441 lines
64 KiB
XML
441 lines
64 KiB
XML
<document id="C2A51175CDCE7FD6B5DCD5FD915A16D0" ID-DOI="10.5281/zenodo.198523" ID-GBIF-Dataset="7ad3b64d-ffc2-432c-a8e9-e32b05de9900" ID-ISSN="1175-5326" ID-Zenodo-Dep="198523" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460303712167" checkinUser="plazi" docAuthor="Tiunova, Tatiana M." docDate="2010" docId="0390CB54787BFFFFFF2786A7D47BFC5E" docLanguage="en" docName="zt02639p018.pdf" docOrigin="Zootaxa 2639" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Rhithrogena klugei Tiunova, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="FFA9B32C787AFFFBFFB08630D704FFC2" masterDocTitle="A new species of Rhithrogena Eaton, 1881 from the Far East of Russia with notes on this genus in this area (Ephemeroptera: Heptageniidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="2" updateTime="1698252947402" updateUser="plazi">
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<mods:mods id="BAABF75E6F1A51B8157C8E9ABED42615" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EF2A6BA7568ED6226C637C318B4B7F89">
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<mods:title id="D608CA0E002B90D3CEA9B311B51F9BED">A new species of Rhithrogena Eaton, 1881 from the Far East of Russia with notes on this genus in this area (Ephemeroptera: Heptageniidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="133ADC96CBF3A7754EF993517DDD2487" type="personal">
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<mods:role id="459ED38828AF35F08005F3430FB08E6C">
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<mods:roleTerm id="D088AF63FD5507B751F74A40E6A94A76">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EFC61E9FC35EA92E5D43E8BF2CF94D4D">Tiunova, Tatiana M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="85507942E7C74683A19A36E2105518E3">text</mods:typeOfResource>
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<mods:relatedItem id="18E5F4AB53A657AC1CFB26FDD6175F42" type="host">
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<mods:titleInfo id="733A6214706BAB5B641F85AEDAF0CC93">
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<mods:title id="179CCAA9975AFD1392AC9593BA1C84BB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A00C04419EBBE4C17EFA70C18F7F66F8">
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<mods:date id="A944EA1122D598791A58FE6CA71C3851">2010</mods:date>
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<mods:detail id="8D333CDB3BC4AB334C15E77DF3C16BD0" type="volume">
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<mods:number id="A3C54E838C93ADB562B6D90255E95DD1">2639</mods:number>
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</mods:detail>
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||
<mods:extent id="23248502FB5D59066C2F8FA213247DE2" unit="page">
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<mods:start id="01B77125ED39B40962F59B9EDD8410F3">1</mods:start>
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<mods:end id="6D87AD8C9447782B234888F071EDEBE5">18</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="85E0FAD4B917D33C89111B9E233DC4AA">journal article</mods:classification>
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<mods:identifier id="054015F06C4986D9EC239C4F9F9D7E86" type="DOI">10.5281/zenodo.198523</mods:identifier>
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<mods:identifier id="7303960C381B50106DC6C5BADC912425" type="GBIF-Dataset">7ad3b64d-ffc2-432c-a8e9-e32b05de9900</mods:identifier>
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<mods:identifier id="4547B77BBD3D74122FDCBF775D7C1896" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="150459A0213649219B06B750F7B36666" type="Zenodo-Dep">198523</mods:identifier>
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</mods:mods>
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<treatment id="0390CB54787BFFFFFF2786A7D47BFC5E" ID-DOI="http://doi.org/10.5281/zenodo.6206666" ID-GBIF-Taxon="119400129" ID-Zenodo-Dep="6206666" LSID="urn:lsid:plazi:treatment:0390CB54787BFFFFFF2786A7D47BFC5E" httpUri="http://treatment.plazi.org/id/0390CB54787BFFFFFF2786A7D47BFC5E" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2">
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<subSubSection id="C32329C9787BFFFAFF2786A7D622FF16" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="8B867A42787BFFFAFF2786A7D6E3FF73" blockId="1.[151,487,151,212]" box="[151,487,151,177]" pageId="1" pageNumber="2">
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<heading id="D0CECD2E787BFFFAFF2786A7D6E3FF73" bold="true" box="[151,487,151,177]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="B94DA650787BFFFAFF2786A7D6E3FF73" bold="true" box="[151,487,151,177]" pageId="1" pageNumber="2">
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<taxonomicName id="4C3901C1787BFFFAFF2786A7D686FF73" ID-CoL="4SBPL" box="[151,386,151,177]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="klugei" status="sp. nov.">
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<emphasis id="B94DA650787BFFFAFF2786A7D686FF73" bold="true" box="[151,386,151,177]" italics="true" pageId="1" pageNumber="2">Rhithrogena klugei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27E1B2B787BFFFAFE3986A7D6E3FF73" box="[393,487,151,177]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B867A42787BFFFAFF27868AD622FF16" blockId="1.[151,487,151,212]" box="[151,294,186,212]" pageId="1" pageNumber="2">
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(
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<figureCitation id="130266C7787BFFFAFF2F868AD619FF16" box="[159,285,186,212]" captionStart-0="FIGURES 1 – 8" captionStart-1="FIGURES 9 – 23" captionStartId-0="1.[151,269,1925,1949]" captionStartId-1="2.[151,272,1737,1761]" captionTargetBox-0="[243,1346,522,1894]" captionTargetBox-1="[197,1355,187,1701]" captionTargetId-0="figure@1.[230,1357,507,1922]" captionTargetId-1="figure@2.[188,1398,166,1754]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." captionText-1="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri-0="https://zenodo.org/record/198525/files/figure.png" httpUri-1="https://zenodo.org/record/198526/files/figure.png" pageId="1" pageNumber="2">Figs. 1–23</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32329C9787BFFF8FF278735D45EFDE5" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph id="8B867A42787BFFFAFF278735D5E6FE86" blockId="1.[151,1436,261,474]" pageId="1" pageNumber="2">
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<emphasis id="B94DA650787BFFFAFF278735D613FEDD" bold="true" box="[151,279,261,287]" pageId="1" pageNumber="2">
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<typeStatus id="5482C4E0787BFFFAFF278735D616FEDD" box="[151,274,261,287]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male imago,
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<emphasis id="B94DA650787BFFFAFE768735D53BFEDD" bold="true" box="[454,575,261,287]" pageId="1" pageNumber="2">
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<collectingCountry id="F32E3AD2787BFFFAFE768735D53CFEDD" box="[454,568,261,287]" name="Russia" pageId="1" pageNumber="2">RUSSIA</collectingCountry>
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,
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</emphasis>
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Khabarovskiy Kray, Bikin River,
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<quantity id="4CC1D7A7787BFFFAFC5E8735D328FEDD" box="[1006,1068,261,287]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="2" unit="m" value="50.0">50 m</quantity>
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below motor-car bridge, line Vladivostok-Khabarovsk,
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<date id="FF875C82787BFFFAFE7C871AD550FE86" box="[460,596,298,324]" pageId="1" pageNumber="2" value="2005-06-16">16.VI 2005</date>
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, T Tiunova.
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</paragraph>
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<paragraph id="8B867A42787BFFFAFF75877FD60DFE18" blockId="1.[151,1436,261,474]" pageId="1" pageNumber="2">
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<emphasis id="B94DA650787BFFFAFF75877FD64AFEAB" bold="true" box="[197,334,335,361]" pageId="1" pageNumber="2">
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<typeStatus id="5482C4E0787BFFFAFF75877FD64DFEAB" box="[197,329,335,361]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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Collected with
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<typeStatus id="5482C4E0787BFFFAFDA28760D585FEA8" box="[530,641,336,362]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
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:
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<collectingCountry id="F32E3AD2787BFFFAFD3E8760D5E6FEA8" box="[654,738,336,362]" name="Russia" pageId="1" pageNumber="2">Russia</collectingCountry>
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, Khabarovskiy Kray, Bikin River,
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<quantity id="4CC1D7A7787BFFFAFB3C8760D3CEFEA8" box="[1164,1226,336,362]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="2" unit="m" value="50.0">50 m</quantity>
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below motor-car bridge, line Vladivostok-Khabarovsk,
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<date id="FF875C82787BFFFAFDDD8745D5FFFE4D" box="[621,763,373,399]" pageId="1" pageNumber="2" value="2005-06-16">16.VI 2005</date>
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, together with
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<typeStatus id="5482C4E0787BFFFAFC038745D325FE4D" box="[947,1057,373,399]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
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,
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<specimenCount id="9D3FB1CB787BFFFAFB9E8745D394FE4D" box="[1070,1168,373,399]" pageId="1" pageNumber="2" type="male">70 male</specimenCount>
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imagines,
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<specimenCount id="9D3FB1CB787BFFFAFAA78745D278FE4D" box="[1303,1404,373,399]" pageId="1" pageNumber="2" type="larva">3 larvae</specimenCount>
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; T Tiunova; same place,
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<date id="FF875C82787BFFFAFE2A87AAD520FE76" box="[410,548,410,436]" pageId="1" pageNumber="2" value="2005-06-18">18.VI 2005</date>
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, T Tiunova,
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<specimenCount id="9D3FB1CB787BFFFAFD0C87AAD408FE76" box="[700,780,410,436]" pageId="1" pageNumber="2" type="male">2 male</specimenCount>
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imagines; same place,
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<date id="FF875C82787BFFFAFB9187AAD3A2FE76" box="[1057,1190,410,436]" pageId="1" pageNumber="2" value="2005-07-02">2.VII 2005</date>
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, T Tiunova,
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<specimenCount id="9D3FB1CB787BFFFAFA8D87AAD298FE76" box="[1341,1436,410,436]" pageId="1" pageNumber="2" type="male">16 male</specimenCount>
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imagines.
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</paragraph>
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<caption id="DF462ACA787BFFFAFF2781B5D5F6F81E" httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="1" pageNumber="2" targetBox="[243,1346,522,1894]" targetPageId="1">
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<paragraph id="8B867A42787BFFFAFF2781B5D5F6F81E" blockId="1.[151,1437,1925,2012]" pageId="1" pageNumber="2">
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<emphasis id="B94DA650787BFFFAFF2781B5D647F85F" bold="true" box="[151,323,1925,1949]" pageId="1" pageNumber="2">FIGURES 1–8.</emphasis>
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<taxonomicName id="4C3901C1787BFFFAFEFA81B6D513F85F" box="[330,535,1926,1949]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="klugei" status="sp. nov.">
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<emphasis id="B94DA650787BFFFAFEFA81B6D513F85F" box="[330,535,1926,1949]" italics="true" pageId="1" pageNumber="2">Rhithrogena klugei</emphasis>
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</taxonomicName>
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<emphasis id="B94DA650787BFFFAFDAE81B5D570F85F" bold="true" box="[542,628,1925,1949]" pageId="1" pageNumber="2">
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<taxonomicNameLabel id="A27E1B2B787BFFFAFDAE81B5D570F85F" box="[542,628,1925,1949]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1=1.0, 2=0.5, 3–8=0.2.
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</paragraph>
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</caption>
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<caption id="DF462ACA7878FFF9FF2780F9D584F8A2" httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="2" pageNumber="3" targetBox="[197,1355,187,1701]" targetPageId="2">
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<paragraph id="8B867A427878FFF9FF2780F9D584F8A2" blockId="2.[151,1436,1737,1888]" pageId="2" pageNumber="3">
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<emphasis id="B94DA6507878FFF9FF2780F9D650F923" bold="true" box="[151,340,1737,1761]" pageId="2" pageNumber="3">FIGURES 9–23.</emphasis>
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<taxonomicName id="4C3901C17878FFF9FEEC80F9D52BF922" box="[348,559,1737,1760]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="klugei" status="sp. nov.">
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<emphasis id="B94DA6507878FFF9FEEC80F9D52BF922" box="[348,559,1737,1760]" italics="true" pageId="2" pageNumber="3">Rhithrogena klugei</emphasis>
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</taxonomicName>
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<emphasis id="B94DA6507878FFF9FD8680F9D58AF923" bold="true" box="[566,654,1737,1761]" pageId="2" pageNumber="3">
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<taxonomicNameLabel id="A27E1B2B7878FFF9FD8680F9D58AF923" box="[566,654,1737,1761]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14=1.0; 10,12–13,15–17=0.5; 11,18=0.01; 19–20=0.05; 21=0.2; 22–23=0.1.
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</paragraph>
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</caption>
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<paragraph id="8B867A427878FFF8FF7581BED45EFDE5" blockId="2.[151,1437,1934,2035]" lastBlockId="3.[151,1437,152,2007]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<emphasis id="B94DA6507878FFF9FF7581BED511F86A" bold="true" box="[197,533,1934,1960]" pageId="2" pageNumber="3">Other material examined.</emphasis>
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<collectingCountry id="F32E3AD27878FFF9FDAE81BFD570F86B" box="[542,628,1935,1961]" name="Russia" pageId="2" pageNumber="3">Russia</collectingCountry>
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:
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<emphasis id="B94DA6507878FFF9FD3081BED495F86A" bold="true" box="[640,913,1934,1960]" pageId="2" pageNumber="3">Chitinskaya Oblast’:</emphasis>
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Selenga River,
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<quantity id="4CC1D7A77878FFF9FBE581BFD3A6F86B" box="[1109,1186,1935,1961]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="3" unit="km" value="65.0">65 km</quantity>
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upper mouth,
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<date id="FF875C827878FFF9FAE681BFD7D3F80C" pageId="2" pageNumber="3" value="2007-06-25">25.VI 2007</date>
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, N Bazova,
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<specimenCount id="9D3FB1CB7878FFF9FEDF8184D6C6F80C" box="[367,450,1972,1998]" pageId="2" pageNumber="3" type="male">1 male</specimenCount>
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imago; Selenga River, Iljinka Village,
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<date id="FF875C827878FFF9FC178184D332F80C" box="[935,1078,1972,1998]" pageId="2" pageNumber="3" value="2007-06-13">13.VI 2007</date>
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, N Bazova,
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<specimenCount id="9D3FB1CB7878FFF9FB7E8184D220F80C" box="[1230,1316,1972,1998]" pageId="2" pageNumber="3" type="male">1 male</specimenCount>
|
||
,
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<specimenCount id="9D3FB1CB7878FFF9FA818184D298F80C" box="[1329,1436,1972,1998]" pageId="2" pageNumber="3" type="female">2 female</specimenCount>
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imagines; same place,
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<date id="FF875C827878FFF9FE1681E9D535F831" box="[422,561,2009,2035]" pageId="2" pageNumber="3" value="2007-06-25">25.VI 2007</date>
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, N Bazova,
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<specimenCount id="9D3FB1CB7878FFF9FD7581E9D412F831" box="[709,790,2009,2035]" pageId="2" pageNumber="3" type="male">2 male</specimenCount>
|
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imagines; Selenga River, Sutoj Village,
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<date id="FF875C827878FFF9FB4D81E9D27DF831" box="[1277,1401,2009,2035]" pageId="2" pageNumber="3" value="2008-06-09">9.VI 2008</date>
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, N Bazova,
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<specimenCount id="9D3FB1CB7879FFF8FF4E86A8D655FF70" box="[254,337,152,178]" pageId="3" pageNumber="4" type="male">3 male</specimenCount>
|
||
,
|
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<specimenCount id="9D3FB1CB7879FFF8FEED86A8D6C0FF70" box="[349,452,152,178]" pageId="3" pageNumber="4" type="female">1 female</specimenCount>
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imagines; Selenga River, Ust’-Kehta Village,
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<date id="FF875C827879FFF8FC5786A8D366FF70" box="[999,1122,152,178]" pageId="3" pageNumber="4" value="2008-06-09">9.VI 2008</date>
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, N Bazova,
|
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<specimenCount id="9D3FB1CB7879FFF8FB4386A8D246FF70" box="[1267,1346,152,178]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
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imago; Selenga River, Utes Tologoj,
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<date id="FF875C827879FFF8FDB9868DD59EFF15" box="[521,666,189,215]" pageId="3" pageNumber="4" value="2009-06-10">10.VI 2009</date>
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, N Bazova,
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<specimenCount id="9D3FB1CB7879FFF8FC84868DD48FFF15" box="[820,907,189,215]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
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,
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<specimenCount id="9D3FB1CB7879FFF8FC29868DD301FF15" box="[921,1029,189,215]" pageId="3" pageNumber="4" type="female">2 female</specimenCount>
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imagines.
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<emphasis id="B94DA6507879FFF8FB3F868DD290FF15" bold="true" box="[1167,1428,189,215]" pageId="3" pageNumber="4">Amurskaya Oblast’</emphasis>
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: Burunda River,
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<quantity id="4CC1D7A77879FFF8FEE186D2D69FFF3E" box="[337,411,226,252]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="4" unit="m" value="800.0">800 m</quantity>
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upper mouth, tributary Nora River (Selemdzha River Basin),
|
||
<date id="FF875C827879FFF8FBC086D2D3FDFF3E" box="[1136,1273,226,252]" pageId="3" pageNumber="4" value="2004-06-16">16.VI 2004</date>
|
||
, T Tiunova,
|
||
<specimenCount id="9D3FB1CB7879FFF8FA3D86D2D7D1FEE0" pageId="3" pageNumber="4" type="male">7 male</specimenCount>
|
||
imagines; Zeya River, near Krasnoyarovo Village,
|
||
<date id="FF875C827879FFF8FCC48738D30FFEE0" box="[884,1035,264,290]" pageId="3" pageNumber="4" value="2004-06-24">24.VI 2004</date>
|
||
, T Tiunova,
|
||
<specimenCount id="9D3FB1CB7879FFF8FB008738D213FEE0" box="[1200,1303,264,290]" pageId="3" pageNumber="4" type="male">27 male</specimenCount>
|
||
imagines.
|
||
<emphasis id="B94DA6507879FFF8FF27871DD67AFE85" bold="true" box="[151,382,301,327]" pageId="3" pageNumber="4">Primorskiy Kray:</emphasis>
|
||
Ussuri River,
|
||
<quantity id="4CC1D7A77879FFF8FD80871DD56AFE85" box="[560,622,301,327]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="4" unit="km" value="3.0">3 km</quantity>
|
||
below Novomikhailovka Village,
|
||
<date id="FF875C827879FFF8FBA3871DD392FE85" box="[1043,1174,301,327]" pageId="3" pageNumber="4" value="1991-05-27">27.V 1991</date>
|
||
, T Tiunova,
|
||
<specimenCount id="9D3FB1CB7879FFF8FA82871DD29CFE85" box="[1330,1432,301,327]" pageId="3" pageNumber="4" type="larva">3 larvae</specimenCount>
|
||
; Ussuri River, Saratovka Village,
|
||
<date id="FF875C827879FFF8FD948762D5A1FEAE" box="[548,677,338,364]" pageId="3" pageNumber="4" value="1991-05-29">29.V 1991</date>
|
||
, T Tiunova,
|
||
<specimenCount id="9D3FB1CB7879FFF8FC8F8762D4B6FEAE" box="[831,946,338,364]" pageId="3" pageNumber="4" type="larva">12 larvae</specimenCount>
|
||
; Ussuri River,
|
||
<quantity id="4CC1D7A77879FFF8FBD68762D3A0FEAE" box="[1126,1188,338,364]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="4" unit="km" value="6.0">6 km</quantity>
|
||
below Koksharovka Village,
|
||
<date id="FF875C827879FFF8FF4C8748D679FE50" box="[252,381,376,402]" pageId="3" pageNumber="4" value="1992-05-28">28.V 1992</date>
|
||
, T Tiunova,
|
||
<specimenCount id="9D3FB1CB7879FFF8FDA78748D56CFE50" box="[535,616,376,402]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
|
||
imago; Ussuri River, Stepanovka Village,
|
||
<date id="FF875C827879FFF8FBD98748D3F0FE50" box="[1129,1268,376,402]" pageId="3" pageNumber="4" value="2005-06-15">15.VI 2005</date>
|
||
, T Tiunova,
|
||
<specimenCount id="9D3FB1CB7879FFF8FA3D8748D7D5FE75" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
|
||
imago; Razdol’naya River, Pokrovka Village,
|
||
<date id="FF875C827879FFF8FD4F87ADD47FFE75" box="[767,891,413,439]" pageId="3" pageNumber="4" value="1991-06-01">1.VI 1991</date>
|
||
, T Vshivkova,
|
||
<specimenCount id="9D3FB1CB7879FFF8FB8287ADD387FE75" box="[1074,1155,413,439]" pageId="3" pageNumber="4" type="male">3 male</specimenCount>
|
||
imagines; Razdol’naya River, Fadeevka Village,
|
||
<date id="FF875C827879FFF8FE6687F2D58FFE1E" box="[470,651,450,476]" pageId="3" pageNumber="4" value="1998-05-30" valueMax="1998-05-31" valueMin="1998-05-30">30–31.V 1998</date>
|
||
, T Tiunova,
|
||
<specimenCount id="9D3FB1CB7879FFF8FC9B87F2D484FE1E" box="[811,896,450,476]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
|
||
imago; Komissarovka River, Komissarovo Village,
|
||
<date id="FF875C827879FFF8FEB887D8D691FDC0" box="[264,405,488,514]" pageId="3" pageNumber="4" value="1998-05-28">28.V 1998</date>
|
||
, T Tiunova,
|
||
<specimenCount id="9D3FB1CB7879FFF8FD8E87D8D5A2FDC0" box="[574,678,488,514]" pageId="3" pageNumber="4" type="male">12 male</specimenCount>
|
||
imagines,
|
||
<specimenCount id="9D3FB1CB7879FFF8FC8787D8D4A1FDC0" box="[823,933,488,514]" pageId="3" pageNumber="4" type="larva">4 larvae</specimenCount>
|
||
; Bolshaya Ussurka River,
|
||
<quantity id="4CC1D7A77879FFF8FAB387D8D240FDC0" box="[1283,1348,488,514]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="4" unit="km" value="2.0">2 km</quantity>
|
||
below Zvenigorodka Village,
|
||
<date id="FF875C827879FFF8FE16843DD524FDE5" box="[422,544,525,551]" pageId="3" pageNumber="4" value="2004-06-08">8.VI 2004</date>
|
||
, T Tiunova,
|
||
<specimenCount id="9D3FB1CB7879FFF8FD03843DD406FDE5" box="[691,770,525,551]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
|
||
imago.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32329C97879FFF8FF758402D6B3F815" pageId="3" pageNumber="4" type="description">
|
||
<paragraph id="8B867A427879FFF8FF758402D3D3FB95" blockId="3.[151,1437,152,2007]" pageId="3" pageNumber="4">
|
||
<emphasis id="B94DA6507879FFF8FF758402D6FBFD8E" bold="true" box="[197,511,562,588]" pageId="3" pageNumber="4">Description. Male imago</emphasis>
|
||
(in alcohol).
|
||
<emphasis id="B94DA6507879FFF8FD138402D444FD8E" bold="true" box="[675,832,562,588]" pageId="3" pageNumber="4">Dimensions:</emphasis>
|
||
length (mm): body 7.7–10.0; forewings 8.7–11.2; cerci 18.5–24.0. Total color of living imago brown.
|
||
<emphasis id="B94DA6507879FFF8FCB88467D451FDB3" bold="true" box="[776,853,599,625]" pageId="3" pageNumber="4">Head:</emphasis>
|
||
upper portion of eyes from pinkish-grey to dark grey. Eyes contiguous dorsally.
|
||
<emphasis id="B94DA6507879FFF8FDA4844DD578FD55" bold="true" box="[532,636,637,663]" pageId="3" pageNumber="4">Thorax:</emphasis>
|
||
medioscutum brown, submedioscutum light brown, scutellum and sublateroscutum dark brown (
|
||
<figureCitation id="130266C77879FFF8FE498492D53BFD7E" box="[505,575,674,700]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1925,1949]" captionTargetBox="[243,1346,522,1894]" captionTargetId="figure@1.[230,1357,507,1922]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
); median longitudinal suture well expressed, light yellow. Mesosternum brown. Femora and tibia of forelegs brown, darker than the middle and hind legs, with dark brown joints. Tarsal segments from light brown to yellowish, last segments and claw brownish. Femora of all legs with a contrasting black coloration. Length (mm) of foreleg segments: femora 1.9–2.3; tibia 2.6–3.2; tarsal segments 0.3–0.4, 1.5–1.9, 1.3–1.8, 1.0–1.5, 0.4–0.5. Wings hyaline; longitudinal veins brownish or whitish; crosssectional veins colorless; pterostigma whitish.
|
||
<emphasis id="B94DA6507879FFF8FD08856DD438FCB5" bold="true" box="[696,828,861,887]" pageId="3" pageNumber="4">Abdomen:</emphasis>
|
||
terga from brown to light brown, hyaline; terga 2–6 having a poorly defined pair of bow-shaped light colored strokes. Sterna 1–7 brownish, sterna 8–9 whitish, opaque. Forceps dark brown, styliger plate brown. Penis lobes from light brown to yellow, having a contrasting pattern with dark forceps and styliger plate. Penis lobes jut out to the edge of the styliger plate; tops of penis lobes are rounded externally, with an inner apical tooth that is long and pointed (
|
||
<figureCitation id="130266C77879FFF8FB5185C2D221FBCE" box="[1249,1317,1010,1036]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1925,1949]" captionTargetBox="[243,1346,522,1894]" captionTargetId="figure@1.[230,1357,507,1922]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
). Laterodorsoventral tooth on the penis lobes is absent (
|
||
<figureCitation id="130266C77879FFF8FD7D8228D445FBF0" box="[717,833,1048,1074]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1925,1949]" captionTargetBox="[243,1346,522,1894]" captionTargetId="figure@1.[230,1357,507,1922]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="3" pageNumber="4">Figs. 5–7</figureCitation>
|
||
). Titillators curved and pointed (
|
||
<figureCitation id="130266C77879FFF8FB758228D23CFBF0" box="[1221,1336,1048,1074]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1925,1949]" captionTargetBox="[243,1346,522,1894]" captionTargetId="figure@1.[230,1357,507,1922]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="3" pageNumber="4">Figs. 4–5</figureCitation>
|
||
).
|
||
<emphasis id="B94DA6507879FFF8FAFE8227D29FFBF3" bold="true" box="[1358,1435,1047,1073]" pageId="3" pageNumber="4">Cerci:</emphasis>
|
||
proximal third brown, distally from light brown to yellow, base of each segment dark brown.
|
||
</paragraph>
|
||
<paragraph id="8B867A427879FFF8FF758252D65EFB05" blockId="3.[151,1437,152,2007]" pageId="3" pageNumber="4">
|
||
<emphasis id="B94DA6507879FFF8FF758252D527FBBE" bold="true" box="[197,547,1122,1148]" pageId="3" pageNumber="4">Female imago. Dimensions:</emphasis>
|
||
length (mm): 6.5–8.8; forewings 7.2–11.0; cerci 11.0–11.5. Total color of living imago yellow. Terga 2–6 light brown with a pair of white spots. Legs yellow, joints of tarsal segments and claw brown.
|
||
</paragraph>
|
||
<paragraph id="8B867A427879FFF8FF7582E2D6EBFB2E" blockId="3.[151,1437,152,2007]" box="[197,495,1234,1260]" pageId="3" pageNumber="4">
|
||
<emphasis id="B94DA6507879FFF8FF7582E2D668FB2E" bold="true" box="[197,364,1234,1260]" pageId="3" pageNumber="4">Subimagines.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8B867A427879FFF8FF7582C7D6B3F815" blockId="3.[151,1437,152,2007]" pageId="3" pageNumber="4">
|
||
<emphasis id="B94DA6507879FFF8FF7582C7D66AFAD3" bold="true" box="[197,366,1271,1297]" pageId="3" pageNumber="4">Mature larva</emphasis>
|
||
.
|
||
<emphasis id="B94DA6507879FFF8FECE82C7D51FFAD3" bold="true" box="[382,539,1271,1297]" pageId="3" pageNumber="4">Dimensions:</emphasis>
|
||
length (mm): body 9–12, cerci 6–8.
|
||
<emphasis id="B94DA6507879FFF8FC6782C7D320FAD3" bold="true" box="[983,1060,1271,1297]" pageId="3" pageNumber="4">Head:</emphasis>
|
||
frons brown, frequently with a light medial longitudinal spot. Posterior part of the head having a pair of small white spots in the middle area (
|
||
<figureCitation id="130266C77879FFF8FF2F8372D7ECFA9E" box="[159,232,1346,1372]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
|
||
). Labrum slightly concave in the middle of the anterior margin, width 3.2 times longer than the length (
|
||
<figureCitation id="130266C77879FFF8FF108358D7F6FA40" box="[160,242,1384,1410]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig.12</figureCitation>
|
||
). Labium with U-shaped separation of glossae; tips of glossae slightly narrowed and rounded (
|
||
<figureCitation id="130266C77879FFF8FADC8358D7BDFA65" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 10</figureCitation>
|
||
). Lower tooth of mandible 1/3 times the length of the upper tooth (
|
||
<figureCitation id="130266C77879FFF8FC4F83BDD35EFA65" box="[1023,1114,1421,1447]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
|
||
). Hypopharynx with apex slightly convex (
|
||
<figureCitation id="130266C77879FFF8FEEA8382D6B4FA0E" box="[346,432,1458,1484]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 13</figureCitation>
|
||
). Maxilla with 9 comb-shaped setae at the apical margin (
|
||
<figureCitation id="130266C77879FFF8FBE18382D3A3FA0E" box="[1105,1191,1458,1484]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 18</figureCitation>
|
||
).
|
||
<emphasis id="B94DA6507879FFF8FB0C8382D227FA0E" bold="true" box="[1212,1315,1458,1484]" pageId="3" pageNumber="4">Thorax:</emphasis>
|
||
pronotum with white anterior margin and lateral corners; mesonotum brown with a light colored median longitudinal stripe and a pair of white spots at the base of the wing pads. Femur of each leg with dark spot in the middle of the light central area; setae on the dorsal surface of the fore-femora of three different
|
||
<typeStatus id="5482C4E07879FFF8FBC68012D3BCF9FE" box="[1142,1208,1570,1596]" pageId="3" pageNumber="4">types</typeStatus>
|
||
: 1—elongated with a rounded apex, 2—with distinct divergent margins and a truncated apex, 3—with distinct divergent margins and a rounded apex (
|
||
<figureCitation id="130266C77879FFF8FE28805DD6F0F945" box="[408,500,1645,1671]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 19</figureCitation>
|
||
). Claws of fore- and middle legs with four subapical denticles (
|
||
<figureCitation id="130266C77879FFF8FAB4805DD28BF945" box="[1284,1423,1645,1671]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 22–23</figureCitation>
|
||
). Length (mm) of leg segments are the following. Foreleg: femur 1.6–1.9, tibia 1.4–1.7, tarsus 0.5–0.6; middle leg: femur 1.6–2.1, tibia 1.4–1.7, tarsus 0.4–0.6; hind leg: femur 2.2–2.8, tibia 1.6–2.1, tarsus 0.5–0.6. Femora of the hind leg is 1.4 times longer than the femur of the middle and hind legs.
|
||
<emphasis id="B94DA6507879FFF8FB9280EDD3A2F935" bold="true" box="[1058,1190,1757,1783]" pageId="3" pageNumber="4">Abdomen:</emphasis>
|
||
terga brown, usually monotonous, sometimes lighter in the posterior area; central portion of the posterior margin of terga 5 having long and pointed teeth of uniform sizes (
|
||
<figureCitation id="130266C77879FFF8FD398118D5E2F880" box="[649,742,1832,1858]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 20</figureCitation>
|
||
); sterna from light brown to brownish-yellow, sternum 9 darker than the others; abdominal sternum
|
||
<quantity id="4CC1D7A77879FFF8FD3B817DD5BCF8A5" box="[651,696,1869,1895]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="4" unit="in" value="9.0">9 in</quantity>
|
||
females with deep posteromedian emargination (
|
||
<figureCitation id="130266C77879FFF8FB47817DD24AF8A5" box="[1271,1358,1869,1895]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 21</figureCitation>
|
||
). Gills white; gill I large with a wavy outer margin (
|
||
<figureCitation id="130266C77879FFF8FD1A8142D406F84E" box="[682,770,1906,1932]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 14</figureCitation>
|
||
); gills II–VI approximately 1.3 times as long as they are wide (
|
||
<figureCitation id="130266C77879FFF8FF5281A8D66DF870" box="[226,361,1944,1970]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 15–16</figureCitation>
|
||
); gill VII broadens in its middle area (
|
||
<figureCitation id="130266C77879FFF8FC8681A8D494F870" box="[822,912,1944,1970]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="3" pageNumber="4">Fig. 17</figureCitation>
|
||
). Cerci light brown, darker on the proximal portion with yellow tips.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32329C9787EFFFFFF7586A7D5A5FF15" pageId="4" pageNumber="5" type="etymology">
|
||
<paragraph id="8B867A42787EFFFFFF7586A7D5A5FF15" blockId="4.[151,1437,151,924]" pageId="4" pageNumber="5">
|
||
<emphasis id="B94DA650787EFFFFFF7586A7D651FF73" bold="true" box="[197,341,151,177]" pageId="4" pageNumber="5">Etymology.</emphasis>
|
||
The new species is named in honor of ephemeropterologist Nikita Kluge, who has been a long-time investigator of mayflies in
|
||
<collectingCountry id="F32E3AD2787EFFFFFDFB868DD599FF15" box="[587,669,189,215]" name="Russia" pageId="4" pageNumber="5">Russia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32329C9787EFFFFFF7586D2D498FEAE" pageId="4" pageNumber="5" type="distribution">
|
||
<paragraph id="8B867A42787EFFFFFF7586D2D498FEAE" blockId="4.[151,1437,151,924]" pageId="4" pageNumber="5">
|
||
<emphasis id="B94DA650787EFFFFFF7586D2D507FF3E" bold="true" box="[197,515,226,252]" pageId="4" pageNumber="5">Distribution and biology.</emphasis>
|
||
This species was found during an excursion to investigate mayfly species in the Russian Far East and in southwestern Siberia. The imagines were collected in light traps and in the grass and leaves of trees near the water. Larvae of this species were found in gravel and pebbles in the centers of large rivers that can be characterized as hyporhithral and potamal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32329C9787EFFFFFF758747D47BFC5E" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="8B867A42787EFFFFFF758747D47BFC5E" blockId="4.[151,1437,151,924]" pageId="4" pageNumber="5">
|
||
<emphasis id="B94DA650787EFFFFFF758747D656FE53" bold="true" box="[197,338,375,401]" pageId="4" pageNumber="5">Discussion.</emphasis>
|
||
Male imago of
|
||
<taxonomicName id="4C3901C1787EFFFFFDA48748D57BFE53" box="[532,639,376,401]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="klugei" status="sp. nov.">
|
||
<emphasis id="B94DA650787EFFFFFDA48748D57BFE53" box="[532,639,376,401]" italics="true" pageId="4" pageNumber="5">R. klugei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DA650787EFFFFFD378747D5E3FE53" bold="true" box="[647,743,375,401]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27E1B2B787EFFFFFD378747D5E3FE53" box="[647,743,375,401]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from all other members of
|
||
<taxonomicName id="4C3901C1787EFFFFFA828748D298FE53" box="[1330,1436,376,401]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lepnevae">
|
||
<emphasis id="B94DA650787EFFFFFA828748D298FE53" box="[1330,1436,376,401]" italics="true" pageId="4" pageNumber="5">lepnevae</emphasis>
|
||
</taxonomicName>
|
||
species group by the following characteristcs: Titillators of
|
||
<taxonomicName id="4C3901C1787EFFFFFCFF87ADD4D3FE74" box="[847,983,413,438]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lepnevae">
|
||
<emphasis id="B94DA650787EFFFFFCFF87ADD4D3FE74" box="[847,983,413,438]" italics="true" pageId="4" pageNumber="5">R. lepnevae</emphasis>
|
||
</taxonomicName>
|
||
are wide, with 3–5 teeth apically (
|
||
<figureCitation id="130266C7787EFFFFFADD87ADD7BDFE1E" captionStart="FIGURES 43 – 47" captionStartId="10.[151,270,1549,1573]" captionTargetBox="[307,1283,576,1519]" captionTargetId="figure@10.[301,1286,567,1527]" captionTargetPageId="10" captionText="FIGURES 43 – 47. Rhithrogena lepnevae, male imago: 43. Mesonotum, dorsal. 44. Penis, ventral. 45. Penis, dorsal. 46. Titillators, lateral. 47. Penis lobes lateral. Scale (mm): 43 = 1.0; 44 – 45,47 = 0.2; 46 = 0.05." httpUri="https://zenodo.org/record/198529/files/figure.png" pageId="4" pageNumber="5">Fig. 46</figureCitation>
|
||
), while titillators of
|
||
<taxonomicName id="4C3901C1787EFFFFFE0287F2D51BFE19" box="[434,543,450,475]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="klugei" status="sp. nov.">
|
||
<emphasis id="B94DA650787EFFFFFE0287F2D51BFE19" box="[434,543,450,475]" italics="true" pageId="4" pageNumber="5">R. klugei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DA650787EFFFFFD9787F2D58CFE1E" bold="true" box="[551,648,450,476]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27E1B2B787EFFFFFD9787F2D58CFE1E" box="[551,648,450,476]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
narrow to a single point apically (
|
||
<figureCitation id="130266C7787EFFFFFB9C87F2D390FE1E" box="[1068,1172,450,476]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1925,1949]" captionTargetBox="[243,1346,522,1894]" captionTargetId="figure@1.[230,1357,507,1922]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="4" pageNumber="5">Fig. 4–5</figureCitation>
|
||
) and are similar to
|
||
<emphasis id="B94DA650787EFFFFFA3287F2D682FDC3" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C3901C1787EFFFFFA3287F2D602FDC3" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bajkovae">R. bajkovae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3901C1787EFFFFFEA287D8D682FDC3" box="[274,390,488,513]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ingalik">R. ingalik</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B94DA650787EFFFFFE7287D8D562FDC3" box="[450,614,488,513]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C3901C1787EFFFFFE7287D8D566FDC3" box="[450,610,488,513]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="piechockii">R. piechockii</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
However
|
||
<taxonomicName id="4C3901C1787EFFFFFD5387D8D474FDC3" box="[739,880,488,513]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bajkovae">
|
||
<emphasis id="B94DA650787EFFFFFD5387D8D474FDC3" box="[739,880,488,513]" italics="true" pageId="4" pageNumber="5">R. bajkovae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130266C7787EFFFFFC3187D8D301FDC0" box="[897,1029,488,514]" captionStart="FIGURES 24 – 29" captionStartId="6.[151,270,1488,1512]" captionTargetBox="[299,1295,200,1462]" captionTargetId="figure@6.[291,1296,194,1467]" captionTargetPageId="6" captionText="FIGURES 24 – 29. Rhithrogena bajkovae, male imago: 24. Mesonotum, dorsal. 25. Penis, ventral. 26. Penis, dorsal. 27. Penis lobes lateral. 28. Penis lobes, view from above. 29. Apex of penis lobes, dorsal. Scale (mm): 24 = 1.0; 25 – 29 = 0.2." httpUri="https://zenodo.org/record/198527/files/figure.png" pageId="4" pageNumber="5">Fig. 25–29</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C3901C1787EFFFFFBF987D8D3BAFDC3" box="[1097,1214,488,513]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ingalik">
|
||
<emphasis id="B94DA650787EFFFFFBF987D8D3BAFDC3" box="[1097,1214,488,513]" italics="true" pageId="4" pageNumber="5">R. ingalik</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="130266C7787EFFFFFB7E87D8D213FDC0" box="[1230,1303,488,514]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1925,1949]" captionTargetBox="[243,1346,522,1894]" captionTargetId="figure@1.[230,1357,507,1922]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
|
||
(
|
||
<bibRefCitation id="EFA807B3787EFFFFFA9787D8D69CFDE5" author="Randolph" pageId="4" pageNumber="5" refString="Randolph, R. P. & McCafferty, W. P. (2005) The mayflies (Ephemeroptera) of Alaska, including a new species of Heptageniidae, Proceedings of Entomological Society of Washington, 107 (1), 190 - 199." type="journal article" year="2005">Randolph and McCafferty 2005</bibRefCitation>
|
||
)] have latero-dorsoventral teeth on penis lobes in contrast to
|
||
<taxonomicName id="4C3901C1787EFFFFFBD8843DD3D5FDE4" box="[1128,1233,525,550]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="klugei" status="sp. nov.">
|
||
<emphasis id="B94DA650787EFFFFFBD8843DD3D5FDE4" box="[1128,1233,525,550]" italics="true" pageId="4" pageNumber="5">R. klugei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DA650787EFFFFFB69843DD233FDE5" bold="true" box="[1241,1335,525,551]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27E1B2B787EFFFFFB69843DD233FDE5" box="[1241,1335,525,551]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="130266C7787EFFFFFAF7843DD7ADFD8E" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1925,1949]" captionTargetBox="[243,1346,522,1894]" captionTargetId="figure@1.[230,1357,507,1922]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="4" pageNumber="5">Fig. 5– 7</figureCitation>
|
||
). Tops of penis lobes of
|
||
<taxonomicName id="4C3901C1787EFFFFFE678402D540FD89" box="[471,580,562,587]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="klugei" status="sp. nov.">
|
||
<emphasis id="B94DA650787EFFFFFE678402D540FD89" box="[471,580,562,587]" italics="true" pageId="4" pageNumber="5">R. klugei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DA650787EFFFFFDFC8402D5AAFD8E" bold="true" box="[588,686,562,588]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27E1B2B787EFFFFFDFC8402D5AAFD8E" box="[588,686,562,588]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are rounded externally (
|
||
<figureCitation id="130266C7787EFFFFFC6A8402D320FD8E" box="[986,1060,562,588]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1925,1949]" captionTargetBox="[243,1346,522,1894]" captionTargetId="figure@1.[230,1357,507,1922]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
|
||
) and smoothly passes an inner apical tooth that is straight and pointed (
|
||
<figureCitation id="130266C7787EFFFFFDC48468D5B8FDB0" box="[628,700,600,626]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1925,1949]" captionTargetBox="[243,1346,522,1894]" captionTargetId="figure@1.[230,1357,507,1922]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
|
||
), whereas the tops of penis lobes of
|
||
<taxonomicName id="4C3901C1787EFFFFFBDE8468D203FDB3" box="[1134,1287,600,625]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="piechockii">
|
||
<emphasis id="B94DA650787EFFFFFBDE8468D203FDB3" box="[1134,1287,600,625]" italics="true" pageId="4" pageNumber="5">R. piechockii</emphasis>
|
||
</taxonomicName>
|
||
have a deep groove in the middle and at the base of inner apical tooth it is curved and arched [
|
||
<figureCitation id="130266C7787EFFFFFBE3844DD39EFD55" box="[1107,1178,637,663]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1925,1949]" captionTargetBox="[243,1346,522,1894]" captionTargetId="figure@1.[230,1357,507,1922]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Rhithrogena klugei sp. nov., male imago: 1. Mesonotum, dorsal. 2. Genitalia, ventral. 3. Penis, ventral. 4. Penis, dorsal. 5. Penis lobes, semi-lateral. 6. Penis lobes, view from above. 7. Apex of penis lobes, dorsal. 8. Apex of penis lobes, ventral. Scale (mm): 1 = 1.0, 2 = 0.5, 3 – 8 = 0.2." httpUri="https://zenodo.org/record/198525/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
|
||
(
|
||
<bibRefCitation id="EFA807B3787EFFFFFB19844DD256FD55" author="Braasch" box="[1193,1362,637,663]" pageId="4" pageNumber="5" refString="Braasch, D. (1977) Rhithrogena piechockii n. sp. Aus der Mongolei (Ephemeroptera: Heptageniidae). Entomologische Nachrichten, Dresden, 21 (9), 140 - 142." type="journal article" year="1977">Braasch 1977</bibRefCitation>
|
||
)]. The larvae of
|
||
<taxonomicName id="4C3901C1787EFFFFFEB58492D669FD79" box="[261,365,674,699]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="klugei" status="sp. nov.">
|
||
<emphasis id="B94DA650787EFFFFFEB58492D669FD79" box="[261,365,674,699]" italics="true" pageId="4" pageNumber="5">R. klugei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DA650787EFFFFFEC48492D6D5FD7E" bold="true" box="[372,465,674,700]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27E1B2B787EFFFFFEC48492D6D5FD7E" box="[372,465,674,700]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differ from those of
|
||
<taxonomicName id="4C3901C1787EFFFFFD748492D44AFD79" box="[708,846,674,699]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bajkovae">
|
||
<emphasis id="B94DA650787EFFFFFD748492D44AFD79" box="[708,846,674,699]" italics="true" pageId="4" pageNumber="5">R. bajkovae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3901C1787EFFFFFC368492D30BFD79" box="[902,1039,674,699]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lepnevae">
|
||
<emphasis id="B94DA650787EFFFFFC368492D30BFD79" box="[902,1039,674,699]" italics="true" pageId="4" pageNumber="5">R. lepnevae</emphasis>
|
||
</taxonomicName>
|
||
the form of gills I and the ratio of the length to the width of gills III–IV. In
|
||
<taxonomicName id="4C3901C1787EFFFFFDC884F8D5E5FD23" box="[632,737,712,737]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="klugei" status="sp. nov.">
|
||
<emphasis id="B94DA650787EFFFFFDC884F8D5E5FD23" box="[632,737,712,737]" italics="true" pageId="4" pageNumber="5">R. klugei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DA650787EFFFFFD5984F7D443FD23" bold="true" box="[745,839,711,737]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27E1B2B787EFFFFFD5984F7D443FD23" box="[745,839,711,737]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, gill I has a wavy inner margin (
|
||
<figureCitation id="130266C7787EFFFFFB7884F8D21BFD20" box="[1224,1311,712,738]" captionStart="FIGURES 9 – 23" captionStartId="2.[151,272,1737,1761]" captionTargetBox="[197,1355,187,1701]" captionTargetId="figure@2.[188,1398,166,1754]" captionTargetPageId="2" captionText="FIGURES 9 – 23. Rhithrogena klugei sp. nov., larva: 9. Head, dorsal. 10. Labium, dorsal. 11. Teeth of left mandible, dorsal. 12. Labrum, dorsal. 13. Hypopharynx, dorsal. 14. Gill I. 15. Gill III. 16. Gill IV. 17. Gill VII. 18. Comb-shaped setae the maxilla. 19. Setae on the dorsal surface of fore femora. 20. Posterior margin of terga 5. 21. Abdominal sternum 9 (female). 22. Tarsal claw of fore leg. 23. Tarsal claw of middle leg. Scale (mm): 9,14 = 1.0; 10,12 – 13,15 – 17 = 0.5; 11,18 = 0.01; 19 – 20 = 0.05; 21 = 0.2; 22 – 23 = 0.1." httpUri="https://zenodo.org/record/198526/files/figure.png" pageId="4" pageNumber="5">Fig. 14</figureCitation>
|
||
); gills III– IV are approximately 1.3 times as long as they are wide; in contrast to
|
||
<taxonomicName id="4C3901C1787EFFFFFC6884DDD367FCC4" box="[984,1123,749,774]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bajkovae">
|
||
<emphasis id="B94DA650787EFFFFFC6884DDD367FCC4" box="[984,1123,749,774]" italics="true" pageId="4" pageNumber="5">R. bajkovae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3901C1787EFFFFFB2D84DDD222FCC4" box="[1181,1318,749,774]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lepnevae">
|
||
<emphasis id="B94DA650787EFFFFFB2D84DDD222FCC4" box="[1181,1318,749,774]" italics="true" pageId="4" pageNumber="5">R. lepnevae</emphasis>
|
||
</taxonomicName>
|
||
, gill I has an inner marginal (
|
||
<figureCitation id="130266C7787EFFFFFEC58522D6D3FCEE" box="[373,471,786,812]" captionStart="FIGURES 30 – 42" captionStartId="7.[151,269,1837,1861]" captionTargetBox="[236,1358,486,1795]" captionTargetId="figure@7.[226,1360,455,1816]" captionTargetPageId="7" captionText="FIGURES 30 – 42. Rhithrogena bajkovae, larva: 30. Head, dorsal. 31. Labrum, dorsal. 32. Teeth of left mandible, dorsal. 33. Comb-shaped setae of the maxilla. 34. Gill I. 35. Gill III. 36. Gill IV. 37. Gill VII. 38. Setae on the dorsal surface of fore femora. 39. Abdominal sternum 9 (female). 40. Tarsal claw of fore leg. 41. Tarsal claw of middle leg. 42. Posterior margin of terga 5. Scale (mm): 30 = 0.1; 31 = 0.2; 32,40 – 41 = 0.01; 33,38,42 = 0.05; 34 = 1.0; 35 – 37,39 = 0.5." httpUri="https://zenodo.org/record/198528/files/figure.png" pageId="4" pageNumber="5">Figs. 34</figureCitation>
|
||
,
|
||
<figureCitation id="130266C7787EFFFFFE538522D507FCEE" box="[483,515,786,812]" captionStart="FIGURES 48 – 59" captionStartId="11.[151,272,1875,1899]" captionTargetBox="[245,1336,455,1830]" captionTargetId="figure@11.[239,1348,418,1854]" captionTargetPageId="11" captionText="FIGURES 48 – 59. Rhithrogena lepnevae, larva: 48. Head, dorsal. 49. Labrum, dorsal. 50. Comb-shaped setae of the maxilla. 51. Gill I. 52. Gill III. 53. Gill IV. 54. Gill VII. 55. Setae on the dorsal surface of fore femora. 56. Tarsal claw of fore leg. 57. Tarsal claw of middle leg. 58. Posterior margin of terga 5. 59. Abdominal sternum 9 (female). Scale (mm): 48,51 = 1.0; 49 = 0.2; 50,55,58 = 0.05; 52 – 54, 59 = 0.5; 56 – 57 = 0.1." httpUri="https://zenodo.org/record/198530/files/figure.png" pageId="4" pageNumber="5">51</figureCitation>
|
||
); gills III–IV are elongated, being approximately 1.5–1.7 times as long as they are wide. The larva of
|
||
<taxonomicName id="4C3901C1787EFFFFFE138508D508FC93" box="[419,524,824,849]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="klugei" status="sp. nov.">
|
||
<emphasis id="B94DA650787EFFFFFE138508D508FC93" box="[419,524,824,849]" italics="true" pageId="4" pageNumber="5">R. klugei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E1B2B787EFFFFFDA38507D575FC93" box="[531,625,823,849]" pageId="4" pageNumber="5" rank="species">
|
||
<emphasis id="B94DA650787EFFFFFDA38507D575FC93" bold="true" box="[531,625,823,849]" pageId="4" pageNumber="5">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differ from those of
|
||
<taxonomicName id="4C3901C1787EFFFFFCD98508D4E0FC93" box="[873,996,824,849]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sibirica">
|
||
<emphasis id="B94DA650787EFFFFFCD98508D4E0FC93" box="[873,996,824,849]" italics="true" pageId="4" pageNumber="5">R. sibirica</emphasis>
|
||
</taxonomicName>
|
||
in the absence of a rounded ledge on the inner margin of gills III–IV (
|
||
<figureCitation id="130266C7787EFFFFFDA6856DD59FFCB5" box="[534,667,861,887]" captionStart="FIGURES 65 – 77" captionStartId="14.[151,270,1500,1524]" captionTargetBox="[241,1358,215,1462]" captionTargetId="figure@14.[226,1360,194,1479]" captionTargetPageId="14" captionText="FIGURES 65 – 77. Rhithrogena sibirica, larva: 65. Head, dorsal. 66. Labrum, dorsal. 67. Teeth of left mandible, dorsal. 68. Comb-shaped setae of the maxilla. 69. Gill I. 70. Gill III. 71. Gill IV. 72. Gill VII. 73. Setae on the dorsal surface of fore femora. 74. Abdominal sternum 9 (female). 75. Tarsal claw of fore leg. 76. Tarsal claw of middle leg. 77. Posterior margin of terga 5. Scale (mm): 65,69 = 1.0; 66,74 = 0.2; 67 – 68,75 – 76 = 0.01; 70 – 72 = 0.5; 73,77 = 0.05." httpUri="https://zenodo.org/record/198532/files/figure.png" pageId="4" pageNumber="5">Fig. 70–71</figureCitation>
|
||
), in the form of the setae on the dorsal surface of the fore-femora (
|
||
<figureCitation id="130266C7787EFFFFFF2F85B2D7F0FC5E" box="[159,244,898,924]" captionStart="FIGURES 65 – 77" captionStartId="14.[151,270,1500,1524]" captionTargetBox="[241,1358,215,1462]" captionTargetId="figure@14.[226,1360,194,1479]" captionTargetPageId="14" captionText="FIGURES 65 – 77. Rhithrogena sibirica, larva: 65. Head, dorsal. 66. Labrum, dorsal. 67. Teeth of left mandible, dorsal. 68. Comb-shaped setae of the maxilla. 69. Gill I. 70. Gill III. 71. Gill IV. 72. Gill VII. 73. Setae on the dorsal surface of fore femora. 74. Abdominal sternum 9 (female). 75. Tarsal claw of fore leg. 76. Tarsal claw of middle leg. 77. Posterior margin of terga 5. Scale (mm): 65,69 = 1.0; 66,74 = 0.2; 67 – 68,75 – 76 = 0.01; 70 – 72 = 0.5; 73,77 = 0.05." httpUri="https://zenodo.org/record/198532/files/figure.png" pageId="4" pageNumber="5">Fig. 73</figureCitation>
|
||
) and in the number of subapical denticles on the claws.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |