198 lines
16 KiB
XML
198 lines
16 KiB
XML
<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. & Jansa, Sharon A." docDate="2022" docId="03EFDD5DF7046916DB28FDF01994FDE4" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.7:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="7" docTitle="Thylophorops chapalmalensis Ameghino 1908" docType="treatment" docVersion="7" lastPageNumber="319" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="319" updateTime="1661891901480" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
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<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. & School of Biological, Earth & Environmental Sciences University of New South Wales, Australia & Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Voss, Robert S.</mods:namePart>
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<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jansa, Sharon A.</mods:namePart>
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<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>2022</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>457</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>353</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974440" ID-Zenodo-Dep="6974440" LSID="urn:lsid:plazi:treatment:03EFDD5DF7046916DB28FDF01994FDE4" httpUri="http://treatment.plazi.org/id/03EFDD5DF7046916DB28FDF01994FDE4" lastPageNumber="319" pageId="319" pageNumber="319">
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<subSubSection box="[291,454,605,630]" pageId="319" pageNumber="319" type="multiple">
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<paragraph blockId="319.[291,454,605,630]" box="[291,454,605,630]" pageId="319" pageNumber="319">
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<heading box="[291,454,605,630]" centered="true" fontSize="10" level="2" pageId="319" pageNumber="319" reason="4">
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<emphasis bold="true" box="[291,454,605,630]" pageId="319" pageNumber="319">
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†
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<taxonomicName box="[303,454,605,629]" class="Mammalia" family="Didelphidae" genus="Thylophorops" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="species" species="undetermined">
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<emphasis bold="true" box="[303,454,605,629]" italics="true" pageId="319" pageNumber="319">Thylophorops</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="319" pageNumber="319" type="nomenclature">
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<paragraph blockId="319.[108,637,655,1738]" pageId="319" pageNumber="319">
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SPECIES SCORED: †
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<taxonomicName authorityName="Ameghino" authorityYear="1908" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="species" species="chapalmalensis">
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<emphasis italics="true" pageId="319" pageNumber="319">Thylophorops chapalmalensis</emphasis>
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</taxonomicName>
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(
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<typeStatus box="[149,198,689,713]" pageId="319" pageNumber="319">type</typeStatus>
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species).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="319" pageNumber="319" type="materials_examined">
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<paragraph blockId="319.[108,637,655,1738]" pageId="319" pageNumber="319">
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GEOLOGICAL PROVENANCE OF SCORED
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<collectionCode box="[108,168,755,779]" pageId="319" pageNumber="319">SPECIMENS</collectionCode>
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: “Barranca Parodi,” Chapadmalal Formation; but several specimens that we examined to score this taxon lack locality data.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="319" pageNumber="319" type="description">
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<paragraph blockId="319.[108,637,655,1738]" pageId="319" pageNumber="319">
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AGE OF SCORED SPECIMENS: As noted above (in the account for †
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<taxonomicName baseAuthorityName="Beck and Taglioretti" baseAuthorityYear="2020" box="[328,480,887,910]" class="Mammalia" family="Didelphidae" genus="Sparassocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[328,480,887,910]" italics="true" pageId="319" pageNumber="319">Sparassocynus</emphasis>
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</taxonomicName>
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), the Chapadmalal Formation is thought to fall within the interval from 3.3 ± 0.2 Mya to about 5 Mya.
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</paragraph>
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<paragraph blockId="319.[108,637,655,1738]" box="[140,595,986,1011]" pageId="319" pageNumber="319">
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ASSIGNED AGE
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<collectionCode box="[311,388,986,1010]" pageId="319" pageNumber="319">RANGE</collectionCode>
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:
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<geoCoordinate box="[402,461,986,1010]" degrees="5.000" direction="north" orientation="latitude" pageId="319" pageNumber="319" precision="55" value="5.0">5.000</geoCoordinate>
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<geoCoordinate box="[461,533,986,1010]" degrees="3.100" direction="west" orientation="longitude" pageId="319" pageNumber="319" precision="55" value="-3.1">–3.100</geoCoordinate>
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Mya.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="319" pageNumber="319" type="discussion">
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<paragraph blockId="319.[108,637,655,1738]" pageId="319" pageNumber="319">
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REMARKS: †
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<taxonomicName authorityName="Ameghino" authorityYear="1908" box="[274,588,1019,1043]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="species" species="chapalmalensis">
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<emphasis box="[274,588,1019,1043]" italics="true" pageId="319" pageNumber="319">Thylophorops chapalmalensis</emphasis>
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</taxonomicName>
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was originally described as
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<taxonomicName box="[356,461,1052,1076]" class="Mammalia" family="Didelphidae" genus="Didelphys" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[356,461,1052,1076]" italics="true" pageId="319" pageNumber="319">Didelphys</emphasis>
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</taxonomicName>
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†
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<emphasis box="[480,636,1052,1076]" italics="true" pageId="319" pageNumber="319">chapalmalense</emphasis>
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by Ameghino (1908), but it was referred to the newly erected genus †
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<taxonomicName box="[381,529,1118,1142]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[381,529,1118,1142]" italics="true" pageId="319" pageNumber="319">Thylophorops</emphasis>
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</taxonomicName>
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by Reig (1952). Goin and Pardiñas (1996) subsequently referred a second species—originally described by Ameghino (1904) as
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<emphasis box="[378,404,1218,1241]" italics="true" pageId="319" pageNumber="319">D.</emphasis>
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†
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<emphasis box="[423,519,1218,1242]" italics="true" pageId="319" pageNumber="319">perplana</emphasis>
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—to †
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<taxonomicName class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="319" pageNumber="319">Thylophorops</emphasis>
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</taxonomicName>
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. Goin et al. (2009b) described a third species, †
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<taxonomicName box="[225,369,1284,1308]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="species" species="lorenzinii">
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<emphasis box="[225,369,1284,1308]" italics="true" pageId="319" pageNumber="319">T. lorenzinii</emphasis>
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</taxonomicName>
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, which is the largest known didelphid described to date, with an estimated body mass of
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<quantity box="[347,465,1350,1374]" metricMagnitude="0" metricUnit="kg" metricValue="6.1" metricValueMax="7.4" metricValueMin="4.8" pageId="319" pageNumber="319" unit="kg" value="6.1" valueMax="7.4" valueMin="4.8">4.8–7.4 kg</quantity>
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. †
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<taxonomicName authorityName="Ameghino" authorityYear="1908" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="species" species="chapalmalensis">
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<emphasis italics="true" pageId="319" pageNumber="319">Thylophorops chapalmalensis</emphasis>
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</taxonomicName>
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, the only species we examined to score character data, had an estimated body mass of
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<quantity box="[204,319,1449,1473]" metricMagnitude="0" metricUnit="kg" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="319" pageNumber="319" unit="kg" value="3.25" valueMax="3.4" valueMin="3.1">3.1–3.4 kg</quantity>
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(Goin et al., 2009b), which overlaps the known range of body mass for the largest living didelphid,
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<taxonomicName authorityName="Kerr" authorityYear="1792" box="[370,587,1515,1539]" class="Mammalia" family="Didelphidae" genus="Didelphis" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="species" species="virginiana">
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<emphasis box="[370,587,1515,1539]" italics="true" pageId="319" pageNumber="319">Didelphis virginiana</emphasis>
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</taxonomicName>
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(see Jones et al., 2009).
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</paragraph>
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<paragraph blockId="319.[108,637,655,1738]" lastBlockId="319.[684,1212,226,547]" pageId="319" pageNumber="319">
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Most authors (Simpson, 1972; Reig et al., 1987; Goin, 1991; Goin et al., 2009b; Voss and Jansa, 2009) have considered †
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<taxonomicName box="[459,604,1647,1671]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[459,604,1647,1671]" italics="true" pageId="319" pageNumber="319">Thylophorops</emphasis>
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</taxonomicName>
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to belong to the extant didelphid tribe
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<taxonomicName authority="sensu Voss and Jansa (2009)" authorityName="sensu Voss and Jansa" authorityYear="2009" class="Mammalia" family="Didelphidae" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="tribe" tribe="Didelphini">Didelphini sensu Voss and Jansa (2009)</taxonomicName>
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, which also includes
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<taxonomicName authorityName="Illiger" authorityYear="1811" box="[785,911,226,250]" class="Mammalia" family="Didelphidae" genus="Chironectes" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[785,911,226,250]" italics="true" pageId="319" pageNumber="319">Chironectes</emphasis>
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</taxonomicName>
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,
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<taxonomicName baseAuthorityName="Nesslinger" baseAuthorityYear="1956" box="[926,1029,226,250]" class="Mammalia" family="Didelphidae" genus="Didelphis" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[926,1029,226,250]" italics="true" pageId="319" pageNumber="319">Didelphis</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Thomas" authorityYear="1910" box="[1043,1155,226,250]" class="Mammalia" family="Didelphidae" genus="Lutreolina" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[1043,1155,226,250]" italics="true" pageId="319" pageNumber="319">Lutreolina</emphasis>
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</taxonomicName>
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, and
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<taxonomicName authorityName="Tiedemann" authorityYear="1808" box="[686,793,259,283]" class="Mammalia" family="Didelphidae" genus="Philander" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[686,793,259,283]" italics="true" pageId="319" pageNumber="319">Philander</emphasis>
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</taxonomicName>
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. Reig et al. (1987: 73) suggested a close relationship between †
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<taxonomicName box="[1013,1159,292,316]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[1013,1159,292,316]" italics="true" pageId="319" pageNumber="319">Thylophorops</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Thomas" authorityYear="1910" box="[684,797,325,349]" class="Mammalia" family="Didelphidae" genus="Lutreolina" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[684,797,325,349]" italics="true" pageId="319" pageNumber="319">Lutreolina</emphasis>
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</taxonomicName>
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, a possibility also acknowledged by Simpson (1972: 15), who wrote that “classification of
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<taxonomicName authorityName="Ameghino" authorityYear="1908" box="[768,933,391,415]" class="Mammalia" family="Didelphidae" genus="Thylophorops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="species" species="chapalmalensis">
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<emphasis box="[768,933,391,415]" italics="true" pageId="319" pageNumber="319">chapalmalensis</emphasis>
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</taxonomicName>
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as a fairly robust species of
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<taxonomicName authorityName="Thomas" authorityYear="1910" box="[716,830,424,448]" class="Mammalia" family="Didelphidae" genus="Lutreolina" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[716,830,424,448]" italics="true" pageId="319" pageNumber="319">Lutreolina</emphasis>
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</taxonomicName>
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is not ruled out by the available information.” By contrast, Goin (1991) and Goin et al. (2009b) argued for closer affinities with
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<taxonomicName baseAuthorityName="Nesslinger" baseAuthorityYear="1956" box="[741,844,523,547]" class="Mammalia" family="Didelphidae" genus="Didelphis" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[741,844,523,547]" italics="true" pageId="319" pageNumber="319">Didelphis</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Tiedemann" authorityYear="1808" box="[903,1008,523,547]" class="Mammalia" family="Didelphidae" genus="Philander" kingdom="Animalia" order="Didelphimorphia" pageId="319" pageNumber="319" phylum="Chordata" rank="genus">
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<emphasis box="[903,1008,523,547]" italics="true" pageId="319" pageNumber="319">Philander</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |