treatments-xml/data/03/E3/29/03E3293EFFA0FF8CFE006C9A46C689C6.xml
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<document id="3C8099A805ED4353C6CE5888063C43E8" ID-DOI="10.1080/00222933.2010.535914" ID-ISSN="1464-5262" ID-Zenodo-Dep="5203342" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629048039960" checkinUser="felipe" docAuthor="Zacharda, Miloslav, Isaia, Marco &amp; Piva, Erminio" docDate="2011" docId="03E3293EFFA0FF8CFE006C9A46C689C6" docLanguage="en" docName="JNatHist.45.11-12.641-666.pdf" docOrigin="Journal of Natural History 45 (11 - 12)" docSource="http://dx.doi.org/10.1080/00222933.2010.535914" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Troglocheles quinquesolenidiata Zacharda &amp; Isaia &amp; Piva 2011, sp. nov." docType="treatment" docVersion="3" lastPageNumber="649" masterDocId="FFDA5146FFA4FF86FF9B6E15477E8D0F" masterDocTitle="New troglobitic species of the genus Troglocheles (Acari: Prostigmata: Rhagidiidae) from caves in northern Italy and Austria, with a key to adult species of the genus" masterLastPageNumber="666" masterPageNumber="641" pageNumber="643" updateTime="1699057451977" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EBAAC754B06095C5364C9BDEA7E80757">New troglobitic species of the genus Troglocheles (Acari: Prostigmata: Rhagidiidae) from caves in northern Italy and Austria, with a key to adult species of the genus</mods:title>
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<mods:namePart id="383B272261E19629849830ED947DB395">Zacharda, Miloslav</mods:namePart>
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<mods:namePart id="A550AA8F5904485579926DCB5F856CF5">Isaia, Marco</mods:namePart>
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<mods:namePart id="6984B6A5DD2B66DC9925560D22018FD5">Piva, Erminio</mods:namePart>
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<mods:title id="B66B67B5795F2CEC51FC263030317B35">Journal of Natural History</mods:title>
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<mods:date id="E891F4EA8DA612F7C2B7B51C8F78E83E">2011</mods:date>
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<mods:number id="3F60715929314B1FE6E9F1BFFA10C0AE">2011-03-07</mods:number>
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<mods:number id="CD2B7C3284B528E68801B00E8093FA07">45</mods:number>
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<mods:number id="ED4F277BFA90AD4B5552FA73B0CC3563">11 - 12</mods:number>
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<mods:identifier id="D9485F5E939F6F00AA81CAE7B373DF61" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03E3293EFFA0FF8CFE006C9A46C689C6" LSID="urn:lsid:plazi:treatment:03E3293EFFA0FF8CFE006C9A46C689C6" httpUri="http://treatment.plazi.org/id/03E3293EFFA0FF8CFE006C9A46C689C6" lastPageId="10" lastPageNumber="649" pageId="4" pageNumber="643">
<subSubSection id="C350CBA3FFA0FF82FE006C9A44B38FA8" box="[411,973,655,679]" pageId="4" pageNumber="643" type="nomenclature">
<paragraph id="8BF59828FFA0FF82FE006C9A44B38FA8" blockId="4.[411,973,655,711]" box="[411,973,655,679]" pageId="4" pageNumber="643">
<heading id="D0BD2F44FFA0FF82FE006C9A44B38FA8" box="[411,973,655,679]" centered="true" fontSize="36" level="2" pageId="4" pageNumber="643" reason="7">
<taxonomicName id="4C4AE3ABFFA0FF82FE006C9A44138FA8" ID-CoL="4d9f4542-0069-41d7-96c2-c51750fa280b" authority="Zacharda" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[411,877,655,679]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="643" phylum="Arthropoda" rank="species" species="quinquesolenidiata" status="sp. nov.">
<emphasis id="B93E443AFFA0FF82FE006C9A45868FA8" bold="true" box="[411,760,655,679]" italics="true" pageId="4" pageNumber="643">Troglocheles quinquesolenidiata</emphasis>
Zacharda
</taxonomicName>
,
<taxonomicNameLabel id="A20DF941FFA0FF82FCE36C8544B38FA8" box="[888,973,656,679]" pageId="4" pageNumber="643" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C350CBA3FFA0FF8CFDF26CBA46C689C6" lastPageId="10" lastPageNumber="649" pageId="4" pageNumber="643" type="description">
<paragraph id="8BF59828FFA0FF82FDF26CBA45808FC8" blockId="4.[411,973,655,711]" box="[617,766,687,711]" pageId="4" pageNumber="643">
(
<figureCitation id="137184ADFFA0FF82FDEA6CBA45888FC8" box="[625,758,687,711]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStartId-0="5.[144,211,1292,1314]" captionStartId-1="6.[213,280,1417,1439]" captionStartId-2="7.[144,211,1420,1442]" captionStartId-3="8.[213,280,1372,1394]" captionTargetBox-0="[142,1102,163,1216]" captionTargetBox-1="[215,1167,163,1340]" captionTargetBox-2="[142,1102,161,1387]" captionTargetBox-3="[211,1171,161,1338]" captionTargetId-0="figure-7@5.[142,1102,162,1216]" captionTargetId-1="figure-6@6.[214,1168,162,1341]" captionTargetId-2="figure-7@7.[142,1072,160,1387]" captionTargetId-3="figure-6@8.[211,1171,161,1339]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Figure 2. Troglocheles quinquesolenidiata sp. nov., adult female; (A) dorsum; (B) venter (for setal notation see Description)." captionText-1="Figure 3. Troglocheles quinquesolenidiata sp. nov., adult female; (A) chelicera, lateral aspect; (B) rhagidial organ I, dorsal aspect; (C) rhagidial organ II, dorsal aspect; (D) apex of tarsus III with pubescent oar-like iteral (it) setae, lateral aspect; (E) dorsodistal lanceolate solenidion (Φ) on tibia II; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." captionText-2="Figure 4. Troglocheles quinquesolenidiata sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg I; (B) leg II (ω, Φ, σ and ε designations for solenidia on tarsus, tibia, genu and for famulus, respectively)." captionText-3="Figure 5. Troglocheles quinquesolenidiata sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg III; (B) leg IV (Φ, σ designations for solenidia on tibia and genu, respectively)." figureDoi-0="http://doi.org/10.5281/zenodo.5203346" figureDoi-1="http://doi.org/10.5281/zenodo.5203349" figureDoi-2="http://doi.org/10.5281/zenodo.5203351" figureDoi-3="http://doi.org/10.5281/zenodo.5203353" httpUri-0="https://zenodo.org/record/5203346/files/figure.png" httpUri-1="https://zenodo.org/record/5203349/files/figure.png" httpUri-2="https://zenodo.org/record/5203351/files/figure.png" httpUri-3="https://zenodo.org/record/5203353/files/figure.png" pageId="4" pageNumber="643">Figures 25</figureCitation>
)
</paragraph>
<paragraph id="8BF59828FFA0FF82FF4E6D1846D98E2A" blockId="4.[213,423,781,805]" box="[213,423,781,805]" pageId="4" pageNumber="643">
<emphasis id="B93E443AFFA0FF82FF4E6D1846D98E2A" box="[213,423,781,805]" italics="true" pageId="4" pageNumber="643">Material examined</emphasis>
</paragraph>
<paragraph id="8BF59828FFA0FF82FF4E6D2045CD8EA3" blockId="4.[213,1170,821,941]" pageId="4" pageNumber="643">
<materialsCitation id="3B229275FFA0FF82FF4E6D2045CD8EA3" collectingDate="1793-05" collectingDateMax="1996-09-01" collectingDateMin="1793-05" collectionCode="VI" collectorName="G. Peretto &amp; E. Piva &amp; Ohio State University &amp; Type No." country="Italy" county="Adult" elevation="918" location="Recoaro Terme" municipality="Gruppo del Carega-Piccole Dolomiti-Pasubio" pageId="4" pageNumber="643" specimenCode="OSAL0007410" specimenCount="1" stateProvince="Veneto" typeStatus="holotype">
<emphasis id="B93E443AFFA0FF82FF4E6D20463D8E42" box="[213,323,821,845]" italics="true" pageId="4" pageNumber="643">
<typeStatus id="54F1268AFFA0FF82FF4E6D2046408E42" box="[213,318,821,845]" pageId="4" pageNumber="643" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="6294E0A4FFA0FF82FED16D2046F28E42" box="[330,396,821,845]" pageId="4" pageNumber="643">Adult</collectingCounty>
female,
<collectingCountry id="F35DD8B8FFA0FF82FE716D2045618E42" box="[490,543,821,845]" name="Italy" pageId="4" pageNumber="643">Italy</collectingCountry>
,
<collectingRegion id="498E56CAFFA0FF82FDB16D2045058E43" box="[554,635,821,844]" country="Italy" name="Veneto" pageId="4" pageNumber="643">Veneto</collectingRegion>
,
<collectingMunicipality id="6B910252FFA0FF82FD1D6D2043F08E42" box="[646,1166,821,845]" pageId="4" pageNumber="643">Gruppo del Carega-Piccole Dolomiti-Pasubio</collectingMunicipality>
,
<location id="8E95CEF3FFA0FF82FF4E6D4046FF8E63" LSID="urn:lsid:plazi:treatment:03E3293EFFA0FF8CFE006C9A46C689C6:8E95CEF3FFA0FF82FF4E6D4046FF8E63" box="[213,385,853,876]" country="Italy" county="Adult" municipality="Gruppo del Carega-Piccole Dolomiti-Pasubio" name="Recoaro Terme" pageId="4" pageNumber="643" stateProvince="Veneto">Recoaro Terme</location>
(Vicenza),
<location id="8E95CEF3FFA0FF82FE656D4045BB8E63" LSID="urn:lsid:plazi:treatment:03E3293EFFA0FF8CFE006C9A46C689C6:8E95CEF3FFA0FF82FE656D4045BB8E63" box="[510,709,852,876]" country="Italy" county="Adult" municipality="Gruppo del Carega-Piccole Dolomiti-Pasubio" name="Buso Brutto Cave" pageId="4" pageNumber="643" stateProvince="Veneto">Buso Brutto Cave</location>
(
<date id="FFF4BEE8FFA0FF82FD496D41445F8E63" box="[722,801,852,876]" pageId="4" pageNumber="643" value="1793-05">
<collectingDate id="EFB04700FFA0FF82FD496D41445F8E63" box="[722,801,852,876]" pageId="4" pageNumber="643" value="1793-05">1793 V</collectingDate>
</date>
<emphasis id="B93E443AFFA0FF82FCBA6D4044538E62" box="[801,813,853,877]" italics="true" pageId="4" pageNumber="643">/</emphasis>
<collectionCode id="ED5B00EDFFA0FF82FCB56D4144318E63" box="[814,847,852,876]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4" pageNumber="643">VI</collectionCode>
),
<elevation id="00677F1BFFA0FF82FCFA6D4144A18E63" box="[865,991,852,876]" metricMagnitude="2" metricUnit="m" metricValue="9.18" pageId="4" pageNumber="643" unit="m" value="918.0">
<quantity id="4CB235CDFFA0FF82FCFA6D4144DB8E63" box="[865,933,852,876]" metricMagnitude="2" metricUnit="m" metricValue="9.18" pageId="4" pageNumber="643" unit="m" value="918.0">918 m</quantity>
a.s.l.
</elevation>
, coll.
<collectorName id="26BFFDFEFFA0FF82FBBB6D4143EC8E63" box="[1056,1170,852,876]" pageId="4" pageNumber="643">G. Peretto</collectorName>
and
<collectorName id="26BFFDFEFFA0FF82FE9D6D6046268E83" box="[262,344,884,908]" pageId="4" pageNumber="643">E. Piva</collectorName>
,
<date id="FFF4BEE8FFA0FF82FEF96D6145548E83" box="[354,554,884,908]" pageId="4" pageNumber="643" value="1996-09-01">
<collectingDate id="EFB04700FFA0FF82FEF96D6145548E83" box="[354,554,884,908]" pageId="4" pageNumber="643" value="1996-09-01">1 September 1996</collectingDate>
</date>
, deposited in the
<location id="8E95CEF3FFA0FF82FD716D6043318E83" LSID="urn:lsid:plazi:treatment:03E3293EFFA0FF8CFE006C9A46C689C6:8E95CEF3FFA0FF82FD716D6043318E83" box="[746,1103,884,908]" country="Italy" county="Adult" municipality="Gruppo del Carega-Piccole Dolomiti-Pasubio" name="Museum of Biological Diversity" pageId="4" pageNumber="643" stateProvince="Veneto">Museum of Biological Diversity</location>
,
<collectorName id="26BFFDFEFFA0FF82FBC26D6146F58EA3" pageId="4" pageNumber="643">Ohio State University</collectorName>
,
<collectorName id="26BFFDFEFFA0FF82FE0D6D8046808EA3" box="[406,510,917,941]" pageId="4" pageNumber="643">Type No.</collectorName>
<specimenCode id="DBEC3053FFA0FF82FD9E6D8145D18EA3" box="[517,687,916,940]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="4" pageNumber="643">OSAL0007410</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF59828FFA0FF82FF4E6DC345928902" blockId="4.[213,1170,981,1037]" pageId="4" pageNumber="643">
<materialsCitation id="3B229275FFA0FF82FF4E6DC343F38EE2" box="[213,1165,981,1005]" collectingDate="1793-05" collectingDateMax="1996-09-01" collectingDateMin="1793-05" collectionCode="VI" collectorName="G. Peretto &amp; E. Piva &amp; Ohio State University &amp; Type No." country="Italy" county="Adult" elevation="918" location="Recoaro Terme" municipality="Gruppo del Carega-Piccole Dolomiti-Pasubio" pageId="4" pageNumber="643" specimenCount="1" specimenCount-adult="1" stateProvince="Veneto" typeStatus="paratype">
<emphasis id="B93E443AFFA0FF82FF4E6DC346348EE2" box="[213,330,982,1005]" italics="true" pageId="4" pageNumber="643">
<typeStatus id="54F1268AFFA0FF82FF4E6DC3463B8EE2" box="[213,325,982,1005]" pageId="4" pageNumber="643" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Deposited with the
<typeStatus id="54F1268AFFA0FF82FDA96DC045E78EE2" box="[562,665,981,1005]" pageId="4" pageNumber="643" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D4C53A1FFA0FF82FD3D6DC0445C8EE2" box="[678,802,981,1005]" pageId="4" pageNumber="643" type="adult">three adult</specimenCount>
females, same data as holotype
</materialsCitation>
,
<materialsCitation id="3B229275FFA0FF82FF4E6DE046FD8902" box="[213,387,1013,1037]" collectionCode="OSAL" country="Italy" pageId="4" pageNumber="643" specimenCode="OSAL0007411" specimenCount="1" stateProvince="Veneto" typeStatus="paratype">
<specimenCode id="DBEC3053FFA0FF82FF4E6DE046018902" box="[213,383,1013,1037]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="4" pageNumber="643">OSAL0007411</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B229275FFA0FF82FE126DE045498902" box="[393,567,1013,1037]" collectionCode="OSAL" country="Italy" pageId="4" pageNumber="643" specimenCode="OSAL0007412" specimenCount="1" stateProvince="Veneto" typeStatus="paratype">
<specimenCode id="DBEC3053FFA0FF82FE126DE0454D8902" box="[393,563,1013,1037]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="4" pageNumber="643">OSAL0007412</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B229275FFA0FF82FDA56DE045968902" box="[574,744,1013,1037]" collectionCode="OSAL" country="Italy" pageId="4" pageNumber="643" specimenCode="OSAL0007413" specimenCount="1" stateProvince="Veneto" typeStatus="paratype">
<specimenCode id="DBEC3053FFA0FF82FDA56DE045968902" box="[574,744,1013,1037]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="4" pageNumber="643">OSAL0007413</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF59828FFA0FF82FF4E6A41463F8964" blockId="4.[213,321,1108,1131]" box="[213,321,1108,1131]" pageId="4" pageNumber="643">
<emphasis id="B93E443AFFA0FF82FF4E6A41463F8964" box="[213,321,1108,1131]" italics="true" pageId="4" pageNumber="643">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF59828FFA0FF82FF4E6A6E430D885D" blockId="4.[213,1171,1147,1365]" pageId="4" pageNumber="643">
Rhagidial organ I comprises five rhagidial solenidia lying obliquely in separate depressions; rhagidial organ II with four rhagidial solenidia lying in separate oblique depressions, and small spiniform famulus (
<emphasis id="B93E443AFFA0FF82FD2D6AAE45BC89DC" box="[694,706,1211,1235]" italics="true" pageId="4" pageNumber="643">ε</emphasis>
) inserted between first and second proximal rhagidial solenidia antiaxially. Spiniform solenidia on tibiae I, II and III minute. Tibia IV with one minute spiniform dorsal medioproximal solenidion. Ratio of tarsus I length to dorsoventral width 7.55; ratio of palpal tarsus length to dorsoventral width 5.23 (5.05.4). Bothridial setae
<emphasis id="B93E443AFFA0FF82FDAF6B2E452D885A" box="[564,595,1339,1365]" italics="true" pageId="4" pageNumber="643">
sc
<subScript id="17CE9A6DFFA0FF82FDD16B51452D885A" attach="left" box="[586,595,1348,1365]" fontSize="7" pageId="4" pageNumber="643">1</subScript>
</emphasis>
overlapping insertions of opisthosomal setae
<emphasis id="B93E443AFFA0FF82FBCD6B2E4315885A" box="[1110,1131,1339,1365]" italics="true" pageId="4" pageNumber="643">
c
<subScript id="17CE9A6DFFA0FF82FBF96B514315885A" attach="left" box="[1122,1131,1348,1365]" fontSize="7" pageId="4" pageNumber="643">1</subScript>
</emphasis>
.
</paragraph>
<paragraph id="8BF59828FFA0FF82FF4E6B6C462A889F" blockId="4.[213,340,1401,1424]" box="[213,340,1401,1424]" pageId="4" pageNumber="643">
<emphasis id="B93E443AFFA0FF82FF4E6B6C462A889F" box="[213,340,1401,1424]" italics="true" pageId="4" pageNumber="643">Description</emphasis>
</paragraph>
<paragraph id="8BF59828FFA0FF82FF4E6BB545C088D7" blockId="4.[213,1169,1440,1496]" pageId="4" pageNumber="643">Adult female (four examined). Length of idiosoma 900 (8001008) µm. Ratio of leg I length to idiosomal length 2.10 (1.962.28).</paragraph>
<paragraph id="8BF59828FFA0FF82FF4E681443EF8B97" blockId="4.[213,1170,1537,1688]" pageId="4" pageNumber="643">
<emphasis id="B93E443AFFA0FF82FF4E681446168B16" box="[213,360,1537,1561]" italics="true" pageId="4" pageNumber="643">
<taxonomicName id="4C4AE3ABFFA0FF82FF4E6814461D8B16" authorityName="Krefft" authorityYear="1871" box="[213,355,1537,1561]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="643" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
.
</emphasis>
Subcapitulum slender, oval (
<figureCitation id="137184ADFFA0FF82FD31681444638B16" box="[682,797,1537,1561]" captionStart="Figure 3" captionStartId="6.[213,280,1417,1439]" captionTargetBox="[215,1167,163,1340]" captionTargetId="figure-6@6.[214,1168,162,1341]" captionTargetPageId="6" captionText="Figure 3. Troglocheles quinquesolenidiata sp. nov., adult female; (A) chelicera, lateral aspect; (B) rhagidial organ I, dorsal aspect; (C) rhagidial organ II, dorsal aspect; (D) apex of tarsus III with pubescent oar-like iteral (it) setae, lateral aspect; (E) dorsodistal lanceolate solenidion (Φ) on tibia II; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203349" httpUri="https://zenodo.org/record/5203349/files/figure.png" pageId="4" pageNumber="643">Figure 3F</figureCitation>
), subtriangular, ratio of length to breadth 1.331.82; distal hypostomal lips with spiniform internal malar and serrate external malar processes; adoral setae nude and overlapping apex of subcapitulum; proximal subcapitular setae pointed, pubescent, external pair slightly longer than internal pair. Dorsal surface of chelicera with saddle-shaped depression slightly distal
</paragraph>
<paragraph id="8BF59828FFA1FF83FDC96AD945F889E5" blockId="5.[594,646,1228,1258]" box="[594,646,1228,1258]" pageId="5" pageNumber="644">A,B</paragraph>
<caption id="DF35C8A0FFA1FF83FF0B6B1946BB8830" ID-DOI="http://doi.org/10.5281/zenodo.5203346" ID-Zenodo-Dep="5203346" httpUri="https://zenodo.org/record/5203346/files/figure.png" pageId="5" pageNumber="644" startId="5.[144,211,1292,1314]" targetBox="[142,1102,163,1216]" targetPageId="5">
<paragraph id="8BF59828FFA1FF83FF0B6B1946BB8830" blockId="5.[144,1101,1292,1343]" pageId="5" pageNumber="644">
Figure 2.
<taxonomicName id="4C4AE3ABFFA1FF83FF606B194549882D" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[251,567,1292,1314]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="644" phylum="Arthropoda" rank="species" species="quinquesolenidiata" status="sp. nov.">
<emphasis id="B93E443AFFA1FF83FF606B194549882D" box="[251,567,1292,1314]" italics="true" pageId="5" pageNumber="644">Troglocheles quinquesolenidiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFA1FF83FDDB6B1845EF882D" box="[576,657,1293,1314]" pageId="5" pageNumber="644" rank="species">sp. nov.</taxonomicNameLabel>
, adult female; (A) dorsum; (B) venter (for setal notation see
<emphasis id="B93E443AFFA1FF83FEDE6B3F46C68830" box="[325,440,1322,1343]" italics="true" pageId="5" pageNumber="644">Description</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BF59828FFA1FF83FF0B6B7743338B97" blockId="5.[144,1101,1377,1688]" pageId="5" pageNumber="644">
to level of bases of digits; cheliceral digits strikingly long, slender; fixed digit terminates in two cusps, smooth along masticatory surface and with small lateral fissure at level of bases of digits (
<figureCitation id="137184ADFFA1FF83FE026BB4456F88B6" box="[409,529,1441,1465]" captionStart="Figure 3" captionStartId="6.[213,280,1417,1439]" captionTargetBox="[215,1167,163,1340]" captionTargetId="figure-6@6.[214,1168,162,1341]" captionTargetPageId="6" captionText="Figure 3. Troglocheles quinquesolenidiata sp. nov., adult female; (A) chelicera, lateral aspect; (B) rhagidial organ I, dorsal aspect; (C) rhagidial organ II, dorsal aspect; (D) apex of tarsus III with pubescent oar-like iteral (it) setae, lateral aspect; (E) dorsodistal lanceolate solenidion (Φ) on tibia II; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203349" httpUri="https://zenodo.org/record/5203349/files/figure.png" pageId="5" pageNumber="644">Figure 3A</figureCitation>
); movable digit smooth along masticatory surface. Chelicera with two setae inserted distad of articulation of movable digit; tip of proximal seta not reaching insertion of distal seta; tip of distal seta slightly overlaps apex of fixed digit. Length of chelicera 316 µm, dorsoventral width 109 µm, length of movable digit 162 (155165) µm, length of proximal and distal cheliceral setae 18 (1620) and 59 (5363) µm, respectively, distance between their insertions 2023 µm. Ratios: cheliceral length to dorsoventral width 2.752.96; length of movable digit to length of chelicera 0.51; length of movable digit to dorsoventral width of chelicera 1.421.51.
</paragraph>
<paragraph id="8BF59828FFA2FF80FE846B5C43188868" blockId="6.[287,1126,1353,1383]" box="[287,1126,1353,1383]" pageId="6" pageNumber="645">A,F B,C D G / E</paragraph>
<caption id="DF35C8A0FFA2FF80FF4E6B9C44DC88F8" ID-DOI="http://doi.org/10.5281/zenodo.5203349" ID-Zenodo-Dep="5203349" httpUri="https://zenodo.org/record/5203349/files/figure.png" pageId="6" pageNumber="645" startId="6.[213,280,1417,1439]" targetBox="[215,1167,163,1340]" targetPageId="6">
<paragraph id="8BF59828FFA2FF80FF4E6B9C44DC88F8" blockId="6.[213,1170,1417,1527]" pageId="6" pageNumber="645">
Figure 3.
<taxonomicName id="4C4AE3ABFFA2FF80FEA46B9C45028890" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[319,636,1417,1439]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="645" phylum="Arthropoda" rank="species" species="quinquesolenidiata" status="sp. nov.">
<emphasis id="B93E443AFFA2FF80FEA46B9C45028890" box="[319,636,1417,1439]" italics="true" pageId="6" pageNumber="645">Troglocheles quinquesolenidiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFA2FF80FD1F6B9F45AB8890" box="[644,725,1418,1439]" pageId="6" pageNumber="645" rank="species">sp. nov.</taxonomicNameLabel>
, adult female; (A) chelicera, lateral aspect; (B) rhagidial organ I, dorsal aspect; (C) rhagidial organ II, dorsal aspect; (D) apex of tarsus III with pubescent oar-like iteral (
<emphasis id="B93E443AFFA2FF80FD976BD1456488D6" box="[524,538,1476,1497]" italics="true" pageId="6" pageNumber="645">it</emphasis>
) setae, lateral aspect; (E) dorsodistal lanceolate solenidion (
<emphasis id="B93E443AFFA2FF80FBE06BD143F788D5" box="[1147,1161,1476,1498]" italics="true" pageId="6" pageNumber="645">Φ</emphasis>
) on tibia II; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect.
</paragraph>
</caption>
<paragraph id="8BF59828FFA2FF80FF4E685443EC8B97" blockId="6.[213,1170,1601,1689]" pageId="6" pageNumber="645">
Palpal tarsus shorter than femorogenu (
<figureCitation id="137184ADFFA2FF80FD0E685444738B56" box="[661,781,1601,1625]" captionStart="Figure 3" captionStartId="6.[213,280,1417,1439]" captionTargetBox="[215,1167,163,1340]" captionTargetId="figure-6@6.[214,1168,162,1341]" captionTargetPageId="6" captionText="Figure 3. Troglocheles quinquesolenidiata sp. nov., adult female; (A) chelicera, lateral aspect; (B) rhagidial organ I, dorsal aspect; (C) rhagidial organ II, dorsal aspect; (D) apex of tarsus III with pubescent oar-like iteral (it) setae, lateral aspect; (E) dorsodistal lanceolate solenidion (Φ) on tibia II; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203349" httpUri="https://zenodo.org/record/5203349/files/figure.png" pageId="6" pageNumber="645">Figure 3G</figureCitation>
); ratio of length to width of tarsus 5.23 (5.05.4). Length of palpal trochanter, femorogenu, tibia and tarsus 40 (3343), 217 (198231), 62 (5966) and 178 (175181) µm, respectively. Number of setae and
</paragraph>
<caption id="DF35C8A0FFA3FF81FF0B6B9946BF88D3" ID-DOI="http://doi.org/10.5281/zenodo.5203351" ID-Zenodo-Dep="5203351" httpUri="https://zenodo.org/record/5203351/files/figure.png" pageId="7" pageNumber="646" startId="7.[144,211,1420,1442]" targetBox="[142,1102,161,1387]" targetPageId="7">
<paragraph id="8BF59828FFA3FF81FF0B6B9946BF88D3" blockId="7.[144,1101,1420,1500]" pageId="7" pageNumber="646">
Figure 4.
<taxonomicName id="4C4AE3ABFFA3FF81FF636B99454C88AD" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[248,562,1420,1442]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="646" phylum="Arthropoda" rank="species" species="quinquesolenidiata" status="sp. nov.">
<emphasis id="B93E443AFFA3FF81FF636B99454C88AD" box="[248,562,1420,1442]" italics="true" pageId="7" pageNumber="646">Troglocheles quinquesolenidiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFA3FF81FDA26B9945F688AE" box="[569,648,1420,1441]" pageId="7" pageNumber="646" rank="species">sp. nov.</taxonomicNameLabel>
, adult female, setal arrangement on legs, lateral aspect; (A) leg I; (B) leg II (
<emphasis id="B93E443AFFA3FF81FE416BBC469488B0" box="[474,490,1449,1471]" italics="true" pageId="7" pageNumber="646">ω</emphasis>
,
<emphasis id="B93E443AFFA3FF81FE636BBC457888B0" box="[504,518,1449,1471]" italics="true" pageId="7" pageNumber="646">Φ</emphasis>
,
<emphasis id="B93E443AFFA3FF81FD8E6BBC455C88B0" box="[533,546,1449,1471]" italics="true" pageId="7" pageNumber="646">σ</emphasis>
and
<emphasis id="B93E443AFFA3FF81FDC26BBC451A88B0" box="[601,612,1449,1471]" italics="true" pageId="7" pageNumber="646">ε</emphasis>
designations for solenidia on tarsus, tibia, genu and for famulus, respectively).
</paragraph>
</caption>
<paragraph id="8BF59828FFA3FF81FF0B6BE845E78B3A" blockId="7.[144,1101,1533,1589]" pageId="7" pageNumber="646">solenidia (in brackets) on palpal trochanter, femorogenu, tibia and tarsus 0-2-3-10(1), respectively; tarsal solenidion spiniform, erect.</paragraph>
<paragraph id="8BF59828FFA3FF81FF0B687443328B97" blockId="7.[144,1100,1632,1691]" pageId="7" pageNumber="646">
<emphasis id="B93E443AFFA3FF81FF0B6874466F8B76" box="[144,273,1633,1657]" italics="true" pageId="7" pageNumber="646">Prodorsum.</emphasis>
Naso well-developed (
<figureCitation id="137184ADFFA3FF81FD8F687545F28B77" box="[532,652,1632,1656]" captionStart="Figure 2" captionStartId="5.[144,211,1292,1314]" captionTargetBox="[142,1102,163,1216]" captionTargetId="figure-7@5.[142,1102,162,1216]" captionTargetPageId="5" captionText="Figure 2. Troglocheles quinquesolenidiata sp. nov., adult female; (A) dorsum; (B) venter (for setal notation see Description)." figureDoi="http://doi.org/10.5281/zenodo.5203346" httpUri="https://zenodo.org/record/5203346/files/figure.png" pageId="7" pageNumber="646">Figure 2A</figureCitation>
), with pair of internal vertical setae
<emphasis id="B93E443AFFA3FF81FBB46877433A8B74" box="[1071,1092,1634,1659]" italics="true" pageId="7" pageNumber="646">
v
<subScript id="17CE9A6DFFA3FF81FBA0687F433A8B74" attach="left" box="[1083,1092,1642,1659]" fontSize="7" pageId="7" pageNumber="646">1</subScript>
</emphasis>
. Bothridial setae
<emphasis id="B93E443AFFA3FF81FED2689446168B94" box="[329,360,1665,1691]" italics="true" pageId="7" pageNumber="646">
sc
<subScript id="17CE9A6DFFA3FF81FEC4689F46168B94" attach="left" box="[351,360,1674,1691]" fontSize="7" pageId="7" pageNumber="646">1</subScript>
</emphasis>
filiform, finely pubescent, overlapping insertions of opisthosomal
</paragraph>
<caption id="DF35C8A0FFACFF8EFF4E6B49462488A3" ID-DOI="http://doi.org/10.5281/zenodo.5203353" ID-Zenodo-Dep="5203353" httpUri="https://zenodo.org/record/5203353/files/figure.png" pageId="8" pageNumber="647" startId="8.[213,280,1372,1394]" targetBox="[211,1171,161,1338]" targetPageId="8">
<paragraph id="8BF59828FFACFF8EFF4E6B49462488A3" blockId="8.[213,1170,1372,1452]" pageId="8" pageNumber="647">
Figure 5.
<taxonomicName id="4C4AE3ABFFACFF8EFEDA6B494501887D" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[321,639,1372,1394]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="647" phylum="Arthropoda" rank="species" species="quinquesolenidiata" status="sp. nov.">
<emphasis id="B93E443AFFACFF8EFEDA6B494501887D" box="[321,639,1372,1394]" italics="true" pageId="8" pageNumber="647">Troglocheles quinquesolenidiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFACFF8EFD116B4945A3887E" box="[650,733,1372,1393]" pageId="8" pageNumber="647" rank="species">sp. nov.</taxonomicNameLabel>
, adult female, setal arrangement on legs, lateral aspect; (A) leg III; (B) leg IV (
<emphasis id="B93E443AFFACFF8EFDED6B6C45FA8880" box="[630,644,1401,1423]" italics="true" pageId="8" pageNumber="647">Φ</emphasis>
,
<emphasis id="B93E443AFFACFF8EFD0C6B6C45DA8880" box="[663,676,1401,1423]" italics="true" pageId="8" pageNumber="647">σ</emphasis>
designations for solenidia on tibia and genu, respectively).
</paragraph>
</caption>
<paragraph id="8BF59828FFACFF8EFF4E6BC346048B02" blockId="8.[213,1170,1493,1549]" pageId="8" pageNumber="647">
setae
<emphasis id="B93E443AFFACFF8EFE836BC3465288FF" box="[280,300,1494,1520]" italics="true" pageId="8" pageNumber="647">
c
<subScript id="17CE9A6DFFACFF8EFEB86BCA465288FF" attach="left" box="[291,300,1503,1520]" fontSize="7" pageId="8" pageNumber="647">1</subScript>
</emphasis>
. Length of setae:
<emphasis id="B93E443AFFACFF8EFD9C6BC3456288FF" box="[519,540,1494,1520]" italics="true" pageId="8" pageNumber="647">
v
<subScript id="17CE9A6DFFACFF8EFD886BCA456288FF" attach="left" box="[531,540,1503,1520]" fontSize="7" pageId="8" pageNumber="647">1</subScript>
</emphasis>
86 (7392),
<emphasis id="B93E443AFFACFF8EFD2F6BC345B688FF" box="[692,712,1494,1520]" italics="true" pageId="8" pageNumber="647">
v
<subScript id="17CE9A6DFFACFF8EFD246BCA45B688FF" attach="left" box="[703,712,1503,1520]" fontSize="7" pageId="8" pageNumber="647">2</subScript>
</emphasis>
53 (4659),
<emphasis id="B93E443AFFACFF8EFCFB6BC3440188FF" box="[864,895,1494,1520]" italics="true" pageId="8" pageNumber="647">
sc
<subScript id="17CE9A6DFFACFF8EFCED6BCA440188FF" attach="left" box="[886,895,1503,1520]" fontSize="7" pageId="8" pageNumber="647">1</subScript>
</emphasis>
130 (122148),
<emphasis id="B93E443AFFACFF8EFBA46BC3432388FF" box="[1087,1117,1494,1520]" italics="true" pageId="8" pageNumber="647">
sc
<subScript id="17CE9A6DFFACFF8EFBCF6BCA432388FF" attach="left" box="[1108,1117,1503,1520]" fontSize="7" pageId="8" pageNumber="647">2</subScript>
</emphasis>
126 (119139) µm.
</paragraph>
<paragraph id="8BF59828FFACFF8FFF4E685446B78C32" blockId="8.[213,1170,1601,1691]" lastBlockId="9.[144,1101,166,320]" lastPageId="9" lastPageNumber="648" pageId="8" pageNumber="647">
<emphasis id="B93E443AFFACFF8EFF4E685445028B56" box="[213,636,1601,1625]" italics="true" pageId="8" pageNumber="647">Opisthosomal dorsum and anal region.</emphasis>
Cupules
<emphasis id="B93E443AFFACFF8EFD7C685745858B56" box="[743,763,1602,1625]" italics="true" pageId="8" pageNumber="647">ia</emphasis>
positioned proximad of
<emphasis id="B93E443AFFACFF8EFB89685743598B54" box="[1042,1063,1602,1627]" italics="true" pageId="8" pageNumber="647">
c
<subScript id="17CE9A6DFFACFF8EFB85685F43598B54" attach="left" box="[1054,1063,1610,1627]" fontSize="7" pageId="8" pageNumber="647">2</subScript>
</emphasis>
, laterally between setae
<emphasis id="B93E443AFFACFF8EFEE1687746F08B74" box="[378,398,1634,1659]" italics="true" pageId="8" pageNumber="647">
c
<subScript id="17CE9A6DFFACFF8EFE1E687F46F08B74" attach="left" box="[389,398,1642,1659]" fontSize="7" pageId="8" pageNumber="647">1</subScript>
</emphasis>
and
<emphasis id="B93E443AFFACFF8EFE506874469C8B74" box="[459,482,1633,1659]" italics="true" pageId="8" pageNumber="647">
d
<subScript id="17CE9A6DFFACFF8EFE42687F469C8B74" attach="left" box="[473,482,1642,1659]" fontSize="7" pageId="8" pageNumber="647">1</subScript>
</emphasis>
,
<emphasis id="B93E443AFFACFF8EFE68687745718B76" box="[499,527,1634,1657]" italics="true" pageId="8" pageNumber="647">im</emphasis>
lateral to setae
<emphasis id="B93E443AFFACFF8EFD5D687745A58B74" box="[710,731,1634,1659]" italics="true" pageId="8" pageNumber="647">
e
<subScript id="17CE9A6DFFACFF8EFD49687F45A58B74" attach="left" box="[722,731,1642,1659]" fontSize="7" pageId="8" pageNumber="647">1</subScript>
</emphasis>
,
<emphasis id="B93E443AFFACFF8EFD776877447E8B76" box="[748,768,1634,1657]" italics="true" pageId="8" pageNumber="647">ip</emphasis>
slightly distad and laterad of setae
<emphasis id="B93E443AFFACFF8EFF4E689447998B94" box="[213,231,1665,1691]" italics="true" pageId="8" pageNumber="647">
f
<subScript id="17CE9A6DFFACFF8EFF45689F47998B94" attach="left" box="[222,231,1674,1691]" fontSize="7" pageId="8" pageNumber="647">1</subScript>
</emphasis>
,
<emphasis id="B93E443AFFACFF8EFF6D689446748B96" box="[246,266,1665,1689]" italics="true" pageId="8" pageNumber="647">ih</emphasis>
positioned ventrolaterally, laterodistad of adanal setae
<emphasis id="B93E443AFFACFF8EFCEE689444E78B94" box="[885,921,1665,1691]" italics="true" pageId="8" pageNumber="647">
ad
<subScript id="17CE9A6DFFACFF8EFC0B689F44E78B94" attach="left" box="[912,921,1674,1691]" fontSize="7" pageId="8" pageNumber="647">1</subScript>
</emphasis>
(
<figureCitation id="137184ADFFACFF8EFC336895435E8B97" box="[936,1056,1664,1688]" captionStart="Figure 2" captionStartId="5.[144,211,1292,1314]" captionTargetBox="[142,1102,163,1216]" captionTargetId="figure-7@5.[142,1102,162,1216]" captionTargetPageId="5" captionText="Figure 2. Troglocheles quinquesolenidiata sp. nov., adult female; (A) dorsum; (B) venter (for setal notation see Description)." figureDoi="http://doi.org/10.5281/zenodo.5203346" httpUri="https://zenodo.org/record/5203346/files/figure.png" pageId="8" pageNumber="647">Figure 2A</figureCitation>
). Setae
<emphasis id="B93E443AFFACFF8EFBEF689443F78B94" box="[1140,1161,1665,1691]" italics="true" pageId="8" pageNumber="647">
c
<subScript id="17CE9A6DFFACFF8EFB1B689F43F78B94" attach="left" box="[1152,1161,1674,1691]" fontSize="7" pageId="8" pageNumber="647">1</subScript>
</emphasis>
,
<emphasis id="B93E443AFFADFF8FFF0B6EB347D88DCE" box="[144,166,166,193]" italics="true" pageId="9" pageNumber="648">
d
<subScript id="17CE9A6DFFADFF8FFF066EA547D88DCE" attach="left" box="[157,166,176,193]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
reach about two-thirds of distance to insertion of next seta, seta
<emphasis id="B93E443AFFADFF8FFC096EB244D98DCE" box="[914,935,167,193]" italics="true" pageId="9" pageNumber="648">
e
<subScript id="17CE9A6DFFADFF8FFC056EA544D98DCE" attach="left" box="[926,935,176,193]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
reaches about one-third of distance to insertion of
<emphasis id="B93E443AFFADFF8FFDBF6ED345488DEE" box="[548,566,198,225]" italics="true" pageId="9" pageNumber="648">
f
<subScript id="17CE9A6DFFADFF8FFDB66EC545488DEE" attach="left" box="[557,566,208,225]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
, seta
<emphasis id="B93E443AFFADFF8FFDE36ED345F78DEE" box="[632,649,198,225]" italics="true" pageId="9" pageNumber="648">
f
<subScript id="17CE9A6DFFADFF8FFD1B6EC545F78DEE" attach="left" box="[640,649,208,225]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
reaches about to insertion of
<emphasis id="B93E443AFFADFF8FFC4C6ED344938DEE" box="[983,1005,198,225]" italics="true" pageId="9" pageNumber="648">
h
<subScript id="17CE9A6DFFADFF8FFC7F6EC544938DEE" attach="left" box="[996,1005,208,225]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
. Length of setae:
<emphasis id="B93E443AFFADFF8FFF6F6EF246768C0E" box="[244,264,231,257]" italics="true" pageId="9" pageNumber="648">
c
<subScript id="17CE9A6DFFADFF8FFF646EE546768C0E" attach="left" box="[255,264,240,257]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
65 (5973),
<emphasis id="B93E443AFFADFF8FFE086EF246D68C0E" box="[403,424,231,257]" italics="true" pageId="9" pageNumber="648">
c
<subScript id="17CE9A6DFFADFF8FFE046EE546D68C0E" attach="left" box="[415,424,240,257]" fontSize="7" pageId="9" pageNumber="648">2</subScript>
</emphasis>
147 (142155),
<emphasis id="B93E443AFFADFF8FFDC06EF3450F8C0E" box="[603,625,230,257]" italics="true" pageId="9" pageNumber="648">
d
<subScript id="17CE9A6DFFADFF8FFDF36EE5450F8C0E" attach="left" box="[616,625,240,257]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
69 (6673),
<emphasis id="B93E443AFFADFF8FFD676EF2446F8C0E" box="[764,785,231,257]" italics="true" pageId="9" pageNumber="648">
e
<subScript id="17CE9A6DFFADFF8FFC936EE5446F8C0E" attach="left" box="[776,785,240,257]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
70 (6373),
<emphasis id="B93E443AFFADFF8FFC076EF344D08C0E" box="[924,942,230,257]" italics="true" pageId="9" pageNumber="648">
f
<subScript id="17CE9A6DFFADFF8FFC3E6EE544D08C0E" attach="left" box="[933,942,240,257]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
109 (96132),
<emphasis id="B93E443AFFADFF8FFF0B6F1347DC8C2F" box="[144,162,262,288]" italics="true" pageId="9" pageNumber="648">
f
<subScript id="17CE9A6DFFADFF8FFF026F1A47DC8C2F" attach="left" box="[153,162,271,288]" fontSize="7" pageId="9" pageNumber="648">2</subScript>
</emphasis>
61 (5366),
<emphasis id="B93E443AFFADFF8FFEA96F1346368C2F" box="[306,328,262,288]" italics="true" pageId="9" pageNumber="648">
h
<subScript id="17CE9A6DFFADFF8FFEA46F1A46368C2F" attach="left" box="[319,328,271,288]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
129 (119138),
<emphasis id="B93E443AFFADFF8FFD9B6F1345698C2F" box="[512,535,262,288]" italics="true" pageId="9" pageNumber="648">
h
<subScript id="17CE9A6DFFADFF8FFD956F1A45698C2F" attach="left" box="[526,535,271,288]" fontSize="7" pageId="9" pageNumber="648">2</subScript>
</emphasis>
73 (6979),
<emphasis id="B93E443AFFADFF8FFD3C6F1245B98C2F" box="[679,711,263,288]" italics="true" pageId="9" pageNumber="648">
ps
<subScript id="17CE9A6DFFADFF8FFD256F1A45B98C2F" attach="left" box="[702,711,271,288]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
74 (6682),
<emphasis id="B93E443AFFADFF8FFCCC6F1244068C2F" box="[855,888,263,288]" italics="true" pageId="9" pageNumber="648">
ps
<subScript id="17CE9A6DFFADFF8FFCF46F1A44068C2F" attach="left" box="[879,888,271,288]" fontSize="7" pageId="9" pageNumber="648">2</subScript>
</emphasis>
55 (4659),
<emphasis id="B93E443AFFADFF8FFB936F1243568C2F" box="[1032,1064,263,288]" italics="true" pageId="9" pageNumber="648">
ps
<subScript id="17CE9A6DFFADFF8FFB846F1A43568C2F" attach="left" box="[1055,1064,271,288]" fontSize="7" pageId="9" pageNumber="648">3</subScript>
</emphasis>
40 (3941),
<emphasis id="B93E443AFFADFF8FFF696F3346688C4F" box="[242,278,294,320]" italics="true" pageId="9" pageNumber="648">
ad
<subScript id="17CE9A6DFFADFF8FFE966F3A46688C4F" attach="left" box="[269,278,303,320]" fontSize="7" pageId="9" pageNumber="648">1</subScript>
</emphasis>
62 (6063) µm.
</paragraph>
<paragraph id="8BF59828FFADFF8FFF0B6F7946678CAB" blockId="9.[144,1017,364,420]" pageId="9" pageNumber="648">
<emphasis id="B93E443AFFADFF8FFF0B6F7946778C8B" box="[144,265,364,388]" italics="true" pageId="9" pageNumber="648">Podosoma.</emphasis>
Coxae I, II, III, IV with 3-1-5-3 finely pubescent setae, respectively (
<figureCitation id="137184ADFFADFF8FFF036F9946728CAB" box="[152,268,396,420]" captionStart="Figure 2" captionStartId="5.[144,211,1292,1314]" captionTargetBox="[142,1102,163,1216]" captionTargetId="figure-7@5.[142,1102,162,1216]" captionTargetPageId="5" captionText="Figure 2. Troglocheles quinquesolenidiata sp. nov., adult female; (A) dorsum; (B) venter (for setal notation see Description)." figureDoi="http://doi.org/10.5281/zenodo.5203346" httpUri="https://zenodo.org/record/5203346/files/figure.png" pageId="9" pageNumber="648">Figure 2B</figureCitation>
).
</paragraph>
<paragraph id="8BF59828FFADFF8FFF0B6FC6461A8F45" blockId="9.[144,1101,466,586]" pageId="9" pageNumber="648">
<emphasis id="B93E443AFFADFF8FFF0B6FC6464C8CE5" box="[144,306,467,491]" italics="true" pageId="9" pageNumber="648">Genital region.</emphasis>
Genital valves each with five finely pubescent genital setae (
<emphasis id="B93E443AFFADFF8FFC5C6FC644AA8CE5" box="[967,980,467,490]" italics="true" pageId="9" pageNumber="648">g</emphasis>
) of similar length, about 26 µm, arranged evenly along medial edge of valve. Five pairs of pointed aggenital (
<emphasis id="B93E443AFFADFF8FFE9C6C06465C8F25" box="[263,290,531,554]" italics="true" pageId="9" pageNumber="648">ag</emphasis>
) setae of similar length, about 66 (5376) µm. Length of genital valves 141 (135148) µm.
</paragraph>
<paragraph id="8BF59828FFADFF8FFF0B6C6C44DF89C1" blockId="9.[144,1101,632,1230]" pageId="9" pageNumber="648">
<emphasis id="B93E443AFFADFF8FFF0B6C6C47B28F9F" box="[144,204,633,656]" italics="true" pageId="9" pageNumber="648">Legs.</emphasis>
Leg
<date id="FFF4BEE8FFADFF8FFE9D6C6C46338F9F" box="[262,333,632,656]" pageId="9" pageNumber="648" value="1892-01">I 1892</date>
(18242032) µm long, about 2.10 (1.962.28) times as long as idiosoma. Empodia of all legs setulose, longer than claws, no ventrobasal clawlets on claws. Trochanters I strikingly elongated. Tarsus II, III, IV each with oar-like subapical setae typical of
<taxonomicName id="4C4AE3ABFFADFF8FFEFF6CCD46938FFF" authorityName="Zacharda" authorityYear="1980" box="[356,493,728,752]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="648" phylum="Arthropoda" rank="genus">
<emphasis id="B93E443AFFADFF8FFEFF6CCD46938FFF" box="[356,493,728,752]" italics="true" pageId="9" pageNumber="648">Troglocheles</emphasis>
</taxonomicName>
(
<figureCitation id="137184ADFFADFF8FFE606CCD450C8FFF" box="[507,626,728,752]" captionStart="Figure 3" captionStartId="6.[213,280,1417,1439]" captionTargetBox="[215,1167,163,1340]" captionTargetId="figure-6@6.[214,1168,162,1341]" captionTargetPageId="6" captionText="Figure 3. Troglocheles quinquesolenidiata sp. nov., adult female; (A) chelicera, lateral aspect; (B) rhagidial organ I, dorsal aspect; (C) rhagidial organ II, dorsal aspect; (D) apex of tarsus III with pubescent oar-like iteral (it) setae, lateral aspect; (E) dorsodistal lanceolate solenidion (Φ) on tibia II; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203349" httpUri="https://zenodo.org/record/5203349/files/figure.png" pageId="9" pageNumber="648">Figure 3D</figureCitation>
). Number of setae and solenidia [solenidia and famulus (
<emphasis id="B93E443AFFADFF8FFEB06CED46498E1F" box="[299,311,760,784]" italics="true" pageId="9" pageNumber="648">ε</emphasis>
) bracketed], respectively, on legs I-II-III-IV (
<figureCitation id="137184ADFFADFF8FFCAA6CED44E38E1F" box="[817,925,760,784]" captionStart="Figure 4" captionStartId="7.[144,211,1420,1442]" captionTargetBox="[142,1102,161,1387]" captionTargetId="figure-7@7.[142,1072,160,1387]" captionTargetPageId="7" captionText="Figure 4. Troglocheles quinquesolenidiata sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg I; (B) leg II (ω, Φ, σ and ε designations for solenidia on tarsus, tibia, genu and for famulus, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203351" httpUri="https://zenodo.org/record/5203351/files/figure.png" pageId="9" pageNumber="648">Figures 4</figureCitation>
,
<figureCitation id="137184ADFFADFF8FFC326CED44C68E1F" box="[937,952,760,784]" captionStart="Figure 5" captionStartId="8.[213,280,1372,1394]" captionTargetBox="[211,1171,161,1338]" captionTargetId="figure-6@8.[211,1171,161,1339]" captionTargetPageId="8" captionText="Figure 5. Troglocheles quinquesolenidiata sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg III; (B) leg IV (Φ, σ designations for solenidia on tibia and genu, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203353" httpUri="https://zenodo.org/record/5203353/files/figure.png" pageId="9" pageNumber="648">5</figureCitation>
): trochanters 1-1-2-2; basifemora + telofemora 4+5-6+5-4+4-3+4; genua 14(1)-8(1)-5(1)-6; tibiae 11(2)-7(2)-6(2)-5(1); tarsi 17(5+
<emphasis id="B93E443AFFADFF8FFE696D2D46808E5F" box="[498,510,824,848]" italics="true" pageId="9" pageNumber="648">ε</emphasis>
)-17(4+
<emphasis id="B93E443AFFADFF8FFDCE6D2D451F8E5F" box="[597,609,824,848]" italics="true" pageId="9" pageNumber="648">ε</emphasis>
)-16-15. Spiniform solenidia on leg segments very small. Genua I and II each with one erect spiniform distoventral solenidion (
<emphasis id="B93E443AFFADFF8FFBB06D4D43478E7F" box="[1067,1081,856,880]" italics="true" pageId="9" pageNumber="648">σ</emphasis>
); genu III with one small spiniform medioventral solenidion. Tibia I with one erect spiniform dorsodistimedial solenidion (
<emphasis id="B93E443AFFADFF8FFDCB6D8D45218EBF" box="[592,607,920,944]" italics="true" pageId="9" pageNumber="648">Φ</emphasis>
), and one dorsodistal rhagidial solenidion; tibia II with one spiniform erect dorsomedial solenidion, and one lanceolate dorsodistal solenidion recessed in deep pit with small surface pore (
<figureCitation id="137184ADFFADFF8FFCB46DC244DA8EE0" box="[815,932,983,1007]" captionStart="Figure 3" captionStartId="6.[213,280,1417,1439]" captionTargetBox="[215,1167,163,1340]" captionTargetId="figure-6@6.[214,1168,162,1341]" captionTargetPageId="6" captionText="Figure 3. Troglocheles quinquesolenidiata sp. nov., adult female; (A) chelicera, lateral aspect; (B) rhagidial organ I, dorsal aspect; (C) rhagidial organ II, dorsal aspect; (D) apex of tarsus III with pubescent oar-like iteral (it) setae, lateral aspect; (E) dorsodistal lanceolate solenidion (Φ) on tibia II; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203349" httpUri="https://zenodo.org/record/5203349/files/figure.png" pageId="9" pageNumber="648">Figure 3E</figureCitation>
); tibia III with two adjacent erect spiniform dorsomedial solenidia; tibia IV with one erect spiniform dorsoproximal solenidion. Tarsus I slender, its tip slightly tapers in lateral view, ratio length to width 7.55, with five rhagidial solenidia (
<emphasis id="B93E443AFFADFF8FFC806A2244528940" box="[795,812,1079,1103]" italics="true" pageId="9" pageNumber="648">ω</emphasis>
) lying obliquely in separate depressions dorsodistally (
<figureCitation id="137184ADFFADFF8FFE776A43451E8961" box="[492,608,1110,1134]" captionStart="Figure 3" captionStartId="6.[213,280,1417,1439]" captionTargetBox="[215,1167,163,1340]" captionTargetId="figure-6@6.[214,1168,162,1341]" captionTargetPageId="6" captionText="Figure 3. Troglocheles quinquesolenidiata sp. nov., adult female; (A) chelicera, lateral aspect; (B) rhagidial organ I, dorsal aspect; (C) rhagidial organ II, dorsal aspect; (D) apex of tarsus III with pubescent oar-like iteral (it) setae, lateral aspect; (E) dorsodistal lanceolate solenidion (Φ) on tibia II; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203349" httpUri="https://zenodo.org/record/5203349/files/figure.png" pageId="9" pageNumber="648">Figure 3B</figureCitation>
), stellate famulus (
<emphasis id="B93E443AFFADFF8FFCAF6A42443E8960" box="[820,832,1111,1135]" italics="true" pageId="9" pageNumber="648">ε</emphasis>
) subtending the second proximal rhagidial solenidion antiaxially; tarsus II with four rhagidial solenidia lying in separate oblique depressions (
<figureCitation id="137184ADFFADFF8FFE676A83450F89A1" box="[508,625,1174,1198]" captionStart="Figure 3" captionStartId="6.[213,280,1417,1439]" captionTargetBox="[215,1167,163,1340]" captionTargetId="figure-6@6.[214,1168,162,1341]" captionTargetPageId="6" captionText="Figure 3. Troglocheles quinquesolenidiata sp. nov., adult female; (A) chelicera, lateral aspect; (B) rhagidial organ I, dorsal aspect; (C) rhagidial organ II, dorsal aspect; (D) apex of tarsus III with pubescent oar-like iteral (it) setae, lateral aspect; (E) dorsodistal lanceolate solenidion (Φ) on tibia II; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203349" httpUri="https://zenodo.org/record/5203349/files/figure.png" pageId="9" pageNumber="648">Figure 3C</figureCitation>
), and small spiniform famulus (
<emphasis id="B93E443AFFADFF8FFC4C6A82449D89A0" box="[983,995,1175,1199]" italics="true" pageId="9" pageNumber="648">ε</emphasis>
) inserted between the first and second proximal rhagidial solenidion antiaxially.
</paragraph>
<paragraph id="8BF59828FFADFF8FFF0B6B0F478B883D" blockId="9.[144,245,1306,1330]" box="[144,245,1306,1330]" pageId="9" pageNumber="648">
<emphasis id="B93E443AFFADFF8FFF0B6B0F478B883D" box="[144,245,1306,1330]" italics="true" pageId="9" pageNumber="648">Affinities</emphasis>
</paragraph>
<paragraph id="8BF59828FFADFF8FFF0B6B57433388B6" blockId="9.[144,1101,1346,1689]" pageId="9" pageNumber="648">
<taxonomicName id="4C4AE3ABFFADFF8FFF0B6B5746938855" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[144,493,1346,1370]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="648" phylum="Arthropoda" rank="species" species="quinquesolenidiata" status="sp. nov.">
<emphasis id="B93E443AFFADFF8FFF0B6B5746938855" box="[144,493,1346,1370]" italics="true" pageId="9" pageNumber="648">Troglocheles quinquesolenidiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFADFF8FFE6F6B5745368856" box="[500,584,1346,1369]" pageId="9" pageNumber="648" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the
<taxonomicName id="4C4AE3ABFFADFF8FFD6F6B5644318855" baseAuthorityName="Willmann" baseAuthorityYear="1932" box="[756,847,1347,1370]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="648" phylum="Arthropoda" rank="species" species="strasseri">
<emphasis id="B93E443AFFADFF8FFD6F6B5644318855" box="[756,847,1347,1370]" italics="true" pageId="9" pageNumber="648">strasseri</emphasis>
</taxonomicName>
species group because it has 3-1-5-3 setae on coxae I-II-III-IV, respectively, the spiniform solenidion on tibia I is positioned dorsomedially and the distance between insertions of the cheliceral setae on the fixed digit is almost the same as the length of the proximal cheliceral seta.
</paragraph>
<paragraph id="8BF59828FFADFF8CFF206BD4433F8DF1" blockId="9.[144,1101,1346,1689]" lastBlockId="10.[213,1170,166,604]" lastPageId="10" lastPageNumber="649" pageId="9" pageNumber="648">
<taxonomicName id="4C4AE3ABFFADFF8FFF206BD4456888D6" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[187,534,1473,1497]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="648" phylum="Arthropoda" rank="species" species="quinquesolenidiata" status="sp. nov.">
<emphasis id="B93E443AFFADFF8FFF206BD4456888D6" box="[187,534,1473,1497]" italics="true" pageId="9" pageNumber="648">Troglocheles quinquesolenidiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFADFF8FFD806BD7451088D6" box="[539,622,1474,1497]" pageId="9" pageNumber="648" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C4AE3ABFFADFF8FFCB46BD744B688D6" authorityName="Zacharda" authorityYear="2000" box="[815,968,1473,1497]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="648" phylum="Arthropoda" rank="species" species="archetypica">
<emphasis id="B93E443AFFADFF8FFCB46BD744B688D6" box="[815,968,1473,1497]" italics="true" pageId="9" pageNumber="648">T. archetypica</emphasis>
</taxonomicName>
from which it can be distinguished by the following principal characteristics. (1) The rhagidial organ I comprises five rhagidial solenidia; in
<taxonomicName id="4C4AE3ABFFADFF8FFD136817445D8B16" authorityName="Zacharda" authorityYear="2000" box="[648,803,1537,1561]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="648" phylum="Arthropoda" rank="species" species="archetypica">
<emphasis id="B93E443AFFADFF8FFD136817445D8B16" box="[648,803,1537,1561]" italics="true" pageId="9" pageNumber="648">T. archetypica</emphasis>
</taxonomicName>
only four rhagidial solenidia are in the rhagidial organ I. (2) Tibia IV with one minute spiniform dorsal medioproximal solenidion; in
<taxonomicName id="4C4AE3ABFFADFF8FFE6A685745EF8B56" authorityName="Zacharda" authorityYear="2000" box="[497,657,1601,1625]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="648" phylum="Arthropoda" rank="species" species="archetypica">
<emphasis id="B93E443AFFADFF8FFE6A685745EF8B56" box="[497,657,1601,1625]" italics="true" pageId="9" pageNumber="648">T. archetypica</emphasis>
</taxonomicName>
the spiniform solenidion on tibia IV is absent. (3) The ratio of leg I length to idiosomal length is 2.10 (1.962.28); in
<taxonomicName id="4C4AE3ABFFADFF8FFF0B689446538B96" authorityName="Zacharda" authorityYear="2000" box="[144,301,1665,1689]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="648" phylum="Arthropoda" rank="species" species="archetypica">
<emphasis id="B93E443AFFADFF8FFF0B689446538B96" box="[144,301,1665,1689]" italics="true" pageId="9" pageNumber="648">T. archetypica</emphasis>
</taxonomicName>
the ratio of leg I length to idiosomal length is 0.81.36. (4) The ratio of tarsus I length to width is 7.55 (6.907.70); in
<taxonomicName id="4C4AE3ABFFAEFF8CFC906EB244D68DB1" authorityName="Zacharda" authorityYear="2000" box="[779,936,166,190]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="archetypica">
<emphasis id="B93E443AFFAEFF8CFC906EB244D68DB1" box="[779,936,166,190]" italics="true" pageId="10" pageNumber="649">T. archetypica</emphasis>
</taxonomicName>
the ratio of tarsus I length to width is only 4.414.72. (5) The ratio of palpal tarsus length to width is 5.23 (5.05.4); in
<taxonomicName id="4C4AE3ABFFAEFF8CFEF96EF246838DF1" authorityName="Zacharda" authorityYear="2000" box="[354,509,230,254]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="archetypica">
<emphasis id="B93E443AFFAEFF8CFEF96EF246838DF1" box="[354,509,230,254]" italics="true" pageId="10" pageNumber="649">T. archetypica</emphasis>
</taxonomicName>
the ratio of palpal tarsus length to width is 2.73.0.
</paragraph>
<paragraph id="8BF59828FFAEFF8CFE9B6F1345708F53" blockId="10.[213,1170,166,604]" pageId="10" pageNumber="649">
<taxonomicName id="4C4AE3ABFFAEFF8CFE9B6F1345238C11" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[256,605,262,286]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="quinquesolenidiata" status="sp. nov.">
<emphasis id="B93E443AFFAEFF8CFE9B6F1345238C11" box="[256,605,262,286]" italics="true" pageId="10" pageNumber="649">Troglocheles quinquesolenidiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFAEFF8CFDFE6F1345C58C12" box="[613,699,262,285]" pageId="10" pageNumber="649" rank="species">sp. nov.</taxonomicNameLabel>
is also similar to
<taxonomicName id="4C4AE3ABFFAEFF8CFC1E6F12448E8C11" authorityName="Zacharda" authorityYear="1987" box="[901,1008,262,286]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B93E443AFFAEFF8CFC1E6F12448E8C11" box="[901,1008,262,286]" italics="true" pageId="10" pageNumber="649">T. vandeli</emphasis>
</taxonomicName>
from which it can be distinguished by the following principal characteristics. (1) The rhagidial organ I comprises five rhagidial solenidia; in
<taxonomicName id="4C4AE3ABFFAEFF8CFDE66F5245988C51" authorityName="Zacharda" authorityYear="1987" box="[637,742,326,350]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B93E443AFFAEFF8CFDE66F5245988C51" box="[637,742,326,350]" italics="true" pageId="10" pageNumber="649">T. vandeli</emphasis>
</taxonomicName>
only four rhagidial solenidia are in the rhagidial organ I. (2) The spiniform solenidia on tibiae I, II and III are short, minute; in
<taxonomicName id="4C4AE3ABFFAEFF8CFF6B6F9346278C92" authorityName="Zacharda" authorityYear="1987" box="[240,345,389,413]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B93E443AFFAEFF8CFF6B6F9346278C92" box="[240,345,389,413]" italics="true" pageId="10" pageNumber="649">T. vandeli</emphasis>
</taxonomicName>
they are strikingly long. (3) The two dorsomedial spiniform solenidia lying in tandem on tibia III are of the same length; in
<taxonomicName id="4C4AE3ABFFAEFF8CFD606FB3441B8CB2" authorityName="Zacharda" authorityYear="1987" box="[763,869,421,445]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B93E443AFFAEFF8CFD606FB3441B8CB2" box="[763,869,421,445]" italics="true" pageId="10" pageNumber="649">T. vandeli</emphasis>
</taxonomicName>
the proximal solenidion is conspicuously longer than the distal one. (4) The spiniform famulus (
<emphasis id="B93E443AFFAEFF8CFC546FD044A58CD2" box="[975,987,453,477]" italics="true" pageId="10" pageNumber="649">ε</emphasis>
) in the rhagidial organ II is positioned laterad of the proximal rhagidial solenidion; in
<taxonomicName id="4C4AE3ABFFAEFF8CFC686FF343218CF2" authorityName="Zacharda" authorityYear="1987" box="[1011,1119,485,509]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B93E443AFFAEFF8CFC686FF343218CF2" box="[1011,1119,485,509]" italics="true" pageId="10" pageNumber="649">T. vandeli</emphasis>
</taxonomicName>
it is proximal, subtending the proximal rhagidial solenidion. (5) The prodorsal bothridial setae
<emphasis id="B93E443AFFAEFF8CFE8C6C3346488F30" box="[279,310,550,575]" italics="true" pageId="10" pageNumber="649">
sc
<subScript id="17CE9A6DFFAEFF8CFEB66C3B46488F30" attach="left" box="[301,310,558,575]" fontSize="7" pageId="10" pageNumber="649">1</subScript>
</emphasis>
overlap insertions of opisthosomal setae
<emphasis id="B93E443AFFAEFF8CFC826C3344508F30" box="[793,814,550,575]" italics="true" pageId="10" pageNumber="649">
c
<subScript id="17CE9A6DFFAEFF8CFCBE6C3B44508F30" attach="left" box="[805,814,558,575]" fontSize="7" pageId="10" pageNumber="649">1</subScript>
</emphasis>
; in
<taxonomicName id="4C4AE3ABFFAEFF8CFCFB6C3344B38F32" authorityName="Zacharda" authorityYear="1987" box="[864,973,549,573]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B93E443AFFAEFF8CFCFB6C3344B38F32" box="[864,973,549,573]" italics="true" pageId="10" pageNumber="649">T. vandeli</emphasis>
</taxonomicName>
they overlap the disjugal suture only slightly.
</paragraph>
<paragraph id="8BF59828FFAEFF8CFF4E6CB146338FB3" blockId="10.[213,333,676,700]" box="[213,333,676,700]" pageId="10" pageNumber="649">
<emphasis id="B93E443AFFAEFF8CFF4E6CB146338FB3" box="[213,333,676,700]" italics="true" pageId="10" pageNumber="649">Etymology</emphasis>
</paragraph>
<paragraph id="8BF59828FFAEFF8CFF4E6CDE43708E0C" blockId="10.[213,1169,715,772]" pageId="10" pageNumber="649">
The epithet
<taxonomicName id="4C4AE3ABFFAEFF8CFEC66CD945578FEB" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[349,553,716,740]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="quinquesolenidiata">
<emphasis id="B93E443AFFAEFF8CFEC66CD945578FEB" box="[349,553,716,740]" italics="true" pageId="10" pageNumber="649">quinquesolenidiata</emphasis>
</taxonomicName>
reflects five rhagidial solenidia in the rhagidial organ I which is a unique morphological character in the genus. Feminine gender.
</paragraph>
<paragraph id="8BF59828FFAEFF8CFF4E6D5E46498E6C" blockId="10.[213,311,843,867]" box="[213,311,843,867]" pageId="10" pageNumber="649">
<emphasis id="B93E443AFFAEFF8CFF4E6D5E46498E6C" box="[213,311,843,867]" italics="true" pageId="10" pageNumber="649">Remarks</emphasis>
</paragraph>
<paragraph id="8BF59828FFAEFF8CFF4E6D6646C689C6" blockId="10.[213,1170,882,1225]" pageId="10" pageNumber="649">
<taxonomicName id="4C4AE3ABFFAEFF8CFF4E6D6645478E84" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[213,569,883,907]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="quinquesolenidiata" status="sp. nov.">
<emphasis id="B93E443AFFAEFF8CFF4E6D6645478E84" box="[213,569,883,907]" italics="true" pageId="10" pageNumber="649">Troglocheles quinquesolenidiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFAEFF8CFDDD6D6645DA8E85" box="[582,676,883,906]" pageId="10" pageNumber="649" rank="species">sp. nov.</taxonomicNameLabel>
, having a higher number of solenidia in rhagidial organ I and relatively elongated appendages, is evidently morphologically adapted to life in the subterranean habitat whereas
<taxonomicName id="4C4AE3ABFFAEFF8CFCD66DA644908EC5" authorityName="Zacharda" authorityYear="2000" box="[845,1006,946,970]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="archetypica">
<emphasis id="B93E443AFFAEFF8CFCD66DA644908EC5" box="[845,1006,946,970]" italics="true" pageId="10" pageNumber="649">T. archetypica</emphasis>
</taxonomicName>
is an epigeic free-living mite that lacks these morphological adaptations. For further details one may consult the discussion in
<bibRefCitation id="EFDBE5D9FFAEFF8CFDDF6DE744768905" author="Zacharda M" box="[580,776,1010,1034]" pageId="10" pageNumber="649" pagination="463 - 478" refId="ref11291" refString="Zacharda M. 2000. New species of the genus Troglocheles (Acari: Prostigmata: Rhagidiidae) from Oetztal Alps, with a key to adult species of the genus. J Nat Hist. 34: 463 - 478." type="journal article" year="2000">Zacharda (2000)</bibRefCitation>
, where the phylogenetic relationships between species of the genus
<taxonomicName id="4C4AE3ABFFAEFF8CFDE96A0745858925" authorityName="Zacharda" authorityYear="1980" box="[626,763,1042,1066]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="genus">
<emphasis id="B93E443AFFAEFF8CFDE96A0745858925" box="[626,763,1042,1066]" italics="true" pageId="10" pageNumber="649">Troglocheles</emphasis>
</taxonomicName>
are also discussed. In light of our current knowledge about the known
<taxonomicName id="4C4AE3ABFFAEFF8CFD136A2744128945" box="[648,876,1074,1098]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93E443AFFAEFF8CFD136A27446F8945" box="[648,785,1074,1098]" italics="true" pageId="10" pageNumber="649">Troglocheles</emphasis>
species
</taxonomicName>
,
<taxonomicName id="4C4AE3ABFFAEFF8CFCE06A2643168945" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[891,1128,1074,1098]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="quinquesolenidiata" status="sp. nov.">
<emphasis id="B93E443AFFAEFF8CFCE06A2643168945" box="[891,1128,1074,1098]" italics="true" pageId="10" pageNumber="649">T. quinquesolenidiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFAEFF8CFBE86A26467B8966" pageId="10" pageNumber="649" rank="species">sp. nov.</taxonomicNameLabel>
together with
<taxonomicName id="4C4AE3ABFFAEFF8CFE346A4645658965" authorityName="Zacharda" authorityYear="1987" box="[431,539,1106,1130]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="B93E443AFFAEFF8CFE346A4645658965" box="[431,539,1106,1130]" italics="true" pageId="10" pageNumber="649">T. vandeli</emphasis>
</taxonomicName>
are morphologically intermediate between the epigean
<taxonomicName id="4C4AE3ABFFAEFF8CFF4E6A6646118985" authorityName="Zacharda" authorityYear="2000" box="[213,367,1138,1162]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="archetypica">
<emphasis id="B93E443AFFAEFF8CFF4E6A6646118985" box="[213,367,1138,1162]" italics="true" pageId="10" pageNumber="649">T. archetypica</emphasis>
</taxonomicName>
and the troglobitic representatives of the
<taxonomicName id="4C4AE3ABFFAEFF8CFCA76A6644E98985" baseAuthorityName="Willmann" baseAuthorityYear="1932" box="[828,919,1139,1162]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="strasseri">
<emphasis id="B93E443AFFAEFF8CFCA76A6644E98985" box="[828,919,1139,1162]" italics="true" pageId="10" pageNumber="649">strasseri</emphasis>
</taxonomicName>
species group inhabiting solely caves in the Alps. However, actual phylogenetic relationships between these taxa are not known.
</paragraph>
</subSubSection>
</treatment>
</document>