<bibRefCitation id="295DB7B5FE9FB2104515F80ACA050CF5" author="Holmes, S. J." box="[339,515,1987,2009]" pageId="259" pageNumber="260" pagination="12 - 62" refId="ref226245" refString="Holmes, S. J. (1900) Synopsis of California stalk-eyed Crustacea. Occasional Papers of the California Academy of Sciences, 7, 12 - 62." type="journal article" year="1900">Holmes 1900: 75</bibRefCitation>
</treatmentCitation>
, pl. 1, figs. 79. —
<treatmentCitation id="CC6DEC55FE9FB210468AF80ACB960CF5" author="Rathbun, M. J." box="[716,912,1987,2009]" page="190" pageId="259" pageNumber="260" year="1904">
<bibRefCitation id="295DB7B5FE9FB210468AF80ACB960CF5" author="Rathbun, M. J." box="[716,912,1987,2009]" pageId="259" pageNumber="260" pagination="1 - 219" refId="ref234490" refString="Rathbun, M. J. (1904) Decapod crustaceans of the northwest coast of North America. Harriman Alaska Expedition, 10, 1 - 219." type="journal article" year="1904">Rathbun 1904: 190</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="CC6DEC55FE9FB21047DDF80ACC050CF5" author="Rathbun, M. J." box="[923,1027,1987,2009]" page="409" pageId="259" pageNumber="260" year="1917">
<bibRefCitation id="295DB7B5FE9FB21047DDF80ACC050CF5" author="Rathbun, M. J." box="[923,1027,1987,2009]" pageId="259" pageNumber="260" pagination="1 - 445" refId="ref234686" refString="Rathbun, M. J. (1917) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, 1 - 445." type="journal article" year="1917">1917: 409</bibRefCitation>
</treatmentCitation>
, pl. 146. —
<treatmentCitation id="CC6DEC55FE9FB2104039F80ACD3A0CF5" author="Schmitt, W. L." box="[1151,1340,1987,2009]" page="279" pageId="259" pageNumber="260" year="1921">
<bibRefCitation id="295DB7B5FE9FB2104039F80ACD3A0CF5" author="Schmitt, W. L." box="[1151,1340,1987,2009]" pageId="259" pageNumber="260" pagination="1 - 470" refId="ref236092" refString="Schmitt, W. L. (1921) The marine decapod Crustacea of California. University of California Publications in Zoology, 23, 1 - 470." type="journal article" year="1921">Schmitt 1921: 279</bibRefCitation>
</treatmentCitation>
, fig. 164. —
<treatmentCitation id="CC6DEC55FE98B21744A1FF51CA0A0B82" author="Johnson, M. E. &amp; Snook, H. J." box="[231,524,152,174]" page="399" pageId="260" pageNumber="261" year="1927">
<bibRefCitation id="295DB7B5FE98B21744A1FF51CA0A0B82" author="Johnson, M. E. &amp; Snook, H. J." box="[231,524,152,174]" pageId="260" pageNumber="261" refId="ref227400" refString="Johnson, M. E. &amp; Snook, H. J. (1927) Seashore Animals of the Pacific Coast. Dover Publications, New York, reprint 1967, 659 pp." type="book" year="1927">Johnson &amp; Snook 1927: 399</bibRefCitation>
</treatmentCitation>
, figs. 357, 358. —
<treatmentCitation id="CC6DEC55FE98B2174694FF51CBC80B82" author="Ricketts, E. F. &amp; Calvin, J. &amp; Hedgpeth, J. W. &amp; Phillips, D. W." box="[722,974,152,174]" page="354" pageId="260" pageNumber="261" year="1985">
<bibRefCitation id="295DB7B5FE98B2174694FF51CBC80B82" author="Ricketts, E. F. &amp; Calvin, J. &amp; Hedgpeth, J. W. &amp; Phillips, D. W." box="[722,974,152,174]" pageId="260" pageNumber="261" refId="ref234966" refString="Ricketts, E. F., Calvin, J., Hedgpeth, J. W. &amp; Phillips, D. W. (1985) Between Pacific Tides. Stanford University Press, Stanford, California, 5 th ed., 652 pp." type="book" year="1985">
Ricketts
<emphasis id="7FB81656FE98B217476DFF50CB5E0B82" box="[811,856,152,174]" italics="true" pageId="260" pageNumber="261">et al</emphasis>
. 1985: 354
</bibRefCitation>
</treatmentCitation>
, fig. 273. —
<treatmentCitation id="CC6DEC55FE98B2174011FF51CD7E0B82" author="Hubbard, D. M. &amp; Dugan, J. E." box="[1111,1400,152,174]" page="55" pageId="260" pageNumber="261" year="1989">
<bibRefCitation id="295DB7B5FE98B2174011FF51CD7E0B82" author="Hubbard, D. M. &amp; Dugan, J. E." box="[1111,1400,152,174]" pageId="260" pageNumber="261" pagination="55 - 57" refId="ref226856" refString="Hubbard, D. M. &amp; Dugan, J. E. (1989) Northern occurrence of two estuarine crabs: the fiddler crab, Uca crenulata, and the burrowing crab, Malacoplax californiensis. California Fish and Game, 75, 55 - 57." type="journal article" year="1989">Hubbard &amp; Dugan 1989: 55</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="CC6DEC55FE98B2174481FF7EC96B0BE0" author="Jensen, G. C." box="[199,365,182,204]" page="34" pageId="260" pageNumber="261" year="1995">
<bibRefCitation id="295DB7B5FE98B2174481FF7EC96B0BE0" author="Jensen, G. C." box="[199,365,182,204]" pageId="260" pageNumber="261" refId="ref227177" refString="Jensen, G. C. (1995) Pacific Coast Crabs and Shrimps. Sea Challengers, Monterey, California, 87 pp." type="book" year="1995">Jensen 1995: 34</bibRefCitation>
</treatmentCitation>
, fig. 51.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4D73CA44FE98B21744D1FF1FC9B40A25" blockId="260.[151,1436,152,266]" pageId="260" pageNumber="261">
<treatmentCitationGroup id="6DDCED6AFE98B21744D1FF1FC9B40A25" pageId="260" pageNumber="261">
<taxonomicName id="8ACCB1C7FE98B21744D1FF1FC98B0BC7" ID-CoL="7DDQV" authority="crenulata" authorityName="crenulata (Lockington" authorityYear="1877" box="[151,397,213,235]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="260" pageNumber="261" phylum="Arthropoda" rank="subSpecies" species="crenulata" subSpecies="crenulata">
<emphasis id="7FB81656FE98B21744D1FF1FC98B0BC7" box="[151,397,213,235]" italics="true" pageId="260" pageNumber="261">Uca crenulata crenulata</emphasis>
</taxonomicName>
. —
<treatmentCitation id="CC6DEC55FE98B21745FFFF1CCA6F0BC7" author="Crane, J." box="[441,617,213,235]" page="232" pageId="260" pageNumber="261" year="1975">
<bibRefCitation id="295DB7B5FE98B21745FFFF1CCA6F0BC7" author="Crane, J." box="[441,617,213,235]" pageId="260" pageNumber="261" refId="ref220205" refString="Crane, J. (1975) Fiddler Crabs of the World Ocypodidae: Genus Uca. Princeton University Press, Princeton, New Jersey, 736 pp." type="book" year="1975">Crane, 1975: 232</bibRefCitation>
</treatmentCitation>
, pl. 30 E-I, figs. 70D, G, 101. —
<treatmentCitation id="CC6DEC55FE98B21747F1FF1CCCC20BC7" author="Garth, J. S. &amp; Abbott, D." box="[951,1220,213,235]" page="622" pageId="260" pageNumber="261" year="1980">
<bibRefCitation id="295DB7B5FE98B21747F1FF1CCCC20BC7" author="Garth, J. S. &amp; Abbott, D." box="[951,1220,213,235]" pageId="260" pageNumber="261" pagination="594 - 630" refId="ref222430" refString="Garth, J. S. &amp; Abbott, D. (1980) Brachyura: the true crabs. In: Morris, D., Abbott, D. &amp; Haderlie, E. (Eds). Intertidal Invertebrates of California. Stanford University Press, Stanford, California, pp. 594 - 630." type="book chapter" year="1980">Garth &amp; Abbott 1980: 622</bibRefCitation>
</treatmentCitation>
, fig. 25.46. —
<treatmentCitation id="CC6DEC55FE98B217411DFF1CC95D0A25" author="Bonfil, R. &amp; Carvacho, A. &amp; Campos, E." page="50" pageId="260" pageNumber="261" year="1992">
<bibRefCitation id="295DB7B5FE98B217411DFF1CC95D0A25" author="Bonfil, R. &amp; Carvacho, A. &amp; Campos, E." pageId="260" pageNumber="261" pagination="37 - 56" refId="ref216822" refString="Bonfil, R., Carvacho, A. &amp; Campos, E. (1992) Los cangrejos de la Bahia de Todos Santos, Baja California. Parte II. Grapsidae, Pinnotheridae y Ocypodidae (Crustacea: Decapoda: Brachyura). Ciencias Marinas (Mexico), 18, 37 - 56." type="journal article" year="1992">
Bonfil
<emphasis id="7FB81656FE98B2174481FF3DC8F50A26" box="[199,243,244,266]" italics="true" pageId="260" pageNumber="261">et al</emphasis>
. 1992: 50
</bibRefCitation>
</treatmentCitation>
, fig. 5B.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="05D699CFFE98B21744D1FEFFC96F08A2" pageId="260" pageNumber="261" type="diagnosis">
<paragraph id="4D73CA44FE98B21744D1FEFFC95F0966" blockId="260.[151,1436,310,1054]" pageId="260" pageNumber="261">
<emphasis id="7FB81656FE98B21744D1FEFFCA650A62" box="[151,611,310,335]" inLineHeading="true" pageId="260" pageNumber="261" reason="1">
<emphasis id="7FB81656FE98B21744D1FEFFC90C0A63" bold="true" box="[151,266,310,335]" pageId="260" pageNumber="261">Diagnosis</emphasis>
(modified from
<bibRefCitation id="295DB7B5FE98B217458BFEFFCA500A62" author="Crane, J." box="[461,598,310,334]" pageId="260" pageNumber="261" refId="ref220205" refString="Crane, J. (1975) Fiddler Crabs of the World Ocypodidae: Genus Uca. Princeton University Press, Princeton, New Jersey, 736 pp." type="book" year="1975">Crane 1975</bibRefCitation>
).
</emphasis>
Frontal region narrow, its width less than 0.3 times width of carapace. Orbits nearly straight, suborbital margins with rounded tubercles. Carapace smooth, moderately convex; lateral margins nearly parallel behind acute anterolateral angles, then converging.
<collectingRegion id="8F0804A6FE98B21747A4FEB7CC1D0ABA" box="[994,1051,382,406]" country="Maldives" name="Maale" pageId="260" pageNumber="261">Male</collectingRegion>
major cheliped with outer surface finely granulated; inner surface with oblique tuberculate ridge running vertically down from dorsal surface, then bending obliquely to run somewhat parallel to gape; two rows of denticles at base of fixed finger. Fingers of old male cheliped longer than those of younger male. Smaller chela of both sexes with fingers equal, gape narrow. Pereopods 25 with pile, merus transversely granulate. Abdomen of male with 7 segments. Carapace length to
<quantity id="8A3467A1FE98B21741C6FDC7C8C40966" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="260" pageNumber="261" unit="mm" value="13.0">13 mm</quantity>
in either sex.
</paragraph>
<paragraph id="4D73CA44FE98B2174481FD9FCCE309F6" blockId="260.[151,1436,310,1054]" pageId="260" pageNumber="261">
<emphasis id="7FB81656FE98B2174481FD9FC9670943" bold="true" box="[199,353,598,623]" inLineHeading="true" pageId="260" pageNumber="261" reason="1">Color in life.</emphasis>
Carapace grayish white to pale brown. Major chela of male with merus bright red on exterior surface, inner surface of merus, fingers of chela bright white. Palm, inner part of chela yellowish to white. Minor chela pinkish. Merus of pereopod 2 crimson red; otherwise cream-colored; other pereopods marked with cream. Female chelae brownish to cream-colored. See
<bibRefCitation id="295DB7B5FE98B21746ECFD0BCB3809F6" author="Crane, J." box="[682,830,706,730]" pageId="260" pageNumber="261" refId="ref220205" refString="Crane, J. (1975) Fiddler Crabs of the World Ocypodidae: Genus Uca. Princeton University Press, Princeton, New Jersey, 736 pp." type="book" year="1975">Crane (1975)</bibRefCitation>
for a more detailed color description.
</paragraph>
<paragraph id="4D73CA44FE98B2174481FD2FCC3E09D2" blockId="260.[151,1436,310,1054]" box="[199,1080,742,767]" pageId="260" pageNumber="261">
<emphasis id="7FB81656FE98B2174481FD2FC9AF09D3" bold="true" box="[199,425,742,767]" inLineHeading="true" pageId="260" pageNumber="261" reason="1">Habitat and depth.</emphasis>
Upper parts of mud flats and salt marshes, intertidal zone.
</paragraph>
<paragraph id="4D73CA44FE98B2174481FCC2C96F08A2" blockId="260.[151,1436,310,1054]" pageId="260" pageNumber="261">
<emphasis id="7FB81656FE98B2174481FCC2C91C080F" bold="true" box="[199,282,779,803]" inLineHeading="true" pageId="260" pageNumber="261" reason="1">Range.</emphasis>
Discontinuous, from Goleta, California to Turtle Bay,
<collectingRegion id="8F0804A6FE98B21747D8FCC3CC56080E" box="[926,1104,778,802]" country="Mexico" name="Baja California" pageId="260" pageNumber="261">Baja California</collectingRegion>
, including Santa
<taxonomicName id="8ACCB1C7FE98B217415BFCC3CD90080E" authority="I." box="[1309,1430,778,802]" class="Insecta" family="Staphylinidae" genus="Catalina" kingdom="Animalia" order="Coleoptera" pageId="260" pageNumber="221" phylum="Arthropoda" rank="genus">Catalina I.</taxonomicName>
, California; and San Felipe to La Paz and Guaymas to Tenacatita Bay, western
<collectingCountry id="35DB8AD4FE98B217404EFCE7CC67086A" box="[1032,1121,814,838]" name="Mexico" pageId="260" pageNumber="261">Mexico</collectingCountry>
.
<typeStatus id="927774E6FE98B217402AFCE6CCA2086B" box="[1132,1188,815,839]" pageId="260" pageNumber="261">Type</typeStatus>
locality Todos Santos Bay,
<collectingRegion id="8F0804A6FE98B2174495FC9BC9830846" box="[211,389,850,874]" country="Mexico" name="Baja California" pageId="260" pageNumber="261">Baja California</collectingRegion>
. A distinct subspecies,
<taxonomicName id="8ACCB1C7FE98B21746D3FC9ACC850846" authority="(Rathbun, 1893)" baseAuthorityName="Rathbun" baseAuthorityYear="1893" box="[661,1155,850,875]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="260" pageNumber="261" phylum="Arthropoda" rank="subSpecies" species="crenulata" subSpecies="coloradensis">
<emphasis id="7FB81656FE98B21746D3FC9ACAAF0846" box="[661,681,851,874]" italics="true" pageId="260" pageNumber="261">U</emphasis>
.
<emphasis id="7FB81656FE98B21746FCFC9ACBBB0847" box="[698,957,851,875]" italics="true" pageId="260" pageNumber="261">crenulata coloradensis</emphasis>
(
<bibRefCitation id="295DB7B5FE98B2174788FC9BCC7C0846" author="Rathbun, M. J." box="[974,1146,850,874]" pageId="260" pageNumber="261" pagination="223 - 260" refId="ref234064" refString="Rathbun, M. J. (1893) Descriptions of new genera and species of crabs from the west coast of America and the Sandwich Islands. Proceedings of the United States National Museum, 16, 223 - 260." type="journal article" year="1893">Rathbun, 1893</bibRefCitation>
)
</taxonomicName>
, lives at the head of the Gulf of California.
</paragraph>
</subSubSection>
<subSubSection id="05D699CFFE98B2174481FC52CCB80F32" pageId="260" pageNumber="261" type="discussion">
<paragraph id="4D73CA44FE98B2174481FC52CCB80F32" blockId="260.[151,1436,310,1054]" pageId="260" pageNumber="261">
<emphasis id="7FB81656FE98B2174481FC52C93D089F" bold="true" box="[199,315,923,947]" inLineHeading="true" pageId="260" pageNumber="261" reason="1">Remarks.</emphasis>
Fiddler crabs dig burrows in the upper reaches of mud flats and salt marshes. They manipulate balls of sediment in their mouthparts as they extract detritus and other food material. The burrows, mud balls and tracks are good indication of their presence even if the crabs themselves are not visible. Fiddler crabs today live in
<collectingRegion id="8F0804A6FE98B21744D1FBCFC90E0F32" box="[151,264,1030,1054]" country="United States of America" name="California" pageId="260" pageNumber="261">California</collectingRegion>
mostly in wildlife refuges and other areas protected from human habitat destruction.
</paragraph>
</subSubSection>
</treatment>
</document>