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<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. &amp; Jansa, Sharon A." docDate="2022" docId="03EFDD5DF7026910DB18FDF51972FE26" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.8:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="8" docTitle="Herpetotherium undetermined" docType="treatment" docVersion="6" lastPageNumber="313" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="313" updateTime="1659987007554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. &amp; School of Biological, Earth &amp; Environmental Sciences University of New South Wales, Australia &amp; Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Voss, Robert S.</mods:namePart>
<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jansa, Sharon A.</mods:namePart>
<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-28</mods:number>
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<mods:detail type="volume">
<mods:number>2022</mods:number>
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<mods:detail type="issue">
<mods:number>457</mods:number>
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<mods:end>353</mods:end>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
<mods:identifier type="ISSN">0003-0090</mods:identifier>
<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974430" ID-Zenodo-Dep="6974430" LSID="urn:lsid:plazi:treatment:03EFDD5DF7026910DB18FDF51972FE26" httpUri="http://treatment.plazi.org/id/03EFDD5DF7026910DB18FDF51972FE26" lastPageNumber="313" pageId="313" pageNumber="313">
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<paragraph blockId="313.[275,469,603,627]" box="[275,469,603,627]" pageId="313" pageNumber="313">
<heading box="[275,469,603,627]" centered="true" fontSize="10" level="2" pageId="313" pageNumber="313" reason="4">
<emphasis bold="true" box="[275,469,603,627]" pageId="313" pageNumber="313">
<taxonomicName box="[289,469,603,627]" class="Mammalia" family="Didelphidae" genus="Herpetotherium" kingdom="Animalia" order="Didelphimorphia" pageId="313" pageNumber="313" phylum="Chordata" rank="species">
<emphasis bold="true" box="[289,469,603,627]" italics="true" pageId="313" pageNumber="313">Herpetotherium</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection pageId="313" pageNumber="313" type="synonymic_list">
<paragraph blockId="313.[108,637,655,1737]" pageId="313" pageNumber="313">
SPECIES SCORED: †
<taxonomicName authorityName="Cope" authorityYear="1873" box="[377,636,655,679]" class="Mammalia" family="Didelphidae" genus="Herpetotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="313" pageNumber="313" phylum="Chordata" rank="species">
<emphasis box="[377,636,655,679]" italics="true" pageId="313" pageNumber="313">Herpetotherium fugax</emphasis>
</taxonomicName>
(
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species), †
<taxonomicName authorityName="Cope" authorityYear="1873" box="[284,411,688,712]" class="Mammalia" family="Didelphidae" genus="Herpetotherium" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Didelphimorphia" pageId="313" pageNumber="313" phylum="Chordata" rank="species">
<emphasis box="[284,304,689,712]" italics="true" pageId="313" pageNumber="313">H</emphasis>
. cf.
<emphasis box="[353,411,688,712]" italics="true" pageId="313" pageNumber="313">fugax</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="313" pageNumber="313" type="materials_examined">
<paragraph blockId="313.[108,637,655,1737]" pageId="313" pageNumber="313">
GEOLOGICAL PROVENANCE OF SCORED
<collectionCode box="[108,167,754,778]" pageId="313" pageNumber="313">SPECIMENS</collectionCode>
: White River Formation,
<collectingRegion box="[450,560,754,778]" country="United States of America" name="Wyoming" pageId="313" pageNumber="313">Wyoming</collectingRegion>
.
</paragraph>
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<subSubSection pageId="313" pageNumber="313" type="description">
<paragraph blockId="313.[108,637,655,1737]" pageId="313" pageNumber="313">AGE OF SCORED SPECIMENS: All examined specimens come from the Orellan North American Land Mammal Age, which is currently estimated as spanning from 33.9 to 32.0 Mya (Prothero and Emry, 2004; Zanazzi et al., 2009).</paragraph>
<paragraph blockId="313.[108,637,655,1737]" box="[140,621,953,977]" pageId="313" pageNumber="313">
ASSIGNED AGE
<collectionCode box="[311,388,953,977]" pageId="313" pageNumber="313">RANGE</collectionCode>
:
<geoCoordinate box="[402,474,953,977]" degrees="33.900" direction="north" orientation="latitude" pageId="313" pageNumber="313" precision="55" value="33.9">33.900</geoCoordinate>
<geoCoordinate box="[473,559,953,977]" degrees="32.000" direction="west" orientation="longitude" pageId="313" pageNumber="313" precision="55" value="-32.0">32.000</geoCoordinate>
Mya.
</paragraph>
</subSubSection>
<subSubSection pageId="313" pageNumber="313" type="discussion">
<paragraph blockId="313.[108,637,655,1737]" lastBlockId="313.[684,1213,226,481]" pageId="313" pageNumber="313">
REMARKS: †
<taxonomicName authorityName="Cope" authorityYear="1873" box="[277,521,986,1010]" class="Mammalia" family="Didelphidae" genus="Herpetotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="313" pageNumber="313" phylum="Chordata" rank="species">
<emphasis box="[277,521,986,1010]" italics="true" pageId="313" pageNumber="313">Herpetotherium fugax</emphasis>
</taxonomicName>
is known from numerous specimens from the White River Formation, including well-preserved skulls and associated postcranial material (Gabbert, 1998; Sánchez-Villagra et al., 2007; Horovitz et al., 2008). Sánchez-Villagra et al. (2007) included †
<taxonomicName box="[122,295,1184,1208]" class="Mammalia" family="Didelphidae" genus="Herpetotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="313" pageNumber="313" phylum="Chordata" rank="genus">
<emphasis box="[122,295,1184,1208]" italics="true" pageId="313" pageNumber="313">Herpetotherium</emphasis>
</taxonomicName>
in a comprehensive morphological phylogenetic analysis of marsupials and relatives, and found that this taxon fell outside (crown-clade)
<taxonomicName box="[282,419,1283,1307]" class="Mammalia" kingdom="Animalia" order="Marsupialia" pageId="313" pageNumber="313" phylum="Chordata" rank="order">Marsupialia</taxonomicName>
. Most subsequent morphological and total-evidence analyses have found a similar result, in which †
<taxonomicName box="[466,637,1349,1373]" class="Mammalia" family="Didelphidae" genus="Herpetotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="313" pageNumber="313" phylum="Chordata" rank="genus">
<emphasis box="[466,637,1349,1373]" italics="true" pageId="313" pageNumber="313">Herpetotherium</emphasis>
</taxonomicName>
either falls outside
<taxonomicName box="[321,450,1382,1406]" class="Mammalia" kingdom="Animalia" order="Marsupialia" pageId="313" pageNumber="313" phylum="Chordata" rank="order">Marsupialia</taxonomicName>
or is not unambiguously a member of this clade (Beck et al., 2008, 2014, 2016; Horovitz et al., 2008, 2009; Beck, 2012, 2017b; Forasiepi et al., 2014a; Suarez et al., 2015; Lorente et al., 2016; Carneiro and Oliveira, 2017a, 2017b; Maga and Beck, 2017; Carneiro, 2018, 2019; Carneiro et al., 2018; Engelman et al., 2020; Muizon et al., 2018; Abello and Candela, 2019; Rangel et al., 2019; Ladevèze et al., 2020; Muizon and Ladevèze, 2020; Zimicz and Goin, 2020). However, a few recent published phylogenetic analyses—namely, the morphological analyses of Forasiepi (2009: fig. 51), Engelman and Croft (2014: fig. 6), and Wilson et al. (2016: fig. 3), and the tip-and-node dated total evidence analysis of Maga and Beck (2017: fig. 39)—have found †
<taxonomicName box="[883,1052,391,415]" class="Mammalia" family="Didelphidae" genus="Herpetotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="313" pageNumber="313" phylum="Chordata" rank="genus">
<emphasis box="[883,1052,391,415]" italics="true" pageId="313" pageNumber="313">Herpetotherium</emphasis>
</taxonomicName>
within
<taxonomicName class="Mammalia" kingdom="Animalia" order="Marsupialia" pageId="313" pageNumber="313" phylum="Chordata" rank="order">Marsupialia</taxonomicName>
, so some questions remain regarding its affinities.
</paragraph>
</subSubSection>
</treatment>
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