297 lines
43 KiB
XML
297 lines
43 KiB
XML
<document id="34AE91222A4E3C3EDBECD9D50CBDCF60" ID-DOI="10.5281/zenodo.5374745" ID-ISSN="1638-9395" ID-Zenodo-Dep="5374745" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630276855747" checkinUser="marcus" docAuthor="Märss, Tiiu & Karatajūtē-Talimaa, Valentina" docDate="2002" docId="03FA0B64FF97E03EFC9EA8A49928BF75" docLanguage="en" docName="Geodiversitas.2002.24.2.381-404.pdf" docOrigin="Geodiversitas 24 (2)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Thelodus calvus Märss & Karatajūtē-Talimaa 2002, n. sp." docType="treatment" docVersion="3" lastPageNumber="403" masterDocId="FFC3731CFF84E028FFC5A94F9879BC36" masterDocTitle="Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)" masterLastPageNumber="404" masterPageNumber="381" pageNumber="400" updateTime="1699210438067" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods id="9D5A1EA9D5FE3BBEE7DACA325C4AB0BF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1142F611BC33561762C0B1E88DAD82F6">
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<mods:title id="57907DA19DBB078D1A1EDFF1468B25DF">Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)</mods:title>
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</mods:titleInfo>
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<mods:name id="3B7BC2C9596162791930C9663057769B" type="personal">
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<mods:role id="DA678FEEC4009C21E88DC56EE1E8D101">
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<mods:roleTerm id="9ED2239B53A8D26E0811BB6C9637D93A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A145F76ACDA55FFB88C62C2BBD80E1D7">Märss, Tiiu</mods:namePart>
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<mods:affiliation id="6D88F1518136E56A2E9CA9EE8531439C">Institute of Geology, Tallinn Technical University, 7 Estonia Avenue, Tallinn 10143 (Estonia) marss @ gi. ee</mods:affiliation>
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<mods:nameIdentifier id="F7883E4E47FA4D4FE311BB63D64EBE48" type="email">marss@gi.ee</mods:nameIdentifier>
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</mods:name>
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<mods:name id="C0B7419DDC61B62B8B22D5D8D52EC413" type="personal">
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<mods:role id="CBCE77EE15AF5C3984159338A0CF63B5">
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<mods:roleTerm id="0241EA44404D5319240988B6EB125F16">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1BC8D08218801D8E9C562B1B069BDF20">Karatajūtē-Talimaa, Valentina</mods:namePart>
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<mods:affiliation id="EDCF4FBEE4CBF7D48519FE2632381F34">Institute of Geology of Lithuania, S ˘ evc enkos 13, Vilnius LR 2600 (Lithuania) vtalimaa @ takas. lt Six new and one earlier known thelodont from the Ordovician and Lower Silurian of Severnaya Zemlya Archipelago are described. Scales of the earliest dated thelodont Stroinolepis maenniki n. gen., n. sp. were obtained from the Strojnaya River section, October Revolution Island, upper Ozernaya and Strojnaya formations, Middle-Upper and Upper Ordovician, correspondingly. Loganellia matura n. sp. was found from the Srednii Island, Golomyannyj Formation, middle Llandovery. Paralogania klubovi n. sp. and Loganelliidae gen. et sp. indet. come from the middle (?) Llandovery of Pioneer Island. L. grossi Fredholm, 1990 in the Matusevich River section, Paralogania consimilis n. sp. and Thelodus calvus n. sp. in the Ushakov River section, and Shielia multispinata n. sp., in the Spokojnaya River section, are distributed in the Samojlovich Formation, Wenlock, Lower Silurian of October Revolution Island.</mods:affiliation>
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<mods:nameIdentifier id="45ABD4E8CC85F75C2A93E47E3CF55D3B" type="email">vtalimaa@takas.lt</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="15EB685FCE6DF8DB6E1FC60A19555573">text</mods:typeOfResource>
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<mods:titleInfo id="1A261D0BA7CC325FAFE5587B57D4EB13">
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<mods:title id="5A1036C0647A5500FCFF3997C89C8215">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="FB57705FB9020B33747BD48980A5E5B2">
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<mods:date id="12639B7A56735A421E82E2260F08D36E">2002</mods:date>
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<mods:detail id="7A22623FB851EF464128E924740253F8" type="volume">
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<mods:number id="B25F5EF53F21D212B5C8F4351DA36FCE">24</mods:number>
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</mods:detail>
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<mods:detail id="8894CE796DDBAEBB2EFE55B000BC4993" type="issue">
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<mods:number id="4F87B01C95E58B8B323B57FD8AEE531E">2</mods:number>
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</mods:detail>
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<mods:extent id="AB2EC33B850CEDD7D9C74AA17562A7C2" unit="page">
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<mods:start id="2357D7B77A961E1900D9DE85E8923C19">381</mods:start>
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<mods:end id="CA070F27B95AC04DDD1943101A5862F0">404</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="6F5FFF09E818390220F4855D221DE609">journal article</mods:classification>
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<mods:identifier id="99E7B85190AF25E7135BCB319262639D" type="DOI">10.5281/zenodo.5374745</mods:identifier>
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<mods:identifier id="9F911512D5EF86DE42C7E37962001BA7" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="C8E45131E71312C49A934E5FD2CCDA70" type="Zenodo-Dep">5374745</mods:identifier>
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</mods:mods>
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<treatment id="03FA0B64FF97E03EFC9EA8A49928BF75" LSID="urn:lsid:plazi:treatment:03FA0B64FF97E03EFC9EA8A49928BF75" httpUri="http://treatment.plazi.org/id/03FA0B64FF97E03EFC9EA8A49928BF75" lastPageId="22" lastPageNumber="403" pageId="19" pageNumber="400">
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<subSubSection id="C349E9F9FF97E03BFC9EA8A49C41BE15" pageId="19" pageNumber="400" type="nomenclature">
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<paragraph id="8BECBA72FF97E03BFC9EA8A49C41BE15" blockId="19.[859,1102,491,547]" pageId="19" pageNumber="400">
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<taxonomicName id="4C53C1F1FF97E03BFC9EA8A49C75BE35" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[859,1036,491,515]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="19" pageNumber="400" phylum="Chordata" rank="species" species="calvus" status="sp. nov.">
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<emphasis id="B9276660FF97E03BFC9EA8A49C75BE35" bold="true" box="[859,1036,491,515]" italics="true" pageId="19" pageNumber="400">Thelodus calvus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A214DB1BFF97E03BFBD6A8A39C37BE35" box="[1043,1102,492,515]" pageId="19" pageNumber="400" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="1368A6F7FF97E03BFCBDAB449BC3BE15" box="[888,954,523,547]" captionStart="FIG" captionStartId="7.[151,162,1524,1541]" captionTargetBox="[221,1196,174,1462]" captionTargetId="figure-25@7.[219,1196,174,1462]" captionTargetPageId="7" captionText="FIG. 5. — A-O, Loganellia matura n. sp.; A, B, head scales; C-E, transitional scales; F-M, O, trunk scales; N, fin? scale; A, C, F-H, J-L, N, scales in crown view; B, D, E, M, scales in lateral view; I, a scale in base view; N, posterior half of a trunk? scale in lateral view; A, Pi 7486; B, Pi 7487; C, Pi 7499; D, Pi 7496; E, Pi 7489; F, Pi 7494; G, holotype Pi 7492; H, Pi 7490; I, Pi 7497; J, Pi 7493; K, Pi 7501; L, M, Pi 7495; N, Pi 7485; O, Pi 7500; sample MF 3-8, Srednii Island, Sedov Archipelago, Golomyannyj Formation, middle Llandovery, Lower Silurian; P-S, Thelodus calvus n. sp.; P, LIG 35-907; Q, holotype Pi 7503; R, Pi 7505; S, Pi 7504; P, a scale in crown view; Q, S, in lateral view; R, in base view; P, sample 32a-208; Q-S, sample MF 46-23; Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374755" httpUri="https://zenodo.org/record/5374755/files/figure.png" pageId="19" pageNumber="400">Figs 5</figureCitation>
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P-S; 14; 15)
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</paragraph>
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</subSubSection>
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<subSubSection id="C349E9F9FF97E03BFD0FAB0A9C6BBE6C" box="[714,1042,580,603]" pageId="19" pageNumber="400" type="materials_examined">
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<paragraph id="8BECBA72FF97E03BFD0FAB0A9C6BBE6C" blockId="19.[714,1247,580,1362]" box="[714,1042,580,603]" pageId="19" pageNumber="400">
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<typeStatus id="54E804D0FF97E03BFD0FAB0A9B41BE6C" box="[714,824,581,602]" pageId="19" pageNumber="400" type="holotype">HOLOTYPE</typeStatus>
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. — Pi 7503 (
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<figureCitation id="1368A6F7FF97E03BFC72AB0A9C7FBE6C" box="[951,1030,580,603]" captionStart="FIG" captionStartId="7.[151,162,1524,1541]" captionTargetBox="[221,1196,174,1462]" captionTargetId="figure-25@7.[219,1196,174,1462]" captionTargetPageId="7" captionText="FIG. 5. — A-O, Loganellia matura n. sp.; A, B, head scales; C-E, transitional scales; F-M, O, trunk scales; N, fin? scale; A, C, F-H, J-L, N, scales in crown view; B, D, E, M, scales in lateral view; I, a scale in base view; N, posterior half of a trunk? scale in lateral view; A, Pi 7486; B, Pi 7487; C, Pi 7499; D, Pi 7496; E, Pi 7489; F, Pi 7494; G, holotype Pi 7492; H, Pi 7490; I, Pi 7497; J, Pi 7493; K, Pi 7501; L, M, Pi 7495; N, Pi 7485; O, Pi 7500; sample MF 3-8, Srednii Island, Sedov Archipelago, Golomyannyj Formation, middle Llandovery, Lower Silurian; P-S, Thelodus calvus n. sp.; P, LIG 35-907; Q, holotype Pi 7503; R, Pi 7505; S, Pi 7504; P, a scale in crown view; Q, S, in lateral view; R, in base view; P, sample 32a-208; Q-S, sample MF 46-23; Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374755" httpUri="https://zenodo.org/record/5374755/files/figure.png" pageId="19" pageNumber="400">Fig. 5Q</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C349E9F9FF97E03BFD0FAB229B70BE80" pageId="19" pageNumber="400" type="etymology">
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<paragraph id="8BECBA72FF97E03BFD0FAB229B70BE80" blockId="19.[714,1247,580,1362]" pageId="19" pageNumber="400">
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ETYMOLOGY. — From the Latin word
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<taxonomicName id="4C53C1F1FF97E03BFB8BAB229CF2BEB4" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[1102,1163,621,642]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="19" pageNumber="400" phylum="Chordata" rank="species" species="calvus">
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<emphasis id="B9276660FF97E03BFB8BAB229CF2BEB4" box="[1102,1163,621,642]" italics="true" pageId="19" pageNumber="400">calvus</emphasis>
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</taxonomicName>
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, “bald”, referring to the smooth unsculptured surface of the scales.
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</paragraph>
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</subSubSection>
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<subSubSection id="C349E9F9FF97E03BFD0FAB869B1DB870" pageId="19" pageNumber="400" type="materials_examined">
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<paragraph id="8BECBA72FF97E03BFD0FAB869CB9BF1C" blockId="19.[714,1247,580,1362]" pageId="19" pageNumber="400">
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<collectionCode id="ED4222B7FF97E03BFD0FAB869B85BEEB" box="[714,1020,713,734]" pageId="19" pageNumber="400">TYPE LOCALITY AND HORIZON</collectionCode>
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. — Sample MF 46-23, 32a-208, Ushakov River, October Revolution Island, Severnaya Zemlya Archipelago; upper part of Samojlovich Formation, Wenlock, Lower Silurian.
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</paragraph>
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<paragraph id="8BECBA72FF97E03BFD0FAA729BFDBF5A" blockId="19.[714,1247,580,1362]" pageId="19" pageNumber="400">MATERIAL EXAMINED. — More than 300 scales of good preservation.</paragraph>
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<paragraph id="8BECBA72FF97E03BFD0FAA319B1DB870" blockId="19.[714,1247,580,1362]" pageId="19" pageNumber="400">OCCURRENCE. — Samples MF 46-23, 32a-204, 32a- 208, Ushakov River, and MF 157-7, Spokojnaya River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian; (?)MF 34-1, Cape October, and MF 46-26, Ushakov River, October Revolution Island, Severnaya Zemlya Archipelago, Ust’-Spokojnaya Formation, Ludlow, Upper Silurian.</paragraph>
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</subSubSection>
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<subSubSection id="C349E9F9FF97E03BFD0FAD169BAEB964" pageId="19" pageNumber="400" type="diagnosis">
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<paragraph id="8BECBA72FF97E03BFD0FAD169BAEB964" blockId="19.[714,1247,580,1362]" pageId="19" pageNumber="400">
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DIAGNOSIS. — Scales small; average length
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<quantity id="4CAB1797FF97E03BFB49AD179CA5B858" box="[1164,1244,1112,1134]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="400" unit="mm" value="0.4">0.4 mm</quantity>
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, width
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<quantity id="4CAB1797FF97E03BFCCEAD3D9B2CB8B1" box="[779,853,1138,1159]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="400" unit="mm" value="0.4">0.4 mm</quantity>
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and height
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<quantity id="4CAB1797FF97E03BFC0EAD3D9C60B8B1" box="[971,1049,1138,1159]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="400" unit="mm" value="0.4">0.4 mm</quantity>
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. Scale crowns squarish to oval, smooth and flat or slightly convex. Neck very distinct, rather high, base placed at anterior of scale. Pulp opening in centre or slightly posteriorly in base. Pulp cavity complicated, having openings of dentine canals. Dentine canals occur in base, neck and lower part of crown. Bunches of dentine tubules arise from dentine canals. Both aspidine “pegs” and tubules of Sharpey’s fibres present.
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</paragraph>
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</subSubSection>
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<subSubSection id="C349E9F9FF97E03EFD0FACC59928BF75" lastPageId="22" lastPageNumber="403" pageId="19" pageNumber="400" type="description">
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<paragraph id="8BECBA72FF97E03BFD0FACC59B22B994" blockId="19.[714,1247,1418,1667]" box="[714,859,1418,1444]" pageId="19" pageNumber="400">DESCRIPTION</paragraph>
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<paragraph id="8BECBA72FF97E03BFD0FACE49B3DB9F3" blockId="19.[714,1247,1418,1667]" box="[714,836,1451,1477]" pageId="19" pageNumber="400">
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<emphasis id="B9276660FF97E03BFD0FACE49B3DB9F3" box="[714,836,1451,1477]" italics="true" pageId="19" pageNumber="400">Morphology</emphasis>
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</paragraph>
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<paragraph id="8BECBA72FF97E03CFD0FAC8499CEB955" blockId="19.[714,1247,1418,1667]" lastBlockId="20.[151,684,1131,1379]" lastPageId="20" lastPageNumber="401" pageId="19" pageNumber="400">
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The morphology of the scales is very simple. The length of the scale ranges from
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<quantity id="4CAB1797FF97E03BFBFCACA49CA2BA35" box="[1081,1243,1515,1539]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="9.0" metricValueMin="3.0" pageId="19" pageNumber="400" unit="mm" value="0.6" valueMax="0.9" valueMin="0.3">0.3 to 0.9 mm</quantity>
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, width
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<quantity id="4CAB1797FF97E03BFCD7AF449BE9BA15" box="[786,912,1547,1571]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="19" pageNumber="400" unit="mm" value="0.44999999999999996" valueMax="0.7" valueMin="0.2">0.2-0.7 mm</quantity>
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and height
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<quantity id="4CAB1797FF97E03BFBD0AF449CEBBA15" box="[1045,1170,1547,1571]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="19" pageNumber="400" unit="mm" value="0.4" valueMax="0.6" valueMin="0.2">0.2-0.6 mm</quantity>
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; in the
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<typeStatus id="54E804D0FF97E03BFD0FAF649B4BBA75" box="[714,818,1579,1603]" pageId="19" pageNumber="400" type="holotype">holotype</typeStatus>
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the length/height measurements are 0.4/
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<quantity id="4CAB1797FF97E03BFD36AF049B32BA55" box="[755,843,1611,1635]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="400" unit="mm" value="0.4">0.4 mm</quantity>
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. The crown is mainly rhombic with slightly rounded anterior corner but they also can be oval. The scales reveal no sculpture. The neck is very distinct, smooth and high. The lateral margins of the crown are slightly elevated (
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<figureCitation id="1368A6F7FF90E03CFF5AAD84996EB8D5" box="[159,279,1227,1251]" captionStart="FIG" captionStartId="7.[151,162,1524,1541]" captionTargetBox="[221,1196,174,1462]" captionTargetId="figure-25@7.[219,1196,174,1462]" captionTargetPageId="7" captionText="FIG. 5. — A-O, Loganellia matura n. sp.; A, B, head scales; C-E, transitional scales; F-M, O, trunk scales; N, fin? scale; A, C, F-H, J-L, N, scales in crown view; B, D, E, M, scales in lateral view; I, a scale in base view; N, posterior half of a trunk? scale in lateral view; A, Pi 7486; B, Pi 7487; C, Pi 7499; D, Pi 7496; E, Pi 7489; F, Pi 7494; G, holotype Pi 7492; H, Pi 7490; I, Pi 7497; J, Pi 7493; K, Pi 7501; L, M, Pi 7495; N, Pi 7485; O, Pi 7500; sample MF 3-8, Srednii Island, Sedov Archipelago, Golomyannyj Formation, middle Llandovery, Lower Silurian; P-S, Thelodus calvus n. sp.; P, LIG 35-907; Q, holotype Pi 7503; R, Pi 7505; S, Pi 7504; P, a scale in crown view; Q, S, in lateral view; R, in base view; P, sample 32a-208; Q-S, sample MF 46-23; Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374755" httpUri="https://zenodo.org/record/5374755/files/figure.png" pageId="20" pageNumber="401">Fig. 5Q, S</figureCitation>
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). The base is anteriorly longer than the crown and protrudes forward. The rounded pulp opening is situated in the centre or in the posterior part of the base.
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<figureCitation id="1368A6F7FF90E03CFE72AC649A7BB975" box="[439,514,1323,1347]" captionStart="FIG" captionStartId="20.[151,162,997,1014]" captionTargetBox="[165,1236,176,961]" captionTargetId="figure-271@20.[162,1242,170,973]" captionTargetPageId="20" captionText="FIG. 14. — Set of scales of Thelodus calvus n. sp.; A, B, LIG 35-180; C, D, LIG 35-181; E, LIG 35-182; F, LIG 35-183; G, LIG 35-184; H, LIG 35-185; I, LIG 35-186; J, LIG 35-187; K, L, LIG 35-188; M, LIG 35-191; N, LIG 35-192; O, P, LIG 35-189; Q, LIG 35-190; R, LIG 35-193; sample 32a-208, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5374773" httpUri="https://zenodo.org/record/5374773/files/figure.png" pageId="20" pageNumber="401">Fig. 14</figureCitation>
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exhibits the set of scales of
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<taxonomicName id="4C53C1F1FF90E03CFED0AC03990FB955" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[277,374,1355,1379]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="20" pageNumber="401" phylum="Chordata" rank="species" species="calvus" status="sp. nov.">
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<emphasis id="B9276660FF90E03CFED0AC03990FB955" box="[277,374,1355,1379]" italics="true" pageId="20" pageNumber="401">T. calvus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A214DB1BFF90E03CFEB9AC0399CEB955" box="[380,439,1356,1379]" pageId="20" pageNumber="401" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
|
||
<caption id="DF2CEAFAFF90E03CFF52AAAA9907B800" ID-DOI="http://doi.org/10.5281/zenodo.5374773" ID-Zenodo-Dep="5374773" httpUri="https://zenodo.org/record/5374773/files/figure.png" pageId="20" pageNumber="401" startId="20.[151,162,997,1014]" targetBox="[165,1236,176,961]" targetPageId="20">
|
||
<paragraph id="8BECBA72FF90E03CFF52AAAA9907B800" blockId="20.[151,1249,997,1078]" pageId="20" pageNumber="401">
|
||
FIG. 14. — Set of scales of
|
||
<taxonomicName id="4C53C1F1FF90E03CFEB2AAAA9985BFC0" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[375,508,997,1014]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="20" pageNumber="401" phylum="Chordata" rank="species" species="calvus" status="sp. nov.">
|
||
<emphasis id="B9276660FF90E03CFEB2AAAA9985BFC0" box="[375,508,997,1014]" italics="true" pageId="20" pageNumber="401">Thelodus calvus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A214DB1BFF90E03CFDC4AAAA9A4ABFC0" box="[513,563,997,1014]" pageId="20" pageNumber="401" rank="species">n. sp.</taxonomicNameLabel>
|
||
;
|
||
<emphasis id="B9276660FF90E03CFDFFAAAA9A3EBFC0" bold="true" box="[570,583,997,1014]" pageId="20" pageNumber="401">A</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF90E03CFD94AAAA9A27BFC0" bold="true" box="[593,606,997,1014]" pageId="20" pageNumber="401">B</emphasis>
|
||
, LIG 35-180;
|
||
<emphasis id="B9276660FF90E03CFD15AAAA9AA7BFC0" bold="true" box="[720,734,997,1014]" pageId="20" pageNumber="401">C</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF90E03CFD2DAAAA9A8FBFC0" bold="true" box="[744,758,997,1014]" pageId="20" pageNumber="401">D</emphasis>
|
||
, LIG 35-181;
|
||
<emphasis id="B9276660FF90E03CFCADAAAA9B0DBFC0" bold="true" box="[872,884,997,1014]" pageId="20" pageNumber="401">E</emphasis>
|
||
, LIG 35-182;
|
||
<emphasis id="B9276660FF90E03CFC22AAAA9B8BBFC0" bold="true" box="[999,1010,997,1014]" pageId="20" pageNumber="401">F</emphasis>
|
||
, LIG 35-183;
|
||
<emphasis id="B9276660FF90E03CFBA1AAAA9C0BBFC0" bold="true" box="[1124,1138,997,1014]" pageId="20" pageNumber="401">G</emphasis>
|
||
, LIG 35-184;
|
||
<emphasis id="B9276660FF90E03CFF52AAB498DCB83A" bold="true" box="[151,165,1019,1036]" pageId="20" pageNumber="401">H</emphasis>
|
||
, LIG 35-185;
|
||
<emphasis id="B9276660FF90E03CFED9AAB4995BB83A" bold="true" box="[284,290,1019,1036]" pageId="20" pageNumber="401">I</emphasis>
|
||
, LIG 35-186;
|
||
<emphasis id="B9276660FF90E03CFE5DAAB499DBB83A" bold="true" box="[408,418,1019,1036]" pageId="20" pageNumber="401">J</emphasis>
|
||
, LIG 35-187;
|
||
<emphasis id="B9276660FF90E03CFDDFAAB49A5EB83A" bold="true" box="[538,551,1019,1036]" pageId="20" pageNumber="401">K</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF90E03CFDF6AAB49A47B83A" bold="true" box="[563,574,1019,1036]" pageId="20" pageNumber="401">L</emphasis>
|
||
, LIG 35-188;
|
||
<emphasis id="B9276660FF90E03CFD70AAB49ABFB83A" bold="true" box="[693,710,1019,1036]" pageId="20" pageNumber="401">M</emphasis>
|
||
, LIG 35-191;
|
||
<emphasis id="B9276660FF90E03CFCF8AAB49B32B83A" bold="true" box="[829,843,1019,1036]" pageId="20" pageNumber="401">N</emphasis>
|
||
, LIG 35-192;
|
||
<emphasis id="B9276660FF90E03CFC07AAB49BA8B83A" bold="true" box="[962,977,1019,1036]" pageId="20" pageNumber="401">O</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF90E03CFC18AAB49B90B83A" bold="true" box="[989,1001,1019,1036]" pageId="20" pageNumber="401">P</emphasis>
|
||
, LIG 35-189;
|
||
<emphasis id="B9276660FF90E03CFBA5AAB49C16B83A" bold="true" box="[1120,1135,1019,1036]" pageId="20" pageNumber="401">Q</emphasis>
|
||
, LIG 35-190;
|
||
<emphasis id="B9276660FF90E03CFF52AD5F98DDB817" bold="true" box="[151,164,1040,1057]" pageId="20" pageNumber="401">R</emphasis>
|
||
, LIG 35-193; sample 32a-208, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Scale bar: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BECBA72FF90E03CFF52ACC49923B992" blockId="20.[151,684,1418,1667]" box="[151,346,1418,1445]" pageId="20" pageNumber="401">
|
||
<emphasis id="B9276660FF90E03CFF52ACC49923B992" box="[151,346,1418,1445]" italics="true" pageId="20" pageNumber="401">
|
||
Histology (
|
||
<figureCitation id="1368A6F7FF90E03CFEC0ACC5992BB992" box="[261,338,1418,1444]" captionStart="FIG" captionStartId="21.[151,162,1588,1605]" captionTargetBox="[178,1212,199,1541]" captionTargetId="figure-25@21.[161,1212,193,1554]" captionTargetPageId="21" captionText="FIG. 15. — Thelodus calvus n. sp., microstructure of the scales; A, Pi 7631; B, Pi 7632; C, D, Pi 7633; E, Pi 7634; F, Pi 7635; G, Pi 7636; sample MF 46-23, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; cr, crown; dc, dentine canal; dt, dentine tubules; n, neck, pc, pulp canal; pcv, pulp cavity. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374775" httpUri="https://zenodo.org/record/5374775/files/figure.png" pageId="20" pageNumber="401">Fig. 15</figureCitation>
|
||
)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BECBA72FF90E03CFF52ACE49BBCBA35" blockId="20.[151,684,1418,1667]" lastBlockId="20.[714,1248,1131,1539]" pageId="20" pageNumber="401">
|
||
The pulp cavity continues posteriorly as a pulp canal at a right angle from pulp cavity (
|
||
<figureCitation id="1368A6F7FF90E03CFD87AC849AD9B9D5" box="[578,672,1483,1507]" captionStart="FIG" captionStartId="21.[151,162,1588,1605]" captionTargetBox="[178,1212,199,1541]" captionTargetId="figure-25@21.[161,1212,193,1554]" captionTargetPageId="21" captionText="FIG. 15. — Thelodus calvus n. sp., microstructure of the scales; A, Pi 7631; B, Pi 7632; C, D, Pi 7633; E, Pi 7634; F, Pi 7635; G, Pi 7636; sample MF 46-23, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; cr, crown; dc, dentine canal; dt, dentine tubules; n, neck, pc, pulp canal; pcv, pulp cavity. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374775" httpUri="https://zenodo.org/record/5374775/files/figure.png" pageId="20" pageNumber="401">Fig. 15F</figureCitation>
|
||
). The walls of the pulp cavity are uneven because of openings of dentine canals (
|
||
<figureCitation id="1368A6F7FF90E03CFDCBAF449AE4BA15" box="[526,669,1547,1571]" captionStart="FIG" captionStartId="21.[151,162,1588,1605]" captionTargetBox="[178,1212,199,1541]" captionTargetId="figure-25@21.[161,1212,193,1554]" captionTargetPageId="21" captionText="FIG. 15. — Thelodus calvus n. sp., microstructure of the scales; A, Pi 7631; B, Pi 7632; C, D, Pi 7633; E, Pi 7634; F, Pi 7635; G, Pi 7636; sample MF 46-23, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; cr, crown; dc, dentine canal; dt, dentine tubules; n, neck, pc, pulp canal; pcv, pulp cavity. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374775" httpUri="https://zenodo.org/record/5374775/files/figure.png" pageId="20" pageNumber="401">Fig. 15A, B</figureCitation>
|
||
). Dentine canals that start in the cavity enter into the base, anterior portion of the crown (
|
||
<figureCitation id="1368A6F7FF90E03CFF5AAF24997ABAB5" box="[159,259,1643,1667]" captionStart="FIG" captionStartId="21.[151,162,1588,1605]" captionTargetBox="[178,1212,199,1541]" captionTargetId="figure-25@21.[161,1212,193,1554]" captionTargetPageId="21" captionText="FIG. 15. — Thelodus calvus n. sp., microstructure of the scales; A, Pi 7631; B, Pi 7632; C, D, Pi 7633; E, Pi 7634; F, Pi 7635; G, Pi 7636; sample MF 46-23, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; cr, crown; dc, dentine canal; dt, dentine tubules; n, neck, pc, pulp canal; pcv, pulp cavity. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374775" httpUri="https://zenodo.org/record/5374775/files/figure.png" pageId="20" pageNumber="401">Fig. 15A</figureCitation>
|
||
) and the neck (
|
||
<figureCitation id="1368A6F7FF90E03CFE72AF249A62BAB5" box="[439,539,1643,1667]" captionStart="FIG" captionStartId="21.[151,162,1588,1605]" captionTargetBox="[178,1212,199,1541]" captionTargetId="figure-25@21.[161,1212,193,1554]" captionTargetPageId="21" captionText="FIG. 15. — Thelodus calvus n. sp., microstructure of the scales; A, Pi 7631; B, Pi 7632; C, D, Pi 7633; E, Pi 7634; F, Pi 7635; G, Pi 7636; sample MF 46-23, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; cr, crown; dc, dentine canal; dt, dentine tubules; n, neck, pc, pulp canal; pcv, pulp cavity. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374775" httpUri="https://zenodo.org/record/5374775/files/figure.png" pageId="20" pageNumber="401">Fig. 15B</figureCitation>
|
||
). The posterior region of the neck has very few canals if any. Numerous dentine tubules are distributed as bunches going out from the dentine canals. Dentine tubules in the posterior crown portion are directed upwards and backwards, then they turn anteriorly and again posteriorly, before running straight up towards the crown surface (
|
||
<figureCitation id="1368A6F7FF90E03CFD17AC049B4EB955" box="[722,823,1355,1379]" captionStart="FIG" captionStartId="21.[151,162,1588,1605]" captionTargetBox="[178,1212,199,1541]" captionTargetId="figure-25@21.[161,1212,193,1554]" captionTargetPageId="21" captionText="FIG. 15. — Thelodus calvus n. sp., microstructure of the scales; A, Pi 7631; B, Pi 7632; C, D, Pi 7633; E, Pi 7634; F, Pi 7635; G, Pi 7636; sample MF 46-23, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; cr, crown; dc, dentine canal; dt, dentine tubules; n, neck, pc, pulp canal; pcv, pulp cavity. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374775" httpUri="https://zenodo.org/record/5374775/files/figure.png" pageId="20" pageNumber="401">Fig. 15B</figureCitation>
|
||
). Horizontal thin sections (
|
||
<figureCitation id="1368A6F7FF90E03CFBB2AC049A99B9B5" captionStart="FIG" captionStartId="21.[151,162,1588,1605]" captionTargetBox="[178,1212,199,1541]" captionTargetId="figure-25@21.[161,1212,193,1554]" captionTargetPageId="21" captionText="FIG. 15. — Thelodus calvus n. sp., microstructure of the scales; A, Pi 7631; B, Pi 7632; C, D, Pi 7633; E, Pi 7634; F, Pi 7635; G, Pi 7636; sample MF 46-23, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; cr, crown; dc, dentine canal; dt, dentine tubules; n, neck, pc, pulp canal; pcv, pulp cavity. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374775" httpUri="https://zenodo.org/record/5374775/files/figure.png" pageId="20" pageNumber="401">Fig. 15E, G</figureCitation>
|
||
) of the uppermost part of the crown, show rather straight dentine tubules. In a little deeper section (
|
||
<figureCitation id="1368A6F7FF90E03CFCE2ACE49BD2B9F5" box="[807,939,1451,1475]" captionStart="FIG" captionStartId="21.[151,162,1588,1605]" captionTargetBox="[178,1212,199,1541]" captionTargetId="figure-25@21.[161,1212,193,1554]" captionTargetPageId="21" captionText="FIG. 15. — Thelodus calvus n. sp., microstructure of the scales; A, Pi 7631; B, Pi 7632; C, D, Pi 7633; E, Pi 7634; F, Pi 7635; G, Pi 7636; sample MF 46-23, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; cr, crown; dc, dentine canal; dt, dentine tubules; n, neck, pc, pulp canal; pcv, pulp cavity. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374775" httpUri="https://zenodo.org/record/5374775/files/figure.png" pageId="20" pageNumber="401">Fig. 15C, D</figureCitation>
|
||
) dentine canals are also visible. In the base, aspidine “pegs” and tubules of Sharpey’s fibres occur.
|
||
</paragraph>
|
||
<paragraph id="8BECBA72FF90E03CFD0FAF649B23BA74" blockId="20.[714,1247,1579,1667]" box="[714,858,1579,1605]" pageId="20" pageNumber="401">COMPARISON</paragraph>
|
||
<paragraph id="8BECBA72FF90E03CFD0FAF049CA6BAB5" blockId="20.[714,1247,1579,1667]" pageId="20" pageNumber="401">
|
||
Scales of
|
||
<taxonomicName id="4C53C1F1FF90E03CFCEAAF039BE8BA55" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[815,913,1611,1635]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="20" pageNumber="401" phylum="Chordata" rank="species" species="calvus" status="sp. nov.">
|
||
<emphasis id="B9276660FF90E03CFCEAAF039BE8BA55" box="[815,913,1611,1635]" italics="true" pageId="20" pageNumber="401">T. calvus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A214DB1BFF90E03CFC52AF039BABBA55" box="[919,978,1612,1635]" pageId="20" pageNumber="401" rank="species">n. sp.</taxonomicNameLabel>
|
||
are to some extent similar to smooth flat small scales of
|
||
<emphasis id="B9276660FF90E03CFBF5AF249CA6BAB5" box="[1072,1247,1643,1667]" italics="true" pageId="20" pageNumber="401">
|
||
<taxonomicName id="4C53C1F1FF90E03CFBF5AF249CECBAB5" authorityName="Agassiz" authorityYear="1839" box="[1072,1173,1643,1667]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="20" pageNumber="401" phylum="Chordata" rank="genus">Thelodus</taxonomicName>
|
||
parvi-
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BECBA72FF91E03DFD97A9E09A1DBCFE" blockId="21.[594,612,175,200]" box="[594,612,175,200]" pageId="21" pageNumber="402">B</paragraph>
|
||
<caption id="DF2CEAFAFF91E03DFF52AF7B9924BAB3" ID-DOI="http://doi.org/10.5281/zenodo.5374775" ID-Zenodo-Dep="5374775" httpUri="https://zenodo.org/record/5374775/files/figure.png" pageId="21" pageNumber="402" startId="21.[151,162,1588,1605]" targetBox="[178,1212,199,1541]" targetPageId="21">
|
||
<paragraph id="8BECBA72FF91E03DFF52AF7B9924BAB3" blockId="21.[151,1249,1588,1669]" pageId="21" pageNumber="402">
|
||
FIG. 15. —
|
||
<taxonomicName id="4C53C1F1FF91E03DFF36AF7B9903BA73" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[243,378,1588,1605]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="21" pageNumber="402" phylum="Chordata" rank="species" species="calvus" status="sp. nov.">
|
||
<emphasis id="B9276660FF91E03DFF36AF7B9903BA73" box="[243,378,1588,1605]" italics="true" pageId="21" pageNumber="402">Thelodus calvus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A214DB1BFF91E03DFE45AF7B99CABA73" box="[384,435,1588,1605]" pageId="21" pageNumber="402" rank="species">n. sp.</taxonomicNameLabel>
|
||
, microstructure of the scales;
|
||
<emphasis id="B9276660FF91E03DFD75AF7B9AC4BA73" bold="true" box="[688,701,1588,1605]" pageId="21" pageNumber="402">A</emphasis>
|
||
, Pi 7631;
|
||
<emphasis id="B9276660FF91E03DFCD7AF7B9B66BA73" bold="true" box="[786,799,1588,1605]" pageId="21" pageNumber="402">B</emphasis>
|
||
, Pi 7632;
|
||
<emphasis id="B9276660FF91E03DFCB0AF7B9BFABA73" bold="true" box="[885,899,1588,1605]" pageId="21" pageNumber="402">C</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF91E03DFC4BAF7B9BE5BA73" bold="true" box="[910,924,1588,1605]" pageId="21" pageNumber="402">D</emphasis>
|
||
, Pi 7633;
|
||
<emphasis id="B9276660FF91E03DFC37AF7B9B87BA73" bold="true" box="[1010,1022,1588,1605]" pageId="21" pageNumber="402">E</emphasis>
|
||
, Pi 7634;
|
||
<emphasis id="B9276660FF91E03DFB91AF7B9C26BA73" bold="true" box="[1108,1119,1588,1605]" pageId="21" pageNumber="402">F</emphasis>
|
||
, Pi 7635;
|
||
<emphasis id="B9276660FF91E03DFB70AF7B9CBABA73" bold="true" box="[1205,1219,1588,1605]" pageId="21" pageNumber="402">G</emphasis>
|
||
, Pi 7636; sample MF 46-23, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations:
|
||
<emphasis id="B9276660FF91E03DFF52AF1098CEBA46" bold="true" box="[151,183,1631,1648]" pageId="21" pageNumber="402">asp</emphasis>
|
||
, aspidine “pegs” in the base;
|
||
<emphasis id="B9276660FF91E03DFE76AF1099C7BA46" bold="true" box="[435,446,1631,1648]" pageId="21" pageNumber="402">b</emphasis>
|
||
, base;
|
||
<emphasis id="B9276660FF91E03DFE39AF109A77BA46" bold="true" box="[508,526,1631,1648]" pageId="21" pageNumber="402">cr</emphasis>
|
||
, crown;
|
||
<emphasis id="B9276660FF91E03DFD92AF109A14BA46" bold="true" box="[599,621,1631,1648]" pageId="21" pageNumber="402">dc</emphasis>
|
||
, dentine canal;
|
||
<emphasis id="B9276660FF91E03DFD36AF109B7CBA46" bold="true" box="[755,773,1631,1648]" pageId="21" pageNumber="402">dt</emphasis>
|
||
, dentine tubules;
|
||
<emphasis id="B9276660FF91E03DFC5EAF109BDFBA46" bold="true" box="[923,934,1631,1648]" pageId="21" pageNumber="402">n</emphasis>
|
||
, neck,
|
||
<emphasis id="B9276660FF91E03DFC26AF109B80BA46" bold="true" box="[995,1017,1631,1648]" pageId="21" pageNumber="402">pc</emphasis>
|
||
, pulp canal;
|
||
<emphasis id="B9276660FF91E03DFBA0AF109CFCBA46" bold="true" box="[1125,1157,1631,1648]" pageId="21" pageNumber="402">pcv</emphasis>
|
||
, pulp cavity. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BECBA72FF92E03EFF52A9E498A2BD95" blockId="22.[151,684,171,419]" pageId="22" pageNumber="403">
|
||
<emphasis id="B9276660FF92E03EFF52A9E498B5BCF5" box="[151,204,171,195]" italics="true" pageId="22" pageNumber="403">dens</emphasis>
|
||
Agassiz, 1839. They differ from other
|
||
<taxonomicName id="4C53C1F1FF92E03EFF52A9849910BCD5" box="[151,361,203,227]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" kingdom="Animalia" order="Phlebolepidiformes" pageId="22" pageNumber="403" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B9276660FF92E03EFF52A9849971BCD5" box="[151,264,203,227]" italics="true" pageId="22" pageNumber="403">Thelodus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in lacking any sculpture. Among
|
||
<taxonomicName id="4C53C1F1FF92E03EFF30A9A499CFBD35" box="[245,438,235,259]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" kingdom="Animalia" order="Phlebolepidiformes" pageId="22" pageNumber="403" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B9276660FF92E03EFF30A9A49925BD35" box="[245,348,235,259]" italics="true" pageId="22" pageNumber="403">Thelodus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, dentine canals occur only in
|
||
<taxonomicName id="4C53C1F1FF92E03EFF32A843991BBD15" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[247,354,267,291]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="22" pageNumber="403" phylum="Chordata" rank="species" species="calvus" status="sp. nov.">
|
||
<emphasis id="B9276660FF92E03EFF32A843991BBD15" box="[247,354,267,291]" italics="true" pageId="22" pageNumber="403">T. calvus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A214DB1BFF92E03EFEA9A84399D5BD15" box="[364,428,268,291]" pageId="22" pageNumber="403" rank="species">n. sp.</taxonomicNameLabel>
|
||
Similar scales with a smooth crown and high smooth neck have been found in the Wenlock of East Baltic (see e.g.,
|
||
<bibRefCitation id="EFC2C783FF92E03EFF52A8239965BDB5" author="MARSS T." box="[151,284,363,387]" pageId="22" pageNumber="403" refId="ref11422" refString="MARSS T. 1986. - Pozvonochnye silura Estoniji Zapadnoj Latviji [= Silurian vertebrates of Estonia and West Latvia]. Fossilia Baltica 1, 104 p. (in Russian)." type="book" year="1986">Märss 1986</bibRefCitation>
|
||
; Karatajūtē- Talimaa & Brazauskas 1995).
|
||
</paragraph>
|
||
<paragraph id="8BECBA72FF92E03EFE93A8849998BE35" blockId="22.[342,493,459,515]" pageId="22" pageNumber="403">
|
||
<taxonomicName id="4C53C1F1FF92E03EFE93A88499C2BDD5" authorityName="Agassiz" authorityYear="1839" box="[342,443,459,483]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="22" pageNumber="403" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9276660FF92E03EFE93A88499C2BDD5" box="[342,443,459,483]" italics="true" pageId="22" pageNumber="403">Thelodus</emphasis>
|
||
</taxonomicName>
|
||
? sp. (
|
||
<figureCitation id="1368A6F7FF92E03EFEAFA8A499D0BE35" box="[362,425,491,515]" captionStart="FIG" captionStartId="10.[151,162,1524,1541]" captionTargetBox="[203,1186,173,1467]" captionTargetId="figure-21@10.[203,1186,173,1467]" captionTargetPageId="10" captionText="FIG. 7. — A-S, Shielia multispinata n. sp.; A, head scale; B-D, F, transitional scales; E, G-P, trunk scales; Q-S, fin scales; A, B, scales in lateral view; C-N, P, Q, S, scales in crown view; O, a scale in oblique view; R, a scale in base view; A, Pi 7517; B, Pi 7519; C, Pi 7518; D, Pi 7520; E, Pi 7521; F, Pi 6804; G, Pi 7523; H, Pi 7522; I, Pi 7524; J, Pi 7525; K, holotype Pi 6803; L, Pi 6805; M, Pi 6801; N, O, Pi 7526; P, Pi 6802; Q, Pi 7527; R, Pi 7529; S, Pi 7528; sample MF 157-7, Spokojnaya River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian; T-V, Thelodus? sp., scales in crown view; T, Pi 7516; U, LIG 35 908; V, Pi 6814; T, V, sample MF 46-23; U, sample 32a-208, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: A-I, K-M, P-V, 0.1 mm; J, N, O, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5374759" httpUri="https://zenodo.org/record/5374759/files/figure.png" pageId="22" pageNumber="403">Fig. 7</figureCitation>
|
||
T-V)
|
||
</paragraph>
|
||
<paragraph id="8BECBA72FF92E03EFF52AB649928BF75" blockId="22.[151,684,555,835]" pageId="22" pageNumber="403">
|
||
Ridged scales without spines (
|
||
<figureCitation id="1368A6F7FF92E03EFE34AB649A4EBE75" box="[497,567,555,579]" captionStart="FIG" captionStartId="10.[151,162,1524,1541]" captionTargetBox="[203,1186,173,1467]" captionTargetId="figure-21@10.[203,1186,173,1467]" captionTargetPageId="10" captionText="FIG. 7. — A-S, Shielia multispinata n. sp.; A, head scale; B-D, F, transitional scales; E, G-P, trunk scales; Q-S, fin scales; A, B, scales in lateral view; C-N, P, Q, S, scales in crown view; O, a scale in oblique view; R, a scale in base view; A, Pi 7517; B, Pi 7519; C, Pi 7518; D, Pi 7520; E, Pi 7521; F, Pi 6804; G, Pi 7523; H, Pi 7522; I, Pi 7524; J, Pi 7525; K, holotype Pi 6803; L, Pi 6805; M, Pi 6801; N, O, Pi 7526; P, Pi 6802; Q, Pi 7527; R, Pi 7529; S, Pi 7528; sample MF 157-7, Spokojnaya River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian; T-V, Thelodus? sp., scales in crown view; T, Pi 7516; U, LIG 35 908; V, Pi 6814; T, V, sample MF 46-23; U, sample 32a-208, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: A-I, K-M, P-V, 0.1 mm; J, N, O, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5374759" httpUri="https://zenodo.org/record/5374759/files/figure.png" pageId="22" pageNumber="403">Figs 7</figureCitation>
|
||
T-V) have been found in the Samojlovich Formation, Wenlock, Lower Silurian, Ushakov River section on October Revolution Island, and have been identified as
|
||
<taxonomicName id="4C53C1F1FF92E03EFEA8ABE499AFBEF5" authorityName="Agassiz" authorityYear="1839" box="[365,470,683,707]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="22" pageNumber="403" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9276660FF92E03EFEA8ABE499AFBEF5" box="[365,470,683,707]" italics="true" pageId="22" pageNumber="403">Thelodus</emphasis>
|
||
</taxonomicName>
|
||
? sp. in this work. The possibility cannot be excluded that such scales might also belong to
|
||
<emphasis id="B9276660FF92E03EFE1CABA49A12BF35" box="[473,619,747,771]" italics="true" pageId="22" pageNumber="403">
|
||
<taxonomicName id="4C53C1F1FF92E03EFE1CABA49A1EBF35" authorityName="Karatajute-Talimaa" authorityYear="1997" box="[473,615,747,771]" class="Pteraspidomorphi" family="Loganelliidae" genus="Paralogania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="22" pageNumber="403" phylum="Chordata" rank="genus">Paralogania</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Both
|
||
<taxonomicName id="4C53C1F1FF92E03EFF52AA449887BF15" authorityName="Agassiz" authorityYear="1839" box="[151,254,779,803]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="22" pageNumber="403" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9276660FF92E03EFF52AA449887BF15" box="[151,254,779,803]" italics="true" pageId="22" pageNumber="403">Thelodus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C53C1F1FF92E03EFEFDAA4499C6BF15" authorityName="Karatajute-Talimaa" authorityYear="1997" box="[312,447,779,803]" class="Pteraspidomorphi" family="Loganelliidae" genus="Paralogania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="22" pageNumber="403" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9276660FF92E03EFEFDAA4499C6BF15" box="[312,447,779,803]" italics="true" pageId="22" pageNumber="403">Paralogania</emphasis>
|
||
</taxonomicName>
|
||
scales are present in the same sample.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |