546 lines
72 KiB
XML
546 lines
72 KiB
XML
<document id="CEE07F6C5D1A9B26ADDC41D32CDD0CFE" ID-DOI="10.37520/aemnp.2023.014" ID-ISSN="1804-6487" ID-Zenodo-Dep="10134093" ID-ZooBank="D982BDAC-CB44-41BE-A35C-92D630F96BF7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699897763845" checkinUser="felipe" docAuthor="Bilton, David T." docDate="2023" docId="03F6746EFFD8B626FC93FC1EFAE7B111" docLanguage="en" docName="ActaEntMusNatPra.63.2.249-264.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 63 (2)" docSource="http://dx.doi.org/10.37520/aemnp.2023.014" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Helophorus (Ropalohelophorus) aethiopicus Regimbart 1907" docType="treatment" docVersion="2" lastPageNumber="253" masterDocId="FFCF0C16FFD9B622FFE4FFB6FFE2B204" masterDocTitle="Out of the Palaearctic: the Helophorus water beetles of the Afrotropical Region (Coleoptera: Helophoridae)" masterLastPageNumber="264" masterPageNumber="249" pageNumber="250" updateTime="1700044289635" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="BEE65650118D9E94EFA3179E1930D96F">Out of the Palaearctic: the Helophorus water beetles of the Afrotropical Region (Coleoptera: Helophoridae)</mods:title>
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<mods:namePart id="E1356F7F947AAF71028294D49BBE769C">Bilton, David T.</mods:namePart>
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<mods:affiliation id="CF7948764336DF19F35B9356027A5A81">Marine Biology and Ecology Research Centre, School of Biological and Marine Sciences, University of Plymouth, Drake Circus Plymouth PL & Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, 2006 Johannesburg, South Africa</mods:affiliation>
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<mods:title id="DEA82C20DCDF9ED73D29C035EFC5E349">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="0FEA6439862A66AA840A3F9D9285D23B">2023</mods:date>
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<mods:title id="2822F4E01F7D64C7BA26C4D54B9A3B93">Acta. Ent. Mus. Natl. Pragae</mods:title>
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<mods:number id="83134F2F72A2352E1216AA82A8D2A936">2023-09-10</mods:number>
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<mods:url id="FBF429C7836BED246148B592260C2097">http://dx.doi.org/10.37520/aemnp.2023.014</mods:url>
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<subSubSection id="C34596F3FFD8B623FC93FC1EFB28B1E4" pageId="1" pageNumber="250" type="nomenclature">
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<paragraph id="8BE0C578FFD8B623FC93FC1EFB28B1E4" blockId="1.[887,1377,936,992]" pageId="1" pageNumber="250">
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<heading id="D0A87214FFD8B623FC93FC1EFB28B1E4" centered="true" fontSize="10" level="2" pageId="1" pageNumber="250" reason="3">
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<taxonomicName id="4C5FBEFBFFD8B623FC93FC1EFB28B1E4" ID-CoL="6LMZX" authority="Regimbart, 1907" authorityName="Regimbart" authorityYear="1907" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopicus" subGenus="Ropalohelophorus">
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<emphasis id="B92B196AFFD8B623FC93FC1EFB28B1E4" bold="true" pageId="1" pageNumber="250">
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<emphasis id="B92B196AFFD8B623FC93FC1EFC1BB1C4" bold="true" box="[887,1017,936,960]" italics="true" pageId="1" pageNumber="250">Helophorus</emphasis>
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(
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<emphasis id="B92B196AFFD8B623FBEDFC1EFB36B1C4" bold="true" box="[1033,1236,936,960]" italics="true" pageId="1" pageNumber="250">Ropalohelophorus</emphasis>
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)
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<emphasis id="B92B196AFFD8B623FB07FC1EFA83B1C4" bold="true" box="[1251,1377,936,960]" italics="true" pageId="1" pageNumber="250">aethiopicus</emphasis>
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<bibRefCitation id="EFCEB889FFD8B623FBE9FC7FFB28B1E4" author="REGIMBART M." box="[1037,1226,969,992]" pageId="1" pageNumber="250" pagination="126 - 128" refId="ref16606" refString="REGIMBART M. 1907: Collections recueillis par M. Maurice de Rothschild dans l'Afrique orientale. Insectes coleopteres: Dytiscides et hydrophilides. Bulletin du Museum National d'Histoire Naturelle Paris 13: 126 - 128." type="journal article" year="1907">Régimbart, 1907</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34596F3FFD8B623FBE3FC51FB33B1FE" box="[1031,1233,999,1018]" pageId="1" pageNumber="250" type="description">
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<paragraph id="8BE0C578FFD8B623FBE3FC51FB33B1FE" blockId="1.[828,1233,999,1052]" box="[1031,1233,999,1018]" pageId="1" pageNumber="250">
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(
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<figureCitation id="1364D9FDFFD8B623FBE9FC51FBB1B1FE" box="[1037,1107,999,1018]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="1" pageNumber="250">Figs 1A</figureCitation>
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,
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<figureCitation id="1364D9FDFFD8B623FBB9FC51FB70B1FE" box="[1117,1170,999,1018]" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="1" pageNumber="250">2A–B</figureCitation>
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,
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<figureCitation id="1364D9FDFFD8B623FB7EFC51FB55B1FE" box="[1178,1207,999,1018]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="1" pageNumber="250">3A</figureCitation>
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,
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<figureCitation id="1364D9FDFFD8B623FB5BFC51FB28B1FE" box="[1215,1226,999,1018]" captionStart="Fig" captionStartId="14.[1132,1165,1128,1147]" captionTargetBox="[152,1098,191,1200]" captionTargetId="figure-437@14.[152,1098,191,1200]" captionTargetPageId="14" captionText="Fig. 5. Afrotropical Helophorus, known distributions.Map by https:// mapswire.com." figureDoi="http://doi.org/10.5281/zenodo.10134103" httpUri="https://zenodo.org/record/10134103/files/figure.png" pageId="1" pageNumber="250">5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34596F3FFD8B623FCD8FBBFFB2DB618" box="[828,1231,1033,1052]" pageId="1" pageNumber="250" type="reference_group">
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<paragraph id="8BE0C578FFD8B623FCD8FBBFFB2DB618" blockId="1.[828,1233,999,1052]" box="[828,1231,1033,1052]" pageId="1" pageNumber="250">
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<treatmentCitationGroup id="AB4FE256FFD8B623FCD8FBBFFB2DB618" box="[828,1231,1033,1052]" pageId="1" pageNumber="250">
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<treatmentCitation id="0AFEE369FFD8B623FCD8FBBFFB28B618" author="REGIMBART M." box="[828,1226,1033,1052]" page="127" pageId="1" pageNumber="250" year="1907">
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<taxonomicName id="4C5FBEFBFFD8B623FCD8FBBFFB28B618" ID-CoL="6LMZX" authority="Regimbart, 1907: 127" authorityName="Regimbart" authorityPageNumber="127" authorityYear="1907" box="[828,1226,1033,1052]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopicus">
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<emphasis id="B92B196AFFD8B623FCD8FBBFFBEAB618" box="[828,1032,1033,1052]" italics="true" pageId="1" pageNumber="250">Helophorus aethiopicus</emphasis>
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<bibRefCitation id="EFCEB889FFD8B623FBE9FBBFFB28B618" author="REGIMBART M." box="[1037,1226,1033,1052]" pageId="1" pageNumber="250" pagination="126 - 128" refId="ref16606" refString="REGIMBART M. 1907: Collections recueillis par M. Maurice de Rothschild dans l'Afrique orientale. Insectes coleopteres: Dytiscides et hydrophilides. Bulletin du Museum National d'Histoire Naturelle Paris 13: 126 - 128." type="journal article" year="1907">Régimbart, 1907: 127</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34596F3FFD8B620FCD8FB85FE1FB78F" lastPageId="2" lastPageNumber="251" pageId="1" pageNumber="250" type="materials_examined">
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<paragraph id="8BE0C578FFD8B623FCD8FB85FB96B692" blockId="1.[828,1437,1075,2081]" pageId="1" pageNumber="250">
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<emphasis id="B92B196AFFD8B623FCD8FB85FC56B642" bold="true" box="[828,948,1075,1094]" pageId="1" pageNumber="250">
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<typeStatus id="54E47BDAFFD8B623FCD8FB85FC8BB642" box="[828,873,1075,1094]" pageId="1" pageNumber="250">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F34885E8FFD8B623FC5CFB85FBE1B642" box="[952,1027,1075,1094]" name="Ethiopia" pageId="1" pageNumber="250">Ethiopia</collectingCountry>
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,
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<collectingRegion id="499B0B9AFFD8B623FBEEFB85FB78B642" box="[1034,1178,1075,1094]" country="Ethiopia" name="Oromiya" pageId="1" pageNumber="250">Oromiya Region</collectingRegion>
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, Karsa (Kersa), approximately
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<geoCoordinate id="EE6BA3BFFFD8B623FCD8FBFBFC97B664" box="[828,885,1101,1120]" degrees="7" direction="south" minutes="45" orientation="latitude" pageId="1" pageNumber="250" precision="925" value="-7.75">7°45′S</geoCoordinate>
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<geoCoordinate id="EE6BA3BFFFD8B623FC9EFBFBFC23B664" box="[890,961,1101,1120]" degrees="37" direction="east" minutes="00" orientation="longitude" pageId="1" pageNumber="250" precision="925" value="37.0">37°00′E</geoCoordinate>
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,>
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<quantity id="4CA7689DFFD8B623FC3FFBFBFBC1B664" box="[987,1059,1101,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="1" pageNumber="250" unit="m" value="1740.0">1,740 m</quantity>
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. Exact details imprecise. Karsa (or Kersa) is a woreda (district) in the
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<collectingRegion id="499B0B9AFFD8B623FBF2FBDEFB44B67F" box="[1046,1190,1128,1147]" country="Ethiopia" name="Oromiya" pageId="1" pageNumber="250">Oromiya Region</collectingRegion>
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of southern
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<collectingCountry id="F34885E8FFD8B623FAF6FBDEFABFB67F" box="[1298,1373,1128,1147]" name="Ethiopia" pageId="1" pageNumber="250">Ethiopia</collectingCountry>
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, whose altitude ranges from
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<quantity id="4CA7689DFFD8B623FC09FB35FB92B692" box="[1005,1136,1155,1174]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.66" metricValueMin="1.7399999999999998" pageId="1" pageNumber="250" unit="m" value="2200.0" valueMax="2660.0" valueMin="1740.0">
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<geoCoordinate id="EE6BA3BFFFD8B623FC09FB35FBFEB692" box="[1005,1052,1155,1174]" degrees="1,740" direction="north" orientation="latitude" pageId="1" pageNumber="250" precision="55" value="1.74">1,740</geoCoordinate>
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<geoCoordinate id="EE6BA3BFFFD8B623FBF8FB35FBBAB692" box="[1052,1112,1155,1174]" degrees="2,660" direction="west" orientation="longitude" pageId="1" pageNumber="250" precision="55" value="-2.66">–2,660</geoCoordinate>
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m
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</quantity>
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.
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</paragraph>
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<paragraph id="8BE0C578FFD8B623FCD8FB2BFB72B512" blockId="1.[828,1437,1075,2081]" pageId="1" pageNumber="250">
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<emphasis id="B92B196AFFD8B623FCD8FB2BFC5EB6B4" bold="true" box="[828,956,1181,1200]" pageId="1" pageNumber="250">Type material.</emphasis>
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<materialsCitation id="3B37CF25FFD8B623FC5BFB2BFC6CB754" collectionCode="MNHN" collectorName="Hw." country="Ethiopia" determinerName="R. B. Angus" location="ETHIOPIE" pageId="1" pageNumber="250" specimenCount="4" typeStatus="lectotype">
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<typeStatus id="54E47BDAFFD8B623FC5BFB2BFBFFB6B4" box="[959,1053,1181,1200]" pageId="1" pageNumber="250" type="lectotype">LECTOTYPE</typeStatus>
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: J, labelled:“J” “
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<typeStatus id="54E47BDAFFD8B623FB52FB2BFAE0B6B4" box="[1206,1282,1181,1200]" pageId="1" pageNumber="250" type="lectotype">LECTO-</typeStatus>
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// TYPE” [circular label with purple margin] “MUSEUM PARIS //
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<collectingCountry id="F34885E8FFD8B623FB3BFB0EFADDB6CF" box="[1247,1343,1208,1227]" name="Ethiopia" pageId="1" pageNumber="250">ÉTHIOPIE</collectingCountry>
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MÉRID // KARSSA // Maurice de Rothschild // 1905” “
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||
<taxonomicName id="4C5FBEFBFFD8B623FB5BFB65FAC7B6E2" authorityName="Fabricius" authorityYear="1775" box="[1215,1317,1235,1254]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="genus">Helophorus</taxonomicName>
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||
//
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||
<taxonomicName id="4C5FBEFBFFD8B623FADFFB65FC87B704" authority="Reg." authorityName="Regimbart" authorityYear="1907" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopicus">aethiopicus Rég.</taxonomicName>
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||
typus” [Hw.] “
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||
<typeStatus id="54E47BDAFFD8B623FC02FB5BFB80B704" box="[998,1122,1261,1280]" pageId="1" pageNumber="250" type="lectotype">LECTOTYPE</typeStatus>
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||
1974 //
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||
<taxonomicName id="4C5FBEFBFFD8B623FB48FB5BFABAB704" authority="Reg." authorityName="Regimbart" authorityYear="1907" box="[1196,1368,1261,1280]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopicus">H. aethiopicus Rég.</taxonomicName>
|
||
//
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||
<determinerName id="7E3C29BCFFD8B623FA8BFB5BFC96B71F" pageId="1" pageNumber="250">R. B. Angus</determinerName>
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||
det.” [
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||
<typeStatus id="54E47BDAFFD8B623FC4AFABEFBCEB71F" box="[942,1068,1288,1307]" pageId="1" pageNumber="250" type="lectotype">LECTOTYPE</typeStatus>
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||
, 74 and name and describer
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||
<collectorName id="26AAA0AEFFD8B623FAC1FABEFAA9B71F" box="[1317,1355,1288,1307]" pageId="1" pageNumber="250">Hw.</collectorName>
|
||
] “1974 //
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||
<taxonomicName id="4C5FBEFBFFD8B623FCD8FA95FC3EB732" authority="Sem." authorityName="Semenow" authorityYear="1900" box="[828,988,1315,1334]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="mervensis">H. mervensis Sem.</taxonomicName>
|
||
//
|
||
<determinerName id="7E3C29BCFFD8B623FC0BFA95FBB4B732" box="[1007,1110,1315,1334]" pageId="1" pageNumber="250">R. B. Angus</determinerName>
|
||
det.” [74 and name and describer
|
||
<collectorName id="26AAA0AEFFD8B623FA96FA95FA7AB732" box="[1394,1432,1315,1334]" pageId="1" pageNumber="250">Hw.</collectorName>
|
||
] (
|
||
<collectionCode id="ED4E5DBDFFD8B623FCA6FA8BFC6AB754" box="[834,904,1341,1360]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="250">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="54E47BDAFFD8B623FC71FA8BFBC6B754" box="[917,1060,1341,1360]" pageId="1" pageNumber="250" type="paralectotype">PARALECTOTYPES</typeStatus>
|
||
: 1J, “
|
||
<typeStatus id="54E47BDAFFD8B623FBBAFA8BFB7CB754" box="[1118,1182,1341,1360]" pageId="1" pageNumber="250" type="paratype">PARA-</typeStatus>
|
||
//
|
||
<typeStatus id="54E47BDAFFD8B623FB56FA8BFB1FB754" box="[1202,1277,1341,1360]" pageId="1" pageNumber="250" type="lectotype">LECTO-</typeStatus>
|
||
// TYPE” [circular label with blue margin] “MUSEUM PARIS //
|
||
<collectingCountry id="F34885E8FFD8B623FB3FFAEEFAD9B76F" box="[1243,1339,1368,1387]" name="Ethiopia" pageId="1" pageNumber="250">ÉTHIOPIE</collectingCountry>
|
||
MÉRID // KARSSA //
|
||
<collectingCountry id="F34885E8FFD8B623FC41FAC5FC0EB782" box="[933,1004,1395,1414]" name="Mauritius" pageId="1" pageNumber="250">Maurice</collectingCountry>
|
||
de Rothschild // 1905” “
|
||
<taxonomicName id="4C5FBEFBFFD8B623FB5BFAC5FAC7B782" authorityName="Fabricius" authorityYear="1775" box="[1215,1317,1395,1414]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="genus">Helophorus</taxonomicName>
|
||
//
|
||
<taxonomicName id="4C5FBEFBFFD8B623FADFFAC5FC87B7A4" authority="Reg." authorityName="Regimbart" authorityYear="1907" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopicus">aethiopicus Rég.</taxonomicName>
|
||
// n. sp. // Type - 1906” [Hw.] “
|
||
<typeStatus id="54E47BDAFFD8B623FB8EFA3BFAFFB7A4" box="[1130,1309,1421,1440]" pageId="1" pageNumber="250" type="paralectotype">PARALECTOTYPE</typeStatus>
|
||
<date id="FFE1E3B8FFD8B623FAC6FA3BFA91B7A4" box="[1314,1395,1421,1440]" pageId="1" pageNumber="250" value="1974-09">Sept1974</date>
|
||
//
|
||
<taxonomicName id="4C5FBEFBFFD8B623FA6CFA3BFC28B7BF" authority="Reg." authorityName="Regimbart" authorityYear="1907" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopicus">H. aethiopicus Rég.</taxonomicName>
|
||
// R. B. Angus det.” [
|
||
<typeStatus id="54E47BDAFFD8B623FB9FFA1EFB1BB7BF" box="[1147,1273,1448,1467]" pageId="1" pageNumber="250" type="lectotype">LECTOTYPE</typeStatus>
|
||
, Sept, 74 and name and describer Hw.] “1974 //
|
||
<taxonomicName id="4C5FBEFBFFD8B623FBC9FA75FB2DB7D2" authority="Sem." authorityName="Semenow" authorityYear="1900" box="[1069,1231,1475,1494]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="mervensis">H. mervensis Sem.</taxonomicName>
|
||
// R. B. Angus det.” [74 and name and describer Hw.] (
|
||
<collectionCode id="ED4E5DBDFFD8B623FBDAFA6BFB66B7F4" box="[1086,1156,1501,1520]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="250">MNHN</collectionCode>
|
||
);
|
||
<specimenCount id="9D590EF1FFD8B623FB76FA6BFB53B7F4" box="[1170,1201,1495,1520]" count="1" pageId="1" pageNumber="250" type="female">1 ♀</specimenCount>
|
||
, “
|
||
<specimenCount id="9D590EF1FFD8B623FB21FA61FB37B7F4" box="[1221,1237,1495,1520]" count="1" pageId="1" pageNumber="250" type="female">♀</specimenCount>
|
||
” “
|
||
<typeStatus id="54E47BDAFFD8B623FB0EFA6BFAC8B7F4" box="[1258,1322,1501,1520]" pageId="1" pageNumber="250" type="paratype">PARA-</typeStatus>
|
||
//
|
||
<typeStatus id="54E47BDAFFD8B623FAA4FA6BFA69B7F4" box="[1344,1419,1501,1520]" pageId="1" pageNumber="250" type="lectotype">LECTO-</typeStatus>
|
||
// TYPE” [circular label with blue margin] “MUSEUM PARIS //
|
||
<collectingCountry id="F34885E8FFD8B623FA81FA4EFC89B422" name="Ethiopia" pageId="1" pageNumber="250">ÉTHIOPIE</collectingCountry>
|
||
MÉRID // KARSSA //
|
||
<collectingCountry id="F34885E8FFD8B623FBDFF9A5FB60B422" box="[1083,1154,1555,1574]" name="Mauritius" pageId="1" pageNumber="250">Maurice</collectingCountry>
|
||
de Rothschild // 1905” “TYPE” [red label] “
|
||
<typeStatus id="54E47BDAFFD8B623FC47F99BFBB4B444" box="[931,1110,1581,1600]" pageId="1" pageNumber="250" type="paralectotype">PARALECTOTYPE</typeStatus>
|
||
<date id="FFE1E3B8FFD8B623FBB8F99BFB4FB444" box="[1116,1197,1581,1600]" pageId="1" pageNumber="250" value="1974-09">Sept1974</date>
|
||
//
|
||
<taxonomicName id="4C5FBEFBFFD8B623FB21F99BFA93B444" authority="Reg." authorityName="Regimbart" authorityYear="1907" box="[1221,1393,1581,1600]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopicus">H. aethiopicus Rég.</taxonomicName>
|
||
// R. B. Angus det.” [
|
||
<typeStatus id="54E47BDAFFD8B623FC23F9FEFBA7B45F" box="[967,1093,1608,1627]" pageId="1" pageNumber="250" type="lectotype">LECTOTYPE</typeStatus>
|
||
, Sept, 74 and name and describer Hw.] (
|
||
<collectionCode id="ED4E5DBDFFD8B623FCA6F9D5FC6AB472" box="[834,904,1635,1654]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="250">MNHN</collectionCode>
|
||
);
|
||
<specimenCount id="9D590EF1FFD8B623FC73F9D5FC5AB471" box="[919,952,1628,1654]" count="1" pageId="1" pageNumber="250" type="female">1 ♀</specimenCount>
|
||
, “
|
||
<typeStatus id="54E47BDAFFD8B623FC2FF9D5FBE9B472" box="[971,1035,1635,1654]" pageId="1" pageNumber="250" type="paratype">PARA-</typeStatus>
|
||
//
|
||
<typeStatus id="54E47BDAFFD8B623FBC0F9D5FB8DB472" box="[1060,1135,1635,1654]" pageId="1" pageNumber="250" type="lectotype">LECTO-</typeStatus>
|
||
// TYPE” [circular label with blue margin] “TYPE” [red printing] “MUSEUM PARIS //
|
||
<collectingCountry id="F34885E8FFD8B623FB12F9CBFAB7B494" box="[1270,1365,1661,1680]" name="Ethiopia" pageId="1" pageNumber="250">ÉTHIOPIE</collectingCountry>
|
||
MÉRID // KARSSA //
|
||
<collectingCountry id="F34885E8FFD8B623FC56F92EFC1BB4AF" box="[946,1017,1688,1707]" name="Mauritius" pageId="1" pageNumber="250">Maurice</collectingCountry>
|
||
de Rothschild // 1905” “
|
||
<taxonomicName id="4C5FBEFBFFD8B623FB27F92EFACAB4AF" authorityName="Fabricius" authorityYear="1775" box="[1219,1320,1688,1707]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="genus">Helophorus</taxonomicName>
|
||
// aethiopicus // Rég. n. sp. // Type // 1906” [Hw.] “
|
||
<typeStatus id="54E47BDAFFD8B623FB8FF905FAFCB4C2" box="[1131,1310,1715,1734]" pageId="1" pageNumber="250" type="paralectotype">PARALECTOTYPE</typeStatus>
|
||
<date id="FFE1E3B8FFD8B623FAC7F905FA91B4C2" box="[1315,1395,1715,1734]" pageId="1" pageNumber="250" value="1974-09">Sept1974</date>
|
||
//
|
||
<taxonomicName id="4C5FBEFBFFD8B623FA6CF905FC28B4E4" authority="Reg." authorityName="Regimbart" authorityYear="1907" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopicus">H. aethiopicus Rég.</taxonomicName>
|
||
// R. B. Angus det.” [
|
||
<typeStatus id="54E47BDAFFD8B623FB9FF97BFB1BB4E4" box="[1147,1273,1741,1760]" pageId="1" pageNumber="250" type="lectotype">LECTOTYPE</typeStatus>
|
||
, Sept, 74 and name and describer Hw.] “1974 //
|
||
<taxonomicName id="4C5FBEFBFFD8B623FBC9F95EFB2DB4FF" authority="Sem." authorityName="Semenow" authorityYear="1900" box="[1069,1231,1768,1787]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="mervensis">H. mervensis Sem.</taxonomicName>
|
||
// R. B. Angus det.” [74 and name and describer Hw.] (
|
||
<collectionCode id="ED4E5DBDFFD8B623FBA4F8B5FB64B512" box="[1088,1158,1795,1814]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="250">MNHN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFD8B620FCD8F8ABFE1FB78F" blockId="1.[828,1437,1075,2081]" lastBlockId="2.[151,760,1346,1419]" lastPageId="2" lastPageNumber="251" pageId="1" pageNumber="250">
|
||
<emphasis id="B92B196AFFD8B623FCD8F8ABFBC1B534" bold="true" box="[828,1059,1821,1840]" pageId="1" pageNumber="250">Other material examined.</emphasis>
|
||
1J, “
|
||
<collectingCountry id="F34885E8FFD8B623FBB1F8ABFB5EB534" box="[1109,1212,1821,1840]" name="Ethiopia" pageId="1" pageNumber="250">ETHIOPIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="499B0B9AFFD8B623FB26F8ABFAE1B534" box="[1218,1283,1821,1840]" country="Ethiopia" name="Oromiya" pageId="1" pageNumber="250">Oromia</collectingRegion>
|
||
// Bale Mountains NP //
|
||
<quantity id="4CA7689DFFD8B623FC94F88EFC7EB54F" box="[880,924,1848,1867]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="250" unit="km" value="5.0">5 km</quantity>
|
||
NW Dinsho // small lake” “
|
||
<geoCoordinate id="EE6BA3BFFFD8B623FB72F88EFAFBB54F" box="[1174,1305,1848,1867]" degrees="07" direction="north" minutes="07" orientation="latitude" pageId="1" pageNumber="250" precision="1" seconds="15.62" value="7.1210055">07°07′15.62′′N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE6BA3BFFFD8B623FAC4F88EFA7EB54F" box="[1312,1436,1848,1867]" degrees="39" direction="east" minutes="43" orientation="longitude" pageId="1" pageNumber="250" precision="1" seconds="10.52" value="39.71959">39°43′10.52′′E</geoCoordinate>
|
||
//
|
||
<quantity id="4CA7689DFFD8B623FCB4F8E5FC71B562" box="[848,915,1875,1894]" metricMagnitude="3" metricUnit="m" metricValue="3.057" pageId="1" pageNumber="250" unit="m" value="3057.0">3057 m</quantity>
|
||
a.s.l. //
|
||
<date id="FFE1E3B8FFD8B623FC3AF8E5FBD5B562" box="[990,1079,1875,1894]" pageId="1" pageNumber="250" value="2014-02-22">22.II.2014</date>
|
||
// leg. M.A. Jäch (12)ˮ (
|
||
<collectionCode id="ED4E5DBDFFD8B623FAFCF8E5FAB1B562" box="[1304,1363,1875,1894]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="250">NMW</collectionCode>
|
||
);
|
||
<specimenCount id="9D590EF1FFD8B623FA81F8E5FA75B561" box="[1381,1431,1868,1894]" count="2" pageId="1" pageNumber="250" type="female">2 ♀♀</specimenCount>
|
||
, “
|
||
<collectingCountry id="F34885E8FFD8B623FCA7F8DBFC4BB584" box="[835,937,1901,1920]" name="Ethiopia" pageId="1" pageNumber="250">ETHIOPIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="499B0B9AFFD8B623FC4BF8DBFC0DB584" box="[943,1007,1901,1920]" country="Ethiopia" name="Oromiya" pageId="1" pageNumber="250">Oromia</collectingRegion>
|
||
// Bale Mountains NP //
|
||
<quantity id="4CA7689DFFD8B623FB5CF8DBFB03B584" box="[1208,1249,1901,1920]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="250" unit="km" value="5.0">5 km</quantity>
|
||
NW Dinsho // stream” “
|
||
<geoCoordinate id="EE6BA3BFFFD8B623FCA0F83EFC24B59F" box="[836,966,1928,1947]" degrees="07" direction="north" minutes="07" orientation="latitude" pageId="1" pageNumber="250" precision="1" seconds="24.47" value="7.123464">07°07′24.47′′N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE6BA3BFFFD8B623FC29F83EFBA8B59F" box="[973,1098,1928,1947]" degrees="39" direction="east" minutes="44" orientation="longitude" pageId="1" pageNumber="250" precision="1" seconds="26.31" value="39.740643">39°44′26.31′′E</geoCoordinate>
|
||
//
|
||
<quantity id="4CA7689DFFD8B623FBBAF83EFB7CB59F" box="[1118,1182,1928,1947]" metricMagnitude="3" metricUnit="m" metricValue="3.0439999999999996" pageId="1" pageNumber="250" unit="m" value="3044.0">3044 m</quantity>
|
||
a.s.l. //
|
||
<date id="FFE1E3B8FFD8B623FB3AF83EFAD5B59F" box="[1246,1335,1928,1947]" pageId="1" pageNumber="250" value="2014-02-20">20.II.2014</date>
|
||
// leg. M.A. Jäch (12)ˮ (
|
||
<collectionCode id="ED4E5DBDFFD8B623FC46F815FC3FB5B2" box="[930,989,1955,1974]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="250">NMW</collectionCode>
|
||
); 1 J, “J” “
|
||
<collectingCountry id="F34885E8FFD8B623FBAFF815FB77B5B2" box="[1099,1173,1955,1974]" name="Ethiopia" pageId="1" pageNumber="250">Ethiopie</collectingCountry>
|
||
: Goba // R. de Meulenaere // 1934-1935” “R. Mus. Hist. Nat. // Belge. I.G 10.738” “A. d’Orchymont det. //
|
||
<taxonomicName id="4C5FBEFBFFD8B623FC97F86EFC35B5EF" authorityName="Fabricius" authorityYear="1775" box="[883,983,2008,2027]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="genus" sensu="stricto">Helophorus</taxonomicName>
|
||
<taxonomicNameLabel id="A218A411FFD8B623FC38F86EFBE8B5EF" box="[988,1034,2008,2027]" pageId="1" pageNumber="250" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
//
|
||
<taxonomicName id="4C5FBEFBFFD8B623FBC5F86EFB50B5EF" authority="Reg." authorityName="Regimbart" authorityYear="1907" box="[1057,1202,2008,2027]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopicus">aethiopicus Rég.</taxonomicName>
|
||
” [name and describer Hw.] (
|
||
<collectionCode id="ED4E5DBDFFD8B623FCA6F845FC67BA02" box="[834,901,2035,2054]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="1" pageNumber="250" type="Museum">RBINS</collectionCode>
|
||
); 9 exx., “
|
||
<collectingCountry id="F34885E8FFD8B623FC3AF845FBCABA02" box="[990,1064,2035,2054]" name="Ethiopia" pageId="1" pageNumber="250">Ethiopie</collectingCountry>
|
||
: Goba // R. de Meulenaere // 1934-1935” “R. Mus. Hist. Nat. // Belge. I.G 10.738” (
|
||
<collectionCode id="ED4E5DBDFFD8B623FB45F7BBFB06BA24" box="[1185,1252,2061,2080]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="1" pageNumber="250" type="Museum">RBINS</collectionCode>
|
||
); 2 JJ
|
||
<specimenCount id="9D590EF1FFD8B623FAC2F7BBFAA7BA24" box="[1318,1349,2055,2080]" count="1" pageId="1" pageNumber="250" type="female">1 ♀</specimenCount>
|
||
“
|
||
<collectingCountry id="F34885E8FFD8B623FAB6F7BBFA7EBA24" box="[1362,1436,2061,2080]" name="Ethiopia" pageId="1" pageNumber="250">Ethiopie</collectingCountry>
|
||
: Goba // R. de Meulenaere // 1934-1935” “
|
||
<collectingCountry id="F34885E8FFDBB620FDF9FAF4FD85B751" box="[541,615,1346,1365]" name="Ethiopia" pageId="2" pageNumber="251">Ethiopie</collectingCountry>
|
||
: Goba // R. de Meulenaere // 1934-1935” “R. Mus. Hist. Nat. // Belge. I.G 10.738” “Coll. I.R.Sc.N.B.” (
|
||
<collectionCode id="ED4E5DBDFFDBB620FEACFACEFE6BB78F" box="[328,393,1400,1419]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="2" pageNumber="251" type="Museum">RBINS</collectionCode>
|
||
) [blue label].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2095F0FFDBB620FF73FB12FC75B6E8" ID-DOI="http://doi.org/10.5281/zenodo.10134095" ID-Zenodo-Dep="10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="2" pageNumber="251" startId="2.[151,182,1188,1207]" targetBox="[167,1388,192,1167]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BE0C578FFDBB620FF73FB12FC75B6E8" blockId="2.[151,1438,1188,1260]" pageId="2" pageNumber="251">
|
||
Fig. 1. Afrotropical
|
||
<taxonomicName id="4C5FBEFBFFDBB620FEA0FB12FE4BB6B3" authorityName="Fabricius" authorityYear="1775" box="[324,425,1188,1207]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92B196AFFDBB620FEA0FB12FE4BB6B3" box="[324,425,1188,1207]" italics="true" pageId="2" pageNumber="251">Helophorus</emphasis>
|
||
</taxonomicName>
|
||
, habitus. A –
|
||
<taxonomicName id="4C5FBEFBFFDBB620FDC6FB12FCDAB6B3" authority="Regimbart, 1907" authorityName="Regimbart" authorityYear="1907" box="[546,824,1188,1207]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="aethiopicus">
|
||
<emphasis id="B92B196AFFDBB620FDC6FB12FD7DB6B3" box="[546,671,1188,1207]" italics="true" pageId="2" pageNumber="251">H. aethiopicus</emphasis>
|
||
<bibRefCitation id="EFCEB889FFDBB620FD41FB12FCDAB6B3" author="REGIMBART M." box="[677,824,1188,1207]" pageId="2" pageNumber="251" pagination="126 - 128" refId="ref16606" refString="REGIMBART M. 1907: Collections recueillis par M. Maurice de Rothschild dans l'Afrique orientale. Insectes coleopteres: Dytiscides et hydrophilides. Bulletin du Museum National d'Histoire Naturelle Paris 13: 126 - 128." type="journal article" year="1907">Régimbart, 1907</bibRefCitation>
|
||
</taxonomicName>
|
||
, lectotype; B –
|
||
<taxonomicName id="4C5FBEFBFFDBB620FC25FB12FAE0B6B3" authority="J. Balfour-Browne, 1954" authorityName="J. Balfour-Browne" authorityYear="1954" box="[961,1282,1188,1207]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="aethiops">
|
||
<emphasis id="B92B196AFFDBB620FC25FB12FBC6B6B3" box="[961,1060,1188,1207]" italics="true" pageId="2" pageNumber="251">H. aethiops</emphasis>
|
||
J.
|
||
<bibRefCitation id="EFCEB889FFDBB620FBDAFB12FAE0B6B3" author="BALFOUR-BROWNE J." box="[1086,1282,1188,1207]" pageId="2" pageNumber="251" pagination="96 - 114" refId="ref16244" refString="BALFOUR-BROWNE J. 1954: A preliminary contribution to the knowledge of the SouthAfrican Palpicornia (Coleoptera). (Professor J. Omer-Cooper's investigation of the inland waters of Africa.) Journal of the Entomolological Society of South Africa 17: 96 - 114." type="journal article" year="1954">Balfour-Browne, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype; C –
|
||
<taxonomicName id="4C5FBEFBFFDBB620FA63FB12FE58B6D5" authority="J. Balfour-Browne, 1954" authorityName="J. Balfour-Browne" authorityYear="1954" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="aethiops">
|
||
<emphasis id="B92B196AFFDBB620FA63FB12FF3DB6D5" italics="true" pageId="2" pageNumber="251">H. aethiops</emphasis>
|
||
J.
|
||
<bibRefCitation id="EFCEB889FFDBB620FF13FB08FE58B6D5" author="BALFOUR-BROWNE J." box="[247,442,1214,1233]" pageId="2" pageNumber="251" pagination="96 - 114" refId="ref16244" refString="BALFOUR-BROWNE J. 1954: A preliminary contribution to the knowledge of the SouthAfrican Palpicornia (Coleoptera). (Professor J. Omer-Cooper's investigation of the inland waters of Africa.) Journal of the Entomolological Society of South Africa 17: 96 - 114." type="journal article" year="1954">Balfour-Browne, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
, Kokstad, South Africa; D –
|
||
<taxonomicName id="4C5FBEFBFFDBB620FD4AFB08FC45B6D5" authority="Orchymont, 1947" authorityName="Orchymont" authorityYear="1947" baseAuthorityName="Orchymont" baseAuthorityYear="1948" box="[686,935,1214,1233]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="cooperi">
|
||
<emphasis id="B92B196AFFDBB620FD4AFB08FCEBB6D5" box="[686,777,1214,1233]" italics="true" pageId="2" pageNumber="251">H. cooperi</emphasis>
|
||
Orchymont, 1947
|
||
</taxonomicName>
|
||
, holotype; E –
|
||
<taxonomicName id="4C5FBEFBFFDBB620FBC2FB08FB23B6D5" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[1062,1217,1214,1233]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="brumopluvialis" status="sp. nov.">
|
||
<emphasis id="B92B196AFFDBB620FBC2FB08FB23B6D5" box="[1062,1217,1214,1233]" italics="true" pageId="2" pageNumber="251">H. brumopluvialis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A218A411FFDBB620FB22FB08FAE8B6D5" box="[1222,1290,1214,1233]" pageId="2" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype; F –
|
||
<taxonomicName id="4C5FBEFBFFDBB620FA63FB08FEFCB6E8" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="nyandaruaensis" status="sp. nov.">
|
||
<emphasis id="B92B196AFFDBB620FA63FB08FEFCB6E8" italics="true" pageId="2" pageNumber="251">H. nyandaruaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A218A411FFDBB620FEC0FB6FFE8AB6E8" box="[292,360,1241,1260]" pageId="2" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype; G –
|
||
<taxonomicName id="4C5FBEFBFFDBB620FE0FFB6FFDB1B6E8" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[491,595,1241,1260]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="simiensis" status="sp. nov.">
|
||
<emphasis id="B92B196AFFDBB620FE0FFB6FFDB1B6E8" box="[491,595,1241,1260]" italics="true" pageId="2" pageNumber="251">H. simiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A218A411FFDBB620FDBCFB6FFD7FB6E8" box="[600,669,1241,1260]" pageId="2" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34596F3FFDBB626FF73FA15FEE2B751" lastPageId="4" lastPageNumber="253" pageId="2" pageNumber="251" type="description">
|
||
<paragraph id="8BE0C578FFDBB620FF73FA15FD1BBA1E" blockId="2.[151,761,1442,2074]" pageId="2" pageNumber="251">
|
||
<emphasis id="B92B196AFFDBB620FF73FA15FE97B7BE" bold="true" box="[151,373,1442,1466]" pageId="2" pageNumber="251">
|
||
Redescription.
|
||
<emphasis id="B92B196AFFDBB620FEDAFA14FE97B7BE" bold="true" box="[318,373,1442,1466]" italics="true" pageId="2" pageNumber="251">Male</emphasis>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1364D9FDFFDBB620FE66FA15FE31B7BE" box="[386,467,1443,1466]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="2" pageNumber="251">Fig. 1A</figureCitation>
|
||
).
|
||
<emphasis id="B92B196AFFDBB620FE02FA15FDFDB7BE" box="[486,543,1443,1466]" italics="true" pageId="2" pageNumber="251">Head</emphasis>
|
||
dark brown to black, with strong green to purplish aeneous reflection; strongly shining. Granules mostly absent, reduced to a central seta-bearing pore over central areas of frons and clypeus; more evident inside compound eyes, laterally on clypeus and immediately in front of Y-groove. Y-groove relatively deep; floor of stem with dense, coarse, rugose punctures, punctures in lateral branches smaller, shining as rest of head; stem evenly diverging anteriorly; lateral branches narrow, particularly over apical 0.5, straight proximally then weakly arcuate laterally in front of eyes. Clypeus strongly elevated in centre; anterior margin almost straight. Labrum with shining, impunctate, basal portion with anterior margin acuminate at centre; anterior portion with coarse, rugose punctures, each puncture bearing a short, stout, whitish semi-erect seta; anterior margin arcuate with shallow anteromedian emargination.Antennae 9-segmented, yellowish brown, with cupule and club segments darker. Maxillary palpi yellow, apex of terminal segment infuscated; moderately elongate, terminal segment longest, curved and clearly asymmetrical.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDBB621FCBEFAF4FD1BB564" blockId="2.[828,1437,1346,2073]" lastBlockId="3.[151,761,1705,2080]" lastPageId="3" lastPageNumber="252" pageId="2" pageNumber="251">
|
||
<emphasis id="B92B196AFFDBB620FCBEFAF4FC26B75D" box="[858,964,1346,1369]" italics="true" pageId="2" pageNumber="251">Pronotum</emphasis>
|
||
(
|
||
<figureCitation id="1364D9FDFFDBB620FC36FAF4FBCAB75D" box="[978,1064,1346,1369]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="2" pageNumber="251">Fig. 3A</figureCitation>
|
||
), reddish brown, with strong green to purple aeneous reflections, particularly on internal intervals; anterior border narrowly and lateral borders more widely paler, yellowish brown; shining over entire dorsal surface. Broadest between middle and anterior margins; weakly arched in frontal and dorsal views and somewhat flattened over internal intervals, particularly anteriorly.Anterior margin slightly concave over central 0.5, then sinuate to broadly rounded front angles; posterior margin broadly bisinuate around centre. Sides straight over posterior 0.4, then weakly arcuate around broadest point to anterior angles. Lateral margins setose and weakly crenulated to serrate. Internal and middle intervals slightly arched, strongly shining; most granules reduced to pore and seta; granules more evident on anterior portion of both intervals, particularly middle. External interval, surface strongly shining with evident granules anteriorly. Grooves smooth and shining, median and submedian with almost coalescent, shallow punctures, giving a somewhat rugose appearance; submarginal and marginal with irregular small bumps. Median groove narrowed anteriorly, less so posteriorly, open at both ends more widely so posteriorly. Submedian and submarginal grooves broadly open anteriorly and posteriorly. Submedian grooves relatively broad, distinctly sinuate. Submarginal grooves weakly arcuate, broader than submedian. Marginal grooves curved parallel to the side of pronotum, width uniform. All grooves relatively deep, with intervals curving down towards their margins, which are well-defined but somewhat sloping rather than vertical.
|
||
</paragraph>
|
||
<caption id="DF2095F0FFDAB621FF7DF9B3FE07B46C" ID-DOI="http://doi.org/10.5281/zenodo.10134097" ID-Zenodo-Dep="10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="3" pageNumber="252" startId="3.[153,184,1541,1560]" targetBox="[172,1396,197,1494]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BE0C578FFDAB621FF7DF9B3FE07B46C" blockId="3.[151,1438,1541,1640]" pageId="3" pageNumber="252">
|
||
Fig. 2. Afrotropical
|
||
<taxonomicName id="4C5FBEFBFFDAB621FEA1F9B3FE48B41C" authorityName="Fabricius" authorityYear="1775" box="[325,426,1541,1560]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92B196AFFDAB621FEA1F9B3FE48B41C" box="[325,426,1541,1560]" italics="true" pageId="3" pageNumber="252">Helophorus</emphasis>
|
||
</taxonomicName>
|
||
, aedeagi. A –
|
||
<taxonomicName id="4C5FBEFBFFDAB621FDCCF9B3FCDDB41C" authority="Regimbart, 1907" authorityName="Regimbart" authorityYear="1907" box="[552,831,1541,1560]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="aethiopicus">
|
||
<emphasis id="B92B196AFFDAB621FDCCF9B3FD47B41C" box="[552,677,1541,1560]" italics="true" pageId="3" pageNumber="252">H. aethiopicus</emphasis>
|
||
<bibRefCitation id="EFCEB889FFDAB621FD48F9B3FCDDB41C" author="REGIMBART M." box="[684,831,1541,1560]" pageId="3" pageNumber="252" pagination="126 - 128" refId="ref16606" refString="REGIMBART M. 1907: Collections recueillis par M. Maurice de Rothschild dans l'Afrique orientale. Insectes coleopteres: Dytiscides et hydrophilides. Bulletin du Museum National d'Histoire Naturelle Paris 13: 126 - 128." type="journal article" year="1907">Régimbart, 1907</bibRefCitation>
|
||
</taxonomicName>
|
||
, lectotype; B –
|
||
<taxonomicName id="4C5FBEFBFFDAB621FC2EF9B3FBA5B41C" authorityName="Regimbart" authorityYear="1907" box="[970,1095,1541,1560]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="aethiopicus">
|
||
<emphasis id="B92B196AFFDAB621FC2EF9B3FBA5B41C" box="[970,1095,1541,1560]" italics="true" pageId="3" pageNumber="252">H. aethiopicus</emphasis>
|
||
</taxonomicName>
|
||
, paralectotype; C –
|
||
<taxonomicName id="4C5FBEFBFFDAB621FB1DF9B3FEDFB437" authority="J. Balfour-Browne, 1954" authorityName="J. Balfour-Browne" authorityYear="1954" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="aethiops">
|
||
<emphasis id="B92B196AFFDAB621FB1DF9B3FABEB41C" box="[1273,1372,1541,1560]" italics="true" pageId="3" pageNumber="252">H. aethiops</emphasis>
|
||
J.
|
||
<bibRefCitation id="EFCEB889FFDAB621FA93F9B3FEDFB437" author="BALFOUR-BROWNE J." pageId="3" pageNumber="252" pagination="96 - 114" refId="ref16244" refString="BALFOUR-BROWNE J. 1954: A preliminary contribution to the knowledge of the SouthAfrican Palpicornia (Coleoptera). (Professor J. Omer-Cooper's investigation of the inland waters of Africa.) Journal of the Entomolological Society of South Africa 17: 96 - 114." type="journal article" year="1954">Balfour-Browne, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype; D–F –
|
||
<taxonomicName id="4C5FBEFBFFDAB621FE32F996FCF6B437" authority="J. Balfour-Browne, 1954" authorityName="J. Balfour-Browne" authorityYear="1954" box="[470,788,1568,1587]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="aethiops">
|
||
<emphasis id="B92B196AFFDAB621FE32F996FDDAB437" box="[470,568,1568,1587]" italics="true" pageId="3" pageNumber="252">H. aethiops</emphasis>
|
||
J.
|
||
<bibRefCitation id="EFCEB889FFDAB621FDB5F996FCF6B437" author="BALFOUR-BROWNE J." box="[593,788,1568,1587]" pageId="3" pageNumber="252" pagination="96 - 114" refId="ref16244" refString="BALFOUR-BROWNE J. 1954: A preliminary contribution to the knowledge of the SouthAfrican Palpicornia (Coleoptera). (Professor J. Omer-Cooper's investigation of the inland waters of Africa.) Journal of the Entomolological Society of South Africa 17: 96 - 114." type="journal article" year="1954">Balfour-Browne, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
, Kokstad, South Africa; G –
|
||
<taxonomicName id="4C5FBEFBFFDAB621FBEFF996FAE6B437" authority="Orchymont, 1947" authorityName="Orchymont" authorityYear="1947" baseAuthorityName="Orchymont" baseAuthorityYear="1948" box="[1035,1284,1568,1587]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="cooperi">
|
||
<emphasis id="B92B196AFFDAB621FBEFF996FB84B437" box="[1035,1126,1568,1587]" italics="true" pageId="3" pageNumber="252">H. cooperi</emphasis>
|
||
Orchymont, 1947
|
||
</taxonomicName>
|
||
, holotype; H –
|
||
<taxonomicName id="4C5FBEFBFFDAB621FA63F996FEFAB449" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="brumopluvialis" status="sp. nov.">
|
||
<emphasis id="B92B196AFFDAB621FA63F996FEFAB449" italics="true" pageId="3" pageNumber="252">H. brumopluvialis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A218A411FFDAB621FEF9F98CFE83B449" box="[285,353,1594,1613]" pageId="3" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype; I–J –
|
||
<taxonomicName id="4C5FBEFBFFDAB621FE08F98CFD64B449" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[492,646,1594,1613]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="brumopluvialis" status="sp. nov.">
|
||
<emphasis id="B92B196AFFDAB621FE08F98CFD64B449" box="[492,646,1594,1613]" italics="true" pageId="3" pageNumber="252">H. brumopluvialis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A218A411FFDAB621FD6FF98CFD2DB449" box="[651,719,1594,1613]" pageId="3" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Hopefield, South Africa; K –
|
||
<taxonomicName id="4C5FBEFBFFDAB621FC2AF98CFB8DB449" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[974,1135,1594,1613]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="nyandaruaensis" status="sp. nov.">
|
||
<emphasis id="B92B196AFFDAB621FC2AF98CFB8DB449" box="[974,1135,1594,1613]" italics="true" pageId="3" pageNumber="252">H. nyandaruaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A218A411FFDAB621FB90F98CFB5AB449" box="[1140,1208,1594,1613]" pageId="3" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype; L –
|
||
<taxonomicName id="4C5FBEFBFFDAB621FAD1F98CFA7EB449" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[1333,1436,1594,1613]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="simiensis" status="sp. nov.">
|
||
<emphasis id="B92B196AFFDAB621FAD1F98CFA7EB449" box="[1333,1436,1594,1613]" italics="true" pageId="3" pageNumber="252">H. simiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A218A411FFDAB621FF73F9E3FF3EB46C" box="[151,220,1621,1640]" pageId="3" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype. Scale bar = 100
|
||
<emphasis id="B92B196AFFDAB621FE20F9E3FE2DB463" box="[452,463,1621,1639]" italics="true" pageId="3" pageNumber="252">μ</emphasis>
|
||
m.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE0C578FFDAB621FF51F8DFFEAABA24" blockId="3.[151,761,1705,2080]" pageId="3" pageNumber="252">
|
||
<emphasis id="B92B196AFFDAB621FF51F8DFFF1AB584" box="[181,248,1897,1920]" italics="true" pageId="3" pageNumber="252">Elytra</emphasis>
|
||
(
|
||
<figureCitation id="1364D9FDFFDAB621FEEDF8DFFEBCB584" box="[265,350,1897,1920]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="3" pageNumber="252">Fig. 1A</figureCitation>
|
||
) brown, with diffuse paler markings, particularly posteriorly. Shape elongate oval, sides rounded to bluntly rounded apex. In lateral profile flat over anterior 0.5, then gradually descending to apex. Puncture rows moderately striate, particularly on disc and laterally; intervals arched.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDAB621FCBEF91FFC68B504" blockId="3.[828,1437,1705,2080]" pageId="3" pageNumber="252">
|
||
<emphasis id="B92B196AFFDAB621FCBEF91FFC6EB4C4" box="[858,908,1705,1728]" italics="true" pageId="3" pageNumber="252">Legs</emphasis>
|
||
yellowish-brown, apex of last tarsal segment and claws infuscated. Swimming hairs distinct and clearly visible.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDAB626FCBEF8BFFEB6B111" blockId="3.[828,1437,1705,2080]" lastBlockId="4.[151,761,190,2069]" lastPageId="4" lastPageNumber="253" pageId="3" pageNumber="252">
|
||
<emphasis id="B92B196AFFDAB621FCBEF8BFFC7FB524" box="[858,925,1801,1824]" italics="true" pageId="3" pageNumber="252">Venter</emphasis>
|
||
black, with pronotal hypomeron, antennal pocket and pseudepipleura and epipleura brown to yellow. Mentum shining, centre without microreticulation, with sparse, medium punctures; lateral marginal areas with weak isodiametric microreticulation centrally; margins with fine yellow setae, denser towards posterolateral angles. Submentum shining, without microreticulation; anterior 0.5 with sparse, shallow, coarse punctures and fine, hair-like vestiture. Gula and genae rugose, with dense hydrofuge vestiture. Gular sutures well marked as pits anteriorly. Vestiture only absent around margin of compound eyes and their posterolateral corners. Prosternum dull, rugose, with dense hydrofuge vestiture and sparse, long, erect setae approximately 2× length of vestiture. Pronotal hypomeron dull, with dense hydrofuge vestiture. Pronotal epipleura shining, glabrous, with sparse shallow punctures. Anterior antennal pocket well developed, with long, dense setae internally and long, curved setae at posterolateral margins. Elytral epipleura dull, glabrous, narrowing posteriorly at level of metaventrite and continuing until posterior margin of ventrite. Elytral flanks very narrowly visible from below. Ventral surface of mesothorax, metathorax and abdomen dull, with dense hydrofuge vestiture and longer, erect setae; longer setae concentrated on centre of metaventrite and abdominal ventrites 2–5; setae on ventrites arising from longitudinal tubercles, which sometimes appear as shining patches. Hind margin of abdominal ventrite 5 weakly crenulate.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDDB626FF51FCA8FEE3B1F1" blockId="4.[151,761,190,2069]" pageId="4" pageNumber="253">
|
||
<emphasis id="B92B196AFFDDB626FF51FCA8FEF9B131" box="[181,283,798,821]" italics="true" pageId="4" pageNumber="253">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="1364D9FDFFDDB626FEC3FCA8FE7EB131" box="[295,412,798,821]" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="4" pageNumber="253">Figs 2A–B</figureCitation>
|
||
), elongate,
|
||
<quantity id="4CA7689DFFDDB626FDF4FCA8FD89B131" box="[528,619,798,821]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="253" unit="mm" value="0.45">0.45 mm</quantity>
|
||
in length. Parameres longer than basal piece; outer margins of parameres straight almost to apex; interior margins of parameres rather straight, apices pointed. Median lobe narrow, with weakly pointed apex; sides diverging slightly posteriorly to base of struts, weakly concave; struts distinctly shorter than tube.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDDB626FF51FC4BFE06B611" blockId="4.[151,761,190,2069]" box="[181,484,1021,1045]" pageId="4" pageNumber="253">
|
||
<emphasis id="B92B196AFFDDB626FF51FC4BFEEDB611" bold="true" box="[181,271,1021,1045]" italics="true" pageId="4" pageNumber="253">Female.</emphasis>
|
||
Externally as male.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDDB626FF51FBA8FD79B691" blockId="4.[151,761,190,2069]" pageId="4" pageNumber="253">
|
||
<emphasis id="B92B196AFFDDB626FF51FBA8FEBFB631" bold="true" box="[181,349,1054,1077]" italics="true" pageId="4" pageNumber="253">Measurements.</emphasis>
|
||
<typeStatus id="54E47BDAFFDDB626FE80FBA8FE31B631" box="[356,467,1054,1077]" pageId="4" pageNumber="253" type="lectotype">Lectotype</typeStatus>
|
||
: BL =
|
||
<quantity id="4CA7689DFFDDB626FDFEFBA8FD9EB631" box="[538,636,1054,1077]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="253" unit="mm" value="2.9">2.90 mm</quantity>
|
||
; EL = 2.00; EW =
|
||
<quantity id="4CA7689DFFDDB626FF39FB88FEDDB651" box="[221,319,1086,1109]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="253" unit="mm" value="1.25">1.25 mm</quantity>
|
||
. Males: BL =
|
||
<quantity id="4CA7689DFFDDB626FE32FB88FD97B651" box="[470,629,1086,1109]" metricMagnitude="-3" metricUnit="m" metricValue="2.775" metricValueMax="2.8" metricValueMin="2.75" pageId="4" pageNumber="253" unit="mm" value="2.775" valueMax="2.8" valueMin="2.75">2.75–2.80 mm</quantity>
|
||
; EL = 1.15– 2.00 mm; EW =
|
||
<quantity id="4CA7689DFFDDB626FEA1FBE8FE01B671" box="[325,483,1118,1141]" metricMagnitude="-3" metricUnit="m" metricValue="1.225" metricValueMax="1.25" metricValueMin="1.2" pageId="4" pageNumber="253" unit="mm" value="1.225" valueMax="1.25" valueMin="1.2">1.20–1.25 mm</quantity>
|
||
. Females: BL =
|
||
<quantity id="4CA7689DFFDDB626FD69FBE8FF26B691" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.5" metricValueMin="2.6" pageId="4" pageNumber="253" unit="mm" value="3.05" valueMax="3.5" valueMin="2.6">2.60–3.50 mm</quantity>
|
||
; EL =
|
||
<quantity id="4CA7689DFFDDB626FEEFFBC8FE4BB691" box="[267,425,1150,1173]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="4" pageNumber="253" unit="mm" value="2.15" valueMax="2.5" valueMin="1.8">1.80–2.50 mm</quantity>
|
||
; EW =
|
||
<quantity id="4CA7689DFFDDB626FE1DFBC8FD75B691" box="[505,663,1150,1173]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="4" pageNumber="253" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.20–1.50 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDDB626FF73FB28FEE2B751" blockId="4.[151,761,190,2069]" pageId="4" pageNumber="253">
|
||
<emphasis id="B92B196AFFDDB626FF73FB28FEE8B6B1" bold="true" box="[151,266,1182,1205]" pageId="4" pageNumber="253">Variation.</emphasis>
|
||
Some variation in colour of elytra, some of the
|
||
<typeStatus id="54E47BDAFFDDB626FF73FB08FED3B6D1" box="[151,305,1214,1237]" pageId="4" pageNumber="253" type="paralectotype">paralectotypes</typeStatus>
|
||
and additional specimens being somewhat paler than the
|
||
<typeStatus id="54E47BDAFFDDB626FEC8FB68FE6CB6F1" box="[300,398,1246,1269]" pageId="4" pageNumber="253" type="lectotype">lectotype</typeStatus>
|
||
illustrated. Punctation of pronotal grooves also varies somewhat, being more pronounced in some specimens, including
|
||
<typeStatus id="54E47BDAFFDDB626FE39FAA8FD98B731" box="[477,634,1310,1333]" pageId="4" pageNumber="253" type="paralectotype">paralectotypes</typeStatus>
|
||
, than in the
|
||
<typeStatus id="54E47BDAFFDDB626FF73FA88FF1EB751" box="[151,252,1342,1365]" pageId="4" pageNumber="253" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34596F3FFDDB626FF73FAE8FD3FB7B1" pageId="4" pageNumber="253" type="diagnosis">
|
||
<paragraph id="8BE0C578FFDDB626FF73FAE8FD3FB7B1" blockId="4.[151,761,190,2069]" pageId="4" pageNumber="253">
|
||
<emphasis id="B92B196AFFDDB626FF73FAE8FE6DB771" bold="true" box="[151,399,1374,1397]" pageId="4" pageNumber="253">Differential diagnosis.</emphasis>
|
||
This species can be distinguished by its 9-segemented antennae, relatively small size, weak pronotal granulation and the structure of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34596F3FFDDB626FF73FA08FF14B491" pageId="4" pageNumber="253" type="distribution">
|
||
<paragraph id="8BE0C578FFDDB626FF73FA08FF14B491" blockId="4.[151,761,190,2069]" pageId="4" pageNumber="253">
|
||
<emphasis id="B92B196AFFDDB626FF73FA08FE48B7D1" bold="true" box="[151,426,1470,1493]" pageId="4" pageNumber="253">Distribution and ecology.</emphasis>
|
||
Only known from
|
||
<collectingCountry id="F34885E8FFDDB626FD8DFA08FD23B7D1" box="[617,705,1470,1493]" name="Ethiopia" pageId="4" pageNumber="253">Ethiopia</collectingCountry>
|
||
(
|
||
<figureCitation id="1364D9FDFFDDB626FD2AFA08FF44B7F1" captionStart="Fig" captionStartId="14.[1132,1165,1128,1147]" captionTargetBox="[152,1098,191,1200]" captionTargetId="figure-437@14.[152,1098,191,1200]" captionTargetPageId="14" captionText="Fig. 5. Afrotropical Helophorus, known distributions.Map by https:// mapswire.com." figureDoi="http://doi.org/10.5281/zenodo.10134103" httpUri="https://zenodo.org/record/10134103/files/figure.png" pageId="4" pageNumber="253">Fig. 5</figureCitation>
|
||
), from three collections from two separate areas of the southern highlands; the
|
||
<collectingRegion id="499B0B9AFFDDB626FE70FA48FDA4B411" box="[404,582,1534,1557]" country="Ethiopia" name="Oromiya" pageId="4" pageNumber="253">Oromiya Region</collectingRegion>
|
||
in the southwest (
|
||
<typeStatus id="54E47BDAFFDDB626FF7AF9A8FF2FB431" box="[158,205,1566,1589]" pageId="4" pageNumber="253">type</typeStatus>
|
||
locality) and the Bale Mountains across the Ethiopian Rift Valley to the East. Potentially widespread in these southern high mountain districts, all localities being over
|
||
<quantity id="4CA7689DFFDDB626FF73F9C8FF10B491" box="[151,242,1662,1685]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="253" unit="m" value="2500.0">2,500 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34596F3FFDDB626FF73F928FAE7B111" pageId="4" pageNumber="253" type="discussion">
|
||
<paragraph id="8BE0C578FFDDB626FF73F928FAE7B111" blockId="4.[151,761,190,2069]" lastBlockId="4.[828,1437,190,789]" pageId="4" pageNumber="253">
|
||
<emphasis id="B92B196AFFDDB626FF73F928FEEDB4B1" bold="true" box="[151,271,1694,1717]" pageId="4" pageNumber="253">Comment.</emphasis>
|
||
Robert Angus had considered that this species could be a synonym of
|
||
<taxonomicName id="4C5FBEFBFFDDB626FE4DF908FD15B4D1" authority="Semenow, 1900" authorityName="Semenow" authorityYear="1900" box="[425,759,1726,1749]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="mervensis">
|
||
<emphasis id="B92B196AFFDDB626FE4DF908FDD9B4D1" box="[425,571,1726,1749]" italics="true" pageId="4" pageNumber="253">H. mervensis</emphasis>
|
||
Semenow, 1900
|
||
</taxonomicName>
|
||
known from
|
||
<collectingCountry id="F34885E8FFDDB626FECDF968FE69B4F1" box="[297,395,1758,1781]" name="Armenia" pageId="4" pageNumber="253">Armenia</collectingCountry>
|
||
, through Arabia to
|
||
<collectingCountry id="F34885E8FFDDB626FD8FF968FD7BB4F1" box="[619,665,1758,1781]" name="Iran" pageId="4" pageNumber="253">Iran</collectingCountry>
|
||
, Central Asia and Kashmir, something reflected by labels on part of the material examined (see above). Despite this, the two were never formally synonymised (
|
||
<bibRefCitation id="EFCEB889FFDDB626FDFDF888FD44B551" author="ANGUS R. B." box="[537,678,1854,1877]" pageId="4" pageNumber="253" pagination="238 - 256" refId="ref15844" refString="ANGUS R. B. 1971: Revisional notes on Helophorus F. (Col., Hydrophilidae). 3. - Species resembling H. strigifrons Thoms. and some further notes on species resembling H. minutus F. Entomologist's Monthly Magazine 107: 238 - 256 + 1 plate." type="book chapter" year="1971">ANGUS 1971</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFCEB889FFDDB626FD57F888FD0FB551" author="ANGUS R. B." box="[691,749,1854,1877]" pageId="4" pageNumber="253" pagination="133 - 163" refId="ref16011" refString="ANGUS R. B. 1986: Revision of the Palaearctic species of the Helophorus minutus group (Coleoptera: Hydrophilidae), with chromosome analysis and hybridization experiments. Systematic Entomology 11: 133 - 163." type="journal article" year="1986">1986</bibRefCitation>
|
||
). My assessment is that, whilst these taxa are similar, they are not conspecific. The aedeagi of these species are very close indeed, but the beetles themselves differ. Based on my examination of specimens from
|
||
<collectingCountry id="F34885E8FFDDB626FDC9F808FDB9B5D1" box="[557,603,1982,2005]" name="Iran" pageId="4" pageNumber="253">Iran</collectingCountry>
|
||
, Kashmir and Central Asia (Golodnya Steppe,
|
||
<collectingCountry id="F34885E8FFDDB626FE1AF868FD94B5F1" box="[510,630,2014,2037]" name="Uzbekistan" pageId="4" pageNumber="253">Uzbekistan</collectingCountry>
|
||
and Shymkent,
|
||
<collectingCountry id="F34885E8FFDDB626FF32F848FEB7BA11" box="[214,341,2046,2069]" name="Kazakhstan" pageId="4" pageNumber="253">Kazakhstan</collectingCountry>
|
||
)
|
||
<taxonomicName id="4C5FBEFBFFDDB626FE83F848FE14BA11" authorityName="Semenow" authorityYear="1900" box="[359,502,2046,2069]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="mervensis">
|
||
<emphasis id="B92B196AFFDDB626FE83F848FE14BA11" box="[359,502,2046,2069]" italics="true" pageId="4" pageNumber="253">H. mervensis</emphasis>
|
||
</taxonomicName>
|
||
is comparatively more elongate and paler that
|
||
<taxonomicName id="4C5FBEFBFFDDB626FBDDFF08FB34B2D1" authorityName="Regimbart" authorityYear="1907" box="[1081,1238,190,213]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="aethiopicus">
|
||
<emphasis id="B92B196AFFDDB626FBDDFF08FB34B2D1" box="[1081,1238,190,213]" italics="true" pageId="4" pageNumber="253">H. aethiopicus</emphasis>
|
||
</taxonomicName>
|
||
(ground colour of pronotum pale brown with aeneous reflections; elytra pale brown to yellow). In addition, the pronotal granulation of
|
||
<taxonomicName id="4C5FBEFBFFDDB626FCD8FEA8FC2AB331" authorityName="Semenow" authorityYear="1900" box="[828,968,286,309]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="mervensis">
|
||
<emphasis id="B92B196AFFDDB626FCD8FEA8FC2AB331" box="[828,968,286,309]" italics="true" pageId="4" pageNumber="253">H. mervensis</emphasis>
|
||
</taxonomicName>
|
||
is typically more reduced than in all the
|
||
<taxonomicName id="4C5FBEFBFFDDB626FA66FEA8FC5AB351" authorityName="Regimbart" authorityYear="1907" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="aethiopicus">
|
||
<emphasis id="B92B196AFFDDB626FA66FEA8FC5AB351" italics="true" pageId="4" pageNumber="253">H. aethiopicus</emphasis>
|
||
</taxonomicName>
|
||
I have seen, internal intervals in particular lacking any evident granules in most specimens. The pronotal grooves also appear shallower in
|
||
<taxonomicName id="4C5FBEFBFFDDB626FB39FEC8FA8BB391" authorityName="Semenow" authorityYear="1900" box="[1245,1385,382,405]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="mervensis">
|
||
<emphasis id="B92B196AFFDDB626FB39FEC8FA8BB391" box="[1245,1385,382,405]" italics="true" pageId="4" pageNumber="253">H. mervensis</emphasis>
|
||
</taxonomicName>
|
||
, and the elytral striae less deep apically, with flatter intervals than in
|
||
<taxonomicName id="4C5FBEFBFFDDB626FC62FE08FBFFB3D1" authorityName="Regimbart" authorityYear="1907" box="[902,1053,446,469]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="aethiopicus">
|
||
<emphasis id="B92B196AFFDDB626FC62FE08FBFFB3D1" box="[902,1053,446,469]" italics="true" pageId="4" pageNumber="253">H. aethiopicus</emphasis>
|
||
</taxonomicName>
|
||
, although, as stated by
|
||
<bibRefCitation id="EFCEB889FFDDB626FAECFE08FA7FB3D1" author="ANGUS R. B." box="[1288,1437,446,469]" pageId="4" pageNumber="253" pagination="133 - 163" refId="ref16011" refString="ANGUS R. B. 1986: Revision of the Palaearctic species of the Helophorus minutus group (Coleoptera: Hydrophilidae), with chromosome analysis and hybridization experiments. Systematic Entomology 11: 133 - 163." type="journal article" year="1986">ANGUS (1986)</bibRefCitation>
|
||
this character varies. Given the similarities, particularly in aedeagi, it may be that
|
||
<taxonomicName id="4C5FBEFBFFDDB626FBDFFE48FB2AB011" authorityName="Semenow" authorityYear="1900" box="[1083,1224,510,533]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="mervensis">
|
||
<emphasis id="B92B196AFFDDB626FBDFFE48FB2AB011" box="[1083,1224,510,533]" italics="true" pageId="4" pageNumber="253">H. mervensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5FBEFBFFDDB626FB1BFE48FA7EB011" authorityName="Regimbart" authorityYear="1907" box="[1279,1436,510,533]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="aethiopicus">
|
||
<emphasis id="B92B196AFFDDB626FB1BFE48FA7EB011" box="[1279,1436,510,533]" italics="true" pageId="4" pageNumber="253">H. aethiopicus</emphasis>
|
||
</taxonomicName>
|
||
share a relatively recent common ancestor, one sublineage of which colonised
|
||
<collectingCountry id="F34885E8FFDDB626FBE7FD88FBBDB051" box="[1027,1119,574,597]" name="Ethiopia" pageId="4" pageNumber="253">Ethiopia</collectingCountry>
|
||
. Clearly this hypothesis should be tested with DNA sequence data when practicable. AN- GUS (1986) lists four species described by David Sharp as synonyms of
|
||
<taxonomicName id="4C5FBEFBFFDDB626FC2EFD28FBB7B0B1" authorityName="Semenow" authorityYear="1900" box="[970,1109,670,693]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="mervensis">
|
||
<emphasis id="B92B196AFFDDB626FC2EFD28FBB7B0B1" box="[970,1109,670,693]" italics="true" pageId="4" pageNumber="253">H. mervensis</emphasis>
|
||
</taxonomicName>
|
||
and notes the variability of the species as currently defined. Whether it represents a single, widespread, Palaearctic species, or a complex, seems impossible to resolve without genetic data.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |