treatments-xml/data/03/EF/4E/03EF4E2E807D9D1590A4F3AEFE95F955.xml

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<mods:title id="7485959C650DAFA2B4B8A4BC15F3C873">Devonian Syringostromatid Stromatoporoids from the Broken River Region, North Queensland</mods:title>
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<mods:namePart id="BAB51F394A00949A8AD2DC4FE19285F2">Zhen, Yong Yi</mods:namePart>
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<paragraph id="8BF9FF38807D9D1590A4F3AEFDA5FE4F" blockId="15.[240,630,403,430]" box="[240,630,403,430]" pageId="15" pageNumber="230">
<heading id="D0B14854807D9D1590A4F3AEFDA5FE4F" box="[240,630,403,430]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="230" reason="2">
<taxonomicName id="4C4684BB807D9D1590A4F3AEFDA5FE4F" authority="Fagerstrom, 1982" authorityName="Fagerstrom" authorityYear="1982" box="[240,630,403,430]" class="Hydrozoa" family="Coenostromatidae" genus="Habrostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="230" phylum="Cnidaria" rank="genus">
<emphasis id="B932232A807D9D1590A4F3AEFE5DFE4C" box="[240,398,404,430]" italics="true" pageId="15" pageNumber="230">Habrostroma</emphasis>
<bibRefCitation id="EFD782C9807D9D1591C2F3A9FDA5FE4F" author="Fagerstrom, J" box="[406,630,403,429]" pageId="15" pageNumber="230" pagination="1 - 81" refId="ref17653" refString="Fagerstrom, J. A., 1982. Stromatoporoids of the Detroit River Group and adjacent rocks in the vicinity of the Michigan Basin. Geological Survey of Canada Bulletin 339: 1 - 81." type="journal article" year="1982">
<emphasis id="B932232A807D9D1591C2F3A9FDA5FE4F" bold="true" box="[406,630,403,429]" pageId="15" pageNumber="230">Fagerstrom, 1982</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BF9FF38807D9D15902DF3F5FF66FDE6" blockId="15.[121,748,463,516]" pageId="15" pageNumber="230">
<emphasis id="B932232A807D9D15902DF3F5FEDCFE05" bold="true" box="[121,271,463,487]" pageId="15" pageNumber="230">
<typeStatus id="54FD419A807D9D15902DF3F5FF60FE05" box="[121,179,463,487]" pageId="15" pageNumber="230">Type</typeStatus>
species
</emphasis>
. 
<taxonomicName id="4C4684BB807D9D159175F3EAFF62FDE6" authority="Fagerstrom, 1961" authorityName="Fagerstrom" authorityYear="1961" class="Hydrozoa" family="Hydractiniidae" genus="Stromatopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="230" phylum="Cnidaria" rank="species" species="proxilaminata">
<emphasis id="B932232A807D9D159175F3EAFD8DFE05" box="[289,606,464,487]" italics="true" pageId="15" pageNumber="230">Stromatopora proxilaminata</emphasis>
<bibRefCitation id="EFD782C9807D9D15923DF3EAFF62FDE6" author="Fagerstrom, J" pageId="15" pageNumber="230" pagination="535 - 580" refId="ref17622" refString="Fagerstrom, J. A., 1961. The Fauna of the Middle Devonian Formosa Reef Limestone of southwestern Ontario. Journal of Paleontology 60: 535 - 580." type="journal article" year="1961">Fagerstrom, 1961</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35CACB3807D9D15902DF01DFE95F955" pageId="15" pageNumber="230" type="discussion">
<paragraph id="8BF9FF38807D9D15902DF01DFEE7FB8E" blockId="15.[121,749,551,1719]" pageId="15" pageNumber="230">
<emphasis id="B932232A807D9D15902DF01DFED0FDDD" bold="true" box="[121,259,551,575]" pageId="15" pageNumber="230">Discussion.</emphasis>
In the 
<typeStatus id="54FD419A807D9D159136F012FE40FDDD" box="[354,403,552,575]" pageId="15" pageNumber="230">type</typeStatus>
species of 
<emphasis id="B932232A807D9D159270F012FD15FDDD" box="[548,710,552,575]" italics="true" pageId="15" pageNumber="230">
<taxonomicName id="4C4684BB807D9D159270F012FD11FDDD" authorityName="Fagerstrom" authorityYear="1982" box="[548,706,552,575]" class="Hydrozoa" family="Coenostromatidae" genus="Habrostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="230" phylum="Cnidaria" rank="genus">Habrostroma</taxonomicName>
,
</emphasis>
<taxonomicName id="4C4684BB807D9D159285F012FDF7FDBE" authority="(Fagerstrom, 1961)" baseAuthorityName="Fagerstrom" baseAuthorityYear="1961" class="Hydrozoa" family="Coenostromatidae" genus="Habrostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="230" phylum="Cnidaria" rank="species" species="proxilaminatum">
<emphasis id="B932232A807D9D159285F012FEE3FDBE" italics="true" pageId="15" pageNumber="230">H. proxilaminatum</emphasis>
(
<bibRefCitation id="EFD782C9807D9D159111F07FFDCFFDBE" author="Fagerstrom, J" box="[325,540,581,604]" pageId="15" pageNumber="230" pagination="535 - 580" refId="ref17622" refString="Fagerstrom, J. A., 1961. The Fauna of the Middle Devonian Formosa Reef Limestone of southwestern Ontario. Journal of Paleontology 60: 535 - 580." type="journal article" year="1961">Fagerstrom, 1961</bibRefCitation>
)
</taxonomicName>
, Prosh &amp; Stearn (1996, p. 36) described the pachystele microstructure as “orthoreticular to clinoreticular” in longitudinal section, and as having a “cellular (=irregular microreticular)” microstructure, presumably the “acosmoreticular” 
<typeStatus id="54FD419A807D9D1592F4F080FD1CFD33" box="[672,719,698,721]" pageId="15" pageNumber="230">type</typeStatus>
of 
<bibRefCitation id="EFD782C9807D9D15902DF0E2FED5FD0D" author="Stock, C" box="[121,262,728,751]" pageId="15" pageNumber="230" pagination="119 - 155" refId="ref19566" refString="Stock, C. W., 1989. Microreticulate microstructure in the Stromatoporoidea. Memoir of the Association of Australasian Palaeontologists 8: 119 - 155." type="journal article" year="1989">Stock (1989)</bibRefCitation>
in tangential section. However, other writers have recognized the genus 
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<emphasis id="B932232A807D9D1591ECF0CFFD9FFCEE" box="[440,588,757,780]" italics="true" pageId="15" pageNumber="230">Habrostroma</emphasis>
</taxonomicName>
as exhibiting only acosmoreticular microstructure (see 
<bibRefCitation id="EFD782C9807D9D15920FF128FD34FCCB" author="Stock, C" box="[603,743,786,809]" pageId="15" pageNumber="230" pagination="897 - 911" refId="ref19594" refString="Stock, C. W., 1991. Lower Devonian (Lochkovian) Stromatoporoidea. Journal of Paleontology 65: 897 - 911." type="journal article" year="1991">Stock, 1991</bibRefCitation>
, 
<bibRefCitation id="EFD782C9807D9D15902DF10AFF61FCA5" author="Stock, C" box="[121,178,816,839]" pageId="15" pageNumber="230" pagination="539 - 553" refId="ref19619" refString="Stock, C. W., 1997. Lower Devonian (Lochkovian) Stromatoporoidea from the Coeymans Formation of central New York. Journal of Paleontology 71: 539 - 553." type="journal article" year="1997">1997</bibRefCitation>
; Webby 
<emphasis id="B932232A807D9D159145F10AFE9BFCA5" box="[273,328,816,839]" italics="true" pageId="15" pageNumber="230">et al.</emphasis>
, 1993). For comparative purposes, the genus 
<taxonomicName id="4C4684BB807D9D1590EAF177FE9AFC86" authorityName="Webby &amp; Zhen" authorityYear="2008" box="[190,329,845,868]" class="Hydrozoa" family="Hydractiniidae" genus="Atopostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="15" pageNumber="230" phylum="Cnidaria" rank="genus">
<emphasis id="B932232A807D9D1590EAF177FE9AFC86" box="[190,329,845,868]" italics="true" pageId="15" pageNumber="230">Atopostroma</emphasis>
</taxonomicName>
, based on 
<taxonomicName id="4C4684BB807D9D1591EFF177FF61FC63" authority="Yang &amp; Dong, 1979" authorityName="Yang &amp; Dong" authorityYear="1979" class="Hydrozoa" family="Hydractiniidae" genus="Atopostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="15" pageNumber="230" phylum="Cnidaria" rank="species" species="tuntouense">
<emphasis id="B932232A807D9D1591EFF177FD9DFC86" box="[443,590,845,868]" italics="true" pageId="15" pageNumber="230">A. tuntouense</emphasis>
Yang &amp; Dong, 1979
</taxonomicName>
, has been described as exhibiting “orthoreticulate to clinoreticulate” microstructure (see 
<bibRefCitation id="EFD782C9807D9D15925EF1B2FD4AFC7D" author="Stearn, C" box="[522,665,904,927]" pageId="15" pageNumber="230" pagination="201 - 229" refId="ref19319" refString="Stearn, C. W., 1993. Revision of the order Stromatoporida. Palaeontology 36: 201 - 229." type="journal article" year="1993">Stearn, 1993</bibRefCitation>
, 
<bibRefCitation id="EFD782C9807D9D1592F3F1B2FF29FC5E" author="Stearn, C &amp; Webby, H" pageId="15" pageNumber="230" pagination="1 - 70" refId="ref19524" refString="Stearn, C. W., B. D. Webby, H. Nestor &amp; C. W. Stock, 1999. Revised classification and terminology of Palaeozoic stromatoporoids. Acta Palaeontologica Polonica 44: 1 - 70." type="journal article" year="1999">
Stearn 
<emphasis id="B932232A807D9D15902DF19FFF79FC5E" box="[121,170,933,956]" italics="true" pageId="15" pageNumber="230">et al</emphasis>
., 1999
</bibRefCitation>
), but also in 
<bibRefCitation id="EFD782C9807D9D1591DFF19FFDC8FC5E" author="Stearn, C" box="[395,539,933,956]" pageId="15" pageNumber="230" pagination="493 - 510" refId="ref19282" refString="Stearn, C. W., 1990. Stromatoporoids from the allochthonous reef facies of the Stuart Bay Formation (Lower Devonian), Arctic Canada. Journal of Paleontology 64: 493 - 510." type="journal article" year="1990">Stearn (1990</bibRefCitation>
, p. 498) and 
<bibRefCitation id="EFD782C9807D9D1592FBF19FFF68FC3B" author="Stock, C" pageId="15" pageNumber="230" pagination="539 - 553" refId="ref19619" refString="Stock, C. W., 1997. Lower Devonian (Lochkovian) Stromatoporoidea from the Coeymans Formation of central New York. Journal of Paleontology 71: 539 - 553." type="journal article" year="1997">Stock (1997</bibRefCitation>
, p. 549), as only exhibiting a clinoreticular 
<typeStatus id="54FD419A807D9D1592CBF1F8FD1DFC3B" box="[671,718,962,985]" pageId="15" pageNumber="230">type</typeStatus>
of microstructure, as in 
<taxonomicName id="4C4684BB807D9D15913EF1DAFDCBFC15" authorityName="Bogoyavlenskaya" authorityYear="1972" box="[362,536,992,1015]" class="Hydrozoa" family="Hydractiniidae" genus="Columnostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="15" pageNumber="230" phylum="Cnidaria" rank="genus">
<emphasis id="B932232A807D9D15913EF1DAFDCBFC15" box="[362,536,992,1015]" italics="true" pageId="15" pageNumber="230">Columnostroma</emphasis>
</taxonomicName>
and 
<emphasis id="B932232A807D9D159200F1DAFD3FFC15" box="[596,748,992,1015]" italics="true" pageId="15" pageNumber="230">
<taxonomicName id="4C4684BB807D9D159200F1DAFD3BFC15" authorityName="Winchell" authorityYear="1867" box="[596,744,992,1015]" class="Hydrozoa" family="Hydractiniidae" genus="Coenostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="230" phylum="Cnidaria" rank="genus">Coenostroma</taxonomicName>
,
</emphasis>
respectively. In the present study, the species of 
<taxonomicName id="4C4684BB807D9D1592F6F1C7FF1BFBD0" authorityName="Webby &amp; Zhen" authorityYear="2008" class="Hydrozoa" family="Hydractiniidae" genus="Atopostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="15" pageNumber="230" phylum="Cnidaria" rank="genus">
<emphasis id="B932232A807D9D1592F6F1C7FF1BFBD0" italics="true" pageId="15" pageNumber="230">Atopostroma</emphasis>
</taxonomicName>
show examples of both acosmoreticular and clinoreticular microstructural 
<typeStatus id="54FD419A807D9D1591E3F602FE20FBAD" box="[439,499,1080,1103]" pageId="15" pageNumber="230">types</typeStatus>
; no orthoreticular 
<typeStatus id="54FD419A807D9D1592E7F602FD3FFBAD" box="[691,748,1080,1103]" pageId="15" pageNumber="230">types</typeStatus>
have been found.
</paragraph>
<paragraph id="8BF9FF38807D9D1590C3F648FE95F955" blockId="15.[121,749,551,1719]" pageId="15" pageNumber="230">
The diagnostic features of the two genera are as follows: 
<taxonomicName id="4C4684BB807D9D15902DF6AAFED9FB45" authorityName="Fagerstrom" authorityYear="1982" box="[121,266,1168,1191]" class="Hydrozoa" family="Coenostromatidae" genus="Habrostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="230" phylum="Cnidaria" rank="genus">
<emphasis id="B932232A807D9D15902DF6AAFED9FB45" box="[121,266,1168,1191]" italics="true" pageId="15" pageNumber="230">Habrostroma</emphasis>
</taxonomicName>
typically has pachysteles that are shorter, more irregular, confined largely within “interlaminar” spaces (rarely superposed) and internally composed of an irregular network of acosmoreticulate skeletal material; and the pachystromes prominent but not thick, with one or more microlaminae, and sometimes microlaminalike dissepiments. 
<taxonomicName id="4C4684BB807D9D159119F77AFE09FAB5" authorityName="Webby &amp; Zhen" authorityYear="2008" box="[333,474,1344,1367]" class="Hydrozoa" family="Hydractiniidae" genus="Atopostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="15" pageNumber="230" phylum="Cnidaria" rank="genus">
<emphasis id="B932232A807D9D159119F77AFE09FAB5" box="[333,474,1344,1367]" italics="true" pageId="15" pageNumber="230">Atopostroma</emphasis>
</taxonomicName>
, on the other hand, has pachysteles that are typically superposed, formed by upward and outward accretion to produce irregularly spread network of acosmoreticular to clinoreticulate skeletal material beneath successive, laterally extensive, microlaminae. The two described species 
<taxonomicName id="4C4684BB807D9D15913DF7E9FE39FA08" authorityName="Webby &amp; Zhen" authorityYear="2008" box="[361,490,1491,1514]" class="Hydrozoa" family="Coenostromatidae" genus="Habrostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="230" phylum="Cnidaria" rank="species" species="alternum">
<emphasis id="B932232A807D9D15913DF7E9FE39FA08" box="[361,490,1491,1514]" italics="true" pageId="15" pageNumber="230">H. alternum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4684BB807D9D15924BF7E9FD79FA08" authorityName="Webby &amp; Zhen" authorityYear="2008" box="[543,682,1491,1514]" class="Hydrozoa" family="Hydractiniidae" genus="Atopostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="15" pageNumber="230" phylum="Cnidaria" rank="species" species="protentum">
<emphasis id="B932232A807D9D15924BF7E9FD79FA08" box="[543,682,1491,1514]" italics="true" pageId="15" pageNumber="230">A. protentum</emphasis>
</taxonomicName>
occur together in the Martins Well Limestone Member and are truly transitional forms, showing some features characteristic of each genus. Some consideration has been given to combining them, even treating them as different species of the one genus. However, between the mid-range and extremes of variation of the two species they are clearly representatives of different genera.
</paragraph>
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