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<document id="B402D3C7B9F47E31B254D0BE46BE1CE5" ID-DOI="10.5281/zenodo.274088" ID-GBIF-Dataset="0488fc0c-9dd6-4030-87d7-9212ab8c7916" ID-ISSN="1175-5326" ID-Zenodo-Dep="274088" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459813587655" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2008" docId="03FAB13B6541FFBCDD94FA78FE31FB84" docLanguage="en" docName="zt01696p036.pdf" docOrigin="Zootaxa 1696" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Cephennomicrus typicus Scott, comb. n." docType="treatment" docVersion="6" lastPageNumber="32" masterDocId="FFC3C9436544FFBBDD03FFFAFFFFFFF6" masterDocTitle="Taxonomic notes on the Cephenniini (Coleoptera, Scydmaenidae): Status of Coatesia Lea, Cephennomicrus Reitter, and Neseuthia Scott" masterLastPageNumber="36" masterPageNumber="25" pageNumber="30" updateTime="1698221300510" updateUser="plazi">
<mods:mods id="AFD3AE06463349327763331CCE9F597D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="31FB306BDFC03EA592FE535F7F3DF3DB">
<mods:title id="C757BCABDACB58A9EF13D0EAA0F34F0B">Taxonomic notes on the Cephenniini (Coleoptera, Scydmaenidae): Status of Coatesia Lea, Cephennomicrus Reitter, and Neseuthia Scott</mods:title>
</mods:titleInfo>
<mods:name id="6777C2272156B78560C9D4D6A5EFA9E4" type="personal">
<mods:role id="211470F1D9B85D3211F90E07D9217938">
<mods:roleTerm id="647D380C90F5E1CE20284E30A7CFAE79">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB643DA258BFD1EFF7E4A3FF59DDBABF">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="F770489A5313DB79918E4CD62766DB06">text</mods:typeOfResource>
<mods:relatedItem id="72296B0EB8901FD999BAF4D9215A5094" type="host">
<mods:titleInfo id="F2125D8022053CC8DCAF20279235C6CF">
<mods:title id="EA12E2D2C0CDDEF901B0448F53214E0A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="918ABD653CEBFC52C01F0F1510FE0102">
<mods:date id="84A4B5DD231525913D968D12A41FA113">2008</mods:date>
<mods:detail id="35C3F470280DB97F318F74CE6CD0336B" type="volume">
<mods:number id="DA2575BF09FC261CC03C95D01DDEEFAA">1696</mods:number>
</mods:detail>
<mods:extent id="E516B49A83B1CE57BB978D5C7DBA9ED0" unit="page">
<mods:start id="764EBA3898F89D401EE537BB6DF9DF5D">25</mods:start>
<mods:end id="594FA6BC7E63C9CB2616FB59F83BA1F2">36</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="54E06C0E8243D09957D3F6D936C697FE">journal article</mods:classification>
<mods:identifier id="9304E56AA7AB334C36234F99AE5F3C27" type="DOI">10.5281/zenodo.274088</mods:identifier>
<mods:identifier id="DAFEAE2517AE1F6C0EE45A551629927B" type="GBIF-Dataset">0488fc0c-9dd6-4030-87d7-9212ab8c7916</mods:identifier>
<mods:identifier id="30401A8B2843B8D692991BDD9995BCA9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2A647DDB7B6F66416D42AC1257AA04CE" type="Zenodo-Dep">274088</mods:identifier>
</mods:mods>
<treatment id="03FAB13B6541FFBCDD94FA78FE31FB84" ID-DOI="http://doi.org/10.5281/zenodo.6228959" ID-GBIF-Taxon="119354338" ID-Zenodo-Dep="6228959" LSID="urn:lsid:plazi:treatment:03FAB13B6541FFBCDD94FA78FE31FB84" httpUri="http://treatment.plazi.org/id/03FAB13B6541FFBCDD94FA78FE31FB84" lastPageId="7" lastPageNumber="32" pageId="5" pageNumber="30">
<subSubSection id="C34953A66541FFBEDD94FA78FD6AFA6A" box="[151,661,1410,1436]" pageId="5" pageNumber="30" type="nomenclature">
<paragraph id="8BEC002D6541FFBEDD94FA78FD6AFA6A" blockId="5.[151,661,1410,1436]" box="[151,661,1410,1436]" pageId="5" pageNumber="30">
<heading id="D0A4B7416541FFBEDD94FA78FD6AFA6A" bold="true" box="[151,661,1410,1436]" fontSize="11" level="1" pageId="5" pageNumber="30" reason="1">
<emphasis id="B927DC3F6541FFBEDD94FA78FD6AFA6A" bold="true" box="[151,661,1410,1436]" pageId="5" pageNumber="30">
<taxonomicName id="4C537BAE6541FFBEDD94FA78FDE2FA6A" ID-CoL="SFQ6" authority="Scott" authorityName="Scott" box="[151,541,1410,1436]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="typicus" status="comb. nov.">
<emphasis id="B927DC3F6541FFBEDD94FA78FE3DFA6A" bold="true" box="[151,450,1410,1436]" italics="true" pageId="5" pageNumber="30">Cephennomicrus typicus</emphasis>
(Scott)
</taxonomicName>
,
<taxonomicNameLabel id="A21461446541FFBEDF29FA78FD6AFA6A" box="[554,661,1410,1436]" pageId="5" pageNumber="30">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34953A66541FFBEDD94FA37FDF9FA12" box="[151,518,1485,1508]" pageId="5" pageNumber="30" type="reference_group">
<paragraph id="8BEC002D6541FFBEDD94FA37FDF9FA12" blockId="5.[151,518,1485,1508]" box="[151,518,1485,1508]" pageId="5" pageNumber="30">
<treatmentCitationGroup id="AB4327036541FFBEDD94FA37FDF9FA12" box="[151,518,1485,1508]" pageId="5" pageNumber="30">
<taxonomicName id="4C537BAE6541FFBEDD94FA37FDFDFA12" ID-CoL="47728" authority="Scott, 1922: 203" authorityName="Scott" authorityPageNumber="203" authorityYear="1922" box="[151,514,1485,1508]" class="Insecta" family="Staphylinidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="B927DC3F6541FFBEDD94FA37FEB6FA12" box="[151,329,1485,1508]" italics="true" pageId="5" pageNumber="30">Neseuthia typica</emphasis>
<treatmentCitation id="0AF2263C6541FFBEDC53FA37FDFDFA12" author="Scott" box="[336,514,1485,1508]" page="203" pageId="5" pageNumber="30" year="1922">
<bibRefCitation id="EFC27DDC6541FFBEDC53FA37FE32FA12" author="Scott" box="[336,461,1485,1508]" pageId="5" pageNumber="36" refString="Scott, H. (1922) Coleoptera: Scydmaenidae, Scaphidiidae, Phalacridae, Cucujidae (supplement), Lathridiidae, Mycetophagidae (including Propalticus), Bostrichidae, Lyctidae. The Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner, M. A. Vol. 7, report No. 4. Transactions of the Linnean Society of London, 18, 195 - 260." type="book chapter" year="1922">Scott, 1922</bibRefCitation>
: 203
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34953A66541FFBDDD94F9EFFDC3FEA4" lastPageId="6" lastPageNumber="31" pageId="5" pageNumber="30" type="materials_examined">
<paragraph id="8BEC002D6541FFBDDD94F9EFFDC3FEA4" blockId="5.[151,1436,1557,2023]" lastBlockId="6.[151,1436,152,338]" lastPageId="6" lastPageNumber="31" pageId="5" pageNumber="30">
<emphasis id="B927DC3F6541FFBEDD94F9EFFE27F9D9" bold="true" box="[151,472,1557,1583]" pageId="5" pageNumber="30">
<typeStatus id="54E8BE8F6541FFBEDD94F9EFFF2DF9D9" box="[151,210,1557,1583]" pageId="5" pageNumber="30">Type</typeStatus>
material.
<typeStatus id="54E8BE8F6541FFBEDC5FF9EFFE27F9D9" box="[348,472,1557,1583]" pageId="5" pageNumber="30" type="lectotype">Lectotype</typeStatus>
</emphasis>
(designated here;
<figureCitation id="13681CA86541FFBEDFB8F9EFFCCBF9D9" box="[699,820,1557,1583]" captionStart="FIGURES 4 – 9" captionStartId="6.[151,269,1585,1609]" captionTargetBox="[166,1424,383,1555]" captionTargetId="figure@6.[160,1427,374,1561]" captionTargetPageId="6" captionText="FIGURES 4 – 9. Type specimens of Cephennomicrus perpunctillum Reitter and Neseuthia typica Scott. 4, Holotype male of Cephennomicrus perpunctillum (0.80 mm). 5, Lectotype male of Neseuthia typica (0.97 mm). 6, Aedeagus of the holotype of Cephennomicrus perpunctillum in dorsal view (0.17 mm). 7, Aedeagus of the paralectotype of Neseuthia typica in dorsal view (0.15 mm). 8. Set of original labels on the holotype of Cephennomicrus perpunctillum. 9, Set of original labels on the holotype of Neseuthia typica." httpUri="https://zenodo.org/record/274090/files/figure.png" pageId="5" pageNumber="30">Figs. 5, 9</figureCitation>
):
<emphasis id="B927DC3F6541FFBEDE4DF9EFFB88F9D9" bold="true" box="[846,1143,1557,1583]" pageId="5" pageNumber="30">
<collectingCountry id="F34440BD6541FFBEDE4DF9EFFBE9F9D9" box="[846,1046,1557,1583]" name="Seychelles" pageId="5" pageNumber="30">SEYCHELLES</collectingCountry>
:
<collectingCountry id="F34440BD6541FFBED924F9EFFB8CF9D9" box="[1063,1139,1557,1583]" name="India" pageId="5" pageNumber="30">Mahé</collectingCountry>
:
</emphasis>
male, mounted on card bearing male symbol and number &quot;117&quot; handwritten in black, &quot;
<collectingCountry id="F34440BD6541FFBEDE83F9C7FC36F9A1" box="[896,969,1597,1623]" name="India" pageId="5" pageNumber="30">Mahe</collectingCountry>
, '08-9
<collectingCountry id="F34440BD6541FFBED91AF9C7FB69F9A1" box="[1049,1174,1597,1623]" name="Seychelles" pageId="5" pageNumber="30">Seychelles</collectingCountry>
Exp.&quot; (typed), &quot;Percy Sladen Trust Exped. Brit.Mus. 1926―246.&quot; (typed), &quot;
<typeStatus id="54E8BE8F6541FFBEDE41F99FFC7CF989" box="[834,899,1637,1663]" pageId="5" pageNumber="30">Type</typeStatus>
&quot; (circular, white with red margin, typed), &quot;
<taxonomicName id="4C537BAE6541FFBEDD9CF977FE98F951" box="[159,359,1677,1703]" class="Insecta" family="Staphylinidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="typica">Neseuthia typica</taxonomicName>
<typeStatus id="54E8BE8F6541FFBEDC6DF977FE44F951" box="[366,443,1677,1703]" pageId="5" pageNumber="30">
<collectionCode id="ED4298E86541FFBEDC6DF977FE44F951" box="[366,443,1677,1703]" pageId="5" pageNumber="30">TYPE</collectionCode>
</typeStatus>
. H.Scott [handwritten in black]
<typeStatus id="54E8BE8F6541FFBEDE32F977FC85F951" box="[817,890,1677,1703]" pageId="5" pageNumber="30">
<collectionCode id="ED4298E86541FFBEDE32F977FC85F951" box="[817,890,1677,1703]" pageId="5" pageNumber="30">TYPE</collectionCode>
</typeStatus>
[printed, highlighted in blue]&quot;, &quot;
<emphasis id="B927DC3F6541FFBED9F9F977FF2FF938" italics="true" pageId="5" pageNumber="30">Figured specimen</emphasis>
&quot; (red, printed), &quot;
<taxonomicName id="4C537BAE6541FFBEDC97F94FFDD1F939" authority="Scott" authorityName="Scott" box="[404,558,1717,1743]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="typicus">typicus Scott</taxonomicName>
[handwritten in black] Cl. Besuchet dét. [printed] X [handwritten in black] 195 [printed] 7 [handwritten in black]&quot;, &quot;
<taxonomicName id="4C537BAE6541FFBEDF75F927FC58F901" authority="Scott" authorityName="Scott" box="[630,935,1757,1783]" class="Insecta" family="Staphylinidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="B927DC3F6541FFBEDF75F927FC9EF900" box="[630,865,1757,1782]" italics="true" pageId="5" pageNumber="30">NESEUTHIA typica</emphasis>
Scott
</taxonomicName>
,
<typeStatus id="54E8BE8F6541FFBEDEB1F927FBA2F901" box="[946,1117,1757,1783]" pageId="5" pageNumber="30" type="lectotype">LECTOTYPE</typeStatus>
; P. JAŁOSZYŃSKI, 2007&quot; (printed), &quot;
<taxonomicName id="4C537BAE6541FFBEDC1BF8FFFD2EF8E9" authority="Scott" authorityName="Scott" box="[280,721,1797,1823]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="B927DC3F6541FFBEDC1BF8FFFD87F8E8" box="[280,632,1797,1822]" italics="true" pageId="5" pageNumber="30">CEPHENNOMICRUS typicus</emphasis>
(Scott)
</taxonomicName>
, det. P. JAŁOSZYŃSKI, 2007&quot; (printed) (
<collectionCode id="ED4298E86541FFBED9C1F8FFFADEF8E9" box="[1218,1313,1797,1823]" pageId="5" pageNumber="30">BMNH</collectionCode>
).
<emphasis id="B927DC3F6541FFBED837F8FFFF0EF8B1" bold="true" pageId="5" pageNumber="30">
<typeStatus id="54E8BE8F6541FFBED837F8FFFF0EF8B1" pageId="5" pageNumber="30" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
(
<specimenCount id="9D55CBA46541FFBEDC02F8D7FEA0F8B1" box="[257,351,1837,1863]" pageId="5" pageNumber="30" type="male">2 males</specimenCount>
,
<specimenCount id="9D55CBA46541FFBEDC6FF8D7FE27F8B1" box="[364,472,1837,1863]" pageId="5" pageNumber="30" type="female">1 female</specimenCount>
):
<emphasis id="B927DC3F6541FFBEDCEFF8D7FCEDF8B1" bold="true" box="[492,786,1837,1863]" pageId="5" pageNumber="30">
<collectingCountry id="F34440BD6541FFBEDCEFF8D7FD4BF8B1" box="[492,692,1837,1863]" name="Seychelles" pageId="5" pageNumber="30">SEYCHELLES</collectingCountry>
:
<collectingCountry id="F34440BD6541FFBEDFC3F8D7FCF3F8B1" box="[704,780,1837,1863]" name="India" pageId="5" pageNumber="30">Mahé</collectingCountry>
:
</emphasis>
<specimenCount id="9D55CBA46541FFBEDE19F8D7FC7CF8B1" box="[794,899,1837,1863]" pageId="5" pageNumber="30" type="female">1 female</specimenCount>
, mounted on card bearing female symbol and number &quot;104&quot;, labels as in
<typeStatus id="54E8BE8F6541FFBEDCD5F8AFFDBEF899" box="[470,577,1877,1903]" pageId="5" pageNumber="30" type="lectotype">lectotype</typeStatus>
but lacking the red label &quot;Figured specimen&quot;;
<specimenCount id="9D55CBA46541FFBED964F8AFFB49F899" box="[1127,1206,1877,1903]" pageId="5" pageNumber="30" type="male">1 male</specimenCount>
mounted on different (newer?) card than previous specimens, &quot;
<typeStatus id="54E8BE8F6541FFBEDFB7F887FCDDF861" box="[692,802,1917,1943]" pageId="5" pageNumber="30" type="paratype">Paratype</typeStatus>
&quot; (circular, white with yellow margin, typed), &quot;Percy Sladen Trust Exped. Brit. Mus. 1926-246&quot; (typed), &quot;
<taxonomicName id="4C537BAE6541FFBEDE08F85FFBBBF849" authority="H. Scott" authorityName="H. Scott" box="[779,1092,1957,1983]" class="Insecta" family="Staphylinidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="typica">Neseuthia typica H. Scott</taxonomicName>
,
<typeStatus id="54E8BE8F6541FFBED951F85FFB43F849" box="[1106,1212,1957,1983]" pageId="5" pageNumber="30" type="paratype">Paratype</typeStatus>
&quot; (printed), &quot;
<collectingCountry id="F34440BD6541FFBED84CF85FFA67F849" box="[1359,1432,1957,1983]" name="India" pageId="5" pageNumber="30">Mahe</collectingCountry>
, 1908-9.
<collectingCountry id="F34440BD6541FFBEDDFAF837FE89F811" box="[249,374,1997,2023]" name="Seychelles" pageId="5" pageNumber="30">Seychelles</collectingCountry>
Exp.&quot; (printed), and similar Besuchet's label as that shown in
<figureCitation id="13681CA86541FFBED952F837FB69F811" box="[1105,1174,1997,2023]" captionStart="FIGURES 4 – 9" captionStartId="6.[151,269,1585,1609]" captionTargetBox="[166,1424,383,1555]" captionTargetId="figure@6.[160,1427,374,1561]" captionTargetPageId="6" captionText="FIGURES 4 – 9. Type specimens of Cephennomicrus perpunctillum Reitter and Neseuthia typica Scott. 4, Holotype male of Cephennomicrus perpunctillum (0.80 mm). 5, Lectotype male of Neseuthia typica (0.97 mm). 6, Aedeagus of the holotype of Cephennomicrus perpunctillum in dorsal view (0.17 mm). 7, Aedeagus of the paralectotype of Neseuthia typica in dorsal view (0.15 mm). 8. Set of original labels on the holotype of Cephennomicrus perpunctillum. 9, Set of original labels on the holotype of Neseuthia typica." httpUri="https://zenodo.org/record/274090/files/figure.png" pageId="5" pageNumber="30">Fig. 9</figureCitation>
and found in all specimens;
<specimenCount id="9D55CBA46542FFBDDDEDFF62FEC0FF44" box="[238,319,152,178]" pageId="6" pageNumber="31" type="male">1 male</specimenCount>
mounted on the same
<typeStatus id="54E8BE8F6542FFBDDF53FF62FD7DFF44" box="[592,642,152,178]" pageId="6" pageNumber="31">type</typeStatus>
of card as the previous specimen, &quot;
<typeStatus id="54E8BE8F6542FFBDD92FFF62FB65FF44" box="[1068,1178,152,178]" pageId="6" pageNumber="31" type="paratype">Paratype</typeStatus>
&quot; (circular, white with yellow margin, typed), &quot;Ma(y?)e,
<date id="FFED26ED6542FFBDDF35FF3AFD65FF2C" box="[566,666,192,218]" pageId="6" pageNumber="31" value="1908" valueMax="1909">1908-09</date>
,
<collectingCountry id="F34440BD6542FFBDDFA9FF3AFCD8FF2C" box="[682,807,192,218]" name="Seychelles" pageId="6" pageNumber="31">Seychelles</collectingCountry>
Exp.&quot; (handwritten in black, in easily recognizable Besuchet's hand). All
<typeStatus id="54E8BE8F6542FFBDDC9FFF12FDBAFEF4" box="[412,581,232,258]" pageId="6" pageNumber="31" type="paralectotype">paralectotypes</typeStatus>
with additional labels &quot;
<taxonomicName id="4C537BAE6542FFBDDE63FF12FB6BFEF4" authority="Scott" authorityName="Scott" box="[864,1172,232,258]" class="Insecta" family="Staphylinidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="B927DC3F6542FFBDDE63FF12FBB2FEF7" box="[864,1101,232,257]" italics="true" pageId="6" pageNumber="31">NESEUTHIA typica</emphasis>
Scott
</taxonomicName>
,
<typeStatus id="54E8BE8F6542FFBDD9A2FF12FA67FEF4" box="[1185,1432,232,258]" pageId="6" pageNumber="31" type="paralectotype">PARALECTOTYPE</typeStatus>
; P. JAŁOSZYŃSKI, 2007&quot; (printed), &quot;
<taxonomicName id="4C537BAE6542FFBDDF5DFEEAFBE1FEDC" authority="Scott" authorityName="Scott" box="[606,1054,272,298]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="B927DC3F6542FFBDDF5DFEEAFC3DFEDF" box="[606,962,272,297]" italics="true" pageId="6" pageNumber="31">CEPHENNOMICRUS typicus</emphasis>
(Scott)
</taxonomicName>
, det. P. JAŁOSZYŃSKI, 2007&quot; (printed) (all specimens in
<collectionCode id="ED4298E86542FFBDDCD1FEC2FDCFFEA4" box="[466,560,312,338]" pageId="6" pageNumber="31">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF2C50A56542FFBDDD94F9CBFD47F925" httpUri="https://zenodo.org/record/274090/files/figure.png" pageId="6" pageNumber="31" targetBox="[166,1424,383,1555]" targetPageId="6">
<paragraph id="8BEC002D6542FFBDDD94F9CBFD47F925" blockId="6.[151,1436,1585,1747]" pageId="6" pageNumber="31">
<emphasis id="B927DC3F6542FFBDDD94F9CBFEBDF9BF" bold="true" box="[151,322,1585,1609]" pageId="6" pageNumber="31">FIGURES 4–9.</emphasis>
Type specimens of
<taxonomicName id="4C537BAE6542FFBDDF16F9C8FC4AF9BF" authority="Reitter" authorityName="Reitter" box="[533,949,1586,1609]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="perpunctillum">
<emphasis id="B927DC3F6542FFBDDF16F9C8FC9AF9BF" box="[533,869,1586,1609]" italics="true" pageId="6" pageNumber="31">Cephennomicrus perpunctillum</emphasis>
Reitter
</taxonomicName>
and
<taxonomicName id="4C537BAE6542FFBDDEEBF9C8FB20F9BF" authority="Scott." authorityName="Scott." box="[1000,1247,1586,1609]" class="Insecta" family="Staphylinidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="B927DC3F6542FFBDDEEBF9C8FB65F9BF" box="[1000,1178,1586,1609]" italics="true" pageId="6" pageNumber="31">Neseuthia typica</emphasis>
Scott.
</taxonomicName>
4, Holotype male of
<taxonomicName id="4C537BAE6542FFBDDDB4F9AEFDF5F99D" box="[183,522,1620,1643]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="perpunctillum">
<emphasis id="B927DC3F6542FFBDDDB4F9AEFDF5F99D" box="[183,522,1620,1643]" italics="true" pageId="6" pageNumber="31">Cephennomicrus perpunctillum</emphasis>
</taxonomicName>
(0.80 mm). 5, Lectotype male of
<taxonomicName id="4C537BAE6542FFBDDE8BF9AEFBC2F99D" box="[904,1085,1620,1643]" class="Insecta" family="Staphylinidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="B927DC3F6542FFBDDE8BF9AEFBC2F99D" box="[904,1085,1620,1643]" italics="true" pageId="6" pageNumber="31">Neseuthia typica</emphasis>
</taxonomicName>
(0.97 mm). 6, Aedeagus of the holotype of
<taxonomicName id="4C537BAE6542FFBDDC1FF98DFD90F978" box="[284,623,1655,1678]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="perpunctillum">
<emphasis id="B927DC3F6542FFBDDC1FF98DFD90F978" box="[284,623,1655,1678]" italics="true" pageId="6" pageNumber="31">Cephennomicrus perpunctillum</emphasis>
</taxonomicName>
in dorsal view (0.17 mm). 7, Aedeagus of the paralectotype of
<taxonomicName id="4C537BAE6542FFBDD832F98DFF27F947" class="Insecta" family="Staphylinidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="B927DC3F6542FFBDD832F98DFF27F947" italics="true" pageId="6" pageNumber="31">Neseuthia typica</emphasis>
</taxonomicName>
in dorsal view (0.15 mm). 8. Set of original labels on the holotype of
<taxonomicName id="4C537BAE6542FFBDDEDEF960FACFF947" box="[989,1328,1690,1713]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="perpunctillum">
<emphasis id="B927DC3F6542FFBDDEDEF960FACFF947" box="[989,1328,1690,1713]" italics="true" pageId="6" pageNumber="31">Cephennomicrus perpunctillum</emphasis>
</taxonomicName>
. 9, Set of original labels on the holotype of
<taxonomicName id="4C537BAE6542FFBDDCFDF946FD4EF925" box="[510,689,1724,1747]" class="Insecta" family="Staphylinidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="B927DC3F6542FFBDDCFDF946FD4EF925" box="[510,689,1724,1747]" italics="true" pageId="6" pageNumber="31">Neseuthia typica</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C34953A66542FFBCDDC5F8D3FE31FB84" lastPageId="7" lastPageNumber="32" pageId="6" pageNumber="31" type="discussion">
<paragraph id="8BEC002D6542FFBCDDC5F8D3FD22FD4C" blockId="6.[151,1436,1833,2020]" lastBlockId="7.[151,1437,152,1138]" lastPageId="7" lastPageNumber="32" pageId="6" pageNumber="31">
<emphasis id="B927DC3F6542FFBDDDC5F8D3FEBFF8B5" bold="true" box="[198,320,1833,1859]" pageId="6" pageNumber="31">Remarks.</emphasis>
Besides the characters given in the diagnosis, all species of
<taxonomicName id="4C537BAE6542FFBDD924F8D0FB11F8B5" box="[1063,1262,1834,1859]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6542FFBDD924F8D0FB11F8B5" box="[1063,1262,1834,1859]" italics="true" pageId="6" pageNumber="31">Cephennomicrus</emphasis>
</taxonomicName>
are extremely small, typically much smaller than
<quantity id="4CABADC86542FFBDDF32F8A8FD8BF89A" box="[561,628,1874,1900]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="31" unit="mm" value="1.0">1 mm</quantity>
in length. Very small species can also be found in the Palearctic genera
<taxonomicName id="4C537BAE6542FFBDDDC7F880FE81F865" box="[196,382,1914,1939]" class="Insecta" family="Staphylinidae" genus="Nanophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6542FFBDDDC7F880FE81F865" box="[196,382,1914,1939]" italics="true" pageId="6" pageNumber="31">Nanophthalmus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C537BAE6542FFBDDCBAF880FDB2F865" box="[441,589,1914,1939]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6542FFBDDCBAF880FDB2F865" box="[441,589,1914,1939]" italics="true" pageId="6" pageNumber="31">Cephennium</emphasis>
</taxonomicName>
, but the body as small as 0.6 mm, not uncommon in
<taxonomicName id="4C537BAE6542FFBDD9CDF880FA6BF865" box="[1230,1428,1914,1939]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6542FFBDD9CDF880FA6BF865" box="[1230,1428,1914,1939]" italics="true" pageId="6" pageNumber="31">Cephennomicrus</emphasis>
</taxonomicName>
, makes this genus exceptional within the tribe. The body shape found in this genus is typical for the Cephenniini, with subquadrate pronotum and stout, short elytra, but the constriction between the pronotum and elytra is usually more distinct than in other genera. The antennae show some degree of variability, and since two relatively distinct variants can be found, in future they may be used for defining species groups or subgenera. Antennae in
<taxonomicName id="4C537BAE6543FFBCDC33FF12FE06FEF7" box="[304,505,232,257]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="perpunctillum">
<emphasis id="B927DC3F6543FFBCDC33FF12FE06FEF7" box="[304,505,232,257]" italics="true" pageId="7" pageNumber="32">C. perpunctillum</emphasis>
</taxonomicName>
, the
<typeStatus id="54E8BE8F6543FFBCDF34FF12FD95FEF4" box="[567,618,232,258]" pageId="7" pageNumber="32">type</typeStatus>
species of
<taxonomicName id="4C537BAE6543FFBCDFF1FF12FC46FEF7" box="[754,953,232,257]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6543FFBCDFF1FF12FC46FEF7" box="[754,953,232,257]" italics="true" pageId="7" pageNumber="32">Cephennomicrus</emphasis>
</taxonomicName>
, and in most of other described species, have two terminal antennomeres enlarged, and at first sight the antennal club appears two-segmented. However, closer examination usually reveals that antennomere IX, though clearly smaller than abruptly enlarged X and XI, is still much larger than VII, and can be regarded as belonging to the club. The other variant can be seen in several species, which have more slender antennal club, with antennomeres X and XI enlarged to a lesser extent than those in
<taxonomicName id="4C537BAE6543FFBCDCCFFE4AFD6EFE3F" box="[460,657,432,457]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="perpunctillum">
<emphasis id="B927DC3F6543FFBCDCCFFE4AFD6EFE3F" box="[460,657,432,457]" italics="true" pageId="7" pageNumber="32">C. perpunctillum</emphasis>
</taxonomicName>
. In such cases (like in
<taxonomicName id="4C537BAE6543FFBCDE9FFE4AFA62FE3C" authority="Jaloszynski &amp; Hoshina, 2003" authorityName="Jaloszynski &amp; Hoshina" authorityYear="2003" box="[924,1437,431,458]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="B927DC3F6543FFBCDE9FFE4AFBDEFE3F" box="[924,1057,432,457]" italics="true" pageId="7" pageNumber="32">C. nomurai</emphasis>
<bibRefCitation id="EFC27DDC6543FFBCD92BFE55FA62FE3C" author="Jaloszynski" box="[1064,1437,431,458]" pageId="7" pageNumber="35" refString="Jaloszynski, P. &amp; Hoshina, H. (2003) Four new species of the genus Neseuthia Scott, 1922 (Coleoptera, Scydmaenidae) from Japan. Japanese Journal of Systematic Entomology, 9, 47 - 62." type="journal article" year="2003">(Jałoszyński &amp; Hoshina, 2003))</bibRefCitation>
</taxonomicName>
the club is very indistinctly separated from the funicle, but it still can be regarded as composed of 3 antennomeres. It can be concluded that
<taxonomicName id="4C537BAE6543FFBCDF1DFDFAFD1BFDEF" box="[542,740,512,537]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6543FFBCDF1DFDFAFD1BFDEF" box="[542,740,512,537]" italics="true" pageId="7" pageNumber="32">Cephennomicrus</emphasis>
</taxonomicName>
has 3-segmented antennal club, and the extreme variants taken alone do not represent a diagnostic character to define separate taxa at the rank of subgenus or genus. However, the slender variant may co-occur with the longitudinal median groove on the pronotum found in some described and undescribed species from
<collectingCountry id="F34440BD6543FFBCDFCEFD82FCEFFD64" box="[717,784,632,658]" name="Japan" pageId="7" pageNumber="32">Japan</collectingCountry>
and
<collectingCountry id="F34440BD6543FFBCDE53FD82FC55FD64" box="[848,938,632,658]" name="Taiwan" pageId="7" pageNumber="32">Taiwan</collectingCountry>
; this combination, when confirmed, may serve as a potential subgeneric diagnostic feature.
</paragraph>
<paragraph id="8BEC002D6543FFBCDDC5FD32FE56FC24" blockId="7.[151,1437,152,1138]" pageId="7" pageNumber="32">
The eyes in all species of
<taxonomicName id="4C537BAE6543FFBCDCF8FD32FD3DFD17" box="[507,706,712,737]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6543FFBCDCF8FD32FD3DFD17" box="[507,706,712,737]" italics="true" pageId="7" pageNumber="32">Cephennomicrus</emphasis>
</taxonomicName>
known to the author are very large, strongly protruding laterally from the silhouette of the head, and usually coarsely faceted. This is an interesting character, as miniaturization in the Cephenniini often affects the eyes, which in such cases are usually very small or even entirely missing in minute subterraneous, lightly pigmented species of
<taxonomicName id="4C537BAE6543FFBCDE70FCBAFBD2FCAF" box="[883,1069,832,857]" class="Insecta" family="Staphylinidae" genus="Nanophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6543FFBCDE70FCBAFBD2FCAF" box="[883,1069,832,857]" italics="true" pageId="7" pageNumber="32">Nanophthalmus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C537BAE6543FFBCD938FCBAFB30FCAF" box="[1083,1231,832,857]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6543FFBCD938FCBAFB30FCAF" box="[1083,1231,832,857]" italics="true" pageId="7" pageNumber="32">Cephennium</emphasis>
</taxonomicName>
or
<taxonomicName id="4C537BAE6543FFBCD9F5FCBAFABEFCAF" box="[1270,1345,832,857]" class="Insecta" family="Staphylinidae" genus="Etelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6543FFBCD9F5FCBAFABEFCAF" box="[1270,1345,832,857]" italics="true" pageId="7" pageNumber="32">Etelea</emphasis>
</taxonomicName>
. Oriental species of
<taxonomicName id="4C537BAE6543FFBCDC3BFC92FE00FC77" box="[312,511,872,897]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6543FFBCDC3BFC92FE00FC77" box="[312,511,872,897]" italics="true" pageId="7" pageNumber="32">Cephennomicrus</emphasis>
</taxonomicName>
are rarely collected by methods based on sifting, but more frequently by flight intercept traps. The large eyes composed of a large number of ommatidia may therefore be important for flight-related activities.
</paragraph>
<paragraph id="8BEC002D6543FFBCDDC5FC1AFE31FB84" blockId="7.[151,1437,152,1138]" pageId="7" pageNumber="32">
A high degree of variability can be found in the shapes and structures of the aedeagus among the known species of
<taxonomicName id="4C537BAE6543FFBCDC12FBF2FE28FBD7" box="[273,471,1032,1057]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6543FFBCDC12FBF2FE28FBD7" box="[273,471,1032,1057]" italics="true" pageId="7" pageNumber="32">Cephennomicrus</emphasis>
</taxonomicName>
; the
<typeStatus id="54E8BE8F6543FFBCDF11FBF2FDBAFBD4" box="[530,581,1032,1058]" pageId="7" pageNumber="32">type</typeStatus>
species of the genus and that of
<taxonomicName id="4C537BAE6543FFBCDEBDFBF2FBCCFBD7" box="[958,1075,1032,1057]" class="Insecta" family="Staphylinidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6543FFBCDEBDFBF2FBCCFBD7" box="[958,1075,1032,1057]" italics="true" pageId="7" pageNumber="32">Neseuthia</emphasis>
</taxonomicName>
can be examples of such a difference (
<figureCitation id="13681CA86543FFBCDC02FBCAFE87FBBC" box="[257,376,1072,1098]" captionStart="FIGURES 4 – 9" captionStartId="6.[151,269,1585,1609]" captionTargetBox="[166,1424,383,1555]" captionTargetId="figure@6.[160,1427,374,1561]" captionTargetPageId="6" captionText="FIGURES 4 – 9. Type specimens of Cephennomicrus perpunctillum Reitter and Neseuthia typica Scott. 4, Holotype male of Cephennomicrus perpunctillum (0.80 mm). 5, Lectotype male of Neseuthia typica (0.97 mm). 6, Aedeagus of the holotype of Cephennomicrus perpunctillum in dorsal view (0.17 mm). 7, Aedeagus of the paralectotype of Neseuthia typica in dorsal view (0.15 mm). 8. Set of original labels on the holotype of Cephennomicrus perpunctillum. 9, Set of original labels on the holotype of Neseuthia typica." httpUri="https://zenodo.org/record/274090/files/figure.png" pageId="7" pageNumber="32">Figs. 6, 7</figureCitation>
). Again, a more comprehensive study is necessary to subdivide
<taxonomicName id="4C537BAE6543FFBCD988FBCAFAADFBBF" box="[1163,1362,1072,1097]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B927DC3F6543FFBCD988FBCAFAADFBBF" box="[1163,1362,1072,1097]" italics="true" pageId="7" pageNumber="32">Cephennomicrus</emphasis>
</taxonomicName>
using structures of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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