treatments-xml/data/03/D8/7B/03D87B73FFCAFFF940E1918FA640FB71.xml

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<mods:title id="587D4F550703D63D512334EF7DF3074C">A revision of the South African riffle beetle genus Leielmis Delève, 1964 (Coleoptera: Elmidae)</mods:title>
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<mods:namePart id="244C36D3D3869991D5FD56B7EDAA2201">Bilton, David T.</mods:namePart>
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<mods:date id="6D05D77B01D2BD08135D2F78B060D921">2017</mods:date>
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<treatment id="03D87B73FFCAFFF940E1918FA640FB71" ID-DOI="http://doi.org/10.5281/zenodo.5999281" ID-GBIF-Taxon="128527413" ID-Zenodo-Dep="5999281" LSID="urn:lsid:plazi:treatment:03D87B73FFCAFFF940E1918FA640FB71" httpUri="http://treatment.plazi.org/id/03D87B73FFCAFFF940E1918FA640FB71" lastPageId="9" lastPageNumber="264" pageId="2" pageNumber="257">
<subSubSection id="C36B99EEFFCAFFF240E1918FA40CFC26" pageId="2" pageNumber="257" type="nomenclature">
<paragraph id="8BCECA65FFCAFFF240E1918FA77CFC07" blockId="2.[151,635,900,959]" box="[151,635,900,926]" pageId="2" pageNumber="257">
<heading id="D0867D09FFCAFFF240E1918FA77CFC07" bold="true" box="[151,635,900,926]" fontSize="11" level="1" pageId="2" pageNumber="257" reason="1">
<taxonomicName id="4C71B1E6FFCAFFF240E1918FA77CFC07" ID-CoL="6P7RQ" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" box="[151,635,900,926]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF240E1918FA77CFC07" bold="true" box="[151,635,900,926]" pageId="2" pageNumber="257">
<emphasis id="B9051677FFCAFFF240E1918FA49BFC07" bold="true" box="[151,412,900,926]" italics="true" pageId="2" pageNumber="257">Leielmis georyssoides</emphasis>
(
<bibRefCitation id="EFE0B794FFCAFFF241DD918FA774FC07" author="Grouvelle" box="[427,627,900,926]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCECA65FFCAFFF240E191ADA40CFC26" blockId="2.[151,635,900,959]" box="[151,267,934,959]" pageId="2" pageNumber="257">
(
<figureCitation id="134AD6E0FFCAFFF240E991ADA405FC26" box="[159,258,934,959]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,250,1980,2002]" captionStartId-1="4.[151,250,1664,1686]" captionStartId-2="5.[151,250,1150,1172]" captionStartId-3="6.[151,250,1964,1986]" captionStartId-4="7.[151,250,1953,1975]" captionStartId-5="8.[151,250,1953,1975]" captionStartId-6="9.[151,250,1952,1974]" captionTargetBox-0="[166,1414,644,1940]" captionTargetBox-1="[155,1431,186,1636]" captionTargetBox-2="[164,1408,193,1126]" captionTargetBox-3="[446,1135,210,1910]" captionTargetBox-4="[298,1285,208,1917]" captionTargetBox-5="[259,1317,215,1910]" captionTargetBox-6="[151,1436,1303,1930]" captionTargetId-0="figure@3.[151,1436,625,1958]" captionTargetId-1="figure@4.[151,1436,183,1659]" captionTargetId-2="figure@5.[164,1408,193,1126]" captionTargetId-3="figure@6.[438,1155,193,1932]" captionTargetId-4="figure@7.[283,1304,193,1932]" captionTargetId-5="figure@8.[244,1344,193,1932]" captionTargetId-6="figure@9.[151,1436,1303,1930]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="FIGURE 1. Leielmis species, male habitus. A) L. georyssoides; B) L. gibbosus sp. nov., holotype; C) L. hirsutus sp. nov., holotype. Scale bar = 1 mm." captionText-1="FIGURE 2. Leielmis species, pronotal sculpture. A) &amp; D) L. georyssiodes; B) &amp; E) L. gibbosus sp. nov.; C) &amp; F) L. hirsutus sp. nov. A) – C) scanning electron micrographs; D) – F) stacked macrosope images." captionText-2="FIGURE 3. Leielmis species, Elytral sculpture, scanning electron micrographs. A) L. georyssoides; B) L. gibbosus sp. nov.; C) L. hirsutus sp. nov." captionText-3="FIGURE 4. Leielmis species, male tibial modifications, dorsal view A) &amp; D) L. georyssoides; B) &amp; E) L. gibbosus sp. nov.; C) &amp; F) L. hirsutus sp. nov. A) – C) mesotibiae; D) – F) metatibiae. Scale bar = 0.25 mm." captionText-4="FIGURE 5. Leielmis species, aedeagi, ventral and lateral views. A) L. georyssoides; B) L. gibbosus sp. nov.; C) L. hirsutus sp. nov. Scale bar = 0.25 mm." captionText-5="FIGURE 6. Leielmis species, ovipositors, distal gonocoxites. A) L. georyssoides; B) L. gibbosus sp. nov.; C) L. hirsutus sp. nov. Scale bar = 0.25 mm." captionText-6="FIGURE 7. Leielmis species, habitats. A) Western Cape, Table Mountain, stream in Echo Valley, L. georyssoides; B) Western Cape, Groote Winterhoekberge, Stream below Sneeugatpiek, L. gibbosus sp. nov. &amp; L. hirsutus sp. nov.; C) Western Cape, stream at Mont Rochelle above Franschhoek, L. gibbosus sp. nov. Photos D. T. Bilton &amp; N. Helme." httpUri-0="https://zenodo.org/record/545902/files/figure.png" httpUri-1="https://zenodo.org/record/545903/files/figure.png" httpUri-2="https://zenodo.org/record/545905/files/figure.png" httpUri-3="https://zenodo.org/record/545906/files/figure.png" httpUri-4="https://zenodo.org/record/545907/files/figure.png" httpUri-5="https://zenodo.org/record/545908/files/figure.png" httpUri-6="https://zenodo.org/record/545909/files/figure.png" pageId="2" pageNumber="257">Figs 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36B99EEFFCAFFF340E191E2A67FFED9" lastPageId="3" lastPageNumber="258" pageId="2" pageNumber="257" type="materials_examined">
<paragraph id="8BCECA65FFCAFFF240E191E2A768FB9B" blockId="2.[151,1437,1001,2034]" box="[151,623,1001,1026]" pageId="2" pageNumber="257">
<emphasis id="B9051677FFCAFFF240E191E2A42AFB9B" bold="true" box="[151,301,1001,1026]" pageId="2" pageNumber="257">Type locality</emphasis>
. “cap de Bonne Espérance”.
</paragraph>
<paragraph id="8BCECA65FFCAFFF240B19606A1C8FABB" blockId="2.[151,1437,1001,2034]" pageId="2" pageNumber="257">
<materialsCitation id="3B19C038FFCAFFF240B19606A1CCFABB" ID-GBIF-Occurrence="1455889605" collectionCode="SANC, PART, OF, SERIES, TYPS, DMHF" collectorName="R. Stals &amp; D T Bilton &amp; Genitalia" country="South Africa" location="Recognised" municipality="Pretoria" pageId="2" pageNumber="257" specimenCode="TYPS00594" specimenCount="7" specimenCount-female="4" specimenCount-male="3" typeStatus="lectotype">
<emphasis id="B9051677FFCAFFF240B19606A469FBBF" bold="true" box="[199,366,1037,1062]" pageId="2" pageNumber="257">
<typeStatus id="54CA74C7FFCAFFF240B19606A405FBBF" box="[199,258,1037,1062]" pageId="2" pageNumber="257">Type</typeStatus>
material
</emphasis>
. The 
<typeStatus id="54CA74C7FFCAFFF241D89605A4D9FBBF" box="[430,478,1038,1062]" pageId="2" pageNumber="257">type</typeStatus>
series (
<collectionCode id="ED6052A0FFCAFFF242429605A785FBBF" box="[564,642,1038,1062]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="257">SANC</collectionCode>
) consists of ten specimens, originally mounted on two cards. One card supported 
<specimenCount id="9D7701ECFFCAFFF2417A9639A431FBD0" box="[268,310,1074,1098]" pageId="2" pageNumber="257" type="male">3 ♂</specimenCount>
and 
<specimenCount id="9D7701ECFFCAFFF2411B9639A490FBD0" box="[365,407,1074,1098]" pageId="2" pageNumber="257" type="female">1 ♀</specimenCount>
, the other 
<specimenCount id="9D7701ECFFCAFFF242799639A73EFBD0" box="[527,569,1074,1098]" pageId="2" pageNumber="257" type="female">3 ♀</specimenCount>
, 3 individuals whose sex could not be determined and one gap, from which a specimen has presumably been dislodged historically. I have removed the third male from the left from the first of these cards, remounted it separately, following genital dissection, and here designate it as the 
<typeStatus id="54CA74C7FFCAFFF244B39672A03CFB0B" box="[1221,1339,1145,1170]" pageId="2" pageNumber="257" type="lectotype">
<emphasis id="B9051677FFCAFFF244B39672A03CFB0B" bold="true" box="[1221,1339,1145,1170]" pageId="2" pageNumber="257">Lectotype</emphasis>
</typeStatus>
: “ 
<collectionCode id="ED6052A0FFCAFFF245209671A5BCFB2F" pageId="2" pageNumber="257">PART OF</collectionCode>
<typeStatus id="54CA74C7FFCAFFF240A69695A449FB2F" box="[208,334,1182,1206]" pageId="2" pageNumber="257" type="syntype">SYNTYPE</typeStatus>
<collectionCode id="ED6052A0FFCAFFF241129695A4C7FB2F" box="[356,448,1182,1206]" pageId="2" pageNumber="257">SERIES</collectionCode>
// 
<taxonomicName id="4C71B1E6FFCAFFF241A79696A698FB2F" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" box="[465,927,1181,1206]" class="Insecta" family="Elmidae" genus="Helmis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF241A79696A7C0FB2C" box="[465,711,1181,1205]" italics="true" pageId="2" pageNumber="257">Helmis georyssoides</emphasis>
<bibRefCitation id="EFE0B794FFCAFFF242AA9696A698FB2F" author="Grouvelle" box="[732,927,1181,1206]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
</taxonomicName>
// 
<location id="8EAE9CBEFFCAFFF243C69696A134FB2F" LSID="urn:lsid:plazi:treatment:03D87B73FFCAFFF940E1918FA640FB71:8EAE9CBEFFCAFFF243C69696A134FB2F" box="[944,1075,1181,1206]" country="South Africa" municipality="Pretoria" name="Recognised" pageId="2" pageNumber="257">Recognised</location>
as such: 
<collectorName id="2684AFB3FFCAFFF244B19695A028FB2F" box="[1223,1327,1181,1206]" pageId="2" pageNumber="257">R. Stals</collectorName>
, 2015.”, “NATIONAL COLLECTION 
<collectionCode id="ED6052A0FFCAFFF2418A96C9A727FB43" box="[508,544,1218,1242]" pageId="2" pageNumber="257">OF</collectionCode>
INSECTS (
<collectionCode id="ED6052A0FFCAFFF242C296C9A607FB43" box="[692,768,1218,1242]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="257">SANC</collectionCode>
)// 
<collectingMunicipality id="6BAA501FFFCAFFF2435096CAA686FB43" box="[806,897,1217,1242]" pageId="2" pageNumber="257">Pretoria</collectingMunicipality>
, 
<collectingCountry id="F3668AF5FFCAFFF243E296CAA12DFB43" box="[916,1066,1217,1242]" name="South Africa" pageId="2" pageNumber="257">South Africa</collectingCountry>
”, “ 
<collectionCode id="ED6052A0FFCAFFF2442396C9A1A5FB43" box="[1109,1186,1218,1242]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="257">SANC</collectionCode>
<specimenCode id="DBD7621EFFCAFFF244C696CAA03AFB43" box="[1200,1341,1217,1242]" collectionCode="TYPS" pageId="2" pageNumber="257">
<collectionCode id="ED6052A0FFCAFFF244C696CAA1F2FB43" box="[1200,1269,1217,1242]" pageId="2" pageNumber="257">TYPS</collectionCode>
00594
</specimenCode>
” [green label], “ 
<taxonomicName id="4C71B1E6FFCAFFF2408496EEA79EFB67" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" box="[242,665,1253,1278]" class="Insecta" family="Elmidae" genus="Helmis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF2408496EEA4DCFB64" box="[242,475,1253,1277]" italics="true" pageId="2" pageNumber="257">Helmis georyssoides</emphasis>
<bibRefCitation id="EFE0B794FFCAFFF2419596EEA79EFB67" author="Grouvelle" box="[483,665,1253,1278]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
</taxonomicName>
// 
<typeStatus id="54CA74C7FFCAFFF242C596EDA625FB67" box="[691,802,1254,1278]" pageId="2" pageNumber="257" type="lectotype">Lectotype</typeStatus>
// 
<collectorName id="2684AFB3FFCAFFF2434D96EDA6B0FB67" box="[827,951,1253,1278]" pageId="2" pageNumber="257">D T Bilton</collectorName>
des. 2017” [red label]. 
<collectorName id="2684AFB3FFCAFFF244B096EEA02AFB67" box="[1222,1325,1253,1278]" pageId="2" pageNumber="257">Genitalia</collectorName>
dissected and mounted in 
<collectionCode id="ED6052A0FFCAFFF2413A9701A49AFAB8" box="[332,413,1290,1313]" pageId="2" pageNumber="257">DMHF</collectionCode>
on same card; left elytron mounted separately beside specimen (
<collectionCode id="ED6052A0FFCAFFF244049701A1C4FABB" box="[1138,1219,1290,1314]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="257">SANC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCECA65FFCAFFF240B19726A067F9DB" blockId="2.[151,1437,1001,2034]" pageId="2" pageNumber="257">
<materialsCitation id="3B19C038FFCAFFF240B19726A7D0FA2B" ID-GBIF-Occurrence="1455889629" collectionCode="PART, OF, SERIES, SANC, TYPS" collectorName="R. Stals &amp; D T Bilton" country="South Africa" county="Pretoria" location="Gr" municipality="Recognised" pageId="2" pageNumber="257" specimenCode="TYPS00594" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paralectotype">
<typeStatus id="54CA74C7FFCAFFF240B19726A470FADF" box="[199,375,1325,1350]" pageId="2" pageNumber="257" type="paralectotype">
<emphasis id="B9051677FFCAFFF240B19726A470FADF" bold="true" box="[199,375,1325,1350]" pageId="2" pageNumber="257">Paralectotypes</emphasis>
</typeStatus>
(9): 
<specimenCount id="9D7701ECFFCAFFF241D99725A4DAFADC" box="[431,477,1326,1350]" pageId="2" pageNumber="257" type="male">2 ♂</specimenCount>
, 
<specimenCount id="9D7701ECFFCAFFF2419C9725A712FADC" box="[490,533,1326,1350]" pageId="2" pageNumber="257" type="female">1 ♀</specimenCount>
[mounted on one card] “ 4 
<location id="8EAE9CBEFFCAFFF243259725A674FADF" LSID="urn:lsid:plazi:treatment:03D87B73FFCAFFF940E1918FA640FB71:8EAE9CBEFFCAFFF243259725A674FADF" box="[851,883,1326,1350]" country="South Africa" county="Pretoria" municipality="Recognised" name="Gr" pageId="2" pageNumber="257">Gr</location>
” [HW], “ 
<collectionCode id="ED6052A0FFCAFFF243A99725A155FADF" box="[991,1106,1326,1350]" pageId="2" pageNumber="257">PART OF</collectionCode>
<typeStatus id="54CA74C7FFCAFFF2442F9725A1D0FADF" box="[1113,1239,1326,1350]" pageId="2" pageNumber="257" type="syntype">SYNTYPE</typeStatus>
<collectionCode id="ED6052A0FFCAFFF244969725A03BFADF" box="[1248,1340,1326,1350]" pageId="2" pageNumber="257">SERIES</collectionCode>
// 
<taxonomicName id="4C71B1E6FFCAFFF2453A9726A4E3FAF3" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" class="Insecta" family="Elmidae" genus="Helmis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF2453A9726A42FFAF0" italics="true" pageId="2" pageNumber="257">Helmis georyssoides</emphasis>
<bibRefCitation id="EFE0B794FFCAFFF24159975AA4E3FAF3" author="Grouvelle" box="[303,484,1361,1386]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
</taxonomicName>
// 
<collectingMunicipality id="6BAA501FFFCAFFF24183975AA77FFAF3" box="[501,632,1361,1386]" pageId="2" pageNumber="257">Recognised</collectingMunicipality>
as such: 
<collectorName id="2684AFB3FFCAFFF242929759A63AFAF3" box="[740,829,1361,1386]" pageId="2" pageNumber="257">R. Stals</collectorName>
, 2015.”, “NATIONAL COLLECTION 
<collectionCode id="ED6052A0FFCAFFF244899759A024FAF3" box="[1279,1315,1362,1386]" pageId="2" pageNumber="257">OF</collectionCode>
INSECTS (
<collectionCode id="ED6052A0FFCAFFF240E8977DA5EDFA17" box="[158,234,1398,1422]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="257">SANC</collectionCode>
)// 
<collectingCounty id="62AFB2E9FFCAFFF24178977EA46DFA17" box="[270,362,1397,1422]" pageId="2" pageNumber="257">Pretoria</collectingCounty>
, 
<collectingCountry id="F3668AF5FFCAFFF2410F977EA70BFA17" box="[377,524,1397,1422]" name="South Africa" pageId="2" pageNumber="257">South Africa</collectingCountry>
”, “ 
<collectionCode id="ED6052A0FFCAFFF24242977DA786FA17" box="[564,641,1398,1422]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="257">SANC</collectionCode>
<specimenCode id="DBD7621EFFCAFFF242FA977EA61EFA17" box="[652,793,1397,1422]" collectionCode="TYPS" pageId="2" pageNumber="257">
<collectionCode id="ED6052A0FFCAFFF242FA977EA7D6FA17" box="[652,721,1397,1422]" pageId="2" pageNumber="257">TYPS</collectionCode>
00594
</specimenCode>
” [green label], “ 
<taxonomicName id="4C71B1E6FFCAFFF243AB977EA08DFA17" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" box="[989,1418,1397,1422]" class="Insecta" family="Elmidae" genus="Helmis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF243AB977EA1CFFA14" box="[989,1224,1397,1421]" italics="true" pageId="2" pageNumber="257">Helmis georyssoides</emphasis>
<bibRefCitation id="EFE0B794FFCAFFF244A5977EA08DFA17" author="Grouvelle" box="[1235,1418,1397,1422]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
</taxonomicName>
// 
<typeStatus id="54CA74C7FFCAFFF240E19792A428FA2B" box="[151,303,1433,1458]" pageId="2" pageNumber="257" type="paralectotype">Paralectotype</typeStatus>
<collectorName id="2684AFB3FFCAFFF2414D9791A4BAFA2B" box="[315,445,1433,1458]" pageId="2" pageNumber="257">D T Bilton</collectorName>
des. 2017” [red labels]
</materialsCitation>
; 
<materialsCitation id="3B19C038FFCAFFF242919791A067F9DB" ID-GBIF-Occurrence="1455889628" collectionCode="PART, OF, SERIES, SANC, TYPS" collectorName="R. Stals &amp; D T Bilton &amp; All &amp; de Bonne Esperance &amp; M. L. Peringuey" country="South Africa" location="Recognised" municipality="Pretoria" pageId="2" pageNumber="257" specimenCode="TYPS00594" specimenCount="3" specimenCount-female="3" typeStatus="syntype">
<specimenCount id="9D7701ECFFCAFFF242919791A611FA28" box="[743,790,1434,1458]" pageId="2" pageNumber="257" type="female">3 ♀</specimenCount>
, 3 inds [mounted on one card] “ 
<collectionCode id="ED6052A0FFCAFFF244EB9791A014FA2B" box="[1181,1299,1434,1458]" pageId="2" pageNumber="257">PART OF</collectionCode>
<typeStatus id="54CA74C7FFCAFFF245689791A09BFA2B" box="[1310,1436,1434,1458]" pageId="2" pageNumber="257" type="syntype">SYNTYPE</typeStatus>
<collectionCode id="ED6052A0FFCAFFF240E197B5A5F4FA4F" box="[151,243,1470,1494]" pageId="2" pageNumber="257">SERIES</collectionCode>
// 
<taxonomicName id="4C71B1E6FFCAFFF2417297B6A7DBFA4F" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" box="[260,732,1469,1494]" class="Insecta" family="Elmidae" genus="Helmis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF2417297B6A4FAFA4C" box="[260,509,1469,1493]" italics="true" pageId="2" pageNumber="257">Helmis georyssoides</emphasis>
<bibRefCitation id="EFE0B794FFCAFFF2426097B6A7DBFA4F" author="Grouvelle" box="[534,732,1469,1494]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
</taxonomicName>
// 
<location id="8EAE9CBEFFCAFFF2429B97B6A677FA4F" LSID="urn:lsid:plazi:treatment:03D87B73FFCAFFF940E1918FA640FB71:8EAE9CBEFFCAFFF2429B97B6A677FA4F" box="[749,880,1469,1494]" country="South Africa" municipality="Pretoria" name="Recognised" pageId="2" pageNumber="257">Recognised</location>
as such: 
<collectorName id="2684AFB3FFCAFFF2447897B5A17EFA4F" box="[1038,1145,1469,1494]" pageId="2" pageNumber="257">R. Stals</collectorName>
, 2015.”, “NATIONAL COLLECTION 
<collectionCode id="ED6052A0FFCAFFF2412497E9A471FA63" box="[338,374,1506,1530]" pageId="2" pageNumber="257">OF</collectionCode>
INSECTS (
<collectionCode id="ED6052A0FFCAFFF2427097E9A755FA63" box="[518,594,1506,1530]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="257">SANC</collectionCode>
)// 
<collectingMunicipality id="6BAA501FFFCAFFF2420197EAA7D4FA63" box="[631,723,1505,1530]" pageId="2" pageNumber="257">Pretoria</collectingMunicipality>
, 
<collectingCountry id="F3668AF5FFCAFFF2429497EAA670FA63" box="[738,887,1505,1530]" name="South Africa" pageId="2" pageNumber="257">South Africa</collectingCountry>
”, “ 
<collectionCode id="ED6052A0FFCAFFF243D697E9A6EAFA63" box="[928,1005,1506,1530]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="257">SANC</collectionCode>
<specimenCode id="DBD7621EFFCAFFF2438E97EAA182FA63" box="[1016,1157,1505,1530]" collectionCode="TYPS" pageId="2" pageNumber="257">
<collectionCode id="ED6052A0FFCAFFF2438E97EAA13AFA63" box="[1016,1085,1505,1530]" pageId="2" pageNumber="257">TYPS</collectionCode>
00594
</specimenCode>
” [green label], “ 
<taxonomicName id="4C71B1E6FFCAFFF2453A97EAA4EDF987" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" class="Insecta" family="Elmidae" genus="Helmis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF2453A97EAA42FF984" italics="true" pageId="2" pageNumber="257">Helmis georyssoides</emphasis>
<bibRefCitation id="EFE0B794FFCAFFF24144940EA4EDF987" author="Grouvelle" box="[306,490,1541,1566]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
</taxonomicName>
// 
<typeStatus id="54CA74C7FFCAFFF24270940EA79AF987" box="[518,669,1541,1566]" pageId="2" pageNumber="257" type="paralectotype">Paralectotype</typeStatus>
// 
<collectorName id="2684AFB3FFCAFFF242CF940DA63CF987" box="[697,827,1541,1566]" pageId="2" pageNumber="257">D T Bilton</collectorName>
des. 2017” [red labels]. 
<collectorName id="2684AFB3FFCAFFF24416940EA183F987" box="[1120,1156,1541,1566]" pageId="2" pageNumber="257">All</collectorName>
specimens lack locality labels, but the original description (
<bibRefCitation id="EFE0B794FFCAFFF242559422A7D2F9DB" author="Grouvelle" box="[547,725,1577,1602]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle 1890</bibRefCitation>
) states “..cap 
<collectorName id="2684AFB3FFCAFFF243049422A151F9DB" box="[882,1110,1577,1602]" pageId="2" pageNumber="257">de Bonne Espérance</collectorName>
par 
<collectorName id="2684AFB3FFCAFFF244FE9421A040F9DB" box="[1160,1351,1577,1602]" pageId="2" pageNumber="257">M. L. Péringuey</collectorName>
..”.
</materialsCitation>
</paragraph>
<paragraph id="8BCECA65FFCAFFF240B19446A09BF868" blockId="2.[151,1437,1001,2034]" pageId="2" pageNumber="257">
<materialsCitation id="3B19C038FFCAFFF240B19446A443F937" ID-GBIF-Occurrence="1455889626" collectingDate="2015-02-12" collectionCode="TMNP, AMG, CDTB, ISAM, SANC" collectorName="D T Bilton" country="South Africa" determinerName="D T Bilton" location="Echo Valley" municipality="Table Mountain" pageId="2" pageNumber="257" specimenCount="19" specimenCount-female="6" specimenCount-male="13">
<emphasis id="B9051677FFCAFFF240B19446A731F9FF" bold="true" box="[199,566,1613,1638]" pageId="2" pageNumber="257">Additional material examined:</emphasis>
<specimenCount id="9D7701ECFFCAFFF2424B9445A770F9FC" box="[573,631,1614,1638]" pageId="2" pageNumber="257" type="male">13 ♂</specimenCount>
, 
<specimenCount id="9D7701ECFFCAFFF242F29446A7A9F9FC" box="[644,686,1613,1638]" pageId="2" pageNumber="257" type="female">6 ♀</specimenCount>
“ 
<collectingDate id="EF8B154DFFCAFFF242C99446A632F9FF" box="[703,821,1613,1638]" pageId="2" pageNumber="257" value="2015-02-12">
<date id="FFCFECA5FFCAFFF242C99446A632F9FF" box="[703,821,1613,1638]" pageId="2" pageNumber="257" value="2015-02-12">12/ii/2015</date>
</collectingDate>
<collectingCountry id="F3668AF5FFCAFFF2434A9446A6CBF9FF" box="[828,972,1613,1638]" name="South Africa" pageId="2" pageNumber="257">South Africa</collectingCountry>
WC// 
<collectionCode id="ED6052A0FFCAFFF244619445A162F9FC" box="[1047,1125,1614,1637]" pageId="2" pageNumber="257">TMNP</collectionCode>
<collectingMunicipality id="6BAA501FFFCAFFF2441A9446A01AF9FF" box="[1132,1309,1613,1638]" pageId="2" pageNumber="257">Table Mountain</collectingMunicipality>
// stream in 
<location id="8EAE9CBEFFCAFFF240E1947AA427F913" LSID="urn:lsid:plazi:treatment:03D87B73FFCAFFF940E1918FA640FB71:8EAE9CBEFFCAFFF240E1947AA427F913" box="[151,288,1649,1674]" country="South Africa" municipality="Table Mountain" name="Echo Valley" pageId="2" pageNumber="257">Echo Valley</location>
// 
<collectorName id="2684AFB3FFCAFFF2414E9479A4B3F913" box="[312,436,1649,1674]" pageId="2" pageNumber="257">D T Bilton</collectorName>
leg.”, “ 
<taxonomicName id="4C71B1E6FFCAFFF24278947AA6D0F913" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" box="[526,983,1649,1674]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF24278947AA601F910" box="[526,774,1649,1673]" italics="true" pageId="2" pageNumber="257">Leielmis georyssoides</emphasis>
(
<bibRefCitation id="EFE0B794FFCAFFF24360947AA6C9F913" author="Grouvelle" box="[790,974,1649,1674]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
)
</taxonomicName>
// 
<determinerName id="7E1226A1FFCAFFF243879479A16AF913" box="[1009,1133,1649,1674]" pageId="2" pageNumber="257">D T Bilton</determinerName>
det. 2016” (
<collectionCode id="ED6052A0FFCAFFF2448D9479A039F913" box="[1275,1342,1650,1674]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="2" pageNumber="257">AMG</collectionCode>
, 
<collectionCode id="ED6052A0FFCAFFF2453C9479A090F913" box="[1354,1431,1650,1674]" pageId="2" pageNumber="257">CDTB</collectionCode>
, 
<collectionCode id="ED6052A0FFCAFFF240E1949DA5E7F937" box="[151,224,1686,1710]" pageId="2" pageNumber="257">ISAM</collectionCode>
, 
<collectionCode id="ED6052A0FFCAFFF2409A949DA43BF937" box="[236,316,1686,1710]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="257">SANC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B19C038FFCAFFF24127949DA189F94B" ID-GBIF-Occurrence="1455889609" collectingDate="2010-09-24" collectionCode="TMNP, CDTB" collectorName="D. T. Bilton" country="South Africa" determinerName="D T Bilton" location="Echo Valley" municipality="Table Mountain" pageId="2" pageNumber="257" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D7701ECFFCAFFF24127949DA479F934" box="[337,382,1686,1710]" pageId="2" pageNumber="257" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D7701ECFFCAFFF241FA949DA4B0F934" box="[396,439,1686,1710]" pageId="2" pageNumber="257" type="female">1 ♀</specimenCount>
“ 
<collectingDate id="EF8B154DFFCAFFF241BF949EA742F937" box="[457,581,1685,1710]" pageId="2" pageNumber="257" value="2010-09-24">
<date id="FFCFECA5FFCAFFF241BF949EA742F937" box="[457,581,1685,1710]" pageId="2" pageNumber="257" value="2010-09-24">24/ix/2010</date>
</collectingDate>
<collectingCountry id="F3668AF5FFCAFFF24238949EA7D8F937" box="[590,735,1685,1710]" name="South Africa" pageId="2" pageNumber="257">South Africa</collectingCountry>
WC// 
<collectionCode id="ED6052A0FFCAFFF2435B949DA67CF934" box="[813,891,1686,1709]" pageId="2" pageNumber="257">TMNP</collectionCode>
<collectingMunicipality id="6BAA501FFFCAFFF243F5949EA131F937" box="[899,1078,1685,1710]" pageId="2" pageNumber="257">Table Mountain</collectingMunicipality>
// stream in 
<location id="8EAE9CBEFFCAFFF244B6949EA04DF937" LSID="urn:lsid:plazi:treatment:03D87B73FFCAFFF940E1918FA640FB71:8EAE9CBEFFCAFFF244B6949EA04DF937" box="[1216,1354,1685,1710]" country="South Africa" municipality="Table Mountain" name="Echo Valley" pageId="2" pageNumber="257">Echo Valley</location>
// 
<collectorName id="2684AFB3FFCAFFF24514949DA5DAF94B" pageId="2" pageNumber="257">D. T. Bilton</collectorName>
leg.”, “ 
<taxonomicName id="4C71B1E6FFCAFFF2414C94B2A600F94B" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" box="[314,775,1721,1746]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF2414C94B2A734F948" box="[314,563,1721,1745]" italics="true" pageId="2" pageNumber="257">Leielmis georyssoides</emphasis>
(
<bibRefCitation id="EFE0B794FFCAFFF2423394B2A7F8F94B" author="Grouvelle" box="[581,767,1721,1746]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
)
</taxonomicName>
// 
<determinerName id="7E1226A1FFCAFFF2435494B1A6A5F94B" box="[802,930,1721,1746]" pageId="2" pageNumber="257">D T Bilton</determinerName>
det. 2016” (
<collectionCode id="ED6052A0FFCAFFF2444394B1A181F94B" box="[1077,1158,1722,1746]" pageId="2" pageNumber="257">CDTB</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B19C038FFCAFFF244EA94B1A18BF883" ID-GBIF-Occurrence="1455889627" collectingDate="1997-03-04" collectionCode="NHMW" collectorName="Hess &amp; Heckes" country="South Africa" county="Table Mtn" determinerName="D T Bilton" elevation="500" latitude="-35.58" location="Skeleton Gorge" longLatPrecision="715" longitude="18.28" municipality="Kir" pageId="2" pageNumber="257" specimenCount="14" specimenCount-female="7" specimenCount-male="7" stateProvince="Cape">
<specimenCount id="9D7701ECFFCAFFF244EA94B1A1CCF948" box="[1180,1227,1722,1745]" pageId="2" pageNumber="257" type="male">7 ♂</specimenCount>
, 
<specimenCount id="9D7701ECFFCAFFF244AC94B1A000F948" box="[1242,1287,1722,1745]" pageId="2" pageNumber="257" type="female">7 ♀</specimenCount>
“ 
<collectingCountry id="F3668AF5FFCAFFF2456D94B1A051F94B" box="[1307,1366,1722,1746]" name="South Africa" pageId="2" pageNumber="257">RSA</collectingCountry>
, West 
<collectingRegion id="49B50487FFCAFFF240E194D5A5D2F96F" box="[151,213,1758,1782]" country="South Africa" name="Western Cape" pageId="2" pageNumber="257">Cape</collectingRegion>
: 
<collectingCounty id="62AFB2E9FFCAFFF240A994D6A452F96F" box="[223,341,1757,1782]" pageId="2" pageNumber="257">Table Mtn</collectingCounty>
//[= 
<geoCoordinate id="EE45ACA2FFCAFFF241F594D6A4D7F96F" box="[387,464,1757,1782]" direction="south" orientation="latitude" pageId="2" pageNumber="257" precision="555" value="-35.58">35.58S</geoCoordinate>
/ 
<geoCoordinate id="EE45ACA2FFCAFFF2419794D5A733F96F" box="[481,564,1758,1782]" direction="east" orientation="longitude" pageId="2" pageNumber="257" precision="555" value="18.28">18.28E</geoCoordinate>
] bei 
<collectingMunicipality id="6BAA501FFFCAFFF2421D94D6A788F96F" box="[619,655,1757,1782]" pageId="2" pageNumber="257">Kir</collectingMunicipality>
// stenbosch, 
<location id="8EAE9CBEFFCAFFF2435294D6A6D2F96F" LSID="urn:lsid:plazi:treatment:03D87B73FFCAFFF940E1918FA640FB71:8EAE9CBEFFCAFFF2435294D6A6D2F96F" box="[804,981,1757,1782]" country="South Africa" county="Table Mtn" latitude="-35.58" longLatPrecision="715" longitude="18.28" municipality="Kir" name="Skeleton Gorge" pageId="2" pageNumber="257" stateProvince="Cape">Skeleton Gorge</location>
,// 
<elevation id="005C2D56FFCAFFF2438794D5A195F96C" box="[1009,1170,1757,1782]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="2" pageNumber="257" unit="m" value="500.0" valueMax="700.0" valueMin="300.0">
<quantity id="4C896780FFCAFFF2438794D5A195F96C" box="[1009,1170,1757,1782]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="2" pageNumber="257" unit="m" value="500.0" valueMax="700.0" valueMin="300.0">300 bis 700 m</quantity>
</elevation>
NN. 
<collectingDate id="EF8B154DFFCAFFF244A694D5A036F96F" box="[1232,1329,1758,1782]" pageId="2" pageNumber="257" value="1997-03-04">
<date id="FFCFECA5FFCAFFF244A694D5A036F96F" box="[1232,1329,1758,1782]" pageId="2" pageNumber="257" value="1997-03-04">4.3.1997</date>
</collectingDate>
// 
<collectorName id="2684AFB3FFCAFFF2453F94D5A078F96C" box="[1353,1407,1758,1781]" pageId="2" pageNumber="257">Hess</collectorName>
&amp; 
<collectorName id="2684AFB3FFCAFFF240E1950AA5EDF883" box="[151,234,1793,1818]" pageId="2" pageNumber="257">Heckes</collectorName>
leg.”, “ 
<taxonomicName id="4C71B1E6FFCAFFF24136950AA602F883" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" box="[320,773,1793,1818]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF24136950AA730F880" box="[320,567,1793,1817]" italics="true" pageId="2" pageNumber="257">Leielmis georyssoides</emphasis>
(
<bibRefCitation id="EFE0B794FFCAFFF24230950AA7FAF883" author="Grouvelle" box="[582,765,1793,1818]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
)
</taxonomicName>
// 
<determinerName id="7E1226A1FFCAFFF243689509A690F883" box="[798,919,1793,1818]" pageId="2" pageNumber="257">D T Bilton</determinerName>
det. 2016” (
<collectionCode id="ED6052A0FFCAFFF244549509A183F883" box="[1058,1156,1794,1818]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="257">NHMW</collectionCode>
)
</materialsCitation>
; 3 ♂ “REP. 
<materialsCitation id="3B19C038FFCAFFF245629509A7F0F8FB" ID-GBIF-Occurrence="1455889616" collectingDate="1997-02-27" collectionCode="OF, NHMW" collectorName="G Challet" country="South Africa" determinerName="D T Bilton" location="Reservoir" municipality="Capetown" pageId="2" pageNumber="257" specimenCount="1">
<collectionCode id="ED6052A0FFCAFFF245629509A03FF883" box="[1300,1336,1794,1818]" pageId="2" pageNumber="257">OF</collectionCode>
<collectingCountry id="F3668AF5FFCAFFF245499509A5FAF8A7" name="South Africa" pageId="2" pageNumber="257">SOUTH AFRICA</collectingCountry>
// 
<collectingMunicipality id="6BAA501FFFCAFFF2416B952DA495F8A7" box="[285,402,1830,1854]" pageId="2" pageNumber="257">Capetown</collectingMunicipality>
, Table Mtn.,// stream nr. 
<location id="8EAE9CBEFFCAFFF242AF952EA642F8A7" LSID="urn:lsid:plazi:treatment:03D87B73FFCAFFF940E1918FA640FB71:8EAE9CBEFFCAFFF242AF952EA642F8A7" box="[729,837,1829,1854]" country="South Africa" municipality="Capetown" name="Reservoir" pageId="2" pageNumber="257">Reservoir</location>
// 
<collectingDate id="EF8B154DFFCAFFF24310952DA6D2F8A7" box="[870,981,1830,1854]" pageId="2" pageNumber="257" value="1997-02-27">
<date id="FFCFECA5FFCAFFF24310952DA6D2F8A7" box="[870,981,1830,1854]" pageId="2" pageNumber="257" value="1997-02-27">27.2.1997</date>
</collectingDate>
// 
<collectorName id="2684AFB3FFCAFFF24380952DA176F8A7" box="[1014,1137,1829,1854]" pageId="2" pageNumber="257">G Challet</collectorName>
”, “ 
<taxonomicName id="4C71B1E6FFCAFFF244EA952EA467F8FB" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF244EA952EA09BF8A4" box="[1180,1436,1829,1853]" italics="true" pageId="2" pageNumber="257">Leielmis georyssoides</emphasis>
(
<bibRefCitation id="EFE0B794FFCAFFF240E89542A45EF8FB" author="Grouvelle" box="[158,345,1865,1890]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
)
</taxonomicName>
// 
<determinerName id="7E1226A1FFCAFFF2410A9541A4FDF8FB" box="[380,506,1865,1890]" pageId="2" pageNumber="257">D T Bilton</determinerName>
det. 2016” (
<collectionCode id="ED6052A0FFCAFFF242FB9541A7E8F8FB" box="[653,751,1866,1890]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="257">NHMW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B19C038FFCAFFF243739541A7DEF857" ID-GBIF-Occurrence="1455889624" collectingDate="1954-02-28" collectionCode="BMNH" collectorName="J. Balfour-Browne" country="South Africa" determinerName="J. Deleve &amp; D T Bilton" location="Cape Province" pageId="2" pageNumber="257" specimenCount="1" specimenCount-male="1" stateProvince="Cape Province">
<specimenCount id="9D7701ECFFCAFFF243739541A635F8F8" box="[773,818,1866,1890]" pageId="2" pageNumber="257" type="male">1 ♂</specimenCount>
“STN 5” [HW], “S AFRICA// 
<collectingRegion id="49B50487FFCAFFF244D49541A04EF8FB" box="[1186,1353,1865,1890]" country="South Africa" name="Western Cape" pageId="2" pageNumber="257">Cape Province</collectingRegion>
// 
<collectingRegion id="49B50487FFCAFFF245159541A09AF8FB" box="[1379,1437,1866,1890]" country="South Africa" name="Western Cape" pageId="2" pageNumber="257">Cape</collectingRegion>
District.// 
<collectorName id="2684AFB3FFCAFFF2417B9565A4D8F81F" box="[269,479,1901,1926]" pageId="2" pageNumber="257">J. Balfour-Browne</collectorName>
// B.M. 1954-797.” [HW], “Kirstenbosch.// 
<collectingDate id="EF8B154DFFCAFFF243A99566A149F81F" box="[991,1102,1901,1926]" pageId="2" pageNumber="257" value="1954-02-28">
<date id="FFCFECA5FFCAFFF243A99566A149F81F" box="[991,1102,1901,1926]" pageId="2" pageNumber="257" value="1954-02-28">28.ii.1954</date>
</collectingDate>
// ‘nearly dry waterfall// and stream.’” [HW], “ 
<determinerName id="7E1226A1FFCAFFF2411E9599A4D3F833" box="[360,468,1937,1962]" pageId="2" pageNumber="257">J. Delève</determinerName>
det., 1965// 
<taxonomicName id="4C71B1E6FFCAFFF2421A9599A649F833" authority="Gr." authorityName="Gr." box="[620,846,1937,1962]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">L. georyssoides Gr.</taxonomicName>
” [5 &amp; determination HW], “ 
<taxonomicName id="4C71B1E6FFCAFFF244D4959AA45AF857" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCAFFF244D4959AA09BF830" box="[1186,1436,1937,1961]" italics="true" pageId="2" pageNumber="257">Leielmis georyssoides</emphasis>
(
<bibRefCitation id="EFE0B794FFCAFFF240E895BEA452F857" author="Grouvelle" box="[158,341,1973,1998]" pageId="2" pageNumber="257" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
)
</taxonomicName>
// 
<determinerName id="7E1226A1FFCAFFF2410395BDA4E9F857" box="[373,494,1973,1998]" pageId="2" pageNumber="257">D T Bilton</determinerName>
det. 2016” (
<collectionCode id="ED6052A0FFCAFFF2420195BDA7D6F857" box="[631,721,1974,1998]" pageId="2" pageNumber="257">BMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B19C038FFCAFFF2429295BDA09BF868" ID-GBIF-Occurrence="1455889630" collectingDate="1954-03-02" collectorName="S. Africa. &amp; District. &amp; J. Balfour-Browne." country="South Africa" elevation="503" location="Blinkwater falls" pageId="2" pageNumber="257" specimenCount="2" specimenCount-female="2" stateProvince="Cape Province">
<specimenCount id="9D7701ECFFCAFFF2429295BDA609F854" box="[740,782,1974,1998]" pageId="2" pageNumber="257" type="female">1 ♀</specimenCount>
“ 
<specimenCount id="9D7701ECFFCAFFF2435595BDA632F857" box="[803,821,1974,1998]" pageId="2" pageNumber="257" type="female">♀</specimenCount>
”, “Stn No.// 9” [9 HW], “ 
<location id="8EAE9CBEFFCAFFF2442095BEA00BF857" LSID="urn:lsid:plazi:treatment:03D87B73FFCAFFF940E1918FA640FB71:8EAE9CBEFFCAFFF2442095BEA00BF857" box="[1110,1292,1973,1998]" country="South Africa" name="Blinkwater falls" pageId="2" pageNumber="257" stateProvince="Cape Province">Blinkwater falls</location>
,// in running water// (dry season) 
<elevation id="005C2D56FFCAFFF2410B95D2A4C8F86B" box="[381,463,2009,2034]" metricMagnitude="2" metricUnit="m" metricValue="5.0291999999999994" pageId="2" pageNumber="257" unit="ft" value="1650.0">
<quantity id="4C896780FFCAFFF2410B95D2A4C8F86B" box="[381,463,2009,2034]" metricMagnitude="2" metricUnit="m" metricValue="5.0291999999999994" pageId="2" pageNumber="257" unit="ft" value="1650.0">1650 ft</quantity>
</elevation>
.// 
<collectingDate id="EF8B154DFFCAFFF2419D95D2A75DF86B" box="[491,602,2009,2034]" pageId="2" pageNumber="257" value="1954-03-02">
<date id="FFCFECA5FFCAFFF2419D95D2A75DF86B" box="[491,602,2009,2034]" pageId="2" pageNumber="257" value="1954-03-02">2.iii.1954</date>
</collectingDate>
”, “ 
<collectorName id="2684AFB3FFCAFFF2420895D1A7EEF86B" box="[638,745,2009,2034]" pageId="2" pageNumber="257">S. Africa.</collectorName>
// 
<collectingRegion id="49B50487FFCAFFF2437695D1A6A0F86B" box="[768,935,2009,2034]" country="South Africa" name="Western Cape" pageId="2" pageNumber="257">Cape Province</collectingRegion>
.// 
<collectingRegion id="49B50487FFCAFFF243B495D1A6FCF86B" box="[962,1019,2010,2034]" country="South Africa" name="Western Cape" pageId="2" pageNumber="257">Cape</collectingRegion>
<collectorName id="2684AFB3FFCAFFF2447595D2A15AF86B" box="[1027,1117,2009,2034]" pageId="2" pageNumber="257">District.</collectorName>
// 
<collectorName id="2684AFB3FFCAFFF2440395D1A04CF86B" box="[1141,1355,2009,2034]" pageId="2" pageNumber="257">J. Balfour-Browne.</collectorName>
// B.M.
</materialsCitation>
</paragraph>
<paragraph id="8BCECA65FFCBFFF340E1929CA67FFED9" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="258">
1954-797.”, “J. Delève det., 1964// L. georryssoides [sic!] Gr.” [4 &amp; determination HW], “ 
<taxonomicName id="4C71B1E6FFCBFFF344D29293A459FF4D" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCBFFF344D29293A09BFF29" box="[1188,1436,152,176]" italics="true" pageId="3" pageNumber="258">Leielmis georyssoides</emphasis>
(
<bibRefCitation id="EFE0B794FFCBFFF340E892B7A451FF4D" author="Grouvelle" box="[158,342,188,213]" pageId="3" pageNumber="258" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
)
</taxonomicName>
// D T Bilton det. 2016” (BMNH); 
<materialsCitation id="3B19C038FFCBFFF3429A92B7A673FED9" ID-GBIF-Occurrence="1455889617" collectingDate="1954-02-28" collectionCode="BMNH" collectorName="J. Balfour-Browne" determinerName="J. Deleve &amp; D T Bilton" location="Cape Province" pageId="3" pageNumber="258" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cape Province">
<specimenCount id="9D7701ECFFCBFFF3429A92B7A611FF4D" box="[748,790,188,212]" pageId="3" pageNumber="258" type="male">1 ♂</specimenCount>
“STN 5” [HW], “Kirstenbosch.// 
<collectingDate id="EF8B154DFFCBFFF344E192B7A002FF4C" box="[1175,1285,188,213]" pageId="3" pageNumber="258" value="1954-02-28">
<date id="FFCFECA5FFCBFFF344E192B7A002FF4C" box="[1175,1285,188,213]" pageId="3" pageNumber="258" value="1954-02-28">28.ii.1954</date>
</collectingDate>
// ‘nearly dry waterfall// and stream.’” [HW], “S AFRICA// 
<collectingRegion id="49B50487FFCBFFF342C792EBA650FF61" box="[689,855,223,248]" country="South Africa" name="Western Cape" pageId="3" pageNumber="258">Cape Province</collectingRegion>
// 
<collectingRegion id="49B50487FFCBFFF3430492EBA6ABFF61" box="[882,940,224,248]" country="South Africa" name="Western Cape" pageId="3" pageNumber="258">Cape</collectingRegion>
District.// 
<collectorName id="2684AFB3FFCBFFF3445C92EBA1FAFF61" box="[1066,1277,223,248]" pageId="3" pageNumber="258">J. Balfour-Browne</collectorName>
// B.M. 1954- 797.” [HW], “ 
<specimenCount id="9D7701ECFFCBFFF3413E930FA45DFE85" box="[328,346,260,284]" pageId="3" pageNumber="258" type="female">♀</specimenCount>
” [sic!], “ 
<determinerName id="7E1226A1FFCBFFF341A3930FA743FE84" box="[469,580,260,285]" pageId="3" pageNumber="258">J. Delève</determinerName>
det., 1965// 
<taxonomicName id="4C71B1E6FFCBFFF3429F930EA6D3FE85" authority="Gr." authorityName="Gr." box="[745,980,260,285]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="georyssoides">L. georyssoides Gr.</taxonomicName>
” [5 &amp; determination HW], “ 
<taxonomicName id="4C71B1E6FFCBFFF3454B930FA4F2FED9" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCBFFF3454B930FA42FFED9" italics="true" pageId="3" pageNumber="258">Leielmis georyssoides</emphasis>
(
<bibRefCitation id="EFE0B794FFCBFFF34140932CA4E9FED9" author="Grouvelle" box="[310,494,295,320]" pageId="3" pageNumber="258" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
)
</taxonomicName>
// 
<determinerName id="7E1226A1FFCBFFF3427B9323A780FED9" box="[525,647,295,320]" pageId="3" pageNumber="258">D T Bilton</determinerName>
det. 2016” (
<collectionCode id="ED6052A0FFCBFFF343649323A66BFED9" box="[786,876,296,320]" pageId="3" pageNumber="258">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36B99EEFFCBFFF940B19347A7D6FE49" lastPageId="9" lastPageNumber="264" pageId="3" pageNumber="258" type="description">
<paragraph id="8BCECA65FFCBFFF340B19347A488FE34" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="258">
<emphasis id="B9051677FFCBFFF340B19347A456FEFC" bold="true" box="[199,337,332,357]" pageId="3" pageNumber="258">Description</emphasis>
. Size: 
<typeStatus id="54CA74C7FFCBFFF341D39346A71EFEFC" box="[421,537,333,357]" pageId="3" pageNumber="258" type="lectotype">Lectotype</typeStatus>
: body length (front of pronotum to elytral apices) 
<quantity id="4C896780FFCBFFF344119347A1D5FEFD" box="[1127,1234,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="258" unit="mm" value="1.55">1.55 mm</quantity>
; maximum width (elytra) 
<quantity id="4C896780FFCBFFF340879364A45FFE1E" box="[241,344,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="258" unit="mm" value="1.05">1.05 mm</quantity>
; elytral length 
<quantity id="4C896780FFCBFFF342769364A760FE1E" box="[512,615,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="258" unit="mm" value="1.15">1.15 mm</quantity>
. Same values for 
<typeStatus id="54CA74C7FFCBFFF343599364A6D2FE11" box="[815,981,367,392]" pageId="3" pageNumber="258" type="paralectotype">paralectotypes</typeStatus>
: 
<quantity id="4C896780FFCBFFF343969364A182FE1E" box="[992,1157,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.55" metricValueMin="1.05" pageId="3" pageNumber="258" unit="mm" value="1.3" valueMax="1.55" valueMin="1.05">1.05–1.55 mm</quantity>
, 1.0– 
<quantity id="4C896780FFCBFFF344B59364A020FE1E" box="[1219,1319,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="258" unit="mm" value="1.05">1.05 mm</quantity>
and 
<quantity id="4C896780FFCBFFF3452B9364A5FCFE35" metricMagnitude="-3" metricUnit="m" metricValue="1.1749999999999998" metricValueMax="1.2" metricValueMin="1.15" pageId="3" pageNumber="258" unit="mm" value="1.1749999999999998" valueMax="1.2" valueMin="1.15">1.15– 1.20 mm</quantity>
respectively.
</paragraph>
<paragraph id="8BCECA65FFCBFFF340B193BCA7ADFDA4" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="258">
Colour: Dorsum (
<figureCitation id="134AD6E0FFCBFFF341E393BCA4D0FE49" box="[405,471,439,464]" captionStart="FIGURE 1" captionStartId="3.[151,250,1980,2002]" captionTargetBox="[166,1414,644,1940]" captionTargetId="figure@3.[151,1436,625,1958]" captionTargetPageId="3" captionText="FIGURE 1. Leielmis species, male habitus. A) L. georyssoides; B) L. gibbosus sp. nov., holotype; C) L. hirsutus sp. nov., holotype. Scale bar = 1 mm." httpUri="https://zenodo.org/record/545902/files/figure.png" pageId="3" pageNumber="258">Fig. 1</figureCitation>
A) dark brown to black, palest at anterior margin of pronotum. Legs dark reddish brown, tarsi paler, yellowish. Antennae straw yellow, apical 3–4 segments slightly infuscated. Maxillary palpi dark blackish brown; cardo and stipes yellow. Labial palpi straw yellow, apical segment infuscated. Venter dark brown to black; mentum reddish brown; ligula yellow.
</paragraph>
<caption id="DF0E9AEDFFCBFFF340E195B7A4B1F868" httpUri="https://zenodo.org/record/545902/files/figure.png" pageId="3" pageNumber="258" targetBox="[166,1414,644,1940]" targetPageId="3">
<paragraph id="8BCECA65FFCBFFF340E195B7A4B1F868" blockId="3.[151,1435,1980,2033]" pageId="3" pageNumber="258">
<emphasis id="B9051677FFCBFFF340E195B7A412F848" bold="true" box="[151,277,1980,2002]" pageId="3" pageNumber="258">FIGURE 1.</emphasis>
<taxonomicName id="4C71B1E6FFCBFFF3416895B7A473F848" box="[286,372,1980,2001]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B9051677FFCBFFF3416895B7A473F848" box="[286,372,1980,2001]" italics="true" pageId="3" pageNumber="258">Leielmis</emphasis>
</taxonomicName>
species, male habitus. A) 
<taxonomicName id="4C71B1E6FFCBFFF342FD95B6A62CF848" box="[651,811,1980,2002]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCBFFF342FD95B6A62CF848" box="[651,811,1980,2002]" italics="true" pageId="3" pageNumber="258">L. georyssoides</emphasis>
</taxonomicName>
; B) 
<taxonomicName id="4C71B1E6FFCBFFF3432B95B6A6D3F848" box="[861,980,1980,2002]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B9051677FFCBFFF3432B95B6A6D3F848" box="[861,980,1980,2002]" italics="true" pageId="3" pageNumber="258">L. gibbosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFCBFFF343AB95B7A137F848" box="[989,1072,1980,2001]" pageId="3" pageNumber="258" rank="species">
<emphasis id="B9051677FFCBFFF343AB95B7A137F848" bold="true" box="[989,1072,1980,2001]" pageId="3" pageNumber="258">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype; C) 
<taxonomicName id="4C71B1E6FFCBFFF344BC95B6A03EF848" box="[1226,1337,1980,2002]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B9051677FFCBFFF344BC95B6A03EF848" box="[1226,1337,1980,2002]" italics="true" pageId="3" pageNumber="258">L. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFCBFFF3453495B7A092F848" box="[1346,1429,1980,2001]" pageId="3" pageNumber="258" rank="species">
<emphasis id="B9051677FFCBFFF3453495B7A092F848" bold="true" box="[1346,1429,1980,2001]" pageId="3" pageNumber="258">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="DF0E9AEDFFCCFFF440E1948BA6EBF92D" httpUri="https://zenodo.org/record/545903/files/figure.png" pageId="4" pageNumber="259" targetBox="[155,1431,186,1636]" targetPageId="4">
<paragraph id="8BCECA65FFCCFFF440E1948BA6EBF92D" blockId="4.[151,1436,1664,1717]" pageId="4" pageNumber="259">
<emphasis id="B9051677FFCCFFF440E1948BA414F90C" bold="true" box="[151,275,1664,1686]" pageId="4" pageNumber="259">FIGURE 2.</emphasis>
<taxonomicName id="4C71B1E6FFCCFFF4416D948BA476F90C" box="[283,369,1664,1685]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="B9051677FFCCFFF4416D948BA476F90C" box="[283,369,1664,1685]" italics="true" pageId="4" pageNumber="259">Leielmis</emphasis>
</taxonomicName>
species, pronotal sculpture. A) &amp; D) 
<taxonomicName id="4C71B1E6FFCCFFF44284948AA696F90C" box="[754,913,1664,1686]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="georyssiodes">
<emphasis id="B9051677FFCCFFF44284948AA696F90C" box="[754,913,1664,1686]" italics="true" pageId="4" pageNumber="259">L. georyssiodes</emphasis>
</taxonomicName>
; B) &amp; E) 
<taxonomicName id="4C71B1E6FFCCFFF4438F948AA168F90C" box="[1017,1135,1664,1686]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B9051677FFCCFFF4438F948AA168F90C" box="[1017,1135,1664,1686]" italics="true" pageId="4" pageNumber="259">L. gibbosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFCCFFF44400948BA1C0F90C" box="[1142,1223,1664,1685]" pageId="4" pageNumber="259" rank="species">
<emphasis id="B9051677FFCCFFF44400948BA1C0F90C" bold="true" box="[1142,1223,1664,1685]" pageId="4" pageNumber="259">sp. nov.</emphasis>
</taxonomicNameLabel>
; C) &amp; F) 
<taxonomicName id="4C71B1E6FFCCFFF44559948AA09BF90C" box="[1327,1436,1664,1686]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B9051677FFCCFFF44559948AA09BF90C" box="[1327,1436,1664,1686]" italics="true" pageId="4" pageNumber="259">L. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFCCFFF440E19494A5E0F92D" box="[151,231,1695,1716]" pageId="4" pageNumber="259" rank="species">
<emphasis id="B9051677FFCCFFF440E19494A5E0F92D" bold="true" box="[151,231,1695,1716]" pageId="4" pageNumber="259">sp. nov.</emphasis>
</taxonomicNameLabel>
A) – C) scanning electron micrographs; D) – F) stacked macrosope images.
</paragraph>
</caption>
<paragraph id="8BCECA65FFCCFFF440B194D7A172F868" blockId="4.[151,1437,1756,2033]" pageId="4" pageNumber="259">
Head: Triangular, narrowed to apex. Labrum triangular, with acuminately rounded apex; approximately 0.7x as wide as clypeus, 0.5x as long; shining, weakly microreticulate and finely granulate and punctate, punctures bearing curved, recumbent setae. Clypeus also shining, weakly microreticulate and finely granulate and punctate, punctures bearing curved, recumbent setae. Frontoclypeal suture feebly impressed; weakly arcuate. Frons shining, with moderate, medium punctures, bearing curved, recumbent setae; moderately, coarsely granulate and microreticulate. Eyes occupying approximately half side of head; rounded anteriorly, relatively flat laterally and posteriorly. Maxillary palpi with segment 3 flat and expanded laterally, approximately 
<date id="FFCFECA5FFCCFFF4439195BEA104F854" box="[999,1027,1973,1997]" pageId="4" pageNumber="259">2x</date>
broader and longer than segment 4. Labial palpi with segment 2 flat and expanded laterally, segment 3 narrower and shorter.
</paragraph>
<caption id="DF0E9AEDFFCDFFF540E19675A45DFB2B" httpUri="https://zenodo.org/record/545905/files/figure.png" pageId="5" pageNumber="260" targetBox="[164,1408,193,1126]" targetPageId="5">
<paragraph id="8BCECA65FFCDFFF540E19675A45DFB2B" blockId="5.[151,1435,1150,1203]" pageId="5" pageNumber="260">
<emphasis id="B9051677FFCDFFF540E19675A415FB0A" bold="true" box="[151,274,1150,1172]" pageId="5" pageNumber="260">FIGURE 3.</emphasis>
<taxonomicName id="4C71B1E6FFCDFFF5416E9675A469FB0A" box="[280,366,1150,1171]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="B9051677FFCDFFF5416E9675A469FB0A" box="[280,366,1150,1171]" italics="true" pageId="5" pageNumber="260">Leielmis</emphasis>
</taxonomicName>
species, Elytral sculpture, scanning electron micrographs. A) 
<taxonomicName id="4C71B1E6FFCDFFF543979674A179FB0A" box="[993,1150,1150,1172]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCDFFF543979674A179FB0A" box="[993,1150,1150,1172]" italics="true" pageId="5" pageNumber="260">L. georyssoides</emphasis>
</taxonomicName>
; B) 
<taxonomicName id="4C71B1E6FFCDFFF544DC9674A019FB0A" box="[1194,1310,1150,1172]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B9051677FFCDFFF544DC9674A019FB0A" box="[1194,1310,1150,1172]" italics="true" pageId="5" pageNumber="260">L. gibbosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFCDFFF545539674A072FB0D" box="[1317,1397,1151,1172]" pageId="5" pageNumber="260" rank="species">
<emphasis id="B9051677FFCDFFF545539674A072FB0D" bold="true" box="[1317,1397,1151,1172]" pageId="5" pageNumber="260">sp. nov.</emphasis>
</taxonomicNameLabel>
; C) 
<taxonomicName id="4C71B1E6FFCDFFF540E19695A403FB2B" box="[151,260,1181,1203]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B9051677FFCDFFF540E19695A403FB2B" box="[151,260,1181,1203]" italics="true" pageId="5" pageNumber="260">L. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFCDFFF5417F9696A45DFB2B" box="[265,346,1181,1202]" pageId="5" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BCECA65FFCDFFF540B196EBA08FFA49" blockId="5.[151,1437,1247,2029]" pageId="5" pageNumber="260">
Pronotum: Transverse, 1.7x broader than long; broadest at posterior margin, narrowing strongly to front angles; posterior margin approximately 1.4x width of anterior. Arched, anterior margin broadly arcuate in middle, forming a hood which shields much of the head. Posterior margin bisinuate around centre. Lateral margins strongly bordered, weakly rounded. Anterior angles acute; posterior angles obtusely rounded. Surface (
<figureCitation id="134AD6E0FFCDFFF544CF9740A1FCFAFD" box="[1209,1275,1355,1380]" captionStart="FIGURE 2" captionStartId="4.[151,250,1664,1686]" captionTargetBox="[155,1431,186,1636]" captionTargetId="figure@4.[151,1436,183,1659]" captionTargetPageId="4" captionText="FIGURE 2. Leielmis species, pronotal sculpture. A) &amp; D) L. georyssiodes; B) &amp; E) L. gibbosus sp. nov.; C) &amp; F) L. hirsutus sp. nov. A) – C) scanning electron micrographs; D) – F) stacked macrosope images." httpUri="https://zenodo.org/record/545903/files/figure.png" pageId="5" pageNumber="260">Fig. 2</figureCitation>
A, D) shining, without evident microreticulation; doubly punctate, with moderate to close, coarse punctures, interspersed with moderate to close, fine punctures, larger punctures approximately 
<date id="FFCFECA5FFCDFFF543FD979FA6A1FA32" box="[907,934,1428,1451]" pageId="5" pageNumber="260">7x</date>
diameter of smaller ones. Coarse punctures bearing long, fine, golden, recumbent to decumbent setae. Coarsely granulate between punctures, except on disc.
</paragraph>
<paragraph id="8BCECA65FFCDFFF540B197D0A1B0F976" blockId="5.[151,1437,1247,2029]" pageId="5" pageNumber="260">
Elytra: Slightly elongate, broadest approximately one third of the distance behind shoulder, narrowing strongly in posterior third, to acuminately rounded apex. Punctures of rows (
<figureCitation id="134AD6E0FFCDFFF543E197F4A6DDF981" box="[919,986,1535,1560]" captionStart="FIGURE 3" captionStartId="5.[151,250,1150,1172]" captionTargetBox="[164,1408,193,1126]" captionTargetId="figure@5.[164,1408,193,1126]" captionTargetPageId="5" captionText="FIGURE 3. Leielmis species, Elytral sculpture, scanning electron micrographs. A) L. georyssoides; B) L. gibbosus sp. nov.; C) L. hirsutus sp. nov." httpUri="https://zenodo.org/record/545905/files/figure.png" pageId="5" pageNumber="260">Fig. 3</figureCitation>
A) relatively large anteriorly, especially on disc, punctures in anterior third approximately 
<date id="FFCFECA5FFCDFFF542AF942FA7F2F9A5" box="[729,757,1572,1596]" pageId="5" pageNumber="260">2x</date>
diameter of those at start of posterior third. Puncture rows weakly to moderately striate impressed over anterior half of elytra (
<figureCitation id="134AD6E0FFCDFFF543EF944CA6DBF9F9" box="[921,988,1607,1632]" captionStart="FIGURE 3" captionStartId="5.[151,250,1150,1172]" captionTargetBox="[164,1408,193,1126]" captionTargetId="figure@5.[164,1408,193,1126]" captionTargetPageId="5" captionText="FIGURE 3. Leielmis species, Elytral sculpture, scanning electron micrographs. A) L. georyssoides; B) L. gibbosus sp. nov.; C) L. hirsutus sp. nov." httpUri="https://zenodo.org/record/545905/files/figure.png" pageId="5" pageNumber="260">Fig. 3</figureCitation>
A), especially rows 4–6. Puncture rows obsolete towards apex. Intervals relatively flat, shining, with microreticulation reduced to transverse wrinkles; finely punctate, punctures bearing fine, golden, recumbent setae, especially laterally and posteriorly; interval punctures of two sizes, fine and very fine, fine punctures arranged in irregular rows. Surface coarsely granulate posterolaterally. Scutellum shining, no evident microreticulation; sparsely and finely punctate.
</paragraph>
<paragraph id="8BCECA65FFCDFFF540B194F7A113F875" blockId="5.[151,1437,1247,2029]" pageId="5" pageNumber="260">Venter: Entire surface with dense plastron vestiture, except posterior genae and central portion of metaventrite and abdominal ventrites 1–4; thorax and abdomen with medium, curved, golden, recumbent setae and rounded to elongate-oval, coarse, shining granules. Anterior prosternal emargination broad, relatively shallow. Prosternal process approximately 1.2x length of anterior portion of prosternum; quadrate, with raised lateral margins and close, coarse punctures; abruptly narrowing to bluntly acuminate apex. Fovea at junction of meso and metaventrites small and circular. Metaventrite shining, with no visible microreticulation and coarse, close punctures. Sternite 9 with posterior portion 1.5x length of anterior median strut.</paragraph>
<caption id="DF0E9AEDFFCEFFF640E195A7A101F879" httpUri="https://zenodo.org/record/545906/files/figure.png" pageId="6" pageNumber="261" targetBox="[446,1135,210,1910]" targetPageId="6">
<paragraph id="8BCECA65FFCEFFF640E195A7A101F879" blockId="6.[151,1435,1964,2017]" pageId="6" pageNumber="261">
<emphasis id="B9051677FFCEFFF640E195A7A415F858" bold="true" box="[151,274,1964,1986]" pageId="6" pageNumber="261">FIGURE 4.</emphasis>
<taxonomicName id="4C71B1E6FFCEFFF6416E95A7A469F858" box="[280,366,1964,1985]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B9051677FFCEFFF6416E95A7A469F858" box="[280,366,1964,1985]" italics="true" pageId="6" pageNumber="261">Leielmis</emphasis>
</taxonomicName>
species, male tibial modifications, dorsal view A) &amp; D) 
<taxonomicName id="4C71B1E6FFCEFFF643DF95A7A141F858" box="[937,1094,1964,1985]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCEFFF643DF95A7A141F858" box="[937,1094,1964,1985]" italics="true" pageId="6" pageNumber="261">L. georyssoides</emphasis>
</taxonomicName>
; B) &amp; E) 
<taxonomicName id="4C71B1E6FFCEFFF644DC95A7A018F858" box="[1194,1311,1964,1985]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B9051677FFCEFFF644DC95A7A018F858" box="[1194,1311,1964,1985]" italics="true" pageId="6" pageNumber="261">L. gibbosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFCEFFF6455395A7A072F858" box="[1317,1397,1964,1985]" pageId="6" pageNumber="261" rank="species">
<emphasis id="B9051677FFCEFFF6455395A7A072F858" bold="true" box="[1317,1397,1964,1985]" pageId="6" pageNumber="261">sp. nov.</emphasis>
</taxonomicNameLabel>
; C) &amp; F) 
<taxonomicName id="4C71B1E6FFCEFFF640BB95C0A43DF879" box="[205,314,1995,2016]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B9051677FFCEFFF640BB95C0A43DF879" box="[205,314,1995,2016]" italics="true" pageId="6" pageNumber="261">L. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFCEFFF6413695C0A496F879" box="[320,401,1995,2016]" pageId="6" pageNumber="261" rank="species">
<emphasis id="B9051677FFCEFFF6413695C0A496F879" bold="true" box="[320,401,1995,2016]" pageId="6" pageNumber="261">sp. nov.</emphasis>
</taxonomicNameLabel>
A) – C) mesotibiae; D) – F) metatibiae. Scale bar = 0.25 mm.
</paragraph>
</caption>
<caption id="DF0E9AEDFFCFFFF740E195AAA4A4F84F" httpUri="https://zenodo.org/record/545907/files/figure.png" pageId="7" pageNumber="262" targetBox="[298,1285,208,1917]" targetPageId="7">
<paragraph id="8BCECA65FFCFFFF740E195AAA4A4F84F" blockId="7.[151,1436,1953,2006]" pageId="7" pageNumber="262">
<emphasis id="B9051677FFCFFFF740E195AAA415F82E" bold="true" box="[151,274,1953,1975]" pageId="7" pageNumber="262">FIGURE 5.</emphasis>
<taxonomicName id="4C71B1E6FFCFFFF7416F95A9A468F82E" box="[281,367,1954,1975]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9051677FFCFFFF7416F95A9A468F82E" box="[281,367,1954,1975]" italics="true" pageId="7" pageNumber="262">Leielmis</emphasis>
</taxonomicName>
species, aedeagi, ventral and lateral views. A) 
<taxonomicName id="4C71B1E6FFCFFFF7433F95A9A6E1F82E" box="[841,998,1954,1975]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFCFFFF7433F95A9A6E1F82E" box="[841,998,1954,1975]" italics="true" pageId="7" pageNumber="262">L. georyssoides</emphasis>
</taxonomicName>
; B) 
<taxonomicName id="4C71B1E6FFCFFFF7446595A9A180F82E" box="[1043,1159,1954,1975]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B9051677FFCFFFF7446595A9A180F82E" box="[1043,1159,1954,1975]" italics="true" pageId="7" pageNumber="262">L. gibbosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFCFFFF744F895A9A1D9F82E" box="[1166,1246,1954,1975]" pageId="7" pageNumber="262" rank="species">
<emphasis id="B9051677FFCFFFF744F895A9A1D9F82E" bold="true" box="[1166,1246,1954,1975]" pageId="7" pageNumber="262">sp. nov.</emphasis>
</taxonomicNameLabel>
; C) 
<taxonomicName id="4C71B1E6FFCFFFF7457D95A9A07FF82E" box="[1291,1400,1954,1975]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B9051677FFCFFFF7457D95A9A07FF82E" box="[1291,1400,1954,1975]" italics="true" pageId="7" pageNumber="262">L. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFCFFFF7450B95A9A5C4F84C" pageId="7" pageNumber="262" rank="species">
<emphasis id="B9051677FFCFFFF7450B95A9A5C4F84C" bold="true" pageId="7" pageNumber="262">sp. nov.</emphasis>
</taxonomicNameLabel>
Scale bar = 0.25 mm.
</paragraph>
</caption>
<caption id="DF0E9AEDFFC0FFF840E195AAA4A4F84F" httpUri="https://zenodo.org/record/545908/files/figure.png" pageId="8" pageNumber="263" targetBox="[259,1317,215,1910]" targetPageId="8">
<paragraph id="8BCECA65FFC0FFF840E195AAA4A4F84F" blockId="8.[151,1436,1953,2006]" pageId="8" pageNumber="263">
<emphasis id="B9051677FFC0FFF840E195AAA413F82F" bold="true" box="[151,276,1953,1975]" pageId="8" pageNumber="263">FIGURE 6.</emphasis>
<taxonomicName id="4C71B1E6FFC0FFF8416A95A9A475F82E" box="[284,370,1954,1975]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B9051677FFC0FFF8416A95A9A475F82E" box="[284,370,1954,1975]" italics="true" pageId="8" pageNumber="263">Leielmis</emphasis>
</taxonomicName>
species, ovipositors, distal gonocoxites. A) 
<taxonomicName id="4C71B1E6FFC0FFF8434F95A9A6DFF82E" box="[825,984,1954,1975]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFC0FFF8434F95A9A6DFF82E" box="[825,984,1954,1975]" italics="true" pageId="8" pageNumber="263">L. georyssoides</emphasis>
</taxonomicName>
; B) 
<taxonomicName id="4C71B1E6FFC0FFF8447E95A9A179F82E" box="[1032,1150,1954,1975]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B9051677FFC0FFF8447E95A9A179F82E" box="[1032,1150,1954,1975]" italics="true" pageId="8" pageNumber="263">L. gibbosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFC0FFF844F095A9A1DFF82E" box="[1158,1240,1954,1975]" pageId="8" pageNumber="263" rank="species">
<emphasis id="B9051677FFC0FFF844F095A9A1DFF82E" bold="true" box="[1158,1240,1954,1975]" pageId="8" pageNumber="263">sp. nov.</emphasis>
</taxonomicNameLabel>
; C) 
<taxonomicName id="4C71B1E6FFC0FFF8457E95A9A071F82E" box="[1288,1398,1954,1975]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B9051677FFC0FFF8457E95A9A071F82E" box="[1288,1398,1954,1975]" italics="true" pageId="8" pageNumber="263">L. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFC0FFF8450B95A9A5C4F84C" pageId="8" pageNumber="263" rank="species">
<emphasis id="B9051677FFC0FFF8450B95A9A5C4F84C" bold="true" pageId="8" pageNumber="263">sp. nov.</emphasis>
</taxonomicNameLabel>
Scale bar = 0.25 mm.
</paragraph>
</caption>
<paragraph id="8BCECA65FFC1FFF940B19293A103FF29" blockId="9.[151,1437,151,1256]" box="[199,1028,151,176]" pageId="9" pageNumber="264">
Legs: Meso and metatibiae with characteristic modifications (
<figureCitation id="134AD6E0FFC1FFF9430D929CA6B9FF29" box="[891,958,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1964,1986]" captionTargetBox="[446,1135,210,1910]" captionTargetId="figure@6.[438,1155,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Leielmis species, male tibial modifications, dorsal view A) &amp; D) L. georyssoides; B) &amp; E) L. gibbosus sp. nov.; C) &amp; F) L. hirsutus sp. nov. A) – C) mesotibiae; D) – F) metatibiae. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/545906/files/figure.png" pageId="9" pageNumber="264">Fig. 4</figureCitation>
A, D).
</paragraph>
<paragraph id="8BCECA65FFC1FFF940B192B7A4ECFE84" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="264">
Aedeagus: Phallobase relatively short, 0.7x length of penis. Penis distinctly longer than parameres, curved apically in lateral view, with truncately acuminate apex. Parameres narrow, with rounded apices and slightly curved interior margins (see 
<figureCitation id="134AD6E0FFC1FFF941F3930FA4C1FE84" box="[389,454,260,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1953,1975]" captionTargetBox="[298,1285,208,1917]" captionTargetId="figure@7.[283,1304,193,1932]" captionTargetPageId="7" captionText="FIGURE 5. Leielmis species, aedeagi, ventral and lateral views. A) L. georyssoides; B) L. gibbosus sp. nov.; C) L. hirsutus sp. nov. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/545907/files/figure.png" pageId="9" pageNumber="264">Fig. 5</figureCitation>
A).
</paragraph>
<paragraph id="8BCECA65FFC1FFF940B1932CA08FFEFC" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="264">
Females: As males, except for slightly shallower mesoventral concavity and lack of tibial modifications. Ovipositor elongate, distal gonocoxite with relatively stout apices (
<figureCitation id="134AD6E0FFC1FFF943F09347A6CEFEFC" box="[902,969,332,357]" captionStart="FIGURE 6" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[259,1317,215,1910]" captionTargetId="figure@8.[244,1344,193,1932]" captionTargetPageId="8" captionText="FIGURE 6. Leielmis species, ovipositors, distal gonocoxites. A) L. georyssoides; B) L. gibbosus sp. nov.; C) L. hirsutus sp. nov. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/545908/files/figure.png" pageId="9" pageNumber="264">Fig. 6</figureCitation>
A); 0.7x length of proximal gonocoxite.
</paragraph>
<paragraph id="8BCECA65FFC1FFF940B19364A7D6FE49" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="264">
Variation: Specimens vary somewhat in colouration, most of the 
<typeStatus id="54CA74C7FFC1FFF943D8937BA6D9FE11" box="[942,990,368,392]" pageId="9" pageNumber="264">type</typeStatus>
series (including the 
<typeStatus id="54CA74C7FFC1FFF944A19364A047FE11" box="[1239,1344,367,392]" pageId="9" pageNumber="264" type="lectotype">lectotype</typeStatus>
) having the elytra more brownish in colour than others. All recently collected material is relatively dark, however, the predominant impression being of a blackish insect.
</paragraph>
</subSubSection>
<subSubSection id="C36B99EEFFC1FFF940B193D7A733FCA1" pageId="9" pageNumber="264" type="diagnosis">
<paragraph id="8BCECA65FFC1FFF940B193D7A733FCA1" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="264">
<emphasis id="B9051677FFC1FFF940B193D7A4C9FE6C" bold="true" box="[199,462,476,501]" pageId="9" pageNumber="264">Differential diagnosis</emphasis>
. Very similar in general habitus to 
<taxonomicName id="4C71B1E6FFC1FFF943E393D6A127FE6D" box="[917,1056,476,500]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B9051677FFC1FFF943E393D6A127FE6D" box="[917,1056,476,500]" italics="true" pageId="9" pageNumber="264">L. gibbosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFC1FFF9445993D7A196FE6C" box="[1071,1169,476,501]" pageId="9" pageNumber="264" rank="species">
<emphasis id="B9051677FFC1FFF9445993D7A196FE6C" bold="true" box="[1071,1169,476,501]" pageId="9" pageNumber="264">sp. nov.</emphasis>
</taxonomicNameLabel>
(see 
<figureCitation id="134AD6E0FFC1FFF944A893D7A023FE6D" box="[1246,1316,476,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1980,2002]" captionTargetBox="[166,1414,644,1940]" captionTargetId="figure@3.[151,1436,625,1958]" captionTargetPageId="3" captionText="FIGURE 1. Leielmis species, male habitus. A) L. georyssoides; B) L. gibbosus sp. nov., holotype; C) L. hirsutus sp. nov., holotype. Scale bar = 1 mm." httpUri="https://zenodo.org/record/545902/files/figure.png" pageId="9" pageNumber="264">Fig. 1</figureCitation>
A, B). 
<taxonomicName id="4C71B1E6FFC1FFF945F393D6A42FFD81" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFC1FFF945F393D6A42FFD81" italics="true" pageId="9" pageNumber="264">L. georyssoides</emphasis>
</taxonomicName>
differs from this species in the denser pronotal punctation, the coarse punctures of which are individually larger (ca. 
<date id="FFCFECA5FFC1FFF941D1902FA4C4FDA5" box="[423,451,548,572]" pageId="9" pageNumber="264">2x</date>
diameter of those of 
<taxonomicName id="4C71B1E6FFC1FFF942B2902EA64FFDA5" box="[708,840,548,572]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B9051677FFC1FFF942B2902EA64FFDA5" box="[708,840,548,572]" italics="true" pageId="9" pageNumber="264">L. gibbosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFC1FFF94325902FA6B7FDA4" box="[851,944,548,573]" pageId="9" pageNumber="264" rank="species">
<emphasis id="B9051677FFC1FFF94325902FA6B7FDA4" bold="true" box="[851,944,548,573]" pageId="9" pageNumber="264">sp. nov.</emphasis>
</taxonomicNameLabel>
– 
<figureCitation id="134AD6E0FFC1FFF943A4902FA111FDA5" box="[978,1046,548,573]" captionStart="FIGURE 2" captionStartId="4.[151,250,1664,1686]" captionTargetBox="[155,1431,186,1636]" captionTargetId="figure@4.[151,1436,183,1659]" captionTargetPageId="4" captionText="FIGURE 2. Leielmis species, pronotal sculpture. A) &amp; D) L. georyssiodes; B) &amp; E) L. gibbosus sp. nov.; C) &amp; F) L. hirsutus sp. nov. A) – C) scanning electron micrographs; D) – F) stacked macrosope images." httpUri="https://zenodo.org/record/545903/files/figure.png" pageId="9" pageNumber="264">Fig. 2</figureCitation>
A &amp; B; D &amp; E); the more striate impressed elytral puncture rows, whose punctures are larger, especially at the base (
<figureCitation id="134AD6E0FFC1FFF94412904CA1A0FDF9" box="[1124,1191,583,608]" captionStart="FIGURE 3" captionStartId="5.[151,250,1150,1172]" captionTargetBox="[164,1408,193,1126]" captionTargetId="figure@5.[164,1408,193,1126]" captionTargetPageId="5" captionText="FIGURE 3. Leielmis species, Elytral sculpture, scanning electron micrographs. A) L. georyssoides; B) L. gibbosus sp. nov.; C) L. hirsutus sp. nov." httpUri="https://zenodo.org/record/545905/files/figure.png" pageId="9" pageNumber="264">Fig. 3</figureCitation>
A &amp; B); the broader, shallower anterior emargination of the mesoventrite, the larger, denser punctures of the metaventrite, and the detail of the modifications to the male meso and metatibiae (
<figureCitation id="134AD6E0FFC1FFF9428F9084A63CFD31" box="[761,827,655,680]" captionStart="FIGURE 4" captionStartId="6.[151,250,1964,1986]" captionTargetBox="[446,1135,210,1910]" captionTargetId="figure@6.[438,1155,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Leielmis species, male tibial modifications, dorsal view A) &amp; D) L. georyssoides; B) &amp; E) L. gibbosus sp. nov.; C) &amp; F) L. hirsutus sp. nov. A) – C) mesotibiae; D) – F) metatibiae. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/545906/files/figure.png" pageId="9" pageNumber="264">Fig. 4</figureCitation>
A &amp; B). The aedeagi of the two species (
<figureCitation id="134AD6E0FFC1FFF945709084A040FD31" box="[1286,1351,655,680]" captionStart="FIGURE 5" captionStartId="7.[151,250,1953,1975]" captionTargetBox="[298,1285,208,1917]" captionTargetId="figure@7.[283,1304,193,1932]" captionTargetPageId="7" captionText="FIGURE 5. Leielmis species, aedeagi, ventral and lateral views. A) L. georyssoides; B) L. gibbosus sp. nov.; C) L. hirsutus sp. nov. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/545907/files/figure.png" pageId="9" pageNumber="264">Fig. 5</figureCitation>
A &amp; B) also differ in detail, 
<taxonomicName id="4C71B1E6FFC1FFF9410E90BEA721FD55" box="[376,550,692,716]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFC1FFF9410E90BEA721FD55" box="[376,550,692,716]" italics="true" pageId="9" pageNumber="264">L. georyssoides</emphasis>
</taxonomicName>
having relatively shorter, narrower parameres, not so strongly incurved at their apices, with the tip of the penis extending further beyond the paramere apices and bearing a longer apical acumination. The distal gonocoxite of the ovipositor of 
<taxonomicName id="4C71B1E6FFC1FFF9437F90F6A6B0FC8D" box="[777,951,764,788]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFC1FFF9437F90F6A6B0FC8D" box="[777,951,764,788]" italics="true" pageId="9" pageNumber="264">L. georyssoides</emphasis>
</taxonomicName>
is stouter than that of 
<taxonomicName id="4C71B1E6FFC1FFF944C590F6A033FC8D" box="[1203,1332,764,788]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B9051677FFC1FFF944C590F6A033FC8D" box="[1203,1332,764,788]" italics="true" pageId="9" pageNumber="264">L. gibbosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236AB0CFFC1FFF9454D90F7A092FC8C" box="[1339,1429,764,789]" pageId="9" pageNumber="264" rank="species">
<emphasis id="B9051677FFC1FFF9454D90F7A092FC8C" bold="true" box="[1339,1429,764,789]" pageId="9" pageNumber="264">sp. nov.</emphasis>
</taxonomicNameLabel>
, especially at the apex (
<figureCitation id="134AD6E0FFC1FFF941E19114A4DFFCA1" box="[407,472,799,824]" captionStart="FIGURE 6" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[259,1317,215,1910]" captionTargetId="figure@8.[244,1344,193,1932]" captionTargetPageId="8" captionText="FIGURE 6. Leielmis species, ovipositors, distal gonocoxites. A) L. georyssoides; B) L. gibbosus sp. nov.; C) L. hirsutus sp. nov. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/545908/files/figure.png" pageId="9" pageNumber="264">Fig. 6</figureCitation>
A &amp; B).
</paragraph>
</subSubSection>
<subSubSection id="C36B99EEFFC1FFF940B1914FA640FB71" pageId="9" pageNumber="264" type="distribution">
<paragraph id="8BCECA65FFC1FFF940B1914FA640FB71" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="264">
<emphasis id="B9051677FFC1FFF940B1914FA4F5FCC4" bold="true" box="[199,498,836,861]" pageId="9" pageNumber="264">Distribution and ecology</emphasis>
. 
<bibRefCitation id="EFE0B794FFC1FFF94272914FA7C4FCC5" author="Grouvelle" box="[516,707,836,861]" pageId="9" pageNumber="264" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle (1890</bibRefCitation>
, 
<bibRefCitation id="EFE0B794FFC1FFF942A4914FA60CFCC4" author="Grouvelle" box="[722,779,836,861]" pageId="9" pageNumber="264" refString="Grouvelle, A. (1895) Voyage de M. E. Simon dans l'Afrique austral (Janvier - Avril 1893), 4 0 Memoire. Nitidulidae, Colydiidae, Cucujidae, Cryptophagidae &amp; Parnidae. Annales de la Societe Entomologique de France, 64, 161 - 168." type="journal article" year="1895">1895</bibRefCitation>
) stated that this species was collected at ‘cap de Bonne Espérance’. 
<bibRefCitation id="EFE0B794FFC1FFF94151916CA4C8FC19" author="Deleve" box="[295,463,871,896]" pageId="9" pageNumber="264" refString="Deleve, J. (1964) Contribution a l'etude des Dryopoidea. VIII. Cinq nouveaux genres d'Elminthidae (Coleoptera) de l'Afrique australe. Bulletin et Annales de la Societe Royale d'Enromologie de Belgique, 100, 155 - 176." type="journal article" year="1964">Delève (1964)</bibRefCitation>
based his description of 
<taxonomicName id="4C71B1E6FFC1FFF943739163A663FC19" box="[773,868,872,896]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B9051677FFC1FFF943739163A663FC19" box="[773,868,872,896]" italics="true" pageId="9" pageNumber="264">Leielmis</emphasis>
</taxonomicName>
, and redescription of 
<taxonomicName id="4C71B1E6FFC1FFF944049163A004FC19" box="[1138,1283,872,896]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFC1FFF944049163A004FC19" box="[1138,1283,872,896]" italics="true" pageId="9" pageNumber="264">georyssoides</emphasis>
</taxonomicName>
, on material collected by A. D. Harrison in the Great Berg River, sent to him by M. H. Bertrand. He attributed this to 
<taxonomicName id="4C71B1E6FFC1FFF940E191BBA42FFC51" box="[151,296,944,968]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFC1FFF940E191BBA42FFC51" box="[151,296,944,968]" italics="true" pageId="9" pageNumber="264">georyssoides</emphasis>
</taxonomicName>
, without study of Grouvelle’s 
<typeStatus id="54CA74C7FFC1FFF9420891BBA7BBFC51" box="[638,700,944,968]" pageId="9" pageNumber="264">types</typeStatus>
, on the basis of its close overall match to the original description. 
<bibRefCitation id="EFE0B794FFC1FFF940E191DFA430FC75" author="Deleve" box="[151,311,980,1005]" pageId="9" pageNumber="264" refString="Deleve, J. (1970) Coleoptera: Dryopidae et Elminthidae. In: Hanstrom, B., Brinck, P. &amp; Rudebeck, G. (Eds.), South African Animal Life. Fol. 14. Swedish Natural Science Research Council, Stockholm, pp. 216 - 227" type="journal article" year="1970">Delève (1970)</bibRefCitation>
noted that a female 
<taxonomicName id="4C71B1E6FFC1FFF9426E91DFA770FC75" box="[536,631,980,1004]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B9051677FFC1FFF9426E91DFA770FC75" box="[536,631,980,1004]" italics="true" pageId="9" pageNumber="264">Leielmis</emphasis>
</taxonomicName>
collected on Table Mountain by Per Brinck differed from his concept of 
<taxonomicName id="4C71B1E6FFC1FFF940E191F3A42FFB89" box="[151,296,1016,1040]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFC1FFF940E191F3A42FFB89" box="[151,296,1016,1040]" italics="true" pageId="9" pageNumber="264">georyssoides</emphasis>
</taxonomicName>
in having ‘le pronotum bien plus fortement ponctué, … les stries élytrales plus marqués’, but refrained from describing a new species in the absence of further material. The discovery of the 
<typeStatus id="54CA74C7FFC1FFF945779616A036FBAC" box="[1281,1329,1053,1077]" pageId="9" pageNumber="264">type</typeStatus>
series in SANC shows that this Table Mountain taxon referred to by Delève is in fact the true 
<taxonomicName id="4C71B1E6FFC1FFF9440D964AA02BFBC1" box="[1147,1324,1088,1112]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFC1FFF9440D964AA02BFBC1" box="[1147,1324,1088,1112]" italics="true" pageId="9" pageNumber="264">L. georyssoides</emphasis>
</taxonomicName>
, all other 
<taxonomicName id="4C71B1E6FFC1FFF940E1966FA5F1FBE5" box="[151,246,1124,1148]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B9051677FFC1FFF940E1966FA5F1FBE5" box="[151,246,1124,1148]" italics="true" pageId="9" pageNumber="264">Leielmis</emphasis>
</taxonomicName>
specimens belonging to other, previously undescribed species. It seems that 
<taxonomicName id="4C71B1E6FFC1FFF94427966EA007FBE5" box="[1105,1280,1124,1148]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="georyssoides">
<emphasis id="B9051677FFC1FFF94427966EA007FBE5" box="[1105,1280,1124,1148]" italics="true" pageId="9" pageNumber="264">L. georyssoides</emphasis>
</taxonomicName>
is endemic to the far southwestern 
<collectingRegion id="49B50487FFC1FFF941F79683A4BAFB39" box="[385,445,1160,1184]" country="South Africa" name="Western Cape" pageId="9" pageNumber="264">Cape</collectingRegion>
, perhaps to Table Mountain itself, where it appears to have been collected exclusively in permanent running waters. In 
<date id="FFCFECA5FFC1FFF9419396A7A789FB5C" box="[485,654,1196,1221]" pageId="9" pageNumber="264" value="2010" valueMax="2015">2010 and 2015</date>
, the species was abundant below stones in small riffles in a permanent mountain stream on the summit of Table Mountain (
<figureCitation id="134AD6E0FFC1FFF9429696C4A624FB71" box="[736,803,1231,1256]" captionStart="FIGURE 7" captionStartId="9.[151,250,1952,1974]" captionTargetBox="[151,1436,1303,1930]" captionTargetId="figure@9.[151,1436,1303,1930]" captionTargetPageId="9" captionText="FIGURE 7. Leielmis species, habitats. A) Western Cape, Table Mountain, stream in Echo Valley, L. georyssoides; B) Western Cape, Groote Winterhoekberge, Stream below Sneeugatpiek, L. gibbosus sp. nov. &amp; L. hirsutus sp. nov.; C) Western Cape, stream at Mont Rochelle above Franschhoek, L. gibbosus sp. nov. Photos D. T. Bilton &amp; N. Helme." httpUri="https://zenodo.org/record/545909/files/figure.png" pageId="9" pageNumber="264">Fig. 7</figureCitation>
A).
</paragraph>
</subSubSection>
</treatment>
</document>