treatments-xml/data/03/E3/29/03E3291C1C4092405C8DFF73C0F852B8.xml

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<mods:title id="C4905A498E3131BF8D8842BBA37657C8">A new leafhopper genus and four new species from the Grassland Biome of South Africa (Hemiptera, Cicadellidae)</mods:title>
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<treatment id="03E3291C1C4092405C8DFF73C0F852B8" ID-DOI="http://doi.org/10.5281/zenodo.5294710" ID-Zenodo-Dep="5294710" LSID="urn:lsid:plazi:treatment:03E3291C1C4092405C8DFF73C0F852B8" httpUri="http://treatment.plazi.org/id/03E3291C1C4092405C8DFF73C0F852B8" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36">
<subSubSection id="C350CB811C4092435C8DFF73C2F354DB" box="[151,402,151,177]" pageId="1" pageNumber="36" type="nomenclature">
<paragraph id="8BF5980A1C4092435C8DFF73C2F354DB" blockId="1.[151,402,151,210]" box="[151,402,151,177]" pageId="1" pageNumber="36">
<heading id="D0BD2F661C4092435C8DFF73C2F354DB" bold="true" box="[151,402,151,177]" fontSize="11" level="1" pageId="1" pageNumber="36" reason="1">
<emphasis id="B93E44181C4092435C8DFF73C2F354DB" bold="true" box="[151,402,151,177]" pageId="1" pageNumber="36">
<taxonomicName id="4C4AE3891C4092435C8DFF73C22054DB" ID-CoL="8RGTH" authority="Stiller, 2011" authorityName="Stiller" authorityYear="2011" box="[151,321,151,177]" class="Insecta" family="Cicadellidae" genus="Tetramelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93E44181C4092435C8DFF73C22054DB" bold="true" box="[151,321,151,177]" italics="true" pageId="1" pageNumber="36">Tetramelasma</emphasis>
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<taxonomicNameLabel id="A20DF9631C4092435D52FF73C2F354DB" box="[328,402,151,177]" pageId="1" pageNumber="36" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C350CB811C4092435C8DFF5EC228555D" pageId="1" pageNumber="36" type="description">
<paragraph id="8BF5980A1C4092435C8DFF5EC21354B8" blockId="1.[151,402,151,210]" box="[151,370,186,210]" pageId="1" pageNumber="36">
(
<figureCitation id="1371848F1C4092435C85FF5EC27054B8" box="[159,273,186,210]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">Figs 9a–g</figureCitation>
; 
<figureCitation id="1371848F1C4092435D07FF5EC20454B8" box="[285,357,186,210]" captionStart="FIGURE 10" captionStartId="15.[151,250,1445,1468]" captionTargetBox="[213,1369,183,1434]" captionTargetId="figure-294@15.[213,1373,183,1437]" captionTargetPageId="15" captionText="FIGURE 10a–j. Tetramelasma tanyphysis sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c-e, sternite 7, ventrally; f, valvula 1; g, valvula 2; h, valvula 1 detail; i, valvula 2 detail; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290245" httpUri="https://zenodo.org/record/5290245/files/figure.png" pageId="1" pageNumber="36">10a–k</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4092435C8DFEE7C1045572" blockId="1.[151,613,258,311]" box="[151,613,258,280]" pageId="1" pageNumber="36">
<typeStatus id="54F126A81C4092435C8DFEE7C3AA5572" box="[151,203,259,280]" pageId="1" pageNumber="36">Type</typeStatus>
species: 
<taxonomicName id="4C4AE3891C4092435D3CFEE6C1475572" authority="Stiller, 2011" authorityName="Stiller" authorityYear="2011" box="[294,550,258,280]" class="Insecta" family="Cicadellidae" genus="Tetramelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="species" species="tanyphysis" status="sp. nov.">
<emphasis id="B93E44181C4092435D3CFEE6C1475572" box="[294,550,258,280]" italics="true" pageId="1" pageNumber="36">Tetramelasma tanyphysis</emphasis>
</taxonomicName>
<emphasis id="B93E44181C4092435E36FEE7C1045572" bold="true" box="[556,613,259,280]" pageId="1" pageNumber="36">
<taxonomicNameLabel id="A20DF9631C4092435E36FEE7C1045572" box="[556,613,259,280]" pageId="1" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BF5980A1C4092435C8DFEC5C228555D" blockId="1.[151,613,258,311]" box="[151,329,289,311]" pageId="1" pageNumber="36">Gender feminine.</paragraph>
</subSubSection>
<subSubSection id="C350CB811C4092435C8DFE80C23D55F5" pageId="1" pageNumber="36" type="etymology">
<paragraph id="8BF5980A1C4092435C8DFE80C23D55F5" blockId="1.[151,1437,356,1892]" pageId="1" pageNumber="36">
<emphasis id="B93E44181C4092435C8DFE80C27C5516" bold="true" box="[151,285,356,380]" inLineHeading="true" pageId="1" pageNumber="36" reason="1">Etymology.</emphasis>
Compound word in Greek, 
<emphasis id="B93E44181C4092435E4FFE81C1EB5516" box="[597,650,357,380]" italics="true" pageId="1" pageNumber="36">tetra</emphasis>
, four, 
<emphasis id="B93E44181C4092435EC9FE80C0575516" box="[723,822,356,380]" italics="true" pageId="1" pageNumber="36">melasma</emphasis>
, a black spot, for the four dark markings on the vertex. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C350CB811C4092405CDFFE4FC0F852B8" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36" type="diagnosis">
<paragraph id="8BF5980A1C4092435CDFFE4FC3D2568E" blockId="1.[151,1437,356,1892]" pageId="1" pageNumber="36">
<emphasis id="B93E44181C4092435CDFFE4FC25E55AE" bold="true" box="[197,319,427,452]" inLineHeading="true" pageId="1" pageNumber="36" reason="1">Diagnosis.</emphasis>
Moderately large leafhoppers (
<quantity id="4CB235EF1C4092435E87FE48C19B55A9" box="[669,762,428,452]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="36" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
long), vertex with paired fuscous markings (
<figureCitation id="1371848F1C40924358F5FE48C62455AE" box="[1263,1349,428,452]" captionStart="FIGURE 2" captionStartId="4.[151,249,1729,1752]" captionTargetBox="[218,1369,195,1698]" captionTargetId="figure-141@4.[218,1369,195,1705]" captionTargetPageId="4" captionText="FIGURE 2a–l. Tetramelasma litopyx sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c–e, sternite 7, ventrally; f, hind wing; g, valvula 1; h, valvula 2; i, valvula 1 detail; j, valvula 3 detail; k, valvula 2 detail; l, valvula 3 (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290223" httpUri="https://zenodo.org/record/5290223/files/figure.png" pageId="1" pageNumber="36">Figs 2a</figureCitation>
; 
<figureCitation id="1371848F1C409243594BFE48C60F55AE" box="[1361,1390,428,452]" captionStart="FIGURE 3" captionStartId="6.[151,250,1647,1670]" captionTargetBox="[151,1425,770,1619]" captionTargetId="figure-251@6.[151,1436,770,1623]" captionTargetPageId="6" captionText="FIGURE 3 a–d. Tetramelasma nodosatha sp. n. male. a, Habitus, in situ (scale bar = 2 mm); b, pygofer, dorsally; c, genital capsule, laterally; d, pygofer, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290227" httpUri="https://zenodo.org/record/5290227/files/figure.png" pageId="1" pageNumber="36">3a</figureCitation>
; 
<figureCitation id="1371848F1C4092435960FE48C6F655AE" box="[1402,1431,428,452]" captionStart="FIGURE 5" captionStartId="8.[151,249,1092,1115]" captionTargetBox="[160,1423,267,1066]" captionTargetId="figure-504@8.[160,1426,267,1069]" captionTargetPageId="8" captionText="FIGURE 5a–f. Tetramelasma nodosatha sp. n. female. a, Habitus (scale bar = 2 mm); b–c, sternite 7, ventrally, in situ; d–f, sternite 7, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290231" httpUri="https://zenodo.org/record/5290231/files/figure.png" pageId="1" pageNumber="36">5a</figureCitation>
; 
<figureCitation id="1371848F1C4092435C8DFE2BC3B9558D" box="[151,216,463,487]" captionStart="FIGURE 6" captionStartId="10.[151,250,1635,1658]" captionTargetBox="[249,1334,184,1610]" captionTargetId="figure-186@10.[249,1339,184,1614]" captionTargetPageId="10" captionText="FIGURE 6a–l. Tetramelasma nodosatha sp. n. female. a, Habitus, submacropter; b, habitus, macropter (scale bar = 2 mm); c– g, sternite 7, ventrally; h, valvula 1; i, valvula 1 detail; j, valvula 2 detail; k, valvula 3; l, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290235" httpUri="https://zenodo.org/record/5290235/files/figure.png" pageId="1" pageNumber="36">6a&amp;b</figureCitation>
; 
<figureCitation id="1371848F1C4092435CF9FE34C261558D" box="[227,256,464,487]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="1" pageNumber="36">7a</figureCitation>
; 
<figureCitation id="1371848F1C4092435D16FE2BC248558D" box="[268,297,463,487]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">9a</figureCitation>
; 
<figureCitation id="1371848F1C4092435D2EFE2BC201558D" box="[308,352,463,487]" captionStart="FIGURE 10" captionStartId="15.[151,250,1445,1468]" captionTargetBox="[213,1369,183,1434]" captionTargetId="figure-294@15.[213,1373,183,1437]" captionTargetPageId="15" captionText="FIGURE 10a–j. Tetramelasma tanyphysis sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c-e, sternite 7, ventrally; f, valvula 1; g, valvula 2; h, valvula 1 detail; i, valvula 2 detail; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290245" httpUri="https://zenodo.org/record/5290245/files/figure.png" pageId="1" pageNumber="36">10a</figureCitation>
). Hind wing about ½ as long and ¾ as wide as tegmina, tegmina reaching apex or beyond apex of abdomen. Tegmina without appendix, or very narrow (
<figureCitation id="1371848F1C4092435F0FFE10C00E5666" box="[789,879,500,524]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="1" pageNumber="36">
Figs 
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</figureCitation>
, 
<figureCitation id="1371848F1C4092435F65FE10C0FB5666" box="[895,922,500,524]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">9f</figureCitation>
). Aedeagus narrow, tubular (
<figureCitation id="1371848F1C40924358F6FE10C60B5666" box="[1260,1386,500,524]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="1" pageNumber="36">
Figs 
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&amp;h
</figureCitation>
, 
<figureCitation id="1371848F1C4092435961FE10C6F65666" box="[1403,1431,500,524]" captionStart="FIGURE 4" captionStartId="7.[151,250,855,878]" captionTargetBox="[154,1421,199,825]" captionTargetId="figure-406@7.[151,1436,195,831]" captionTargetPageId="7" captionText="FIGURE 4 a–e. Tetramelasma nodosatha sp. n. male. a, Genital capsule, ventrally; b, genital capsule, laterally; c, style, posteroventrally, 5–6 teeth; d, style, posteroventrally, 1–2 teeth; e, aedeagus &amp; connective, laterally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290229" httpUri="https://zenodo.org/record/5290229/files/figure.png" pageId="1" pageNumber="36">4e</figureCitation>
, 
<figureCitation id="1371848F1C4092435C8DFDF3C3B25645" box="[151,211,535,559]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="1" pageNumber="36">7e&amp;f</figureCitation>
, 
<figureCitation id="1371848F1C4092435CC5FDF3C39D5645" box="[223,252,535,559]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">9e</figureCitation>
), base fused to connective (e.g. 
<figureCitation id="1371848F1C4092435E73FDF3C1DF5645" box="[617,702,535,559]" captionStart="FIGURE 4" captionStartId="7.[151,250,855,878]" captionTargetBox="[154,1421,199,825]" captionTargetId="figure-406@7.[151,1436,195,831]" captionTargetPageId="7" captionText="FIGURE 4 a–e. Tetramelasma nodosatha sp. n. male. a, Genital capsule, ventrally; b, genital capsule, laterally; c, style, posteroventrally, 5–6 teeth; d, style, posteroventrally, 1–2 teeth; e, aedeagus &amp; connective, laterally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290229" httpUri="https://zenodo.org/record/5290229/files/figure.png" pageId="1" pageNumber="36">Figs 4e</figureCitation>
, 
<figureCitation id="1371848F1C4092435ED0FDF3C1865645" box="[714,743,535,559]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">9e</figureCitation>
), shaft with subapical dorsal gonopore. Plate variable, generally truncate, length either up to ½ as long as pygofer (
<figureCitation id="1371848F1C4092435F3AFDD8C0FC563E" box="[800,925,572,596]" captionStart="FIGURE 3" captionStartId="6.[151,250,1647,1670]" captionTargetBox="[151,1425,770,1619]" captionTargetId="figure-251@6.[151,1436,770,1623]" captionTargetPageId="6" captionText="FIGURE 3 a–d. Tetramelasma nodosatha sp. n. male. a, Habitus, in situ (scale bar = 2 mm); b, pygofer, dorsally; c, genital capsule, laterally; d, pygofer, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290227" httpUri="https://zenodo.org/record/5290227/files/figure.png" pageId="1" pageNumber="36">Figs 3c&amp;d</figureCitation>
, 
<figureCitation id="1371848F1C4092435FB6FDD8C08C563E" box="[940,1005,572,596]" captionStart="FIGURE 4" captionStartId="7.[151,250,855,878]" captionTargetBox="[154,1421,199,825]" captionTargetId="figure-406@7.[151,1436,195,831]" captionTargetPageId="7" captionText="FIGURE 4 a–e. Tetramelasma nodosatha sp. n. male. a, Genital capsule, ventrally; b, genital capsule, laterally; c, style, posteroventrally, 5–6 teeth; d, style, posteroventrally, 1–2 teeth; e, aedeagus &amp; connective, laterally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290229" httpUri="https://zenodo.org/record/5290229/files/figure.png" pageId="1" pageNumber="36">4a&amp;b</figureCitation>
, 
<figureCitation id="1371848F1C4092435FE6FDD8C75F563E" box="[1020,1086,572,596]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">9b&amp;c</figureCitation>
), or about as long as pygofer, viewed laterally (
<figureCitation id="1371848F1C4092435D40FDBBC2B1561D" box="[346,464,607,631]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="1" pageNumber="36">Fig. 1c&amp;d</figureCitation>
). Style either with apophysis elongate, with ventral tooth (
<figureCitation id="1371848F1C4092435847FDBBC7CE561D" box="[1117,1199,607,631]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="1" pageNumber="36">Figs 1f</figureCitation>
, 
<figureCitation id="1371848F1C40924358A6FDBBC7B8561D" box="[1212,1241,607,631]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">9d</figureCitation>
) or short, invaginated posteriorly, variably dentate (
<figureCitation id="1371848F1C4092435E3AFD60C1F856F6" box="[544,665,644,668]" captionStart="FIGURE 4" captionStartId="7.[151,250,855,878]" captionTargetBox="[154,1421,199,825]" captionTargetId="figure-406@7.[151,1436,195,831]" captionTargetPageId="7" captionText="FIGURE 4 a–e. Tetramelasma nodosatha sp. n. male. a, Genital capsule, ventrally; b, genital capsule, laterally; c, style, posteroventrally, 5–6 teeth; d, style, posteroventrally, 1–2 teeth; e, aedeagus &amp; connective, laterally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290229" httpUri="https://zenodo.org/record/5290229/files/figure.png" pageId="1" pageNumber="36">Figs 4c&amp;d</figureCitation>
; 
<figureCitation id="1371848F1C4092435EBEFD60C1A256F6" box="[676,707,644,668]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="1" pageNumber="36">7d</figureCitation>
). Female valvulae exposed basally (
<figureCitation id="1371848F1C409243584EFD60C7CA56F6" box="[1108,1195,644,668]" captionStart="FIGURE 2" captionStartId="4.[151,249,1729,1752]" captionTargetBox="[218,1369,195,1698]" captionTargetId="figure-141@4.[218,1369,195,1705]" captionTargetPageId="4" captionText="FIGURE 2a–l. Tetramelasma litopyx sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c–e, sternite 7, ventrally; f, hind wing; g, valvula 1; h, valvula 2; i, valvula 1 detail; j, valvula 3 detail; k, valvula 2 detail; l, valvula 3 (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290223" httpUri="https://zenodo.org/record/5290223/files/figure.png" pageId="1" pageNumber="36">Figs 2b</figureCitation>
, 
<figureCitation id="1371848F1C40924358ACFD60C79656F6" box="[1206,1271,644,668]" captionStart="FIGURE 5" captionStartId="8.[151,249,1092,1115]" captionTargetBox="[160,1423,267,1066]" captionTargetId="figure-504@8.[160,1426,267,1069]" captionTargetPageId="8" captionText="FIGURE 5a–f. Tetramelasma nodosatha sp. n. female. a, Habitus (scale bar = 2 mm); b–c, sternite 7, ventrally, in situ; d–f, sternite 7, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290231" httpUri="https://zenodo.org/record/5290231/files/figure.png" pageId="1" pageNumber="36">5b&amp;c</figureCitation>
, 
<figureCitation id="1371848F1C4092435918FD60C67C56F6" box="[1282,1309,644,668]" captionStart="FIGURE 8" captionStartId="13.[151,250,1545,1568]" captionTargetBox="[212,1370,382,1518]" captionTargetId="figure-343@13.[212,1374,382,1521]" captionTargetPageId="13" captionText="FIGURE 8a–k. Tetramelasma scolosatha sp. n. female. a, Sternite 7 in situ, ventrally; b–d, sternite 7, ventrally; e, valvula 1; f, valvula 1 detail; g, valvula 2; h, valvula 2 detail; i, valvifer 2; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290241" httpUri="https://zenodo.org/record/5290241/files/figure.png" pageId="1" pageNumber="36">8a</figureCitation>
, 
<figureCitation id="1371848F1C4092435933FD60C63656F6" box="[1321,1367,644,668]" captionStart="FIGURE 10" captionStartId="15.[151,250,1445,1468]" captionTargetBox="[213,1369,183,1434]" captionTargetId="figure-294@15.[213,1373,183,1437]" captionTargetPageId="15" captionText="FIGURE 10a–j. Tetramelasma tanyphysis sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c-e, sternite 7, ventrally; f, valvula 1; g, valvula 2; h, valvula 1 detail; i, valvula 2 detail; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290245" httpUri="https://zenodo.org/record/5290245/files/figure.png" pageId="1" pageNumber="36">10b</figureCitation>
); sternite 7 variable, with wide notch of variable depth; variably produced laterally (
<figureCitation id="1371848F1C409243580FFD43C7E956D5" box="[1045,1160,679,703]" captionStart="FIGURE 2" captionStartId="4.[151,249,1729,1752]" captionTargetBox="[218,1369,195,1698]" captionTargetId="figure-141@4.[218,1369,195,1705]" captionTargetPageId="4" captionText="FIGURE 2a–l. Tetramelasma litopyx sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c–e, sternite 7, ventrally; f, hind wing; g, valvula 1; h, valvula 2; i, valvula 1 detail; j, valvula 3 detail; k, valvula 2 detail; l, valvula 3 (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290223" httpUri="https://zenodo.org/record/5290223/files/figure.png" pageId="1" pageNumber="36">Figs 2b–e</figureCitation>
; 
<figureCitation id="1371848F1C409243588FFD43C7AA56D5" box="[1173,1227,679,703]" captionStart="FIGURE 5" captionStartId="8.[151,249,1092,1115]" captionTargetBox="[160,1423,267,1066]" captionTargetId="figure-504@8.[160,1426,267,1069]" captionTargetPageId="8" captionText="FIGURE 5a–f. Tetramelasma nodosatha sp. n. female. a, Habitus (scale bar = 2 mm); b–c, sternite 7, ventrally, in situ; d–f, sternite 7, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290231" httpUri="https://zenodo.org/record/5290231/files/figure.png" pageId="1" pageNumber="36">5b–f</figureCitation>
; 
<figureCitation id="1371848F1C40924358C2FD43C67056D5" box="[1240,1297,679,703]" captionStart="FIGURE 6" captionStartId="10.[151,250,1635,1658]" captionTargetBox="[249,1334,184,1610]" captionTargetId="figure-186@10.[249,1339,184,1614]" captionTargetPageId="10" captionText="FIGURE 6a–l. Tetramelasma nodosatha sp. n. female. a, Habitus, submacropter; b, habitus, macropter (scale bar = 2 mm); c– g, sternite 7, ventrally; h, valvula 1; i, valvula 1 detail; j, valvula 2 detail; k, valvula 3; l, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290235" httpUri="https://zenodo.org/record/5290235/files/figure.png" pageId="1" pageNumber="36">6c–g</figureCitation>
; 
<figureCitation id="1371848F1C4092435904FD43C63656D5" box="[1310,1367,679,703]" captionStart="FIGURE 8" captionStartId="13.[151,250,1545,1568]" captionTargetBox="[212,1370,382,1518]" captionTargetId="figure-343@13.[212,1374,382,1521]" captionTargetPageId="13" captionText="FIGURE 8a–k. Tetramelasma scolosatha sp. n. female. a, Sternite 7 in situ, ventrally; b–d, sternite 7, ventrally; e, valvula 1; f, valvula 1 detail; g, valvula 2; h, valvula 2 detail; i, valvifer 2; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290241" httpUri="https://zenodo.org/record/5290241/files/figure.png" pageId="1" pageNumber="36">8a–d</figureCitation>
; 
<figureCitation id="1371848F1C409243597EFD43C3C7568E" captionStart="FIGURE 10" captionStartId="15.[151,250,1445,1468]" captionTargetBox="[213,1369,183,1434]" captionTargetId="figure-294@15.[213,1373,183,1437]" captionTargetPageId="15" captionText="FIGURE 10a–j. Tetramelasma tanyphysis sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c-e, sternite 7, ventrally; f, valvula 1; g, valvula 2; h, valvula 1 detail; i, valvula 2 detail; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290245" httpUri="https://zenodo.org/record/5290245/files/figure.png" pageId="1" pageNumber="36">10b– e</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4092435CDFFD0BC22B578A" blockId="1.[151,1437,356,1892]" pageId="1" pageNumber="36">
<emphasis id="B93E44181C4092435CDFFD0BC2825762" bold="true" box="[197,483,751,776]" pageId="1" pageNumber="36">
<emphasis id="B93E44181C4092435CDFFD0BC27C5762" box="[197,285,751,776]" inLineHeading="true" pageId="1" pageNumber="36" reason="1">Colour.</emphasis>
Male &amp; female.
</emphasis>
Base colour yellowish-brown. Vertex dorsoapically with two paired fuscous markings: apical pair usually triangular, close together; basal pair close to anterior margin of compound eyes, roughly circular; markings of variable size. Vertex, pronotum and rarely scutellum sometimes dorsally with faint to darkbrown longitudinal stripes. Clypeus yellowish overlain by pale to dark-brown horizontal arcs, sometimes extending dorsally into vertex. Tegmina translucent smoky-yellowish-brown or almost colourless, veins pale to transparent; costal cell pale yellow, other cells sometimes bordered with light-brown markings (especially in apical cells) (e.g. 
<figureCitation id="1371848F1C4092435C8DFC2CC38B578A" box="[151,234,968,992]" captionStart="FIGURE 3" captionStartId="6.[151,250,1647,1670]" captionTargetBox="[151,1425,770,1619]" captionTargetId="figure-251@6.[151,1436,770,1623]" captionTargetPageId="6" captionText="FIGURE 3 a–d. Tetramelasma nodosatha sp. n. male. a, Habitus, in situ (scale bar = 2 mm); b, pygofer, dorsally; c, genital capsule, laterally; d, pygofer, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290227" httpUri="https://zenodo.org/record/5290227/files/figure.png" pageId="1" pageNumber="36">Figs 3a</figureCitation>
, 
<figureCitation id="1371848F1C4092435CECFC2CC27257B5" box="[246,275,968,991]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="1" pageNumber="36">7a</figureCitation>
, 
<figureCitation id="1371848F1C4092435D05FC2CC25D578A" box="[287,316,968,992]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">9a</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4092435CDFFC0FC010501A" blockId="1.[151,1437,356,1892]" pageId="1" pageNumber="36">
<emphasis id="B93E44181C4092435CDFFC0FC1F4506E" bold="true" box="[197,661,1003,1028]" pageId="1" pageNumber="36">Tegmina &amp; hind wings. Male &amp; female</emphasis>
. Tegmina macropterous (
<figureCitation id="1371848F1C4092435FAEFC08C76B506E" box="[948,1034,1004,1028]" captionStart="FIGURE 3" captionStartId="6.[151,250,1647,1670]" captionTargetBox="[151,1425,770,1619]" captionTargetId="figure-251@6.[151,1436,770,1623]" captionTargetPageId="6" captionText="FIGURE 3 a–d. Tetramelasma nodosatha sp. n. male. a, Habitus, in situ (scale bar = 2 mm); b, pygofer, dorsally; c, genital capsule, laterally; d, pygofer, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290227" httpUri="https://zenodo.org/record/5290227/files/figure.png" pageId="1" pageNumber="36">Figs 3a</figureCitation>
, 
<figureCitation id="1371848F1C409243580DFC08C752506E" box="[1047,1075,1004,1028]" captionStart="FIGURE 5" captionStartId="8.[151,249,1092,1115]" captionTargetBox="[160,1423,267,1066]" captionTargetId="figure-504@8.[160,1426,267,1069]" captionTargetPageId="8" captionText="FIGURE 5a–f. Tetramelasma nodosatha sp. n. female. a, Habitus (scale bar = 2 mm); b–c, sternite 7, ventrally, in situ; d–f, sternite 7, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290231" httpUri="https://zenodo.org/record/5290231/files/figure.png" pageId="1" pageNumber="36">5a</figureCitation>
, 
<figureCitation id="1371848F1C409243585AFC08C73F506E" box="[1088,1118,1004,1028]" captionStart="FIGURE 6" captionStartId="10.[151,250,1635,1658]" captionTargetBox="[249,1334,184,1610]" captionTargetId="figure-186@10.[249,1339,184,1614]" captionTargetPageId="10" captionText="FIGURE 6a–l. Tetramelasma nodosatha sp. n. female. a, Habitus, submacropter; b, habitus, macropter (scale bar = 2 mm); c– g, sternite 7, ventrally; h, valvula 1; i, valvula 1 detail; j, valvula 2 detail; k, valvula 3; l, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290235" httpUri="https://zenodo.org/record/5290235/files/figure.png" pageId="1" pageNumber="36">6b</figureCitation>
, 
<figureCitation id="1371848F1C4092435870FC08C7E65069" box="[1130,1159,1004,1027]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="1" pageNumber="36">7a</figureCitation>
, 
<figureCitation id="1371848F1C4092435889FC08C7CF506E" box="[1171,1198,1004,1028]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">9a</figureCitation>
) or submacropterous (
<figureCitation id="1371848F1C4092435C85FBF4C3955042" box="[159,244,1040,1064]" captionStart="FIGURE 2" captionStartId="4.[151,249,1729,1752]" captionTargetBox="[218,1369,195,1698]" captionTargetId="figure-141@4.[218,1369,195,1705]" captionTargetPageId="4" captionText="FIGURE 2a–l. Tetramelasma litopyx sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c–e, sternite 7, ventrally; f, hind wing; g, valvula 1; h, valvula 2; i, valvula 1 detail; j, valvula 3 detail; k, valvula 2 detail; l, valvula 3 (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290223" httpUri="https://zenodo.org/record/5290223/files/figure.png" pageId="1" pageNumber="36">Figs 2a</figureCitation>
, 
<figureCitation id="1371848F1C4092435D1AFBF4C27C5042" box="[256,285,1040,1064]" captionStart="FIGURE 6" captionStartId="10.[151,250,1635,1658]" captionTargetBox="[249,1334,184,1610]" captionTargetId="figure-186@10.[249,1339,184,1614]" captionTargetPageId="10" captionText="FIGURE 6a–l. Tetramelasma nodosatha sp. n. female. a, Habitus, submacropter; b, habitus, macropter (scale bar = 2 mm); c– g, sternite 7, ventrally; h, valvula 1; i, valvula 1 detail; j, valvula 2 detail; k, valvula 3; l, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290235" httpUri="https://zenodo.org/record/5290235/files/figure.png" pageId="1" pageNumber="36">6a</figureCitation>
, 
<figureCitation id="1371848F1C4092435D32FBF4C2355042" box="[296,340,1040,1064]" captionStart="FIGURE 10" captionStartId="15.[151,250,1445,1468]" captionTargetBox="[213,1369,183,1434]" captionTargetId="figure-294@15.[213,1373,183,1437]" captionTargetPageId="15" captionText="FIGURE 10a–j. Tetramelasma tanyphysis sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c-e, sternite 7, ventrally; f, valvula 1; g, valvula 2; h, valvula 1 detail; i, valvula 2 detail; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290245" httpUri="https://zenodo.org/record/5290245/files/figure.png" pageId="1" pageNumber="36">10a</figureCitation>
). Hind wing usually reduced, ½ as wide and ¾ as long as tegmina, jugal lobe present, small. (
<figureCitation id="1371848F1C4092435970FBF4C3CC5026" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="1" pageNumber="36">Figs 1i</figureCitation>
, 
<figureCitation id="1371848F1C4092435CA0FBD0C3B55026" box="[186,212,1076,1100]" captionStart="FIGURE 2" captionStartId="4.[151,249,1729,1752]" captionTargetBox="[218,1369,195,1698]" captionTargetId="figure-141@4.[218,1369,195,1705]" captionTargetPageId="4" captionText="FIGURE 2a–l. Tetramelasma litopyx sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c–e, sternite 7, ventrally; f, hind wing; g, valvula 1; h, valvula 2; i, valvula 1 detail; j, valvula 3 detail; k, valvula 2 detail; l, valvula 3 (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290223" httpUri="https://zenodo.org/record/5290223/files/figure.png" pageId="1" pageNumber="36">2f</figureCitation>
, 
<figureCitation id="1371848F1C4092435CC5FBD0C27D5026" box="[223,284,1076,1100]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="1" pageNumber="36">7h&amp;i</figureCitation>
, 
<figureCitation id="1371848F1C4092435D3DFBD0C2245026" box="[295,325,1076,1100]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">
<quantity id="4CB235EF1C4092435D3DFBD0C2245026" box="[295,325,1076,1100]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="1" pageNumber="36" unit="g" value="9.0">9g</quantity>
</figureCitation>
), M1+2 sometimes fused to R4+5. Rarely hind wing fully developed (256 specimen’s wings examined, 
<specimenCount id="9D4C53831C4092435CCFFBBCC20D501A" box="[213,364,1112,1136]" pageId="1" pageNumber="36" type="generic">12 specimens</specimenCount>
with hind wing as long and wide as tegmina).
</paragraph>
<paragraph id="8BF5980A1C4092435CDFFB98C221522E" blockId="1.[151,1437,356,1892]" pageId="1" pageNumber="36">
<emphasis id="B93E44181C4092435CDFFB98C1EC50FE" bold="true" box="[197,653,1147,1172]" pageId="1" pageNumber="36">
<emphasis id="B93E44181C4092435CDFFB98C2AC50FE" box="[197,461,1148,1172]" inLineHeading="true" pageId="1" pageNumber="36" reason="1">External morphology.</emphasis>
Male &amp; female
</emphasis>
. Vertex 1.3–2.0 times longer medially than length of vertex next to eye, width of head 1.0–1.1 times wider than width across pronotum, diameter of ocellus 28 µm, ocellocular distance up to 48 µm. Vertex narrowly rounded to face, shagreened, disc of vertex smooth. Pronotum smooth. Clypellus rectangular, lateral convergent, subparallel margins. Gena extending slightly beyond posterior margin clypellus, slightly incised below eye. Spinulation of protibia 1+4. Profemur setae in row AV short, relatively thick, intercalary row setae about three times longer than those in AV, AV1 seta slightly longer and thicker than intercalary setae. Mesotibial setal formula 4+4. Metafemoral setal formula 2+2+1, apical paired setae slightly shorter than subapical seta, medial pair of different length and thickness. Metathoracic tibis with four rows of setae, as defined by 
<bibRefCitation id="EFDBE5FB1C4092435941FA9CC38451DE" author="Davis, R. B." pageId="1" pageNumber="36" refId="ref11211" refString="Davis, R. B. (1975) Classifiction of selected higher cateories of Auchenorrhynchous Homoptera (Cicadellidae and Aetalionidae). United States Department of Agriculture, Agricultural Research Services, Technical Bulleting No. 1494, 52 pp." type="book" year="1975">Davis (1975)</bibRefCitation>
. In all species of 
<emphasis id="B93E44181C4092435DA9FA78C13651DE" box="[435,599,1436,1460]" italics="true" pageId="1" pageNumber="36">
<taxonomicName id="4C4AE3891C4092435DA9FA78C13251DE" authorityName="Stiller" authorityYear="2011" box="[435,595,1436,1460]" class="Insecta" family="Cicadellidae" genus="Tetramelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus">Tetramelasma</taxonomicName>
,
</emphasis>
except in 
<taxonomicName id="4C4AE3891C4092435EC8FA79C05D51DE" authorityName="Stiller" authorityYear="2011" box="[722,828,1436,1460]" class="Insecta" family="Cicadellidae" genus="Tetramelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="species" species="litopyx">
<emphasis id="B93E44181C4092435EC8FA79C05D51DE" box="[722,828,1436,1460]" italics="true" pageId="1" pageNumber="36">T. litopyx</emphasis>
</taxonomicName>
row III with uniform row of setae, reaching apex of tibia. In 
<taxonomicName id="4C4AE3891C4092435CEDFA25C20151B2" authorityName="Stiller" authorityYear="2011" box="[247,352,1472,1496]" class="Insecta" family="Cicadellidae" genus="Tetramelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="species" species="litopyx">
<emphasis id="B93E44181C4092435CEDFA25C20151B2" box="[247,352,1472,1496]" italics="true" pageId="1" pageNumber="36">T. litopyx</emphasis>
</taxonomicName>
with row III without apical seta, but two subapical, heavily sclerotized setae, with tibia around base of these two setae sclerotized, both or one sometimes asymmetrical, about three times longer than basal setae. 
<emphasis id="B93E44181C4092435C8DF9ECC1CA524A" bold="true" box="[151,683,1543,1568]" pageId="1" pageNumber="36">
<emphasis id="B93E44181C4092435C8DF9ECC2FC524A" box="[151,413,1544,1568]" inLineHeading="true" pageId="1" pageNumber="36" reason="1">Internal morphology.</emphasis>
Abdominal apodeme
</emphasis>
. Anterior and posterior apodeme narrow, without significantly enlarged lobes.
</paragraph>
<paragraph id="8BF5980A1C4092435CDFF9B4C0E452DA" blockId="1.[151,1437,356,1892]" pageId="1" pageNumber="36">
<emphasis id="B93E44181C4092435CDFF9B4C2605202" bold="true" box="[197,257,1616,1640]" pageId="1" pageNumber="36">Male</emphasis>
. 
<emphasis id="B93E44181C4092435D0AF9ABC2C35202" bold="true" box="[272,418,1615,1640]" pageId="1" pageNumber="36">Dimensions.</emphasis>
Apex of vertex to tips of tegmina 
<quantity id="4CB235EF1C4092435F36F9B4C0D7520D" box="[812,950,1616,1640]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="4.5" metricValueMin="3.9" pageId="1" pageNumber="36" unit="mm" value="4.2" valueMax="4.5" valueMin="3.9">3.9–4.5 mm</quantity>
, vertex 1.1–1.5 times longer medially than length next to eye, head 1.1–1.3 times wider than pronotum, vertex 0.7–1.3 times longer than medial length of pronotum, vertex 0.3–0.5 times longer than width of head across eyes.
</paragraph>
<paragraph id="8BF5980A1C4092435CDFF95FC213530E" blockId="1.[151,1437,356,1892]" pageId="1" pageNumber="36">
<emphasis id="B93E44181C4092435CDFF95FC2E852BE" bold="true" box="[197,393,1723,1748]" inLineHeading="true" pageId="1" pageNumber="36" reason="1">Genital capsule.</emphasis>
Segment 10 weakly sclerotized, lateral margins concave (
<figureCitation id="1371848F1C4092435824F958C7F252BE" box="[1086,1171,1724,1748]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="1" pageNumber="36">Fig. 1b</figureCitation>
) or straight (
<figureCitation id="1371848F1C409243592CF958C6EC52BE" box="[1334,1421,1724,1748]" captionStart="FIGURE 3" captionStartId="6.[151,250,1647,1670]" captionTargetBox="[151,1425,770,1619]" captionTargetId="figure-251@6.[151,1436,770,1623]" captionTargetPageId="6" captionText="FIGURE 3 a–d. Tetramelasma nodosatha sp. n. male. a, Habitus, in situ (scale bar = 2 mm); b, pygofer, dorsally; c, genital capsule, laterally; d, pygofer, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290227" httpUri="https://zenodo.org/record/5290227/files/figure.png" pageId="1" pageNumber="36">Fig. 3b</figureCitation>
), incised ⅓ or ½ way into pygofer. Pygofer, viewed laterally, with posterior lobe bluntly triangular or rounded (
<figureCitation id="1371848F1C4092435970F93BC3D55376" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="1" pageNumber="36">Figs 1d</figureCitation>
, 
<figureCitation id="1371848F1C4092435CD8F8E0C3BF5376" box="[194,222,1796,1820]" captionStart="FIGURE 3" captionStartId="6.[151,250,1647,1670]" captionTargetBox="[151,1425,770,1619]" captionTargetId="figure-251@6.[151,1436,770,1623]" captionTargetPageId="6" captionText="FIGURE 3 a–d. Tetramelasma nodosatha sp. n. male. a, Habitus, in situ (scale bar = 2 mm); b, pygofer, dorsally; c, genital capsule, laterally; d, pygofer, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290227" httpUri="https://zenodo.org/record/5290227/files/figure.png" pageId="1" pageNumber="36">3c</figureCitation>
, 
<figureCitation id="1371848F1C4092435CF6F8E0C26B5376" box="[236,266,1796,1820]" captionStart="FIGURE 4" captionStartId="7.[151,250,855,878]" captionTargetBox="[154,1421,199,825]" captionTargetId="figure-406@7.[151,1436,195,831]" captionTargetPageId="7" captionText="FIGURE 4 a–e. Tetramelasma nodosatha sp. n. male. a, Genital capsule, ventrally; b, genital capsule, laterally; c, style, posteroventrally, 5–6 teeth; d, style, posteroventrally, 1–2 teeth; e, aedeagus &amp; connective, laterally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290229" httpUri="https://zenodo.org/record/5290229/files/figure.png" pageId="1" pageNumber="36">4b</figureCitation>
, 
<figureCitation id="1371848F1C4092435D0DF8E0C2545376" box="[279,309,1796,1820]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="1" pageNumber="36">7b</figureCitation>
, 
<figureCitation id="1371848F1C4092435D58F8E0C23E5376" box="[322,351,1796,1820]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">9c</figureCitation>
); ventral margin usually bulbous, of variable size, sometimes larger than pygofer posterior lobe (
<figureCitation id="1371848F1C4092435C85F8C3C3905355" box="[159,241,1831,1855]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="1" pageNumber="36">Fig. 9c</figureCitation>
). Medial surface of ventral pygofer lobe with spine-like process of variable length; apex of spine diagnostic in some species:
</paragraph>
<paragraph id="8BF5980A1C4092435C8DF87EC15E53DB" blockId="1.[151,649,1945,2005]" box="[151,575,1945,1969]" pageId="1" pageNumber="36">
a. Narrowly acuminate (
<figureCitation id="1371848F1C4092435DA1F87DC15353DB" box="[443,562,1945,1969]" captionStart="FIGURE 4" captionStartId="7.[151,250,855,878]" captionTargetBox="[154,1421,199,825]" captionTargetId="figure-406@7.[151,1436,195,831]" captionTargetPageId="7" captionText="FIGURE 4 a–e. Tetramelasma nodosatha sp. n. male. a, Genital capsule, ventrally; b, genital capsule, laterally; c, style, posteroventrally, 5–6 teeth; d, style, posteroventrally, 1–2 teeth; e, aedeagus &amp; connective, laterally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290229" httpUri="https://zenodo.org/record/5290229/files/figure.png" pageId="1" pageNumber="36">Fig. 4a&amp;b</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4092435C8DF859C1E853BF" blockId="1.[151,649,1945,2005]" box="[151,649,1981,2005]" pageId="1" pageNumber="36">
b. Wide with subapical tooth (
<figureCitation id="1371848F1C4092435DE1F859C13153BF" box="[507,592,1981,2005]" captionStart="FIGURE 3" captionStartId="6.[151,250,1647,1670]" captionTargetBox="[151,1425,770,1619]" captionTargetId="figure-251@6.[151,1436,770,1623]" captionTargetPageId="6" captionText="FIGURE 3 a–d. Tetramelasma nodosatha sp. n. male. a, Habitus, in situ (scale bar = 2 mm); b, pygofer, dorsally; c, genital capsule, laterally; d, pygofer, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290227" httpUri="https://zenodo.org/record/5290227/files/figure.png" pageId="1" pageNumber="36">Figs 3c</figureCitation>
, 
<figureCitation id="1371848F1C4092435E46F859C11A53BF" box="[604,635,1981,2005]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="1" pageNumber="36">7b</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4392405C8DFF7CC2C354C5" blockId="2.[151,418,151,212]" box="[151,418,151,175]" pageId="2" pageNumber="37">
c. Bipinnate (
<figureCitation id="1371848F1C4392405D59FF73C2F454C5" box="[323,405,151,175]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="2" pageNumber="37">Fig. 9c</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4392405C8DFF58C2F954BE" blockId="2.[151,418,151,212]" box="[151,408,188,212]" pageId="2" pageNumber="37">
d. Reduced (
<figureCitation id="1371848F1C4392405D22FF58C2EB54BE" box="[312,394,188,212]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="2" pageNumber="37">Fig. 1d</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4392405CDDFEE0C77D55A5" blockId="2.[151,1436,260,464]" pageId="2" pageNumber="37">
Ventral pygofer lobe sometimes reduced (
<figureCitation id="1371848F1C4392405EBBFEE0C1955576" box="[673,756,260,284]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="2" pageNumber="37">Fig. 1d</figureCitation>
). Plate variable: commonly short, about one fifth the length of pygofer, posterior margin truncate, lateral and medial margins rounded or straight (
<figureCitation id="1371848F1C4392405866FEC3C7995555" box="[1148,1272,295,319]" captionStart="FIGURE 3" captionStartId="6.[151,250,1647,1670]" captionTargetBox="[151,1425,770,1619]" captionTargetId="figure-251@6.[151,1436,770,1623]" captionTargetPageId="6" captionText="FIGURE 3 a–d. Tetramelasma nodosatha sp. n. male. a, Habitus, in situ (scale bar = 2 mm); b, pygofer, dorsally; c, genital capsule, laterally; d, pygofer, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290227" httpUri="https://zenodo.org/record/5290227/files/figure.png" pageId="2" pageNumber="37">Figs 3c&amp;d</figureCitation>
, 
<figureCitation id="1371848F1C439240591DFEC3C6295555" box="[1287,1352,295,319]" captionStart="FIGURE 4" captionStartId="7.[151,250,855,878]" captionTargetBox="[154,1421,199,825]" captionTargetId="figure-406@7.[151,1436,195,831]" captionTargetPageId="7" captionText="FIGURE 4 a–e. Tetramelasma nodosatha sp. n. male. a, Genital capsule, ventrally; b, genital capsule, laterally; c, style, posteroventrally, 5–6 teeth; d, style, posteroventrally, 1–2 teeth; e, aedeagus &amp; connective, laterally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290229" httpUri="https://zenodo.org/record/5290229/files/figure.png" pageId="2" pageNumber="37">4a&amp;b</figureCitation>
, 
<figureCitation id="1371848F1C439240594DFEC3C6F95555" box="[1367,1432,295,319]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="2" pageNumber="37">7b&amp;c</figureCitation>
, 
<figureCitation id="1371848F1C4392405C8DFEA8C3B8550E" box="[151,217,332,356]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="2" pageNumber="37">9b&amp;c</figureCitation>
); rarely about as long as length of pygofer (
<figureCitation id="1371848F1C4392405ED1FEA8C07C550E" box="[715,797,332,356]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="2" pageNumber="37">Fig. 1d</figureCitation>
); posterior, medial and lateral margins rounded, sinuous subapically (
<figureCitation id="1371848F1C4392405D32FE8BC21B55ED" box="[296,378,367,391]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="2" pageNumber="37">Fig. 1c</figureCitation>
); dorsolateral margin with sclerotized process apically, process digitate or rounded, variably extended beyond posterior margin of plate. Macrosetae grouped lateromedially, 
<quantity id="4CB235EF1C439240582BFE70C71B55C6" box="[1073,1146,404,428]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="2.032" metricValueMin="1.016" pageId="2" pageNumber="37" unit="in" value="6.0" valueMax="8.0" valueMin="4.0">4–8 in</quantity>
number. Style apophysis generally short (about 1–2 times as long as greatest basal width), shape variable:
</paragraph>
<paragraph id="8BF5980A1C4392405C8DFDE4C2155635" blockId="2.[151,1436,511,644]" pageId="2" pageNumber="37">
a. Expanded, barrel-shaped, posteriorly excavated, with mediolateral margin variably toothed, either with series of 5–6 distinct, uniseriate teeth (
<figureCitation id="1371848F1C4392405E2AFDC0C1E45656" box="[560,645,548,572]" captionStart="FIGURE 4" captionStartId="7.[151,250,855,878]" captionTargetBox="[154,1421,199,825]" captionTargetId="figure-406@7.[151,1436,195,831]" captionTargetPageId="7" captionText="FIGURE 4 a–e. Tetramelasma nodosatha sp. n. male. a, Genital capsule, ventrally; b, genital capsule, laterally; c, style, posteroventrally, 5–6 teeth; d, style, posteroventrally, 1–2 teeth; e, aedeagus &amp; connective, laterally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290229" httpUri="https://zenodo.org/record/5290229/files/figure.png" pageId="2" pageNumber="37">Figs 4c</figureCitation>
, 
<figureCitation id="1371848F1C4392405E8BFDC0C1CC5656" box="[657,685,548,572]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="2" pageNumber="37">7d</figureCitation>
) or serration reduced to rounded protrusion and cluster of 2–4 denticles (
<figureCitation id="1371848F1C4392405D09FDA3C2065635" box="[275,359,583,607]" captionStart="FIGURE 4" captionStartId="7.[151,250,855,878]" captionTargetBox="[154,1421,199,825]" captionTargetId="figure-406@7.[151,1436,195,831]" captionTargetPageId="7" captionText="FIGURE 4 a–e. Tetramelasma nodosatha sp. n. male. a, Genital capsule, ventrally; b, genital capsule, laterally; c, style, posteroventrally, 5–6 teeth; d, style, posteroventrally, 1–2 teeth; e, aedeagus &amp; connective, laterally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290229" httpUri="https://zenodo.org/record/5290229/files/figure.png" pageId="2" pageNumber="37">Fig. 4d</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4392405C8DFD88C08356EE" blockId="2.[151,1436,511,644]" box="[151,994,620,644]" pageId="2" pageNumber="37">
b. Narrowly produced with medioventral or lateral serration (
<figureCitation id="1371848F1C4392405F42FD88C0C856EE" box="[856,937,620,644]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="2" pageNumber="37">Figs 1f</figureCitation>
, 
<figureCitation id="1371848F1C4392405FACFD88C0B456EE" box="[950,981,620,644]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="2" pageNumber="37">9d</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4392405CDDFD48C2AD5745" blockId="2.[151,1436,684,960]" pageId="2" pageNumber="37">
Style basally with lateral lobe short, posteriad; medial lobe narrow, elongate, perpendicular to base. Aedeagus with shaft prominent, tubular, straight in dorsal view, curved or undulating in lateral view, gonopore subapical, dorsal (
<figureCitation id="1371848F1C4392405CDEFD10C2215766" box="[196,320,756,780]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="2" pageNumber="37">
Figs 
<quantity id="4CB235EF1C4392405CE7FD10C2775766" box="[253,278,756,780]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="2" pageNumber="37" unit="g" value="1.0">1g</quantity>
&amp;h
</figureCitation>
; 
<figureCitation id="1371848F1C4392405D51FD10C2085766" box="[331,361,756,780]" captionStart="FIGURE 4" captionStartId="7.[151,250,855,878]" captionTargetBox="[154,1421,199,825]" captionTargetId="figure-406@7.[151,1436,195,831]" captionTargetPageId="7" captionText="FIGURE 4 a–e. Tetramelasma nodosatha sp. n. male. a, Genital capsule, ventrally; b, genital capsule, laterally; c, style, posteroventrally, 5–6 teeth; d, style, posteroventrally, 1–2 teeth; e, aedeagus &amp; connective, laterally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290229" httpUri="https://zenodo.org/record/5290229/files/figure.png" pageId="2" pageNumber="37">4e</figureCitation>
; 
<figureCitation id="1371848F1C4392405D6EFD10C2D05766" box="[372,433,756,780]" captionStart="FIGURE 7" captionStartId="12.[151,250,1466,1489]" captionTargetBox="[155,1419,195,1439]" captionTargetId="figure-277@12.[155,1431,195,1442]" captionTargetPageId="12" captionText="FIGURE 7a–i. Tetramelasma scolosatha sp. n. male. a, Habitus (scale bar = 2 mm); b, pygofer, laterally; c, pygofer, ventrally; d, style, posteroventrally; e, aedeagus, laterally; f, aedeagus, dorsally; g, tegmina; h–i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290237" httpUri="https://zenodo.org/record/5290237/files/figure.png" pageId="2" pageNumber="37">7e&amp;f</figureCitation>
; 
<figureCitation id="1371848F1C4392405DA6FD10C2B85766" box="[444,473,756,780]" captionStart="FIGURE 9" captionStartId="14.[151,250,1979,2002]" captionTargetBox="[262,1324,1050,1959]" captionTargetId="figure-359@14.[262,1328,1050,1961]" captionTargetPageId="14" captionText="FIGURE 9a–g. Tetramelasma tanyphysis sp. n. male. a, Habitus, male (scale bar = 2 mm); b, genital capsule, ventrally; c, pygofer, laterally; d, style; e, aedeagus &amp; connective, laterally; f, tegmina; g, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290243" httpUri="https://zenodo.org/record/5290243/files/figure.png" pageId="2" pageNumber="37">9e</figureCitation>
); dorsal apodeme reduced. Connective linear, of uniform width, arms fused anteriorly, fused to aedeagus (
<figureCitation id="1371848F1C4392405D77FCF3C2DC5745" box="[365,445,791,815]" captionStart="FIGURE 1" captionStartId="3.[151,249,1946,1969]" captionTargetBox="[273,1327,1026,1933]" captionTargetId="figure-697@3.[273,1328,1026,1933]" captionTargetPageId="3" captionText="FIGURE 1a–i. Tetramelasma litopyx sp. n. male. a, genital capsule, ventrally, in situ; b, pygofer, dorsally; c, plate, ventrally; d, genital capsule, laterally; e, connective; f, style; g, aedeagus,dorsally; h, aedeagus, laterally; i, hind wing (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290221" httpUri="https://zenodo.org/record/5290221/files/figure.png" pageId="2" pageNumber="37">Fig. 1e</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4392405CDFFCD8C0E457F6" blockId="2.[151,1436,684,960]" pageId="2" pageNumber="37">
<emphasis id="B93E44181C4392405CDFFCD8C2DA573E" bold="true" box="[197,443,827,852]" pageId="2" pageNumber="37">
<emphasis id="B93E44181C4392405CDFFCD8C242573E" box="[197,291,828,852]" inLineHeading="true" pageId="2" pageNumber="37" reason="1">Female.</emphasis>
Dimensions.
</emphasis>
4.2–5.0 mm from apex of vertex to tips of tegmina, vertex 1.3–1.5 times longer medially than length next to eye, head 1.1 times wider than pronotum, vertex 1.1–1.5 times longer than medial length of pronotum, vertex 0.4–0.6 times longer than width of head across eyes.
</paragraph>
<paragraph id="8BF5980A1C4392405CDDFC43C0FC57D5" blockId="2.[151,1436,684,960]" box="[199,925,935,960]" pageId="2" pageNumber="37">
<emphasis id="B93E44181C4392405CDDFC43C25D57AA" bold="true" box="[199,316,935,960]" inLineHeading="true" pageId="2" pageNumber="37" reason="1">Genitalia.</emphasis>
Abdominal sternite 7 narrow, with three basic shapes:
</paragraph>
<paragraph id="8BF5980A1C4392405C8DFC0FC0DF5068" blockId="2.[151,1435,1002,1134]" box="[151,958,1002,1026]" pageId="2" pageNumber="37">
a. Falcate, with posterolateral margin produced uniformly (
<figureCitation id="1371848F1C4392405F58FC0EC0D05068" box="[834,945,1002,1026]" captionStart="FIGURE 6" captionStartId="10.[151,250,1635,1658]" captionTargetBox="[249,1334,184,1610]" captionTargetId="figure-186@10.[249,1339,184,1614]" captionTargetPageId="10" captionText="FIGURE 6a–l. Tetramelasma nodosatha sp. n. female. a, Habitus, submacropter; b, habitus, macropter (scale bar = 2 mm); c– g, sternite 7, ventrally; h, valvula 1; i, valvula 1 detail; j, valvula 2 detail; k, valvula 3; l, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290235" httpUri="https://zenodo.org/record/5290235/files/figure.png" pageId="2" pageNumber="37">Fig. 6c–g</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4392405C8DFBEAC700504C" blockId="2.[151,1435,1002,1134]" box="[151,1121,1038,1062]" pageId="2" pageNumber="37">
b. Falcate, with posterolateral margin folded under lateral margins (
<figureCitation id="1371848F1C4392405F84FBEAC76F504C" box="[926,1038,1038,1062]" captionStart="FIGURE 5" captionStartId="8.[151,249,1092,1115]" captionTargetBox="[160,1423,267,1066]" captionTargetId="figure-504@8.[160,1426,267,1069]" captionTargetPageId="8" captionText="FIGURE 5a–f. Tetramelasma nodosatha sp. n. female. a, Habitus (scale bar = 2 mm); b–c, sternite 7, ventrally, in situ; d–f, sternite 7, ventrally (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290231" httpUri="https://zenodo.org/record/5290231/files/figure.png" pageId="2" pageNumber="37">Figs 5b–f</figureCitation>
; 
<figureCitation id="1371848F1C4392405803FBEAC735504C" box="[1049,1108,1038,1062]" captionStart="FIGURE 8" captionStartId="13.[151,250,1545,1568]" captionTargetBox="[212,1370,382,1518]" captionTargetId="figure-343@13.[212,1374,382,1521]" captionTargetPageId="13" captionText="FIGURE 8a–k. Tetramelasma scolosatha sp. n. female. a, Sternite 7 in situ, ventrally; b–d, sternite 7, ventrally; e, valvula 1; f, valvula 1 detail; g, valvula 2; h, valvula 2 detail; i, valvifer 2; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290241" httpUri="https://zenodo.org/record/5290241/files/figure.png" pageId="2" pageNumber="37">8a–d</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4392405C8DFBD7C7505004" blockId="2.[151,1435,1002,1134]" pageId="2" pageNumber="37">
c. Trapezoid, posterior margin with medial, narrow V-shaped notch, lateral margins straight, angled slightly posteriad, medial disc membranous, laterally more or less sclerotized (
<figureCitation id="1371848F1C4392405FACFBB2C7445004" box="[950,1061,1110,1134]" captionStart="FIGURE 2" captionStartId="4.[151,249,1729,1752]" captionTargetBox="[218,1369,195,1698]" captionTargetId="figure-141@4.[218,1369,195,1705]" captionTargetPageId="4" captionText="FIGURE 2a–l. Tetramelasma litopyx sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c–e, sternite 7, ventrally; f, hind wing; g, valvula 1; h, valvula 2; i, valvula 1 detail; j, valvula 3 detail; k, valvula 2 detail; l, valvula 3 (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290223" httpUri="https://zenodo.org/record/5290223/files/figure.png" pageId="2" pageNumber="37">Fig. 2b–e</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4392405CDFFB7AC65250B0" blockId="2.[151,1436,1182,1747]" pageId="2" pageNumber="37">
Valvula 3 without major setae, only fine setae (
<figureCitation id="1371848F1C4392405EC5FB7AC03150DC" box="[735,848,1182,1206]" captionStart="FIGURE 2" captionStartId="4.[151,249,1729,1752]" captionTargetBox="[218,1369,195,1698]" captionTargetId="figure-141@4.[218,1369,195,1705]" captionTargetPageId="4" captionText="FIGURE 2a–l. Tetramelasma litopyx sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c–e, sternite 7, ventrally; f, hind wing; g, valvula 1; h, valvula 2; i, valvula 1 detail; j, valvula 3 detail; k, valvula 2 detail; l, valvula 3 (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290223" httpUri="https://zenodo.org/record/5290223/files/figure.png" pageId="2" pageNumber="37">Figs 2j&amp;l</figureCitation>
; 
<figureCitation id="1371848F1C4392405F47FB7AC0FB50DC" box="[861,922,1182,1206]" captionStart="FIGURE 6" captionStartId="10.[151,250,1635,1658]" captionTargetBox="[249,1334,184,1610]" captionTargetId="figure-186@10.[249,1339,184,1614]" captionTargetPageId="10" captionText="FIGURE 6a–l. Tetramelasma nodosatha sp. n. female. a, Habitus, submacropter; b, habitus, macropter (scale bar = 2 mm); c– g, sternite 7, ventrally; h, valvula 1; i, valvula 1 detail; j, valvula 2 detail; k, valvula 3; l, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290235" httpUri="https://zenodo.org/record/5290235/files/figure.png" pageId="2" pageNumber="37">6k&amp;l</figureCitation>
; 
<figureCitation id="1371848F1C4392405FBDFB7AC08250DC" box="[935,995,1182,1206]" captionStart="FIGURE 8" captionStartId="13.[151,250,1545,1568]" captionTargetBox="[212,1370,382,1518]" captionTargetId="figure-343@13.[212,1374,382,1521]" captionTargetPageId="13" captionText="FIGURE 8a–k. Tetramelasma scolosatha sp. n. female. a, Sternite 7 in situ, ventrally; b–d, sternite 7, ventrally; e, valvula 1; f, valvula 1 detail; g, valvula 2; h, valvula 2 detail; i, valvifer 2; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290241" httpUri="https://zenodo.org/record/5290241/files/figure.png" pageId="2" pageNumber="37">8j&amp;k</figureCitation>
; 
<figureCitation id="1371848F1C4392405FEBFB7AC75D50DC" box="[1009,1084,1182,1206]" captionStart="FIGURE 10" captionStartId="15.[151,250,1445,1468]" captionTargetBox="[213,1369,183,1434]" captionTargetId="figure-294@15.[213,1373,183,1437]" captionTargetPageId="15" captionText="FIGURE 10a–j. Tetramelasma tanyphysis sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c-e, sternite 7, ventrally; f, valvula 1; g, valvula 2; h, valvula 1 detail; i, valvula 2 detail; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290245" httpUri="https://zenodo.org/record/5290245/files/figure.png" pageId="2" pageNumber="37">10j&amp;k</figureCitation>
). Valvula 2 finely serrate (
<figureCitation id="1371848F1C4392405970FB7AC3BB50B0" captionStart="FIGURE 2" captionStartId="4.[151,249,1729,1752]" captionTargetBox="[218,1369,195,1698]" captionTargetId="figure-141@4.[218,1369,195,1705]" captionTargetPageId="4" captionText="FIGURE 2a–l. Tetramelasma litopyx sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c–e, sternite 7, ventrally; f, hind wing; g, valvula 1; h, valvula 2; i, valvula 1 detail; j, valvula 3 detail; k, valvula 2 detail; l, valvula 3 (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290223" httpUri="https://zenodo.org/record/5290223/files/figure.png" pageId="2" pageNumber="37">Figs 2h&amp;k</figureCitation>
; 
<figureCitation id="1371848F1C4392405CFFFB26C39F50B0" box="[229,254,1218,1242]" captionStart="FIGURE 6" captionStartId="10.[151,250,1635,1658]" captionTargetBox="[249,1334,184,1610]" captionTargetId="figure-186@10.[249,1339,184,1614]" captionTargetPageId="10" captionText="FIGURE 6a–l. Tetramelasma nodosatha sp. n. female. a, Habitus, submacropter; b, habitus, macropter (scale bar = 2 mm); c– g, sternite 7, ventrally; h, valvula 1; i, valvula 1 detail; j, valvula 2 detail; k, valvula 3; l, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290235" httpUri="https://zenodo.org/record/5290235/files/figure.png" pageId="2" pageNumber="37">6j</figureCitation>
; 
<figureCitation id="1371848F1C4392405D10FB26C22C50B0" box="[266,333,1218,1242]" captionStart="FIGURE 8" captionStartId="13.[151,250,1545,1568]" captionTargetBox="[212,1370,382,1518]" captionTargetId="figure-343@13.[212,1374,382,1521]" captionTargetPageId="13" captionText="FIGURE 8a–k. Tetramelasma scolosatha sp. n. female. a, Sternite 7 in situ, ventrally; b–d, sternite 7, ventrally; e, valvula 1; f, valvula 1 detail; g, valvula 2; h, valvula 2 detail; i, valvifer 2; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290241" httpUri="https://zenodo.org/record/5290241/files/figure.png" pageId="2" pageNumber="37">
<quantity id="4CB235EF1C4392405D10FB26C24750B0" box="[266,294,1218,1242]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="2" pageNumber="37" unit="g" value="8.0">8g</quantity>
&amp;h
</figureCitation>
; 
<figureCitation id="1371848F1C4392405D42FB26C2C550B0" box="[344,420,1218,1242]" captionStart="FIGURE 10" captionStartId="15.[151,250,1445,1468]" captionTargetBox="[213,1369,183,1434]" captionTargetId="figure-294@15.[213,1373,183,1437]" captionTargetPageId="15" captionText="FIGURE 10a–j. Tetramelasma tanyphysis sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c-e, sternite 7, ventrally; f, valvula 1; g, valvula 2; h, valvula 1 detail; i, valvula 2 detail; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290245" httpUri="https://zenodo.org/record/5290245/files/figure.png" pageId="2" pageNumber="37">
<quantity id="4CB235EF1C4392405D42FB26C2E550B0" box="[344,388,1218,1242]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="2" pageNumber="37" unit="g" value="10.0">10g</quantity>
&amp;i
</figureCitation>
). Valvula 1 lanceolate, with imbricate sculpture (
<figureCitation id="1371848F1C4392405FDDFB26C75C50B0" box="[967,1085,1218,1242]" captionStart="FIGURE 2" captionStartId="4.[151,249,1729,1752]" captionTargetBox="[218,1369,195,1698]" captionTargetId="figure-141@4.[218,1369,195,1705]" captionTargetPageId="4" captionText="FIGURE 2a–l. Tetramelasma litopyx sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c–e, sternite 7, ventrally; f, hind wing; g, valvula 1; h, valvula 2; i, valvula 1 detail; j, valvula 3 detail; k, valvula 2 detail; l, valvula 3 (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290223" httpUri="https://zenodo.org/record/5290223/files/figure.png" pageId="2" pageNumber="37">
Figs 
<quantity id="4CB235EF1C439240581AFB26C77F50B0" box="[1024,1054,1218,1242]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="2" pageNumber="37" unit="g" value="2.0">2g</quantity>
&amp;i
</figureCitation>
; 
<figureCitation id="1371848F1C4392405852FB26C7E450B0" box="[1096,1157,1218,1242]" captionStart="FIGURE 6" captionStartId="10.[151,250,1635,1658]" captionTargetBox="[249,1334,184,1610]" captionTargetId="figure-186@10.[249,1339,184,1614]" captionTargetPageId="10" captionText="FIGURE 6a–l. Tetramelasma nodosatha sp. n. female. a, Habitus, submacropter; b, habitus, macropter (scale bar = 2 mm); c– g, sternite 7, ventrally; h, valvula 1; i, valvula 1 detail; j, valvula 2 detail; k, valvula 3; l, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290235" httpUri="https://zenodo.org/record/5290235/files/figure.png" pageId="2" pageNumber="37">6h&amp;i</figureCitation>
; 
<figureCitation id="1371848F1C439240588BFB26C7AC50B0" box="[1169,1229,1218,1242]" captionStart="FIGURE 8" captionStartId="13.[151,250,1545,1568]" captionTargetBox="[212,1370,382,1518]" captionTargetId="figure-343@13.[212,1374,382,1521]" captionTargetPageId="13" captionText="FIGURE 8a–k. Tetramelasma scolosatha sp. n. female. a, Sternite 7 in situ, ventrally; b–d, sternite 7, ventrally; e, valvula 1; f, valvula 1 detail; g, valvula 2; h, valvula 2 detail; i, valvifer 2; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290241" httpUri="https://zenodo.org/record/5290241/files/figure.png" pageId="2" pageNumber="37">8e&amp;f</figureCitation>
; 
<figureCitation id="1371848F1C43924058C3FB26C64750B0" box="[1241,1318,1218,1242]" captionStart="FIGURE 10" captionStartId="15.[151,250,1445,1468]" captionTargetBox="[213,1369,183,1434]" captionTargetId="figure-294@15.[213,1373,183,1437]" captionTargetPageId="15" captionText="FIGURE 10a–j. Tetramelasma tanyphysis sp. n. female. a, Habitus (scale bar = 2 mm); b, sternite 7, ventrally, in situ; c-e, sternite 7, ventrally; f, valvula 1; g, valvula 2; h, valvula 1 detail; i, valvula 2 detail; j, valvula 3; k, valvula 3 detail (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5290245" httpUri="https://zenodo.org/record/5290245/files/figure.png" pageId="2" pageNumber="37">10f&amp;h</figureCitation>
).
</paragraph>
<paragraph id="8BF5980A1C4392405CDFFB02C388512C" blockId="2.[151,1436,1182,1747]" pageId="2" pageNumber="37">
<emphasis id="B93E44181C4392405CDFFB02C20F5095" bold="true" box="[197,366,1254,1279]" inLineHeading="true" pageId="2" pageNumber="37" reason="1">Relationships.</emphasis>
<taxonomicName id="4C4AE3891C4392405D6CFB02C1755094" authorityName="Stiller" authorityYear="2011" box="[374,532,1254,1278]" class="Insecta" family="Cicadellidae" genus="Tetramelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B93E44181C4392405D6CFB02C1755094" box="[374,532,1254,1278]" italics="true" pageId="2" pageNumber="37">Tetramelasma</emphasis>
</taxonomicName>
is placed in the tribe 
<taxonomicName id="4C4AE3891C4392405F14FB02C0D25094" authorityName="Dallas" authorityYear="1870" box="[782,947,1254,1278]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="tribe" tribe="Deltocephalini">Deltocephalini</taxonomicName>
based on the linear shaped connective and fusion between the connective and aedeagus, as defined by 
<bibRefCitation id="EFDBE5FB1C4392405F26FAEEC71E5148" author="Webb, M. D. &amp; Viraktamath, C. A." box="[828,1151,1290,1314]" pageId="2" pageNumber="37" pagination="1 - 64" refId="ref11436" refString="Webb, M. D. &amp; Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera, Auchenorrhyncha, Cicadellidae) Zootaxa, 2163, 1 - 64." type="journal article" year="2009">Webb &amp; Viraktamath (2009)</bibRefCitation>
and 
<bibRefCitation id="EFDBE5FB1C43924058A2FAEEC384512C" author="Zahniser, J. N. &amp; Dietrich, C. H." pageId="2" pageNumber="37" pagination="489 - 511" refId="ref11539" refString="Zahniser, J. N. &amp; Dietrich, C. H. (2010) Phylogeny of the leafhopper subfamily Deltocephalinae (Hemiptera: Cicadellidae) based on molecular and morphological data with a revised family-group classification. Systematic Entomology, 35, 489 - 511." type="journal article" year="2010">Zahniser &amp; Dietrich (2010)</bibRefCitation>
.
</paragraph>
<paragraph id="8BF5980A1C4392405CDFFAB6C0BE5274" blockId="2.[151,1436,1182,1747]" pageId="2" pageNumber="37">
Other 
<taxonomicName id="4C4AE3891C4392405D11FAB6C2D15100" authorityName="Dallas" authorityYear="1870" box="[267,432,1362,1386]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="tribe" tribe="Deltocephalini">Deltocephalini</taxonomicName>
in 
<collectingCountry id="F35DD89A1C4392405DC9FAB6C1035100" box="[467,610,1362,1386]" name="South Africa" pageId="2" pageNumber="37">South Africa</collectingCountry>
include 
<taxonomicName id="4C4AE3891C4392405ED8FAB7C0DC5100" authority="Stiller, 2009" authorityName="Stiller" authorityYear="2009" box="[706,957,1362,1387]" class="Insecta" family="Cicadellidae" genus="Theronus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B93E44181C4392405ED8FAB7C04A5101" box="[706,811,1363,1387]" italics="true" pageId="2" pageNumber="37">Theronus</emphasis>
Stiller, 2009
</taxonomicName>
, 
<taxonomicName id="4C4AE3891C4392405FD2FAB7C7AB5100" authority="Theron, 1988" authorityName="Theron" authorityYear="1988" box="[968,1226,1362,1387]" class="Insecta" family="Cicadellidae" genus="Heidinus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B93E44181C4392405FD2FAB7C74C5101" box="[968,1069,1363,1387]" italics="true" pageId="2" pageNumber="37">Heidinus</emphasis>
Theron, 1988
</taxonomicName>
and 
<taxonomicName id="4C4AE3891C439240591AFAB7C26D51E4" authority="Distant. None" authorityName="Distant. None" authorityYear="1917" class="Insecta" family="Cicadellidae" genus="Maiestas" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B93E44181C439240591AFAB7C6045100" box="[1280,1381,1363,1386]" italics="true" pageId="2" pageNumber="37">Maiestas</emphasis>
Distant. None
</taxonomicName>
of these genera are significantly similar to 
<emphasis id="B93E44181C4392405EEEFA92C0F951E4" box="[756,920,1398,1422]" italics="true" pageId="2" pageNumber="37">
<taxonomicName id="4C4AE3891C4392405EEEFA92C0F451E4" authorityName="Stiller" authorityYear="2011" box="[756,917,1398,1422]" class="Insecta" family="Cicadellidae" genus="Tetramelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">Tetramelasma</taxonomicName>
,
</emphasis>
except possibly the simple, tubular aedeagus. The fusion between the aedeagus and connective is distinct in these four genera, with the arms of the connective contiguous. The fusion of the arms of the connective is a distinguishing feature of the 
<taxonomicName id="4C4AE3891C4392405841FA5AC78151BC" authorityName="Distant" authorityYear="1908" box="[1115,1248,1470,1494]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="tribe" tribe="Paralimnini">Paralimnini</taxonomicName>
, but members of this tribe have the connective articulated with the aedeagus. In some instances this distinction between fusion and articulation becomes vague as pointed out by 
<bibRefCitation id="EFDBE5FB1C4392405E8DF9E2C0BB5274" author="Webb, M. D. &amp; Viraktamath, C. A." box="[663,986,1542,1566]" pageId="2" pageNumber="37" pagination="1 - 64" refId="ref11436" refString="Webb, M. D. &amp; Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera, Auchenorrhyncha, Cicadellidae) Zootaxa, 2163, 1 - 64." type="journal article" year="2009">Webb &amp; Viraktamath (2009)</bibRefCitation>
.
</paragraph>
<paragraph id="8BF5980A1C4392405CDFF9CEC0F852B8" blockId="2.[151,1436,1182,1747]" pageId="2" pageNumber="37">
When compared to Palaearctic genera, 
<taxonomicName id="4C4AE3891C4392405E93F9CFC0475229" authorityName="Stiller" authorityYear="2011" box="[649,806,1579,1603]" class="Insecta" family="Cicadellidae" genus="Tetramelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B93E44181C4392405E93F9CFC0475229" box="[649,806,1579,1603]" italics="true" pageId="2" pageNumber="37">Tetramelasma</emphasis>
</taxonomicName>
has similar colour and markings to that of 
<taxonomicName id="4C4AE3891C4392405905F9CFC23B520C" authority="Osborne and Ball" authorityName="Osborne and Ball" class="Insecta" family="Cicadellidae" genus="Conosanus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B93E44181C4392405905F9CFC6FA5229" box="[1311,1435,1579,1603]" italics="true" pageId="2" pageNumber="37">Conosanus</emphasis>
Osborne and Ball
</taxonomicName>
(
<taxonomicName id="4C4AE3891C4392405D70F9AAC145520C" authorityName="Dallas" authorityYear="1870" box="[362,548,1614,1638]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="subFamily" subFamily="Deltocephalinae">Deltocephalinae</taxonomicName>
, 
<taxonomicName id="4C4AE3891C4392405E35F9AAC1CD520C" authorityName="Van Duzee" authorityYear="1892" box="[559,684,1614,1638]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="tribe" tribe="Athysanini">Athysanini</taxonomicName>
) and 
<taxonomicName id="4C4AE3891C4392405EF0F9ABC0FD520C" authority="Ribaut" authorityName="Ribaut" box="[746,924,1614,1638]" class="Insecta" family="Cicadellidae" genus="Turrutus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B93E44181C4392405EF0F9ABC02B520C" box="[746,842,1615,1638]" italics="true" pageId="2" pageNumber="37">Turrutus</emphasis>
Ribaut
</taxonomicName>
(
<taxonomicName id="4C4AE3891C4392405FB1F9AAC704520C" authorityName="Dallas" authorityYear="1870" box="[939,1125,1614,1638]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="subFamily" subFamily="Deltocephalinae">Deltocephalinae</taxonomicName>
, 
<taxonomicName id="4C4AE3891C439240586AF9AAC67A520C" authorityName="Dallas" authorityYear="1870" box="[1136,1307,1614,1638]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="tribe" tribe="Deltocephalini">Deltocephalini</taxonomicName>
). The excavated apophysis of the style of 
<taxonomicName id="4C4AE3891C4392405DE0F997C1D152E0" authority="Ribaut" authorityName="Ribaut" box="[506,688,1650,1675]" class="Insecta" family="Cicadellidae" genus="Ebarrius" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B93E44181C4392405DE0F997C13C52E1" box="[506,605,1651,1675]" italics="true" pageId="2" pageNumber="37">Ebarrius</emphasis>
Ribaut
</taxonomicName>
(
<taxonomicName id="4C4AE3891C4392405EA5F996C01852E0" authorityName="Dallas" authorityYear="1870" box="[703,889,1650,1674]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="subFamily" subFamily="Deltocephalinae">Deltocephalinae</taxonomicName>
, 
<taxonomicName id="4C4AE3891C4392405F9CF996C76B52E0" authorityName="Distant" authorityYear="1908" box="[902,1034,1650,1674]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="tribe" tribe="Paralimnini">Paralimnini</taxonomicName>
) bears some resemblance to that of 
<taxonomicName id="4C4AE3891C4392405C8DF972C25452C4" authorityName="Stiller" authorityYear="2011" box="[151,309,1686,1710]" class="Insecta" family="Cicadellidae" genus="Tetramelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B93E44181C4392405C8DF972C25452C4" box="[151,309,1686,1710]" italics="true" pageId="2" pageNumber="37">Tetramelasma</emphasis>
</taxonomicName>
. In the Neotropical Region the shape of the plate of 
<taxonomicName id="4C4AE3891C4392405F9EF972C74352C4" authorityName="Stiller" authorityYear="2011" box="[900,1058,1686,1710]" class="Insecta" family="Cicadellidae" genus="Tetramelasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B93E44181C4392405F9EF972C74352C4" box="[900,1058,1686,1710]" italics="true" pageId="2" pageNumber="37">Tetramelasma</emphasis>
</taxonomicName>
somewhat resembles that of 
<taxonomicName id="4C4AE3891C4392405973F972C16E52B8" authority="Linnavuori" authorityName="Linnavuori" class="Insecta" family="Cicadellidae" genus="Haldorus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="subGenus" subGenus="Parahaldorus">
<emphasis id="B93E44181C4392405973F972C3B652B9" italics="true" pageId="2" pageNumber="37">Haldorus</emphasis>
(
<emphasis id="B93E44181C4392405CFDF95FC2E352B9" box="[231,386,1723,1747]" italics="true" pageId="2" pageNumber="37">Parahaldorus</emphasis>
) Linnavuori
</taxonomicName>
(
<taxonomicName id="4C4AE3891C4392405E07F95EC1B952B8" authorityName="Dallas" authorityYear="1870" box="[541,728,1722,1746]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="subFamily" subFamily="Deltocephalinae">Deltocephalinae</taxonomicName>
, 
<taxonomicName id="4C4AE3891C4392405EF9F95EC0EC52B8" authorityName="Dallas" authorityYear="1870" box="[739,909,1722,1746]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="tribe" tribe="Deltocephalini">Deltocephalini</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>