treatments-xml/data/03/E3/13/03E3133CFFB15B4FD55F52B987469614.xml
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<document id="C0F71F8739A288C207EB83677145ADB8" ID-DOI="10.11646/zootaxa.1094.1.3" ID-ISSN="1175­5334" ID-Zenodo-Dep="5051350" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625109582879" checkinUser="felipe" docAuthor="Barraclough, David A." docDate="2005" docId="03E3133CFFB15B4FD55F52B987469614" docLanguage="en" docName="zt01094p051.pdf" docOrigin="Zootaxa 1094 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1094.1.3" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Prosoeca difficile Barraclough 2005, comb. n." docType="treatment" docVersion="4" lastPageNumber="49" masterDocId="FFDA6B44FFB95B47D457512F85259177" masterDocTitle="A review of the type material of the Southern African genus Stenobasipteron Lichtwardt, 1910 (Diptera: Nemestrinidae), with transfer of two species to Prosoeca Schiner, 1867" masterLastPageNumber="51" masterPageNumber="41" pageNumber="49" updateTime="1698981470411" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8BD33976F07443C205A94DE76B3A4CA0">A review of the type material of the Southern African genus Stenobasipteron Lichtwardt, 1910 (Diptera: Nemestrinidae), with transfer of two species to Prosoeca Schiner, 1867</mods:title>
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<mods:namePart id="13FBAF5EB265AE3B634555919C96B69C">Barraclough, David A.</mods:namePart>
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<mods:date id="A79B70DF6662FEF9E985D2D0F5FE6983">2005</mods:date>
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<treatment id="03E3133CFFB15B4FD55F52B987469614" ID-DOI="http://doi.org/10.5281/zenodo.5054227" ID-Zenodo-Dep="5054227" LSID="urn:lsid:plazi:treatment:03E3133CFFB15B4FD55F52B987469614" httpUri="http://treatment.plazi.org/id/03E3133CFFB15B4FD55F52B987469614" lastPageNumber="49" pageId="8" pageNumber="49">
<subSubSection id="C350F1A1FFB15B4FD55F52B987C692C7" box="[264,739,918,944]" pageId="8" pageNumber="49" type="nomenclature">
<paragraph id="8BF5A22AFFB15B4FD55F52B987C692C7" blockId="8.[264,739,918,944]" box="[264,739,918,944]" pageId="8" pageNumber="49">
<heading id="D0BD1546FFB15B4FD55F52B987C692C7" bold="true" box="[264,739,918,944]" fontSize="11" level="1" pageId="8" pageNumber="49" reason="1">
<emphasis id="B93E7E38FFB15B4FD55F52B987C692C7" bold="true" box="[264,739,918,944]" pageId="8" pageNumber="49">
<taxonomicName id="4C4AD9A9FFB15B4FD55F52B9874F92C7" ID-CoL="4MVX4" authority="(Bequaert)" authorityName="Barraclough" authorityYear="2005" baseAuthorityName="Bequaert" baseAuthorityYear="1925" box="[264,618,918,944]" class="Insecta" family="Nemestrinidae" genus="Prosoeca" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="difficile" status="comb. nov.">
<emphasis id="B93E7E38FFB15B4FD55F52B984FF92C7" bold="true" box="[264,474,918,944]" italics="true" pageId="8" pageNumber="49">Prosoeca difficile</emphasis>
(Bequaert)
</taxonomicName>
,
<taxonomicNameLabel id="A20DC343FFB15B4FD62F52B987C692C7" box="[632,739,918,944]" pageId="8" pageNumber="49" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<paragraph id="8BF5A22AFFB15B4FD55F52CD87D4928F" blockId="8.[264,753,993,1017]" box="[264,753,993,1017]" pageId="8" pageNumber="49">
<taxonomicName id="4C4AD9A9FFB15B4FD55F52CD87C8928F" authority="Bequaert, 1925: 11" authorityName="Bequaert" authorityPageNumber="11" authorityYear="1925" box="[264,749,993,1017]" class="Insecta" family="Nemestrinidae" genus="Stenobasiopteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="difficile">
<emphasis id="B93E7E38FFB15B4FD55F52CD873D928E" box="[264,536,994,1017]" italics="true" pageId="8" pageNumber="49">Stenobasiopteron difficile</emphasis>
Bequaert, 1925: 11
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C350F1A1FFB15B4FD55F550687469614" pageId="8" pageNumber="49" type="materials_examined">
<paragraph id="8BF5A22AFFB15B4FD55F550687D395CC" blockId="8.[264,1323,1065,1891]" pageId="8" pageNumber="49">
<materialsCitation id="3B22A877FFB15B4FD55F550687D395CC" collectingDate="1929-04-06" collectingDateMax="1929-04-17" collectingDateMin="1929-04-06" collectionCode="TYPE, NMBZ" collectorName="Museum &amp; J. Beq. &amp; The &amp; There" country="Zimbabwe" determinerName="J. Bequaert" location="Cloudlands" municipality="Vumba Mts" pageId="8" pageNumber="49" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
<typeStatus id="54F11C88FFB15B4FD55F550684509534" box="[264,373,1065,1091]" pageId="8" pageNumber="49" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4C69A3FFB15B4FD529550684B7954A" box="[382,402,1065,1085]" pageId="8" pageNumber="49" type="female"></specimenCount>
:
<collectingCountry id="F35DE2BAFFB15B4FD5F4550687009534" box="[419,549,1065,1091]" name="Zimbabwe" pageId="8" pageNumber="49">Zimbabwe</collectingCountry>
:
<location id="8E95F4F1FFB15B4FD6655506879E9534" LSID="urn:lsid:plazi:treatment:03E3133CFFB15B4FD55F52B987469614:8E95F4F1FFB15B4FD6655506879E9534" box="[562,699,1065,1091]" country="Zimbabwe" municipality="Vumba Mts" name="Cloudlands" pageId="8" pageNumber="49">Cloudlands</location>
, 6000 [high altitude grassland] /
<collectingMunicipality id="6B913850FFB15B4FD006550681F99534" box="[1105,1244,1065,1091]" pageId="8" pageNumber="49">Vumba Mts</collectingMunicipality>
/
<specimenCount id="9D4C69A3FFB15B4FD0AF55068029954A" box="[1272,1292,1065,1085]" pageId="8" pageNumber="49" type="female"></specimenCount>
S. Rhodesia /
<date id="FFF484EAFFB15B4FD5C6557E877D951C" box="[401,600,1105,1131]" pageId="8" pageNumber="49" value="1929-04-06" valueMax="1929-04-17" valueMin="1929-04-06">
<collectingDate id="EFB07D02FFB15B4FD5C6557E877D951C" box="[401,600,1105,1131]" pageId="8" pageNumber="49" value="1929-04-06" valueMax="1929-04-17" valueMin="1929-04-06">617 April 1929</collectingDate>
</date>
/
<collectingCountry id="F35DE2BAFFB15B4FD622557E87C7951C" box="[629,738,1105,1131]" name="Zimbabwe" pageId="8" pageNumber="49">Rhodesia</collectingCountry>
<collectorName id="26BFC7FCFFB15B4FD6BB557E8673951C" box="[748,854,1105,1131]" pageId="8" pageNumber="49">Museum</collectorName>
;
<taxonomicName id="4C4AD9A9FFB15B4FD732557E8104951D" authorityName="Lichtwardt" authorityYear="1910" box="[869,1057,1105,1130]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B93E7E38FFB15B4FD732557E8104951D" box="[869,1057,1105,1130]" italics="true" pageId="8" pageNumber="49">Stenobasipteron</emphasis>
</taxonomicName>
/
<taxonomicName id="4C4AD9A9FFB15B4FD069557E81B2951D" authorityName="Bequaert" authorityYear="1925" box="[1086,1175,1105,1130]" class="Insecta" family="Nemestrinidae" genus="Stenobasiopteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="difficile">
<emphasis id="B93E7E38FFB15B4FD069557E81B2951D" box="[1086,1175,1105,1130]" italics="true" pageId="8" pageNumber="49">difficile</emphasis>
</taxonomicName>
/ type
<specimenCount id="9D4C69A3FFB15B4FD0A7557E80219512" box="[1264,1284,1105,1125]" pageId="8" pageNumber="49" type="female"></specimenCount>
<collectorName id="26BFC7FCFFB15B4FD14F557E841A95E4" pageId="8" pageNumber="49">J. Beq.</collectorName>
/
<determinerName id="7E294EEEFFB15B4FD500555684FE95E4" box="[343,475,1145,1171]" pageId="8" pageNumber="49">J. Bequaert</determinerName>
det.;
<collectionCode id="ED5B3AEFFFB15B4FD6775556874C95E4" box="[544,617,1145,1171]" pageId="8" pageNumber="49">TYPE</collectionCode>
[rectangular red card].
<collectorName id="26BFC7FCFFB15B4FD7D65556868B95E4" box="[897,942,1145,1171]" pageId="8" pageNumber="49">The</collectorName>
<typeStatus id="54F11C88FFB15B4FD7E15556813995E4" box="[950,1052,1145,1171]" pageId="8" pageNumber="49" type="holotype">holotype</typeStatus>
(in
<collectionCode id="ED5B3AEFFFB15B4FD0195556818095E4" box="[1102,1189,1145,1171]" country="Zimbabwe" httpUri="http://grbio.org/cool/yne7-xrqc" name="Natural History Museum of Zimbabwe" pageId="8" pageNumber="49">NMBZ</collectionCode>
) is in good condition.
<collectorName id="26BFC7FCFFB15B4FD5D2558E84EC95CC" box="[389,457,1185,1211]" pageId="8" pageNumber="49">There</collectorName>
is no other type material.
</materialsCitation>
</paragraph>
<paragraph id="8BF5A22AFFB15B4FD56F55E681D494F4" blockId="8.[264,1323,1065,1891]" pageId="8" pageNumber="49">
Discussion: In his original description, Bequaert remarked on the very close similarity between
<taxonomicName id="4C4AD9A9FFB15B4FD52D55DE8784947D" authorityName="Bequaert" authorityYear="1925" box="[378,673,1265,1290]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="difficile">
<emphasis id="B93E7E38FFB15B4FD52D55DE8784947D" box="[378,673,1265,1290]" italics="true" pageId="8" pageNumber="49">Stenobasipteron difficile</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4AD9A9FFB15B4FD6BB55DE8643947D" authorityName="Lichtwardt" authorityYear="1910" box="[748,870,1265,1290]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B93E7E38FFB15B4FD6BB55DE8643947D" box="[748,870,1265,1290]" italics="true" pageId="8" pageNumber="49">S. gracile</emphasis>
</taxonomicName>
, and listed seven characters which supposedly separated the species. This comparison is however incorrect, as Bequaert appears not to have examined the
<typeStatus id="54F11C88FFB15B4FD6C0546E87EC942C" box="[663,713,1345,1371]" pageId="8" pageNumber="49">type</typeStatus>
material of
<taxonomicName id="4C4AD9A9FFB15B4FD700546E86ED942D" authorityName="Lichtwardt" authorityYear="1910" box="[855,968,1345,1370]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B93E7E38FFB15B4FD700546E86ED942D" box="[855,968,1345,1370]" italics="true" pageId="8" pageNumber="49">S. gracile</emphasis>
</taxonomicName>
, and the material of
<taxonomicName id="4C4AD9A9FFB15B4FD0ED546E800F942D" authorityName="Lichtwardt" authorityYear="1910" box="[1210,1322,1345,1370]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B93E7E38FFB15B4FD0ED546E800F942D" box="[1210,1322,1345,1370]" italics="true" pageId="8" pageNumber="49">S. gracile</emphasis>
</taxonomicName>
used for the comparison was misidentified as
<taxonomicName id="4C4AD9A9FFB15B4FD777544686B294F5" authorityName="Bequaert" authorityYear="1925" box="[800,919,1385,1410]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="difficile">
<emphasis id="B93E7E38FFB15B4FD777544686B294F5" box="[800,919,1385,1410]" italics="true" pageId="8" pageNumber="49">S. difficile</emphasis>
</taxonomicName>
(see above under
<taxonomicName id="4C4AD9A9FFB15B4FD03B544681C594F5" authorityName="Bequaert" authorityYear="1925" box="[1132,1248,1385,1410]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="arnoldi">
<emphasis id="B93E7E38FFB15B4FD03B544681C594F5" box="[1132,1248,1385,1410]" italics="true" pageId="8" pageNumber="49">S. arnoldi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BF5A22AFFB15B4FD56F54BE87469614" blockId="8.[264,1323,1065,1891]" pageId="8" pageNumber="49">
I have examined the male
<typeStatus id="54F11C88FFB15B4FD62B54BE87F294DC" box="[636,727,1425,1451]" pageId="8" pageNumber="49" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C4AD9A9FFB15B4FD75354BE865D94DD" authorityName="Lichtwardt" authorityYear="1910" box="[772,888,1425,1450]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B93E7E38FFB15B4FD75354BE865D94DD" box="[772,888,1425,1450]" italics="true" pageId="8" pageNumber="49">S. gracile</emphasis>
</taxonomicName>
(a female
<typeStatus id="54F11C88FFB15B4FD7A954BE817C94DC" box="[1022,1113,1425,1451]" pageId="8" pageNumber="49" type="syntype">syntype</typeStatus>
is the only other specimen in the type series), and there are several major characters separating it from
<taxonomicName id="4C4AD9A9FFB15B4FD14354968444948D" authorityName="Bequaert" authorityYear="1925" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="difficile">
<emphasis id="B93E7E38FFB15B4FD14354968444948D" italics="true" pageId="8" pageNumber="49">S. difficile</emphasis>
</taxonomicName>
. The most important character is the development of vestiture on abdominal terga 3 and 4.
<taxonomicName id="4C4AD9A9FFB15B4FD527572684CD9755" authorityName="Bequaert" authorityYear="1925" box="[368,488,1545,1570]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="difficile">
<emphasis id="B93E7E38FFB15B4FD527572684CD9755" box="[368,488,1545,1570]" italics="true" pageId="8" pageNumber="49">S. difficile</emphasis>
</taxonomicName>
has obvious erect vestiture on these terga, unlike
<taxonomicName id="4C4AD9A9FFB15B4FD06D5726818E9755" authorityName="Lichtwardt" authorityYear="1910" box="[1082,1195,1545,1570]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B93E7E38FFB15B4FD06D5726818E9755" box="[1082,1195,1545,1570]" italics="true" pageId="8" pageNumber="49">S. gracile</emphasis>
</taxonomicName>
which has inconspicuous, recumbent vestiture, characteristic of other
<taxonomicName id="4C4AD9A9FFB15B4FD7EF571E81FD973C" box="[952,1240,1585,1611]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93E7E38FFB15B4FD7EF571E8151973D" box="[952,1140,1585,1610]" italics="true" pageId="8" pageNumber="49">Stenobasipteron</emphasis>
species.
</taxonomicName>
As my diagnosis of
<taxonomicName id="4C4AD9A9FFB15B4FD5F3577687459705" authorityName="Lichtwardt" authorityYear="1910" box="[420,608,1625,1650]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B93E7E38FFB15B4FD5F3577687459705" box="[420,608,1625,1650]" italics="true" pageId="8" pageNumber="49">Stenobasipteron</emphasis>
</taxonomicName>
includes only species with recumbent vestiture, I herewith transfer
<taxonomicName id="4C4AD9A9FFB15B4FD53E57AE84FA97ED" authorityName="Bequaert" authorityYear="1925" box="[361,479,1665,1690]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="difficile">
<emphasis id="B93E7E38FFB15B4FD53E57AE84FA97ED" box="[361,479,1665,1690]" italics="true" pageId="8" pageNumber="49">S. difficile</emphasis>
</taxonomicName>
to
<taxonomicName id="4C4AD9A9FFB15B4FD65257AE875797ED" authorityName="Schiner" authorityYear="1867" box="[517,626,1665,1690]" class="Insecta" family="Nemestrinidae" genus="Prosoeca" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B93E7E38FFB15B4FD65257AE875797ED" box="[517,626,1665,1690]" italics="true" pageId="8" pageNumber="49">Prosoeca</emphasis>
</taxonomicName>
. It should be noted though that
<taxonomicName id="4C4AD9A9FFB15B4FD7BF57AE819097ED" authorityName="Barraclough" authorityYear="2005" baseAuthorityName="Bequaert" baseAuthorityYear="1925" box="[1000,1205,1665,1690]" class="Insecta" family="Nemestrinidae" genus="Prosoeca" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="difficile" status="comb. nov.">
<emphasis id="B93E7E38FFB15B4FD7BF57AE819097ED" box="[1000,1205,1665,1690]" italics="true" pageId="8" pageNumber="49">Prosoeca difficile</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20DC343FFB15B4FD09257AE800F97EC" box="[1221,1322,1665,1691]" pageId="8" pageNumber="49" rank="species">comb. n.</taxonomicNameLabel>
does have a reduced alula, which was previously considered to characterise only
<taxonomicName id="4C4AD9A9FFB15B4FD55F57FE8716979C" box="[264,563,1745,1771]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93E7E38FFB15B4FD55F57FE84E1979D" box="[264,452,1745,1770]" italics="true" pageId="8" pageNumber="49">Stenobasipteron</emphasis>
species.
</taxonomicName>
The species are otherwise easily distinguished by colour characters.
<taxonomicName id="4C4AD9A9FFB15B4FD5C557D687229665" authorityName="Lichtwardt" authorityYear="1910" box="[402,519,1785,1810]" class="Insecta" family="Nemestrinidae" genus="Stenobasipteron" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B93E7E38FFB15B4FD5C557D687229665" box="[402,519,1785,1810]" italics="true" pageId="8" pageNumber="49">S. gracile</emphasis>
</taxonomicName>
is a pallid species with reddish­brown legs and very pale brown infuscated wings, whilst
<taxonomicName id="4C4AD9A9FFB15B4FD67D560E87D2964D" authorityName="Barraclough" authorityYear="2005" baseAuthorityName="Bequaert" baseAuthorityYear="1925" box="[554,759,1825,1850]" class="Insecta" family="Nemestrinidae" genus="Prosoeca" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="difficile">
<emphasis id="B93E7E38FFB15B4FD67D560E87D2964D" box="[554,759,1825,1850]" italics="true" pageId="8" pageNumber="49">Prosoeca difficile</emphasis>
</taxonomicName>
is a dark species with yellow legs and relatively dark brown infuscated wings.
</paragraph>
</subSubSection>
</treatment>
</document>